treatments-xml/data/03/F3/87/03F3878EFFD1FFB563B90000FB1D6F8C.xml
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<document id="8024ADF6EC57831961456D6FB8960A0E" ID-DOI="10.11646/zootaxa.4802.2.7" ID-ISSN="1175-5326" ID-Zenodo-Dep="3907558" ID-ZooBank="41E57AB6-CBA9-490A-9540-20DF787EAD3C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1593080386831" checkinUser="plazi" docAuthor="Zhang, Wei, Zhang, Min, Han, Na &amp; Zhou, Chang-Fa" docDate="2020" docId="03F3878EFFD1FFB563B90000FB1D6F8C" docLanguage="en" docName="zootaxa.4802.2.7.pdf" docOrigin="Zootaxa 4802 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Notacanthurus maculatus Zhang &amp; Zhang &amp; Han &amp; Zhou 2020, sp. n." docType="treatment" docVersion="3" lastPageNumber="341" masterDocId="FFCAFFF6FFD0FFB3632E026DFFBC6D0B" masterDocTitle="Two new species of the genus Notacanthurus from China (Ephemeroptera: Heptageniidae, Ecdyonurinae)" masterLastPageNumber="348" masterPageNumber="335" pageNumber="336" updateTime="1700276942354" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="73E58BDB7ACEA8E5FC2AF3D8690ACCDD">Two new species of the genus Notacanthurus from China (Ephemeroptera: Heptageniidae, Ecdyonurinae)</mods:title>
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<mods:namePart id="1A0A856FCE92D7F57578EB2BA4F26576">Zhang, Wei</mods:namePart>
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<mods:namePart id="D5598C52F5C4F3BC20A4F356CFE396EA">Zhang, Min</mods:namePart>
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<mods:namePart id="58CFBF7732DEF2B0E89A2F461A1BCDDC">Han, Na</mods:namePart>
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<mods:namePart id="8219EB29B27FF850D6BC250A6DA5D18C">Zhou, Chang-Fa</mods:namePart>
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<mods:title id="F547BFDA5B74B20177BA5735FC76D455">Zootaxa</mods:title>
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<mods:part id="81B1FCBABA0E589A2EA0D590FA59576F">
<mods:date id="D6A2F5AADAD534F3EF0EFE51FCC8BFD7">2020</mods:date>
<mods:detail id="C8040B54E5FD58E1CDACC8BCB340BBA7" type="pubDate">
<mods:number id="797D35EAC7B428E7115FDB5087954926">2020-06-23</mods:number>
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<mods:number id="1CB979C50D826318B112112795FEFC2F">4802</mods:number>
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<mods:identifier id="03A45E5C19B161201CE6B0E51F39E475" type="DOI">10.11646/zootaxa.4802.2.7</mods:identifier>
<mods:identifier id="F2224A26691FE40654B872D163A9A06C" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03F3878EFFD1FFB563B90000FB1D6F8C" LSID="urn:lsid:plazi:treatment:03F3878EFFD1FFB563B90000FB1D6F8C" httpUri="http://treatment.plazi.org/id/03F3878EFFD1FFB563B90000FB1D6F8C" lastPageId="6" lastPageNumber="341" pageId="1" pageNumber="336">
<subSubSection id="C3406513FFD1FFB263B90000FDD26F8C" box="[151,622,621,648]" pageId="1" pageNumber="336" type="nomenclature">
<paragraph id="8BE53698FFD1FFB263B90000FDD26F8C" blockId="1.[151,622,621,648]" box="[151,622,621,648]" pageId="1" pageNumber="336">
<heading id="D0AD81F4FFD1FFB263B90000FDD26F8C" bold="true" box="[151,622,621,648]" fontSize="11" level="1" pageId="1" pageNumber="336" reason="1">
<emphasis id="B92EEA8AFFD1FFB263B90000FDD26F8C" bold="true" box="[151,622,621,648]" pageId="1" pageNumber="336">
<taxonomicName id="4C5A4D1BFFD1FFB263B90000FDA26F8C" authority="Zhou" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[151,542,621,648]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD1FFB263B90000FE6F6F83" bold="true" box="[151,467,621,648]" italics="true" pageId="1" pageNumber="336">Notacanthurus maculatus</emphasis>
Zhou
</taxonomicName>
,
<taxonomicNameLabel id="A21D57F1FFD1FFB261050000FDD26F8C" box="[555,622,621,647]" pageId="1" pageNumber="336" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3406513FFD1FFB263B900D9FD9B6950" pageId="1" pageNumber="336" type="materials_examined">
<paragraph id="8BE53698FFD1FFB263B900D9FED56E1C" blockId="1.[151,1437,692,1259]" pageId="1" pageNumber="336">
<emphasis id="B92EEA8AFFD1FFB263B900D9FE296FC4" bold="true" box="[151,405,692,719]" pageId="1" pageNumber="336">Materials examined.</emphasis>
<materialsCitation id="3B323CC5FFD1FFB2628800D8FED56E1C" collectingDate="2000-07-25" collectionCode="ZHOU" country="China" county="Qiang" latitude="33.547462" location="Fo-Ping county" longLatPrecision="1" longitude="107.82482" municipality="Han-Zhong" pageId="1" pageNumber="336" specimenCount="1" stateProvince="Shaanxi" typeStatus="holotype">
<typeStatus id="54E1883AFFD1FFB2628800D8FDAF6FC4" box="[422,531,693,719]" pageId="1" pageNumber="336" type="holotype">Holotype</typeStatus>
: male imago (reared from mature nymph), Yue-Ba town (
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,
<geoCoordinate id="EE6E505FFFD1FFB263B900B5FEF26FF8" box="[151,334,728,755]" degrees="107" direction="east" minutes="49" orientation="longitude" pageId="1" pageNumber="336" precision="1" seconds="29.36" value="107.82482">107°4929.36″E</geoCoordinate>
),
<location id="8E856043FFD1FFB2624F00B4FDB36FF8" LSID="urn:lsid:plazi:treatment:03F3878EFFD1FFB563B90000FB1D6F8C:8E856043FFD1FFB2624F00B4FDB36FF8" box="[353,527,729,755]" country="China" county="Qiang" latitude="33.547462" longLatPrecision="1" longitude="107.82482" municipality="Han-Zhong" name="Fo-Ping county" pageId="1" pageNumber="336" stateProvince="Shaanxi">Fo-Ping county</location>
,
<collectingMunicipality id="6B81ACE2FFD1FFB2613400B4FD276FF8" box="[538,667,729,755]" pageId="1" pageNumber="336">Han-Zhong</collectingMunicipality>
city,
<collectingRegion id="499EF87AFFD1FFB261F900B4FC226FF8" box="[727,926,729,755]" country="China" name="Shaanxi" pageId="1" pageNumber="336">Shaanxi Province</collectingRegion>
,
<collectingCountry id="F34D7608FFD1FFB2608700B4FC536FF8" box="[937,1007,729,755]" name="China" pageId="1" pageNumber="336">China</collectingCountry>
, collected by Chang-Fa
<collectionCode id="ED4BAE5DFFD1FFB267D200B4FAF16FF8" box="[1276,1357,729,755]" pageId="1" pageNumber="336">ZHOU</collectionCode>
,
<collectingCounty id="62844E14FFD1FFB2667600B4FA206FF8" box="[1368,1436,729,755]" pageId="1" pageNumber="336">Qiang</collectingCounty>
XIE,
<date id="FFE41058FFD1FFB263FA0091FED96E1C" box="[212,357,764,791]" pageId="1" pageNumber="336" value="2000-07-25">
<collectingDate id="EFA0E9B0FFD1FFB263FA0091FED96E1C" box="[212,357,764,791]" pageId="1" pageNumber="336" value="2000-07-25">25-VII-2000</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BE53698FFD1FFB263E9014CFD9B6950" blockId="1.[151,1437,692,1259]" pageId="1" pageNumber="336">
<typeStatus id="54E1883AFFD1FFB263E9014CFE846E30" box="[199,312,801,827]" pageId="1" pageNumber="336" type="paratype">Paratypes</typeStatus>
:
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imagoes
<specimenCount id="9D5CFD11FFD1FFB262DD014CFDEE6E30" box="[499,594,801,827]" pageId="1" pageNumber="336" type="female">1 female</specimenCount>
imago
<specimenCount id="9D5CFD11FFD1FFB2618D014CFCBE6E30" box="[675,770,801,827]" pageId="1" pageNumber="336" type="female">3 female</specimenCount>
subimagoes (reared from mature nymphs) 50 nymphs, same as the
