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<document id="74FCC3737FE66C6BF8D8A62AA25D85E6" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2019" docId="03F32A40FF90B14C51BB9F69F561FA29" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion katatonos Hyman &amp; Köhler 2019, sp. nov." docType="treatment" docVersion="5" lastPageNumber="427" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="424" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="A1DF46C1FE332AA28EC96F6F7434AE2B">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="EB6722B00210FB78AAD76E6A6C95D85B">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="62BB86DC3A7DAD758A22ADBEAC8A4FF4">Köhler, Frank</mods:namePart>
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<mods:title id="5E08678539A2A0795ECF547D1619350A">Contributions to Zoology</mods:title>
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<mods:date id="8CF15B42031EE1065BF013B8F86FDCB6">2019</mods:date>
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<mods:number id="861D691D5019E2A49CBB1D792DF324FF">2019-10-03</mods:number>
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<treatment id="03F32A40FF90B14C51BB9F69F561FA29" LSID="urn:lsid:plazi:treatment:03F32A40FF90B14C51BB9F69F561FA29" httpUri="http://treatment.plazi.org/id/03F32A40FF90B14C51BB9F69F561FA29" lastPageId="76" lastPageNumber="427" pageId="73" pageNumber="424">
<subSubSection id="C340C8DDFF90B14951BB9F69F7ABFC6F" box="[641,988,889,916]" pageId="73" pageNumber="424" type="nomenclature">
<paragraph id="8BE59B56FF90B14951BB9F69F7ABFC6F" blockId="73.[641,988,889,916]" box="[641,988,889,916]" pageId="73" pageNumber="424">
<heading id="D0AD2C3AFF90B14951BB9F69F7ABFC6F" box="[641,988,889,916]" fontSize="11" level="2" pageId="73" pageNumber="424" reason="2">
<emphasis id="B92E4744FF90B14951BB9F69F7ABFC6F" bold="true" box="[641,988,889,916]" pageId="73" pageNumber="424">
<taxonomicName id="4C5AE0D5FF90B14951BB9F69F7F5FC6F" ID-CoL="7RBRK" ID-ENA="2658025" authority="Hyman &amp; Köhler, 2019" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[641,898,889,916]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="katatonos" status="sp. nov.">
<emphasis id="B92E4744FF90B14951BB9F69F7F5FC6F" bold="true" box="[641,898,889,916]" italics="true" pageId="73" pageNumber="424">Fastosarion katatonos</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21DFA3FFF90B14950BD9F6AF7ABFC6F" box="[903,988,890,916]" pageId="73" pageNumber="424" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF90B14951BC9F8DF75EFC4C" box="[646,809,925,951]" pageId="73" pageNumber="424" type="description">
<paragraph id="8BE59B56FF90B14951BC9F8DF75EFC4C" blockId="73.[646,809,925,951]" box="[646,809,925,951]" pageId="73" pageNumber="424">
<figureCitation id="136187D3FF90B14951BC9F8DF6A1FC4C" box="[646,726,925,951]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="73" pageNumber="424">Figs. 18</figureCitation>
,
<figureCitation id="136187D3FF90B14951DA9F8DF772FC4C" box="[736,773,925,951]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="73" pageNumber="424">19F</figureCitation>
,
<figureCitation id="136187D3FF90B14950349F8DF75EFC4C" box="[782,809,925,951]" captionStart="FIGURE 26" captionStartId="77.[98,171,1660,1681]" captionTargetBox="[98,915,186,1627]" captionTargetId="figure-35@77.[98,915,186,1627]" captionTargetPageId="77" captionText="FIGURE 26 Genitalia of F.katatonos, MO85837, Mt Archer.A. Reproductive system.Arrow shows the junction of the penis and epiphallus.B. Penis interior. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8343113" httpUri="https://zenodo.org/record/8343113/files/figure.png" pageId="73" pageNumber="424">26</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF90B14951BA9FF5F714FB90" pageId="73" pageNumber="424" type="etymology">