<typeStatus id="54E1883AFFD1FFB263F00128FEF86E54" box="[222,324,837,863]" pageId="1" pageNumber="336" type="holotype">holotype</typeStatus>
;
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imago
<materialsCitation id="3B323CC5FFD1FFB262C20128FC596E88" collectingDate="2000-08-08" collectionCode="ZHOU" latitude="32.87935" location="Wen county" longLatPrecision="1" longitude="104.40621" municipality="Tie-Lou village" pageId="1" pageNumber="336" specimenCount="3" specimenCount-female="3" stateProvince="Gansu" typeStatus="paratype">
<specimenCount id="9D5CFD11FFD1FFB262C20128FDF06E54" box="[492,588,837,863]" pageId="1" pageNumber="336" type="female">3 female</specimenCount>
imagoes, Sai-Ke bridge (
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,
<geoCoordinate id="EE6E505FFFD1FFB267300129FB6A6E54" box="[1054,1238,836,863]" degrees="104" direction="east" minutes="24" orientation="longitude" pageId="1" pageNumber="336" precision="1" seconds="22.36" value="104.40621">104°2422.36″E</geoCoordinate>
),
<collectingMunicipality id="6B81ACE2FFD1FFB267C60128FA246E54" box="[1256,1432,837,863]" pageId="1" pageNumber="336">Tie-Lou village</collectingMunicipality>
,
<location id="8E856043FFD1FFB263B90104FE9C6E88" LSID="urn:lsid:plazi:treatment:03F3878EFFD1FFB563B90000FB1D6F8C:8E856043FFD1FFB263B90104FE9C6E88" box="[151,288,873,899]" latitude="32.87935" longLatPrecision="1" longitude="104.40621" municipality="Tie-Lou village" name="Wen county" pageId="1" pageNumber="336" stateProvince="Gansu">Wen county</location>
,
<collectingRegion id="499EF87AFFD1FFB262030104FE5F6E88" box="[301,483,873,899]" country="China" name="Gansu" pageId="1" pageNumber="336">Gansu Province</collectingRegion>
, collected by Chang-Fa
<collectionCode id="ED4BAE5DFFD1FFB261D50104FCF06E88" box="[763,844,873,899]" pageId="1" pageNumber="336">ZHOU</collectionCode>
,
<date id="FFE41058FFD1FFB260760104FC596E88" box="[856,997,873,899]" pageId="1" pageNumber="336" value="2000-08-08">
<collectingDate id="EFA0E9B0FFD1FFB260760104FC596E88" box="[856,997,873,899]" pageId="1" pageNumber="336" value="2000-08-08">8-VIII-2000</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D5CFD11FFD1FFB260DC0104FB836E88" box="[1010,1087,873,899]" pageId="1" pageNumber="336" type="male">4 male</specimenCount>
imagoes
<materialsCitation id="3B323CC5FFD1FFB267800104FE7C6EC0" collectionCode="WANG" collectorName="Xi-Xia" county="Xia-Chuan town" latitude="32.87935" location="Yan-Xia" longLatPrecision="1" longitude="104.40621" municipality="Jin-Cheng" pageId="1" pageNumber="336" specimenCount="2" specimenCount-female="2" stateProvince="Shanxi" typeStatus="paratype">
<specimenCount id="9D5CFD11FFD1FFB267800104FAAD6E88" box="[1198,1297,873,899]" pageId="1" pageNumber="336" type="female">2 female</specimenCount>
imagoes 50 nymphs,
<collectorName id="26AF534EFFD1FFB263D001E0FEF06EAC" box="[254,332,909,935]" pageId="1" pageNumber="336">Xi-Xia</collectorName>
stream (
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,
<geoCoordinate id="EE6E505FFFD1FFB2614B01E1FCA06EAC" box="[613,796,908,935]" degrees="104" direction="east" minutes="24" orientation="longitude" pageId="1" pageNumber="336" precision="1" seconds="22.36" value="104.40621">104°2422.36″E</geoCoordinate>
),
<collectingCounty id="62844E14FFD1FFB2601E01E0FC516EAC" box="[816,1005,909,935]" pageId="1" pageNumber="336">Xia-Chuan town</collectingCounty>
,
<collectingMunicipality id="6B81ACE2FFD1FFB260D701E0FBD06EAC" box="[1017,1132,909,935]" pageId="1" pageNumber="336">Jin-Cheng</collectingMunicipality>
city,
<collectingRegion id="499EF87AFFD1FFB2678401E0FAD96EAC" box="[1194,1381,909,935]" country="China" name="Shanxi" pageId="1" pageNumber="336">Shanxi Province</collectingRegion>
, col- lected by
<location id="8E856043FFD1FFB2622A01DCFEDE6EC0" LSID="urn:lsid:plazi:treatment:03F3878EFFD1FFB563B90000FB1D6F8C:8E856043FFD1FFB2622A01DCFEDE6EC0" box="[260,354,945,971]" county="Xia-Chuan town" latitude="32.87935" longLatPrecision="1" longitude="104.40621" municipality="Jin-Cheng" name="Yan-Xia" pageId="1" pageNumber="336" stateProvince="Shanxi">Yan-Xia</location>
<collectionCode id="ED4BAE5DFFD1FFB2624701DCFE7C6EC0" box="[361,448,945,971]" pageId="1" pageNumber="336">WANG</collectionCode>
</materialsCitation>
,
<materialsCitation id="3B323CC5FFD1FFB262E501DCFCB06EC0" box="[459,780,945,971]" collectingDate="2012-07-23" collectionCode="WANG" pageId="1" pageNumber="336" specimenCount="1" stateProvince="Shanxi" typeStatus="paratype">
Ze-Yu
<collectionCode id="ED4BAE5DFFD1FFB2613601DCFDD36EC0" box="[536,623,945,971]" pageId="1" pageNumber="336">WANG</collectionCode>
,
<date id="FFE41058FFD1FFB2615501DCFCB06EC0" box="[635,780,945,971]" pageId="1" pageNumber="336" value="2012-07-23">
<collectingDate id="EFA0E9B0FFD1FFB2615501DCFCB06EC0" box="[635,780,945,971]" pageId="1" pageNumber="336" value="2012-07-23">23-VII-2012</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D5CFD11FFD1FFB2603901DCFCDF6EC0" box="[791,867,945,971]" pageId="1" pageNumber="336" type="male">3 male</specimenCount>
imagoes
<specimenCount id="9D5CFD11FFD1FFB260FE01DCFBA76EC0" box="[976,1051,945,971]" pageId="1" pageNumber="336" type="male">6 male</specimenCount>
subimagoes
<materialsCitation id="3B323CC5FFD1FFB2678101DCFC416918" collectingDate="2013-09-13" collectionCode="WANG" collectorName="Chang-Hai &amp; Ji-Hua XU &amp; Zi-Wei" county="Great-River village" latitude="35.73997" location="Lin-Fen" longLatPrecision="1" longitude="111.03984" municipality="Yi-Cheng county" pageId="1" pageNumber="336" specimenCount="4" specimenCount-female="4" stateProvince="Shanxi" typeStatus="paratype">
<specimenCount id="9D5CFD11FFD1FFB2678101DCFAAC6EC0" box="[1199,1296,945,971]" pageId="1" pageNumber="336" type="female">4 female</specimenCount>
subimagoes 30 nymphs,
<collectingCounty id="62844E14FFD1FFB2620801B8FDBE6EE4" box="[294,514,981,1007]" pageId="1" pageNumber="336">Great-River village</collectingCounty>
(
<geoCoordinate id="EE6E505FFFD1FFB2613D01B9FD7C6EE4" box="[531,704,980,1007]" degrees="35" direction="north" minutes="44" orientation="latitude" pageId="1" pageNumber="336" precision="1" seconds="23.90" value="35.73997">35°4423.90″N</geoCoordinate>
,
<geoCoordinate id="EE6E505FFFD1FFB261E001B9FC3F6EE4" box="[718,899,980,1007]" degrees="111" direction="east" minutes="02" orientation="longitude" pageId="1" pageNumber="336" precision="1" seconds="23.43" value="111.03984">111°0223.43″E</geoCoordinate>
),
<collectingMunicipality id="6B81ACE2FFD1FFB260B601B8FBE06EE4" box="[920,1116,981,1007]" pageId="1" pageNumber="336">Yi-Cheng county</collectingMunicipality>
,
<location id="8E856043FFD1FFB2674401B8FB786EE4" LSID="urn:lsid:plazi:treatment:03F3878EFFD1FFB563B90000FB1D6F8C:8E856043FFD1FFB2674401B8FB786EE4" box="[1130,1220,981,1007]" county="Great-River village" latitude="35.73997" longLatPrecision="1" longitude="111.03984" municipality="Yi-Cheng county" name="Lin-Fen" pageId="1" pageNumber="336" stateProvince="Shanxi">Lin-Fen</location>
city,
<collectingRegion id="499EF87AFFD1FFB2662801B8FAE86EE4" box="[1286,1364,981,1007]" country="China" name="Shanxi" pageId="1" pageNumber="336">Shanxi</collectingRegion>
Prov- ince, collected by
<collectorName id="26AF534EFFD1FFB262490194FE5E6918" box="[359,482,1017,1043]" pageId="1" pageNumber="336">Chang-Hai</collectorName>
SUN,
<collectorName id="26AF534EFFD1FFB2611F0194FD136918" box="[561,687,1017,1043]" pageId="1" pageNumber="336">Ji-Hua XU</collectorName>
,
<collectorName id="26AF534EFFD1FFB261920194FCB76918" box="[700,779,1017,1043]" pageId="1" pageNumber="336">Zi-Wei</collectorName>
<collectionCode id="ED4BAE5DFFD1FFB2603C0194FCD56918" box="[786,873,1017,1043]" pageId="1" pageNumber="336">WANG</collectionCode>
,
<date id="FFE41058FFD1FFB260580194FC416918" box="[886,1021,1017,1043]" pageId="1" pageNumber="336" value="2013-09-13">
<collectingDate id="EFA0E9B0FFD1FFB260580194FC416918" box="[886,1021,1017,1043]" pageId="1" pageNumber="336" value="2013-09-13">13-IX-2013</collectingDate>
</date>
</materialsCitation>
;
<specimenCount id="9D5CFD11FFD1FFB267240194FBEA6918" box="[1034,1110,1017,1043]" pageId="1" pageNumber="336" type="male">2 male</specimenCount>
imagoes
<materialsCitation id="3B323CC5FFD1FFB267EB0194FD9F6950" collectingDate="2019-08-02" collectionCode="ZHANG" collectorName="Ran LI &amp; Wei" latitude="30.7479" location="Yi-Chang county" longLatPrecision="1" longitude="110.76607" pageId="1" pageNumber="336" specimenCount="1" specimenCount-female="1" stateProvince="Hubei" typeStatus="paratype">
<specimenCount id="9D5CFD11FFD1FFB267EB0194FA9B6918" box="[1221,1319,1017,1043]" pageId="1" pageNumber="336" type="female">1 female</specimenCount>
subimago 2 nymphs, Yang-Lin-Qiao town (
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,
<geoCoordinate id="EE6E505FFFD1FFB261EC0671FCC4693C" box="[706,888,1052,1079]" degrees="110" direction="east" minutes="45" orientation="longitude" pageId="1" pageNumber="336" precision="1" seconds="57.85" value="110.76607">110°4557.85″E</geoCoordinate>
),
<location id="8E856043FFD1FFB260A40670FBF7693C" LSID="urn:lsid:plazi:treatment:03F3878EFFD1FFB563B90000FB1D6F8C:8E856043FFD1FFB260A40670FBF7693C" box="[906,1099,1053,1079]" latitude="30.7479" longLatPrecision="1" longitude="110.76607" name="Yi-Chang county" pageId="1" pageNumber="336" stateProvince="Hubei">Yi-Chang county</location>
,
<collectingRegion id="499EF87AFFD1FFB267790670FAB4693C" box="[1111,1288,1053,1079]" country="China" name="Hubei" pageId="1" pageNumber="336">Hubei Province</collectingRegion>
, collected by
<collectorName id="26AF534EFFD1FFB263B9062CFF546950" box="[151,232,1089,1115]" pageId="1" pageNumber="336">Ran LI</collectorName>
,
<collectorName id="26AF534EFFD1FFB263DD062CFEA36950" box="[243,287,1089,1115]" pageId="1" pageNumber="336">Wei</collectorName>
<collectionCode id="ED4BAE5DFFD1FFB26209062CFE306950" box="[295,396,1089,1115]" pageId="1" pageNumber="336">ZHANG</collectionCode>
,
<date id="FFE41058FFD1FFB262B9062CFD9F6950" box="[407,547,1089,1115]" pageId="1" pageNumber="336" value="2019-08-02">
<collectingDate id="EFA0E9B0FFD1FFB262B9062CFD9F6950" box="[407,547,1089,1115]" pageId="1" pageNumber="336" value="2019-08-02">2-VIII-2019</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3406513FFD1FFB663E90609FBAB6A12" lastPageId="5" lastPageNumber="340" pageId="1" pageNumber="336" type="description">
<paragraph id="8BE53698FFD1FFB263E90609FE6169E0" blockId="1.[151,1437,692,1259]" pageId="1" pageNumber="336">
<emphasis id="B92EEA8AFFD1FFB263E90609FE056974" bold="true" box="[199,441,1124,1151]" pageId="1" pageNumber="336">Description. Nymph</emphasis>
(in alcohol,
<figureCitation id="13612A1DFFD1FFB261680608FD176974" box="[582,683,1125,1151]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1810,1835]" captionStartId-1="2.[151,250,1910,1935]" captionStartId-2="3.[151,250,991,1016]" captionStartId-3="3.[151,250,1428,1453]" captionTargetBox-0="[152,1436,1278,1782]" captionTargetBox-1="[151,1436,885,1885]" captionTargetBox-2="[151,1436,181,965]" captionTargetBox-3="[152,1436,1110,1403]" captionTargetId-0="figure@1.[152,1436,1278,1782]" captionTargetId-1="figure@2.[151,1436,885,1885]" captionTargetId-2="figure@3.[151,1436,181,965]" captionTargetId-3="figure@3.[152,1436,1110,1403]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="3" captionText-0="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." captionText-1="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." captionText-2="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." captionText-3="FIGURE 4. SEM photos of Notacanthurus maculatus sp. n. maxilla:A. Dentisetae enlarged (letters show canines, distal dentiseta and proximal dentiseta respectively); B. Enlarged proximal dentiseta; C. Scattered setae on ventral surface." figureDoi-0="http://doi.org/10.5281/zenodo.3907564" figureDoi-1="http://doi.org/10.5281/zenodo.3907566" figureDoi-2="http://doi.org/10.5281/zenodo.3907568" figureDoi-3="http://doi.org/10.5281/zenodo.3907570" httpUri-0="https://zenodo.org/record/3907564/files/figure.png" httpUri-1="https://zenodo.org/record/3907566/files/figure.png" httpUri-2="https://zenodo.org/record/3907568/files/figure.png" httpUri-3="https://zenodo.org/record/3907570/files/figure.png" pageId="1" pageNumber="336">Figs 14</figureCitation>
): body length 9.013.0 mm, caudal filaments 13.015.0 mm (mean of 14 nymphs). Head width 1.41.8x of length, yellow brown to dark brown. Anterior margin of head not thickened; anterior and lateral margins slightly convex while posterior margin nearly straight; vertex with thin pale stripe above compound eyes (
<figureCitation id="13612A1DFFD1FFB2627B06BCFE2869E0" box="[341,404,1233,1259]" captionStart="FIGURE 1" captionStartId="1.[151,250,1810,1835]" captionTargetBox="[152,1436,1278,1782]" captionTargetId="figure@1.[152,1436,1278,1782]" captionTargetPageId="1" captionText="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." figureDoi="http://doi.org/10.5281/zenodo.3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="1" pageNumber="336">Fig. 1</figureCitation>
AB).
</paragraph>
<caption id="DF256610FFD1FFB263B9057FFD026A44" ID-DOI="http://doi.org/10.5281/zenodo.3907564" ID-Zenodo-Dep="3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="1" pageNumber="336" startId="1.[151,250,1810,1835]" targetBox="[152,1436,1278,1782]" targetPageId="1">
<paragraph id="8BE53698FFD1FFB263B9057FFD026A44" blockId="1.[151,1436,1810,1871]" pageId="1" pageNumber="336">
<emphasis id="B92EEA8AFFD1FFB263B9057FFEAF6A20" bold="true" box="[151,275,1810,1835]" pageId="1" pageNumber="336">FIGURE 1.</emphasis>
Nymph of
<taxonomicName id="4C5A4D1BFFD1FFB262A6057EFD326A20" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[392,654,1811,1835]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="1" pageNumber="336" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD1FFB262A6057EFD326A20" box="[392,654,1811,1835]" italics="true" pageId="1" pageNumber="336">Notacanthurus maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD1FFB261BA057EFD726A20" bold="true" box="[660,718,1811,1835]" pageId="1" pageNumber="336">
<taxonomicNameLabel id="A21D57F1FFD1FFB261BA057EFD726A20" box="[660,718,1811,1835]" pageId="1" pageNumber="336" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen.