<paragraph id="8BE59B56FF90B14951BA9FF5F68FFC04" blockId="73.[640,1156,997,1131]" box="[640,760,997,1023]" pageId="73" pageNumber="424">Etymology</paragraph>
<paragraph id="8BE59B56FF90B14951BC981AF714FB90" blockId="73.[640,1156,997,1131]" pageId="73" pageNumber="424">
From
<taxonomicName id="4C5AE0D5FF90B14951F79819F74EFBD8" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[717,825,1033,1059]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="73" pageNumber="424" phylum="Mollusca" rank="species" species="katatonos">
<emphasis id="B92E4744FF90B14951F79819F74EFBD8" box="[717,825,1033,1059]" italics="true" pageId="73" pageNumber="424">katatonos</emphasis>
</taxonomicName>
(Greek: broader than high), referring to the wide aperture of the shell; noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF90B14951BA9889F798FA55" pageId="73" pageNumber="424" type="materials_examined">
<paragraph id="8BE59B56FF90B14951BA9889F723FB48" blockId="73.[640,1159,1177,1454]" box="[640,852,1177,1203]" pageId="73" pageNumber="424">Material examined</paragraph>
<paragraph id="8BE59B56FF90B14951BC98ADF7AEFAB8" blockId="73.[640,1159,1177,1454]" pageId="73" pageNumber="424">
<emphasis id="B92E4744FF90B14951BC98ADF6B2FB2D" box="[646,709,1213,1238]" italics="true" pageId="73" pageNumber="424">Types</emphasis>
:
<materialsCitation id="3B32910BFF90B14951EC98ACF7AEFAB8" collectingDate="1994-07-25" collectionCode="QM" collectorName="J. Stanisic &amp; D. Potter &amp; G. Ingram &amp; C. Eddie" latitude="-23.344444" location="Rockhampton" longLatPrecision="21" longitude="150.58055" municipality="Mt Archer" pageId="73" pageNumber="424" specimenCode="MO85837" specimenCount="1" typeStatus="holotype">
<typeStatus id="54E125F4FF90B14951EC98ACF736FB2D" box="[726,833,1212,1238]" pageId="73" pageNumber="424">Holotype</typeStatus>
:
<collectionCode id="ED4B0393FF90B149507498ADF70EFB2C" box="[846,889,1213,1239]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF90B14950BE98ACF798FB2C" box="[900,1007,1212,1239]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">MO85837</specimenCode>
(
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, c.
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NE at
<collectingMunicipality id="6B81012CFF90B149506C98F1F7BCFB01" box="[854,971,1248,1275]" pageId="73" pageNumber="424">Mt Archer</collectingMunicipality>
, nr base,
<geoCoordinate id="EE6EFD91FF90B149570398F1F6C8FAE5" degrees="23" direction="south" minutes="20" orientation="latitude" pageId="73" pageNumber="424" precision="15" seconds="40" value="-23.344444">23° 20' 40&quot; S</geoCoordinate>
,
<geoCoordinate id="EE6EFD91FF90B14951F09914F72CFAE4" box="[714,859,1284,1311]" degrees="150" direction="east" minutes="34" orientation="longitude" pageId="73" pageNumber="424" precision="15" seconds="50" value="150.58055">150° 34' 50&quot; E</geoCoordinate>
, coll.
<date id="FFE4BD96FF90B14950A29914F079FAE5" box="[920,1038,1284,1311]" pageId="73" pageNumber="424" value="1994-07-25">
<collectingDate id="EFA0447EFF90B14950A29914F079FAE5" box="[920,1038,1284,1311]" pageId="73" pageNumber="424" value="1994-07-25">25 Jul 1994</collectingDate>
</date>
,
<collectorName id="26AFFE80FF90B149572D9915F0F4FAE5" box="[1047,1155,1284,1311]" pageId="73" pageNumber="424">J. Stanisic</collectorName>
,
<collectorName id="26AFFE80FF90B14951BC9939F690FAB8" box="[646,743,1321,1347]" pageId="73" pageNumber="424">D. Potter</collectorName>
,
<collectorName id="26AFFE80FF90B14951CB9938F715FAB8" box="[753,866,1320,1347]" pageId="73" pageNumber="424">G. Ingram</collectorName>
,
<collectorName id="26AFFE80FF90B14950569938F7BAFAB8" box="[876,973,1320,1347]" pageId="73" pageNumber="424">C. Eddie</collectorName>
).