</paragraph>
</caption>
<paragraph id="8BE53698FFD1FFB163E90510FA926F10" blockId="1.[151,1436,1916,2016]" lastBlockId="2.[151,1437,153,863]" lastPageId="2" lastPageNumber="337" pageId="1" pageNumber="336">
<emphasis id="B92EEA8AFFD1FFB263E90510FEEA6A9C" bold="true" box="[199,342,1917,1943]" pageId="1" pageNumber="336">Mouthparts</emphasis>
: labrum 0.5x width of head, laterally tapering and slightly curved, relatively long setae on dorsal surface and free margin, ventral surface with shallow median emargination and ventral groove, row of bristles along the margin of groove (
<figureCitation id="13612A1DFFD1FFB262BA05A8FE6A6AD4" box="[404,470,1989,2015]" captionStart="FIGURE 2" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1436,885,1885]" captionTargetId="figure@2.[151,1436,885,1885]" captionTargetPageId="2" captionText="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." figureDoi="http://doi.org/10.5281/zenodo.3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="1" pageNumber="336">Fig. 2</figureCitation>
AB). Both mandibles covered with numerous long setae on outer margins; prostheca with 46 fimbriate bristles; outer incisor of left mandible with serrated margin and one larger terminal denticle; inner one with trifid apex and row of sparse hair-like setae below the molar; outer incisor of right mandible serrated with 2 apical terminal denticles; inner one divided into 3 sharp denticles (
<figureCitation id="13612A1DFFD2FFB16050028CFC7C6DF0" box="[894,960,225,251]" captionStart="FIGURE 2" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1436,885,1885]" captionTargetId="figure@2.[151,1436,885,1885]" captionTargetPageId="2" captionText="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." figureDoi="http://doi.org/10.5281/zenodo.3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="2" pageNumber="337">Fig. 2</figureCitation>
CD, HI). Maxillae with scattered simple setae on ventral surface, with 1921 comb-shaped setae on crown of galea-lacinia, distal dentisetae branched, proximal dentisetae bifid and fimbriate (
<figureCitation id="13612A1DFFD2FFB161330344FDC56C48" box="[541,633,297,323]" captionStart="FIGURE 2" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1436,885,1885]" captionTargetId="figure@2.[151,1436,885,1885]" captionTargetPageId="2" captionText="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." figureDoi="http://doi.org/10.5281/zenodo.3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="2" pageNumber="337">Figs 2G</figureCitation>
,
<figureCitation id="13612A1DFFD2FFB161AA0344FD2E6C48" box="[644,658,297,323]" captionStart="FIGURE 4" captionStartId="3.[151,250,1428,1453]" captionTargetBox="[152,1436,1110,1403]" captionTargetId="figure@3.[152,1436,1110,1403]" captionTargetPageId="3" captionText="FIGURE 4. SEM photos of Notacanthurus maculatus sp. n. maxilla:A. Dentisetae enlarged (letters show canines, distal dentiseta and proximal dentiseta respectively); B. Enlarged proximal dentiseta; C. Scattered setae on ventral surface." figureDoi="http://doi.org/10.5281/zenodo.3907570" httpUri="https://zenodo.org/record/3907570/files/figure.png" pageId="2" pageNumber="337">4</figureCitation>
AC); maxillary palp 3-segmented, basal segment with sporadic setae on outer margin; outer margin of second segment with longer setae and terminal segment with dense setae brush (
<figureCitation id="13612A1DFFD2FFB1638E031CFF466C80" box="[160,250,369,395]" captionStart="FIGURE 2" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1436,885,1885]" captionTargetId="figure@2.[151,1436,885,1885]" captionTargetPageId="2" captionText="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." figureDoi="http://doi.org/10.5281/zenodo.3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="2" pageNumber="337">Fig. 2G</figureCitation>
). Hypopharynx: apex of superlinguae strongly curved and extended into round lobe-like structure, row of hair-like setae on the outer margin from base to apex; lingua bell-like, with tuft of setae on apex (
<figureCitation id="13612A1DFFD2FFB167FD03F8FA966CA4" box="[1235,1322,405,431]" captionStart="FIGURE 2" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1436,885,1885]" captionTargetId="figure@2.[151,1436,885,1885]" captionTargetPageId="2" captionText="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." figureDoi="http://doi.org/10.5281/zenodo.3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="2" pageNumber="337">Fig. 2E</figureCitation>
). Labium: glossae subquadrangular, inner margin of glossae slightly expanded, with tuft of long setae; paraglossae expanded into distinct lobes, with dense setae and bristles on dorsal and free margins; labial palp 2-segmented, ventral surface and free margin of second segment with setae brush, dorsal surface with scattered golden setae (
<figureCitation id="13612A1DFFD2FFB167E5006CFA9D6F10" box="[1227,1313,513,539]" captionStart="FIGURE 2" captionStartId="2.[151,250,1910,1935]" captionTargetBox="[151,1436,885,1885]" captionTargetId="figure@2.[151,1436,885,1885]" captionTargetPageId="2" captionText="FIGURE 2. Mouthparts of Notacanthurus maculatus sp. n. nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible." figureDoi="http://doi.org/10.5281/zenodo.3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="2" pageNumber="337">Fig. 2F</figureCitation>
).
</paragraph>
<paragraph id="8BE53698FFD2FFB163E90048FCA96E54" blockId="2.[151,1437,153,863]" pageId="2" pageNumber="337">
<emphasis id="B92EEA8AFFD2FFB163E90048FE9C6F34" bold="true" box="[199,288,549,575]" pageId="2" pageNumber="337">Thorax</emphasis>
: pronotum extended laterally, slightly wider than head (
<figureCitation id="13612A1DFFD2FFB160B10048FC636F34" box="[927,991,549,575]" captionStart="FIGURE 1" captionStartId="1.[151,250,1810,1835]" captionTargetBox="[152,1436,1278,1782]" captionTargetId="figure@1.[152,1436,1278,1782]" captionTargetPageId="1" captionText="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." figureDoi="http://doi.org/10.5281/zenodo.3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="2" pageNumber="337">Fig. 1</figureCitation>
AB). Supracoxal spurs rounded. Legs yellowish with two brown bands on femora; ventral surface of femora with spots on middle and apex (
<figureCitation id="13612A1DFFD2FFB166370024FAE76F68" box="[1305,1371,585,611]" captionStart="FIGURE 1" captionStartId="1.[151,250,1810,1835]" captionTargetBox="[152,1436,1278,1782]" captionTargetId="figure@1.[152,1436,1278,1782]" captionTargetPageId="1" captionText="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." figureDoi="http://doi.org/10.5281/zenodo.3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="2" pageNumber="337">Figs 1</figureCitation>
AC, 3AC). Femora of all legs with row of long setae on outer margin, dorsal surface with scattered spatulate setae, apex of femora with blunt apical projection (
<figureCitation id="13612A1DFFD2FFB1616300FCFD316FA0" box="[589,653,657,683]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="2" pageNumber="337">Fig. 3</figureCitation>
AC, 3F); foretibiae subequal to femora in length, with rows of spinelike setae on inner and dorsal margins, outer margin with few tiny and short setae on base, foretarsi ca. half length of tibiae (
<figureCitation id="13612A1DFFD2FFB163CB00B4FE836FF8" box="[229,319,729,755]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="2" pageNumber="337">Fig. 3A</figureCitation>
); midlegs similar to forelegs, except tibiae 0.8x length of femora, with dense setae on outer margin, tarsi 1/3 length of tibiae (
<figureCitation id="13612A1DFFD2FFB162820090FDBF6E1C" box="[428,515,765,791]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="2" pageNumber="337">Fig. 3B</figureCitation>
); hindlegs similar to midlegs except tibiae 0.77x length of femora and outer margin with denser setae, hindtarsi with row of hair-like setae on outer margin (
<figureCitation id="13612A1DFFD2FFB16091014CFBBC6E30" box="[959,1024,801,827]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="2" pageNumber="337">Fig. 3</figureCitation>
CD). Claws of all legs similar, with a large basal denticle and 2 subapical denticles (
<figureCitation id="13612A1DFFD2FFB1619F0128FCB46E54" box="[689,776,837,863]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="2" pageNumber="337">Fig. 3E</figureCitation>
).
</paragraph>
<caption id="DF256610FFD2FFB163B9051BFB496ADC" ID-DOI="http://doi.org/10.5281/zenodo.3907566" ID-Zenodo-Dep="3907566" httpUri="https://zenodo.org/record/3907566/files/figure.png" pageId="2" pageNumber="337" startId="2.[151,250,1910,1935]" targetBox="[151,1436,885,1885]" targetPageId="2">
<paragraph id="8BE53698FFD2FFB163B9051BFB496ADC" blockId="2.[151,1437,1910,2007]" pageId="2" pageNumber="337">
<emphasis id="B92EEA8AFFD2FFB163B9051BFEAA6A84" bold="true" box="[151,278,1910,1935]" pageId="2" pageNumber="337">FIGURE 2.</emphasis>
Mouthparts of
<taxonomicName id="4C5A4D1BFFD2FFB16292051AFD786A84" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[444,708,1911,1935]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="337" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD2FFB16292051AFD786A84" box="[444,708,1911,1935]" italics="true" pageId="2" pageNumber="337">Notacanthurus maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD2FFB161E0051AFCB66A84" bold="true" box="[718,778,1911,1935]" pageId="2" pageNumber="337">
<taxonomicNameLabel id="A21D57F1FFD2FFB161E0051AFCB66A84" box="[718,778,1911,1935]" pageId="2" pageNumber="337" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
nymph (digital photos): A. Labrum (ventral view); B. Median bristles of ventral labrum; C. Right mandible; D. Left mandible; E. Hypopharynx (ventral view); F. Labium (ventral view); G. Maxilla (ventral view); H. Incisors and prostheca of right mandible; I. Incisors and prostheca of left mandible.