</materialsCitation>
</paragraph>
<paragraph id="8BE59B56FF90B149519F995DF68DFA70" blockId="73.[640,1159,1177,1454]" pageId="73" pageNumber="424">
<materialsCitation id="3B32910BFF90B149519F995DF681FA70" collectingDate="1994-07-25" collectionCode="QM" collectorName="J. Stanisic &amp; D. Potter &amp; G. Ingram &amp; C. Eddie" latitude="-23.344444" location="Rockhampton" longLatPrecision="21" longitude="150.58055" municipality="Mt Archer" pageId="73" pageNumber="424" specimenCode="MO54515" specimenCount="1" typeStatus="paratype">
<typeStatus id="54E125F4FF90B149519F995DF763FA9C" box="[677,788,1357,1383]" pageId="73" pageNumber="424">Paratypes</typeStatus>
:
<collectionCode id="ED4B0393FF90B149501D995CF725FA9D" box="[807,850,1356,1382]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF90B1495058995CF7BFFA9C" box="[866,968,1356,1383]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="73" pageNumber="424" type="Museum">MO54515</specimenCode>
(same data as holotype)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE59B56FF90B149519F9985F798FA55" blockId="73.[640,1159,1177,1454]" box="[677,1007,1428,1454]" pageId="73" pageNumber="424">
<emphasis id="B92E4744FF90B149519F9985F707FA55" box="[677,880,1428,1454]" italics="true" pageId="73" pageNumber="424">Non-type material:</emphasis>
See
<tableCitation id="C6D8AEEDFF90B14950989984F79CFA55" box="[930,1003,1428,1454]" captionStart="TABLE 1" captionStartId="3.[56,116,402,428]" captionTargetBox="[56,1638,477,1528]" captionTargetPageId="3" captionText="TABLE 1 Material examined. Abbreviations: AM, Australian Museum, Sydney; H, holotype; NMV,Museums Victoria, Melbourne; P, Paratype; QM, Queensland Museum, Brisbane; S, syntype; NHRM, Naturhistoriska Riksmuseet, Stockholm, Sweden" httpUri="http://table.plazi.org/id/DF25CBDEFFDAB10353029D82F71DFE37" pageId="73" pageNumber="424" tableUuid="DF25CBDEFFDAB10353029D82F71DFE37">table 1</tableCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C340C8DDFF90B14C51BA99CCF4A0FE66" lastPageId="76" lastPageNumber="427" pageId="73" pageNumber="424" type="diagnosis">
<paragraph id="8BE59B56FF90B14951BA99CCF69AFA0D" blockId="73.[640,1157,1500,1741]" box="[640,749,1500,1526]" pageId="73" pageNumber="424">Diagnosis</paragraph>
<paragraph id="8BE59B56FF90B14A51BC9A10F583F99A" blockId="73.[640,1157,1500,1741]" lastBlockId="74.[129,639,1499,1740]" lastPageId="74" lastPageNumber="425" pageId="73" pageNumber="424">
<emphasis id="B92E4744FF90B14951BC9A10F707F9E1" box="[646,880,1536,1562]" italics="true" pageId="73" pageNumber="424">External morphology</emphasis>
: Shell (fig. 19F) small to medium-sized (
<quantity id="4CA236B3FF90B14950049A34F7A0F9C5" box="[830,983,1572,1598]" metricMagnitude="-2" metricUnit="m" metricValue="1.055" metricValueMax="1.23" metricValueMin="0.88" pageId="73" pageNumber="424" unit="mm" value="10.55" valueMax="12.3" valueMin="8.8">8.812.3 mm</quantity>
), pale golden, 3.03.9 whorls, subglobose with a low spire, last whorl very large. Body
<quantity id="4CA236B3FF90B14950FC9A7BF061F97D" box="[966,1046,1643,1670]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="73" pageNumber="424" unit="mm" value="25.0">25 mm</quantity>
long, colour (in ethanol) beige, darker on tail with a pale keel. Mantle lobes and shell lappets large, lappets with dark streaks, right lappet with a row of five warts at base and few smaller pustules scattered randomly, left lappet with eight warts arranged in two rows.