</paragraph>
</caption>
<caption id="DF256610FFD3FFB063B901B2FEDF6934" ID-DOI="http://doi.org/10.5281/zenodo.3907568" ID-Zenodo-Dep="3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="3" pageNumber="338" startId="3.[151,250,991,1016]" targetBox="[151,1436,181,965]" targetPageId="3">
<paragraph id="8BE53698FFD3FFB063B901B2FEDF6934" blockId="3.[151,1436,991,1088]" pageId="3" pageNumber="338">
<emphasis id="B92EEA8AFFD3FFB063B901B2FEAF6EF3" bold="true" box="[151,275,991,1016]" pageId="3" pageNumber="338">FIGURE 3.</emphasis>
Structures of
<taxonomicName id="4C5A4D1BFFD3FFB0628A018DFD156EF3" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[420,681,992,1016]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD3FFB0628A018DFD156EF3" box="[420,681,992,1016]" italics="true" pageId="3" pageNumber="338">Notacanthurus maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD3FFB0619E018DFD566EF3" bold="true" box="[688,746,992,1016]" pageId="3" pageNumber="338">
<taxonomicNameLabel id="A21D57F1FFD3FFB0619E018DFD566EF3" box="[688,746,992,1016]" pageId="3" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII.
</paragraph>
</caption>
<caption id="DF256610FFD3FFB063B907F9FABE68DA" ID-DOI="http://doi.org/10.5281/zenodo.3907570" ID-Zenodo-Dep="3907570" httpUri="https://zenodo.org/record/3907570/files/figure.png" pageId="3" pageNumber="338" startId="3.[151,250,1428,1453]" targetBox="[152,1436,1110,1403]" targetPageId="3">
<paragraph id="8BE53698FFD3FFB063B907F9FABE68DA" blockId="3.[151,1436,1428,1489]" pageId="3" pageNumber="338">
<emphasis id="B92EEA8AFFD3FFB063B907F9FEAF68A6" bold="true" box="[151,275,1428,1453]" pageId="3" pageNumber="338">FIGURE 4.</emphasis>
SEM photos of
<taxonomicName id="4C5A4D1BFFD3FFB0629907F8FD0068A6" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[439,700,1429,1453]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="338" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD3FFB0629907F8FD0068A6" box="[439,700,1429,1453]" italics="true" pageId="3" pageNumber="338">Notacanthurus maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD3FFB061EC07F8FD4768A6" bold="true" box="[706,763,1429,1453]" pageId="3" pageNumber="338">
<taxonomicNameLabel id="A21D57F1FFD3FFB061EC07F8FD4768A6" box="[706,763,1429,1453]" pageId="3" pageNumber="338" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
maxilla: A. Dentisetae enlarged (letters show canines, distal dentiseta and proximal dentiseta respectively); B. Enlarged proximal dentiseta; C. Scattered setae on ventral surface.
</paragraph>
</caption>
<paragraph id="8BE53698FFD3FFB063E90792FB816A55" blockId="3.[151,1437,1535,2030]" pageId="3" pageNumber="338">
<emphasis id="B92EEA8AFFD3FFB063E90792FE806B11" bold="true" box="[199,316,1535,1562]" pageId="3" pageNumber="338">Abdomen</emphasis>
: abdominal terga IIX each with median black dot on anterior margin and a longitudinal ridge, the ridge protrudes towards posterior of tergum, forming a spine; submedian part of terga IIX each with pair of brown oblique stripes, an additional small pair of pale dots nearby (
<figureCitation id="13612A1DFFD3FFB0606A042AFC716B69" box="[836,973,1607,1634]" captionStart="FIGURE 1" captionStartId="1.[151,250,1810,1835]" captionTargetBox="[152,1436,1278,1782]" captionTargetId="figure@1.[152,1436,1278,1782]" captionTargetPageId="1" captionText="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." figureDoi="http://doi.org/10.5281/zenodo.3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, 1C</figureCitation>
). Each tergum with row of small pointed spines on posterior margin and black stripe close to these spines (
<figureCitation id="13612A1DFFD3FFB060410406FC496B8D" box="[879,1013,1643,1670]" captionStart="FIGURE 1" captionStartId="1.[151,250,1810,1835]" captionTargetBox="[152,1436,1278,1782]" captionTargetId="figure@1.[152,1436,1278,1782]" captionTargetPageId="1" captionText="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." figureDoi="http://doi.org/10.5281/zenodo.3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="3" pageNumber="338">Fig. 1A, 1C</figureCitation>
); posterolateral angles of terga IVIII extended into small acute projections; posterior margin of sternum IX straight to slightly convex (
<figureCitation id="13612A1DFFD3FFB067DF04E2FAF66BA1" box="[1265,1354,1679,1706]" captionStart="FIGURE 1" captionStartId="1.[151,250,1810,1835]" captionTargetBox="[152,1436,1278,1782]" captionTargetId="figure@1.[152,1436,1278,1782]" captionTargetPageId="1" captionText="FIGURE 1. Nymph of Notacanthurus maculatus sp. n. (digital photos): A. Habitus; B. Head capsule and pronotum; C. dorsal view of abdominal terga; D. ventral view of abdomen." figureDoi="http://doi.org/10.5281/zenodo.3907564" httpUri="https://zenodo.org/record/3907564/files/figure.png" pageId="3" pageNumber="338">Fig. 1D</figureCitation>
), some mature nymphs washed with median dark brown stripes on sternum IX. Gills IVI with both lamellae and fibrilliform portions and gill VII with lamellae only; gill I leaf-like, inner margin curved, lamellae longer than fibrilliform portion (
<figureCitation id="13612A1DFFD3FFB063D60496FEE86A1D" box="[248,340,1787,1814]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="3" pageNumber="338">Fig. 3H</figureCitation>
); gills IIVI similar to each other, lamellae broad (
<figureCitation id="13612A1DFFD3FFB060B80496FC646A1D" box="[918,984,1787,1814]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="3" pageNumber="338">Fig. 3</figureCitation>
IJ); gills VVI each with an additional arrow-like accessory lobe (
<figureCitation id="13612A1DFFD3FFB062EA0572FDAE6A31" box="[452,530,1823,1850]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="3" pageNumber="338">Fig. 3J</figureCitation>
); gill VII slightly asymmetrical, with fine marginal setae (
<figureCitation id="13612A1DFFD3FFB067B90572FB4C6A31" box="[1175,1264,1823,1850]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="3" pageNumber="338">Fig. 3K</figureCitation>
). Terminal fila- ment subequal to cerci, all of them with whorls of spines on articulations (
<figureCitation id="13612A1DFFD3FFB060F8052EFB8C6A55" box="[982,1072,1859,1886]" captionStart="FIGURE 3" captionStartId="3.[151,250,991,1016]" captionTargetBox="[151,1436,181,965]" captionTargetId="figure@3.[151,1436,181,965]" captionTargetPageId="3" captionText="FIGURE 3. Structures of Notacanthurus maculatus sp. n. nymph (digital photos): A. Foreleg; B. Midleg; C. Hindleg; D. Part of hind tibiae; E. Foreleg claw; F. Projection of hind femur; G. Segments of caudal filaments near base; H. Gill I; I. Gill II; J. Gill VI; K. Gill VII." figureDoi="http://doi.org/10.5281/zenodo.3907568" httpUri="https://zenodo.org/record/3907568/files/figure.png" pageId="3" pageNumber="338">Fig. 3G</figureCitation>
).
</paragraph>
<paragraph id="8BE53698FFD3FFB763E90505FB826CA4" blockId="3.[151,1437,1535,2030]" lastBlockId="4.[151,1437,152,431]" lastPageId="4" lastPageNumber="339" pageId="3" pageNumber="338">
<emphasis id="B92EEA8AFFD3FFB063E90505FEEA6A89" bold="true" box="[199,342,1895,1922]" pageId="3" pageNumber="338">Male imago</emphasis>
(in alcohol,
<figureCitation id="13612A1DFFD3FFB062C2050AFDE86A89" box="[492,596,1895,1922]" captionStart-0="FIGURE 5" captionStart-1="FIGURE 6" captionStartId-0="4.[151,250,1463,1488]" captionStartId-1="5.[151,250,1648,1673]" captionTargetBox-0="[151,1436,453,1438]" captionTargetBox-1="[151,1436,453,1623]" captionTargetId-0="figure@4.[151,1436,453,1438]" captionTargetId-1="figure@5.[151,1436,453,1623]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionText-0="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." captionText-1="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi-0="http://doi.org/10.5281/zenodo.3907572" figureDoi-1="http://doi.org/10.5281/zenodo.3907574" httpUri-0="https://zenodo.org/record/3907572/files/figure.png" httpUri-1="https://zenodo.org/record/3907574/files/figure.png" pageId="3" pageNumber="338">Figs 56</figureCitation>
): body length 10.012.0 mm, forewing 10.013.0 mm, hindwing 4.05.0 mm, cerci 25.030.0 mm. Body pale to yellowish with brown dots and stripes; anterior margin of head capsule not greatly produced, compound eyes contiguous (
<figureCitation id="13612A1DFFD3FFB061E005C2FCAC6AC1" box="[718,784,1967,1994]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="3" pageNumber="338">Fig. 5</figureCitation>
AB). Mesonotum with apparent transverse suture, lateral parapsidal sutures slightly bent, joining posterior transverse suture (
<figureCitation id="13612A1DFFD3FFB060B805BEFC536AE5" box="[918,1007,2003,2030]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="3" pageNumber="338">Fig. 5B</figureCitation>
). Prosternum relatively flat; mesotho- racic furcasternum parallel (
<figureCitation id="13612A1DFFD4FFB762FD02F4FD976DB8" box="[467,555,152,179]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="4" pageNumber="339">Fig. 5C</figureCitation>
). Femora of all legs pale, with 3 brown bands; tibiae and tarsi pale with brown apex (
<figureCitation id="13612A1DFFD4FFB763F302D0FE856DDC" box="[221,313,188,215]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="4" pageNumber="339">Fig. 5A</figureCitation>
). Femora: tibiae: tarsi of forelegs=3.0: 4.0: 6.4, tarsal segments from basal to apical=1.2: 2.0: 1.8: 1.0: 0.4; midlegs and hindlegs similar to forelegs except femora: tibiae: tarsi=3.2: 2.5: 1.4, tarsal segments arranged in decreasing order=5, 1, 2, 3, 4. Forewing hyaline, crossveins pigmented dark brown between C, Sc, R
<subScript id="17DE34DDFFD4FFB76617037BFAFD6C2F" attach="left" box="[1337,1345,278,292]" fontSize="6" pageId="4" pageNumber="339">1</subScript>
and Rs; several crowded crossveins between Sc, R
<subScript id="17DE34DDFFD4FFB761400357FDCA6C43" attach="left" box="[622,630,314,328]" fontSize="6" pageId="4" pageNumber="339">1</subScript>
and Rs; Rs and MP forked sub-equidistant from base of wing to margin, MA forked over 1/2 of distance from base of wing to margin (
<figureCitation id="13612A1DFFD4FFB760480320FC7E6C63" box="[870,962,333,360]" captionStart="FIGURE 6" captionStartId="5.[151,250,1648,1673]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure@5.[151,1436,453,1623]" captionTargetPageId="5" captionText="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="4" pageNumber="339">Fig. 6A</figureCitation>
). Hindwing transparent, with basal costal projection, MA forked near middle but MP forked basally (
<figureCitation id="13612A1DFFD4FFB76015031CFC2A6C87" box="[827,918,369,396]" captionStart="FIGURE 6" captionStartId="5.[151,250,1648,1673]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure@5.[151,1436,453,1623]" captionTargetPageId="5" captionText="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="4" pageNumber="339">Fig. 6B</figureCitation>
). Each tergum of abdomen IIX with median brown dots on anterior margin and pair of oblique brown stripes laterally (
<figureCitation id="13612A1DFFD4FFB760F903F8FB8D6CA4" box="[983,1073,404,431]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="4" pageNumber="339">Fig. 5D</figureCitation>
).
</paragraph>
<caption id="DF256610FFD4FFB763B907DAFF6D6B13" ID-DOI="http://doi.org/10.5281/zenodo.3907572" ID-Zenodo-Dep="3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="4" pageNumber="339" startId="4.[151,250,1463,1488]" targetBox="[151,1436,453,1438]" targetPageId="4">
<paragraph id="8BE53698FFD4FFB763B907DAFF6D6B13" blockId="4.[151,1437,1463,1560]" pageId="4" pageNumber="339">
<emphasis id="B92EEA8AFFD4FFB763B907DAFEA968C4" bold="true" box="[151,277,1463,1488]" pageId="4" pageNumber="339">FIGURE 5.</emphasis>
Adults of
<taxonomicName id="4C5A4D1BFFD4FFB762A907DAFD3268C4" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[391,654,1463,1487]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="339" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD4FFB762A907DAFD3268C4" box="[391,654,1463,1487]" italics="true" pageId="4" pageNumber="339">Notacanthurus maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD4FFB761B907DAFD6E68C4" bold="true" box="[663,722,1463,1487]" pageId="4" pageNumber="339">
<taxonomicNameLabel id="A21D57F1FFD4FFB761B907DAFD6E68C4" box="[663,722,1463,1487]" pageId="4" pageNumber="339" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult.
</paragraph>
</caption>
<paragraph id="8BE53698FFD4FFB763E90428FB636BC7" blockId="4.[151,1437,1605,2028]" pageId="4" pageNumber="339">
<emphasis id="B92EEA8AFFD4FFB763E90428FE896B6B" bold="true" box="[199,309,1605,1632]" pageId="4" pageNumber="339">Genitalia</emphasis>
: styliger plate almost straight with 2 small lateral projections (
<figureCitation id="13612A1DFFD4FFB760D1042BFBE66B6A" box="[1023,1114,1606,1633]" captionStart="FIGURE 6" captionStartId="5.[151,250,1648,1673]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure@5.[151,1436,453,1623]" captionTargetPageId="5" captionText="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="4" pageNumber="339">Fig. 6D</figureCitation>
). Forceps segments 34 less than half length of segment 2, segment 4 oval, slightly shorter than segment 3. Penes fused but with apical incision and distinct brown spermatic duct on ventral surface, stem base of penes sclerotized laterally; with median spinelike titillators subapically and mesally (
<figureCitation id="13612A1DFFD4FFB7617F04DFFD2F6BC6" box="[593,659,1714,1741]" captionStart="FIGURE 6" captionStartId="5.[151,250,1648,1673]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure@5.[151,1436,453,1623]" captionTargetPageId="5" captionText="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="4" pageNumber="339">Fig. 6</figureCitation>
DE). Cerci pale with brown articulations (
<figureCitation id="13612A1DFFD4FFB7675604DFFB6E6BC7" box="[1144,1234,1713,1740]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="4" pageNumber="339">Fig. 5A</figureCitation>
).
</paragraph>
<paragraph id="8BE53698FFD4FFB763E904BBFB4F6A57" blockId="4.[151,1437,1605,2028]" pageId="4" pageNumber="339">
<emphasis id="B92EEA8AFFD4FFB763E904BBFEC16BFB" bold="true" box="[199,381,1749,1776]" pageId="4" pageNumber="339">Male subimago</emphasis>
(in alcohol): body length 10.012.0 mm, forewing 11.012.0 mm, hindwing 4.05.0 mm, caudal filaments 9.012.0 mm. Resembles the male imago except dull. Femora: tibiae: tarsi of forelegs=3.0: 2.8: 2.4, tarsal segments arranged in decreasing order=2, 3, 1, 4, 5; those of midlegs=3.0: 2.6: 1.2, tarsal segments arranged in decreasing order=5, 1, 2, 3, 4; those of hindlegs=3.2: 3.6: 1.2, tarsal segments similar to midlegs.