</paragraph>
<caption id="DF25CBDEFF93B14A53BB9932F52BFA89" ID-DOI="http://doi.org/10.5281/zenodo.8343109" ID-Zenodo-Dep="8343109" httpUri="https://zenodo.org/record/8343109/files/figure.png" pageId="74" pageNumber="425" startId="74.[129,202,1314,1335]" targetBox="[129,1177,192,1282]" targetPageId="74" targetType="figure">
<paragraph id="8BE59B56FF93B14A53BB9932F52BFA89" blockId="74.[129,1176,1313,1394]" pageId="74" pageNumber="425">
FIGURE 24 Genitalia of
<taxonomicName id="4C5AE0D5FF93B14A52519932F5A8FACC" box="[363,479,1314,1335]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="74" pageNumber="425" phylum="Mollusca" rank="species" species="hannianus">
<emphasis id="B92E4744FF93B14A52519932F5A8FACC" box="[363,479,1314,1335]" italics="true" pageId="74" pageNumber="425">F. hannianus</emphasis>
</taxonomicName>
. A, B. MO72277, Bakers Blue Mountain. C. MO76564, Hann Tableland. A. Reproductive system. Arrow shows the junction of the penis and epiphallus. BC. Penis interior. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BE59B56FF93B14C53A59A7BF4A0FE66" blockId="74.[129,639,1499,1740]" lastBlockId="76.[127,641,171,413]" lastPageId="76" lastPageNumber="427" pageId="74" pageNumber="425">
<emphasis id="B92E4744FF93B14A53A59A7BF515F97F" box="[159,354,1643,1669]" italics="true" pageId="74" pageNumber="425">Genital anatomy:</emphasis>
Genitalia (fig. 26) with moderately short vagina; bursa copulatrix short; duct swollen, bursa spherical. Penis moderately large, mostly cylindrical, slender, swollen proximally, inner penial wall sculptured with very fine transverse lamellae, becoming coarser proximally; penis interior with two longitudinal pilasters, major pilaster branches distally, then proximally becomes swollen and irregular; 60% of penis contained in penial tunica. Epiphallus approx. same length as penis, entering through simple pore; epiphallus 2 equal in length to epiphallus 1; epiphallic caecum moderate length, terminating with a swollen region consisting of multiple small outpocketings; flagellum short, slender.
</paragraph>
</subSubSection>
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<paragraph id="8BE59B56FF92B14B535899FCF682F9E4" blockId="75.[98,1137,1516,1567]" pageId="75" pageNumber="426">
FIGURE 25 Genitalia of
<taxonomicName id="4C5AE0D5FF92B14B527699FCF5AEF9F9" authority=", MO" authorityName="MO" box="[332,473,1516,1538]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="75" pageNumber="426" phylum="Mollusca" rank="species" species="insularis">
<emphasis id="B92E4744FF92B14B527699FCF5DBF9FA" box="[332,428,1516,1537]" italics="true" pageId="75" pageNumber="426">F.insularis</emphasis>
, MO
</taxonomicName>
85836, Palm Island. A. Reproductive system. Arrow shows the junction of the penis and epiphallus. B. Penis interior. Scale bars 1 mm.