</paragraph>
<paragraph id="8BE53698FFD4FFB663E9050BFEC56DF0" blockId="4.[151,1437,1605,2028]" lastBlockId="5.[151,1437,152,431]" lastPageId="5" lastPageNumber="340" pageId="4" pageNumber="339">
<emphasis id="B92EEA8AFFD4FFB763E9050BFED16A8B" bold="true" box="[199,365,1893,1920]" pageId="4" pageNumber="339">Female imago</emphasis>
(in alcohol,
<figureCitation id="13612A1DFFD4FFB762D6050BFDEC6A8B" box="[504,592,1893,1920]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="4" pageNumber="339">Figs 5E</figureCitation>
,
<figureCitation id="13612A1DFFD4FFB76175050BFDC36A8A" box="[603,639,1894,1921]" captionStart="FIGURE 6" captionStartId="5.[151,250,1648,1673]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure@5.[151,1436,453,1623]" captionTargetPageId="5" captionText="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="4" pageNumber="339">6C</figureCitation>
): body length 10.014.0 mm, caudal filaments 18.027.0 mm, forewing 12.015.0 mm, hindwing 4.05.0 mm. Similar to male imago except as follows: lengths of femora: tibiae: tarsi of forelegs=3.0: 3.2: 2.2, tarsal segments arranged in decreasing order=2, 3, 1, 4, 5; those of midlegs=3.6: 3.0: 1.4, length order of tarsal segments in decreasing order=5, 1, 2, 3, 4; those of hindlegs=4.0: 3.0: 1.4, tarsal segments similar to midlegs. Lateral stripes on abdomen terga reddish and much broader (
<figureCitation id="13612A1DFFD5FFB6670802F4FBC26DB8" box="[1062,1150,152,179]" captionStart="FIGURE 5" captionStartId="4.[151,250,1463,1488]" captionTargetBox="[151,1436,453,1438]" captionTargetId="figure@4.[151,1436,453,1438]" captionTargetPageId="4" captionText="FIGURE 5. Adults of Notacanthurus maculatus sp. n. (digital photos): A. Male adult; B. Transverse suture of mesonotum; C. Medial depression of furcasternum (shown in red arrow); D. Abdominal terga of male adult; E. Abdominal terga of female adult." figureDoi="http://doi.org/10.5281/zenodo.3907572" httpUri="https://zenodo.org/record/3907572/files/figure.png" pageId="5" pageNumber="340">Fig. 5E</figureCitation>
). Subgenital plate almost close to posterior margin of sternum VIII; subanal plate with convex posterior margin (
<figureCitation id="13612A1DFFD5FFB6675202D0FB6A6DD3" box="[1148,1238,189,216]" captionStart="FIGURE 6" captionStartId="5.[151,250,1648,1673]" captionTargetBox="[151,1436,453,1623]" captionTargetId="figure@5.[151,1436,453,1623]" captionTargetPageId="5" captionText="FIGURE 6. Male and female structures of Notacanthurus maculatus sp. n. (digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view)." figureDoi="http://doi.org/10.5281/zenodo.3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="5" pageNumber="340">Fig. 6C</figureCitation>
). Articulations of cerci brown to dark.
</paragraph>
<paragraph id="8BE53698FFD5FFB663E90368FF706CA4" blockId="5.[151,1437,152,431]" pageId="5" pageNumber="340">
<emphasis id="B92EEA8AFFD5FFB663E90368FE266C14" bold="true" box="[199,410,260,287]" pageId="5" pageNumber="340">Female subimago</emphasis>
(in alcohol): body length 10.012.0 mm, forewing 12.014.0 mm, hindwing 4.05.0 mm, caudal filaments 10.015.0 mm. Color pattern resembles female imago except dull. Femora: tibiae: tarsi of forelegs=3.0: 2.6: 2.4, tarsal segments arranged in decreasing order=2, 3, 5, 1, 4; those of midlegs=3.0: 2.4: 1.2, length order of tarsal segments in decreasing=5, 1, 2, 3, 4; those of hindlegs=3.2: 2.8: 1.2, tarsal segments similar to midlegs.
</paragraph>
<caption id="DF256610FFD5FFB663B9041DFAEA6BA6" ID-DOI="http://doi.org/10.5281/zenodo.3907574" ID-Zenodo-Dep="3907574" httpUri="https://zenodo.org/record/3907574/files/figure.png" pageId="5" pageNumber="340" startId="5.[151,250,1648,1673]" targetBox="[151,1436,453,1623]" targetPageId="5">
<paragraph id="8BE53698FFD5FFB663B9041DFAEA6BA6" blockId="5.[151,1436,1648,1709]" pageId="5" pageNumber="340">
<emphasis id="B92EEA8AFFD5FFB663B9041DFEA96B82" bold="true" box="[151,277,1648,1673]" pageId="5" pageNumber="340">FIGURE 6.</emphasis>
Male and female structures of
<taxonomicName id="4C5A4D1BFFD5FFB66174041CFCDD6B82" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[602,865,1649,1673]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD5FFB66174041CFCDD6B82" box="[602,865,1649,1673]" italics="true" pageId="5" pageNumber="340">Notacanthurus maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD5FFB66047041CFC186B82" bold="true" box="[873,932,1649,1673]" pageId="5" pageNumber="340">
<taxonomicNameLabel id="A21D57F1FFD5FFB66047041CFC186B82" box="[873,932,1649,1673]" pageId="5" pageNumber="340" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
(digital photos): A. Forewing of male imago; B. Hindwing of male imago; C. Abdominal sterna of female; D. Genitalia (ventral view); E. Penes enlarged (ventral view).
</paragraph>
</caption>
<paragraph id="8BE53698FFD5FFB663E904B6FBAB6A12" blockId="5.[151,1436,1755,2033]" pageId="5" pageNumber="340">
<emphasis id="B92EEA8AFFD5FFB663E904B6FF4A6BFE" bold="true" box="[199,246,1755,1781]" pageId="5" pageNumber="340">Egg</emphasis>
(
<figureCitation id="13612A1DFFD5FFB6622804B6FEF76BFE" box="[262,331,1755,1781]" captionStart="FIGURE 7" captionStartId="6.[151,250,1191,1216]" captionTargetBox="[152,1436,669,1166]" captionTargetId="figure@6.[152,1436,669,1166]" captionTargetPageId="6" captionText="FIGURE 7. Egg of Notacanthurus maculatus sp. n.: A. Whole picture; B. Micropyle enlarged." figureDoi="http://doi.org/10.5281/zenodo.3907576" httpUri="https://zenodo.org/record/3907576/files/figure.png" pageId="5" pageNumber="340">Fig. 7</figureCitation>
): Egg ovoid, chorion decorated with irregular tubercle-like projections and small knob-terminated coiled threads (KCTs) on polar (
<figureCitation id="13612A1DFFD5FFB6612F0492FDE76A12" box="[513,603,1791,1817]" captionStart="FIGURE 7" captionStartId="6.[151,250,1191,1216]" captionTargetBox="[152,1436,669,1166]" captionTargetId="figure@6.[152,1436,669,1166]" captionTargetPageId="6" captionText="FIGURE 7. Egg of Notacanthurus maculatus sp. n.: A. Whole picture; B. Micropyle enlarged." figureDoi="http://doi.org/10.5281/zenodo.3907576" httpUri="https://zenodo.org/record/3907576/files/figure.png" pageId="5" pageNumber="340">Fig. 7A</figureCitation>
). Micropyle round to oval (
<figureCitation id="13612A1DFFD5FFB660A10492FC6D6A12" box="[911,977,1791,1817]" captionStart="FIGURE 7" captionStartId="6.[151,250,1191,1216]" captionTargetBox="[152,1436,669,1166]" captionTargetId="figure@6.[152,1436,669,1166]" captionTargetPageId="6" captionText="FIGURE 7. Egg of Notacanthurus maculatus sp. n.: A. Whole picture; B. Micropyle enlarged." figureDoi="http://doi.org/10.5281/zenodo.3907576" httpUri="https://zenodo.org/record/3907576/files/figure.png" pageId="5" pageNumber="340">Fig. 7</figureCitation>
AB).
</paragraph>
</subSubSection>
<subSubSection id="C3406513FFD5FFB663E9054EFCBF6A6A" pageId="5" pageNumber="340" type="etymology">
<paragraph id="8BE53698FFD5FFB663E9054EFCBF6A6A" blockId="5.[151,1436,1755,2033]" pageId="5" pageNumber="340">
<emphasis id="B92EEA8AFFD5FFB663E9054EFEF46A36" bold="true" box="[199,328,1827,1853]" pageId="5" pageNumber="340">Etymology</emphasis>
: The Latin specific name
<taxonomicName id="4C5A4D1BFFD5FFB66145054EFD626A36" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[619,734,1827,1853]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="340" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B92EEA8AFFD5FFB66145054EFD626A36" box="[619,734,1827,1853]" italics="true" pageId="5" pageNumber="340">maculatus</emphasis>
</taxonomicName>
means marks, spots or having spots. It indicates the nymphs of this species having more spots on both sides of femora.
</paragraph>
</subSubSection>
<subSubSection id="C3406513FFD5FFB563E90507FB1D6F8C" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340" type="diagnosis">
<paragraph id="8BE53698FFD5FFB563E90507FAB46DB8" blockId="5.[151,1436,1755,2033]" lastBlockId="6.[151,1437,153,647]" lastPageId="6" lastPageNumber="341" pageId="5" pageNumber="340">
<emphasis id="B92EEA8AFFD5FFB663E90507FE866A8E" bold="true" box="[199,314,1898,1925]" pageId="5" pageNumber="340">Diagnosis</emphasis>
. This species can be distinguished from others
<taxonomicName id="4C5A4D1BFFD5FFB660660506FC526A8E" box="[840,1006,1899,1925]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="5" pageNumber="340" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEA8AFFD5FFB660660506FC526A8E" box="[840,1006,1899,1925]" italics="true" pageId="5" pageNumber="340">Notacanthurus</emphasis>
</taxonomicName>
by the following combination of characters. In nymph: 1) femora of all legs with ventral spots; 2) tibiae of mid- and hindleg with rows of spine-like setae on dorsal and inner margins; 3) claws with 2 subapical denticles; 4) gills VVI with additional arrow-like lobes. In imagoes: 1) round penis lobes with visible spermatic duct and basal sclerites on ventral surface; 2) anterior margin of head not expanded; 3) posterior margin of subgenital plate almost reaching the tip of sternum VIII.
</paragraph>
<paragraph id="8BE53698FFD6FFB563E902D0FE4D6F10" blockId="6.[151,1437,153,647]" pageId="6" pageNumber="341">
In the seven known species of the genus
<taxonomicName id="4C5A4D1BFFD6FFB561A402D0FC8C6DDC" box="[650,816,189,215]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="genus">
<emphasis id="B92EEA8AFFD6FFB561A402D0FC8C6DDC" box="[650,816,189,215]" italics="true" pageId="6" pageNumber="341">Notacanthurus</emphasis>
</taxonomicName>
, the nymph of
<taxonomicName id="4C5A4D1BFFD6FFB560F902D3FBD66DDC" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[983,1130,189,215]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD6FFB560F902D3FBD66DDC" box="[983,1130,189,215]" italics="true" pageId="6" pageNumber="341">N. maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD6FFB5675E02D0FB136DDC" bold="true" box="[1136,1199,189,215]" pageId="6" pageNumber="341">
<taxonomicNameLabel id="A21D57F1FFD6FFB5675E02D0FB136DDC" box="[1136,1199,189,215]" pageId="6" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
appears to be closely related to
<taxonomicName id="4C5A4D1BFFD6FFB56225028FFD016DF0" authority="Braasch &amp; Boonsoong (2009)" authorityName="Braasch &amp; Boonsoong" authorityYear="2009" box="[267,701,225,251]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="baei">
<emphasis id="B92EEA8AFFD6FFB56225028FFEE26DF0" box="[267,350,225,251]" italics="true" pageId="6" pageNumber="341">N. baei</emphasis>
<bibRefCitation id="EFCB4B69FFD6FFB56249028CFD016DF0" author="Braasch, D. &amp; Boonsoong, B." box="[359,701,225,251]" pageId="6" pageNumber="341" pagination="33 - 44" refId="ref7464" refString="Braasch, D. &amp; Boonsoong, B. (2009) A new Notacanthurus Tshernova, 1974 and a new Rhithrogena Eaton, 1881 (subgenus Tumungula Zhou &amp; Peters, 2004) from Thailand (Heptageniidae, Ephemeroptera). Zootaxa, 2166 (1), 33 - 44. https: // doi. org / 10.11646 / zootaxa. 2166.1.3" type="journal article" year="2009">Braasch &amp; Boonsoong (2009)</bibRefCitation>
</taxonomicName>
because of their similar color patterns of abdominal terga, head capsule with nearly straight posterior margins, gills VVI with accessory lobes and sternum IX with prospective median dark stripe. However, compared to
<taxonomicName id="4C5A4D1BFFD6FFB561590347FD7B6C48" authorityName="Braasch &amp; Boonsoong" authorityYear="2009" box="[631,711,297,323]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="baei">
<emphasis id="B92EEA8AFFD6FFB561590347FD7B6C48" box="[631,711,297,323]" italics="true" pageId="6" pageNumber="341">N. baei</emphasis>
</taxonomicName>
, the nymph of
<taxonomicName id="4C5A4D1BFFD6FFB560410347FBBF6C48" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[879,1027,297,323]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD6FFB560410347FBBF6C48" box="[879,1027,297,323]" italics="true" pageId="6" pageNumber="341">N. maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD6FFB567270344FBF46C48" bold="true" box="[1033,1096,297,323]" pageId="6" pageNumber="341">
<taxonomicNameLabel id="A21D57F1FFD6FFB567270344FBF46C48" box="[1033,1096,297,323]" pageId="6" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has larger body size (9.013.0 mm), head width more than 1.4 of length (1.41.8), a thin white stripe above compound eyes, pronotum slightly wider than head, posterior margin of sternum IX nearly straight. In contrast, the nymph of
<taxonomicName id="4C5A4D1BFFD6FFB56792031FFAAC6C80" authorityName="Braasch &amp; Boonsoong" authorityYear="2009" box="[1212,1296,369,395]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="baei">
<emphasis id="B92EEA8AFFD6FFB56792031FFAAC6C80" box="[1212,1296,369,395]" italics="true" pageId="6" pageNumber="341">N. baei</emphasis>
</taxonomicName>
has smaller size (8.09.0 mm), head width 1.4 of length, head capsule with irregular pale yellow markings, pronotum slightly smaller than head, posterior margin of sternum IX with notch. In addition, the gills shape of them are also different;
<taxonomicName id="4C5A4D1BFFD6FFB563E603B3FEE76CFC" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[200,347,477,503]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD6FFB563E603B3FEE76CFC" box="[200,347,477,503]" italics="true" pageId="6" pageNumber="341">N. maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD6FFB5624C03B0FE1D6CFC" bold="true" box="[354,417,477,503]" pageId="6" pageNumber="341">
<taxonomicNameLabel id="A21D57F1FFD6FFB5624C03B0FE1D6CFC" box="[354,417,477,503]" pageId="6" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
has gill I leaf-like, lamellae of gills IIVII broad while
<taxonomicName id="4C5A4D1BFFD6FFB5672003B3FBE36CFC" authorityName="Braasch &amp; Boonsoong" authorityYear="2009" box="[1038,1119,477,503]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="baei">
<emphasis id="B92EEA8AFFD6FFB5672003B3FBE36CFC" box="[1038,1119,477,503]" italics="true" pageId="6" pageNumber="341">N. baei</emphasis>
</taxonomicName>
has gill I knife-shaped, gills IIVII with narrower lamellae.
</paragraph>
<paragraph id="8BE53698FFD6FFB563E90048FB1D6F8C" blockId="6.[151,1437,153,647]" pageId="6" pageNumber="341">
In adults, male and female imagoes of
<taxonomicName id="4C5A4D1BFFD6FFB56158004BFCB66F34" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[630,778,549,575]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD6FFB56158004BFCB66F34" box="[630,778,549,575]" italics="true" pageId="6" pageNumber="341">N. maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD6FFB5603E0048FCEC6F34" bold="true" box="[784,848,549,575]" pageId="6" pageNumber="341">
<taxonomicNameLabel id="A21D57F1FFD6FFB5603E0048FCEC6F34" box="[784,848,549,575]" pageId="6" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
have pigmented crossveins in fields of C, Sc, R
<subScript id="17DE34DDFFD6FFB5664B005BFAD16F4F" attach="left" box="[1381,1389,566,580]" fontSize="6" pageId="6" pageNumber="341">1</subScript>
and Rs. In contrast to other congeners (
<taxonomicName id="4C5A4D1BFFD6FFB5610E0027FD1D6F68" baseAuthorityName="Braasch" baseAuthorityYear="1980" box="[544,673,586,611]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="cristatus">
<emphasis id="B92EEA8AFFD6FFB5610E0027FD1D6F68" box="[544,673,586,611]" italics="true" pageId="6" pageNumber="341">N. cristatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5A4D1BFFD6FFB561810027FCE86F68" authorityName="Braasch" authorityYear="1986" box="[687,852,585,611]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="ladakhensis">
<emphasis id="B92EEA8AFFD6FFB561810027FCE86F68" box="[687,852,585,611]" italics="true" pageId="6" pageNumber="341">N. ladakhensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5A4D1BFFD6FFB560A40027FBA76F68" baseAuthorityName="Tshernova" baseAuthorityYear="1972" box="[906,1051,585,611]" class="Insecta" family="Heptageniidae" genus="Ecdyonurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="zhiltzovae">
<emphasis id="B92EEA8AFFD6FFB560A40027FBA76F68" box="[906,1051,585,611]" italics="true" pageId="6" pageNumber="341">N. zhiltzovae</emphasis>
</taxonomicName>
),
<taxonomicName id="4C5A4D1BFFD6FFB5671C0027FB7A6F68" authority="Zhang &amp; Zhang &amp; Han &amp; Zhou, 2020" authorityName="Zhang &amp; Zhang &amp; Han &amp; Zhou" authorityYear="2020" box="[1074,1222,585,611]" class="Insecta" family="Heptageniidae" genus="Notacanthurus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="6" pageNumber="341" phylum="Arthropoda" rank="species" species="maculatus" status="sp. nov.">
<emphasis id="B92EEA8AFFD6FFB5671C0027FB7A6F68" box="[1074,1222,585,611]" italics="true" pageId="6" pageNumber="341">N. maculatus</emphasis>
</taxonomicName>
<emphasis id="B92EEA8AFFD6FFB567E30024FAB16F68" bold="true" box="[1229,1293,585,611]" pageId="6" pageNumber="341">
<taxonomicNameLabel id="A21D57F1FFD6FFB567E30024FAB16F68" box="[1229,1293,585,611]" pageId="6" pageNumber="341" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is unique by having unexpanded anterior margin of head capsule and simple teardrop shaped penis lobes.
</paragraph>
</subSubSection>
</treatment>
</document>