</paragraph>
</caption>
<subSubSection id="C340C8DDFF95B14C53419DDAF561FA29" pageId="76" pageNumber="427" type="discussion">
<paragraph id="8BE59B56FF95B14C53419DDAF4AAFE1F" blockId="76.[123,642,458,1490]" box="[123,221,458,484]" pageId="76" pageNumber="427">Remarks</paragraph>
<paragraph id="8BE59B56FF95B14C53BB9DFFF5EBFBFB" blockId="76.[123,642,458,1490]" pageId="76" pageNumber="427">
<taxonomicName id="4C5AE0D5FF95B14C53BB9DFFF50FFDF3" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[129,376,494,520]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="species" species="katatonos">
<emphasis id="B92E4744FF95B14C53BB9DFFF50FFDF3" box="[129,376,494,520]" italics="true" pageId="76" pageNumber="427">Fastosarion katatonos</emphasis>
</taxonomicName>
(referred to above as
<taxonomicName id="4C5AE0D5FF95B14C53BF9E02F55BFDD7" box="[133,300,530,556]" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
sp. Mt Archer) is found in vine thickets at Mt Archer (northeast of Rockhampton) and also further to the northeast at Hidden Valley, near Yeppoon (fig. 18). It has been previously considered to be part of
<taxonomicName id="4C5AE0D5FF95B14C51579EB3F585FD1B" authority="(Stanisic et al., 2010)" baseAuthorityName="Stanisic" baseAuthorityYear="2010" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="species" species="pustulosus">
<emphasis id="B92E4744FF95B14C51579EB3F481FD1B" italics="true" pageId="76" pageNumber="427">F. pustulosus</emphasis>
(
<bibRefCitation id="EFCBE6A7FF95B14C52309ED6F59FFD1B" author="Stanisic, J. &amp; Shea, M. &amp; Potter, D. &amp; Griffiths, O." box="[266,488,710,736]" pageId="76" pageNumber="427" refId="ref43055" refString="Stanisic, J., Shea, M., Potter, D. &amp; Griffiths, O. (2010) Australian Land Snails. 1. A Field Guide to Eastern Australian Species. Bioculture Press, Riviere des Anguilles, Mauritius." type="book" year="2010">Stanisic et al., 2010</bibRefCitation>
)
</taxonomicName>
, and shares with this species a very similar shell and the presence of warts and dark streaks on the shell lappets. However,
<taxonomicName id="4C5AE0D5FF95B14C526E9F22F5ABFCB7" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[340,476,818,844]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="species" species="katatonos">
<emphasis id="B92E4744FF95B14C526E9F22F5ABFCB7" box="[340,476,818,844]" italics="true" pageId="76" pageNumber="427">F. katatonos</emphasis>
</taxonomicName>
has a slightly lower spire and more numerous warts on the shell lappets, and anatomically the two species can be very easily distinguished by their very different penial anatomy and the shape of the epiphallic caecum.
</paragraph>
<paragraph id="8BE59B56FF95B14C53A59819F561FA29" blockId="76.[123,642,458,1490]" pageId="76" pageNumber="427">
This species forms the sister taxon to
<taxonomicName id="4C5AE0D5FF95B14C5157981AF4ABFBBC" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="species" species="minerva">
<emphasis id="B92E4744FF95B14C5157981AF4ABFBBC" italics="true" pageId="76" pageNumber="427">F. minerva</emphasis>
</taxonomicName>
,
<taxonomicName id="4C5AE0D5FF95B14C53D0983EF523FBBC" box="[234,340,1069,1095]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="species" species="deensis">
<emphasis id="B92E4744FF95B14C53D0983EF523FBBC" box="[234,340,1069,1095]" italics="true" pageId="76" pageNumber="427">F. deensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF95B14C52AA983EF581FBBC" box="[400,502,1070,1095]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="76" pageNumber="427" phylum="Mollusca" rank="species" species="rowani">
<emphasis id="B92E4744FF95B14C52AA983EF581FBBC" box="[400,502,1070,1095]" italics="true" pageId="76" pageNumber="427">F. rowani</emphasis>
</taxonomicName>
in our phylogenetic analysis. It shares some similarities with these species, in particular the subglobose shell with a relatively high whorl count and the presence of rows of warts on its shell lappets, but is smaller in size with a unique genital anatomy. This species is set apart by its distinctive epiphallic caecum shape and short flagellum, its proximally swollen penis with fine transverse lamellae, equal length epiphallus arms and its short bursa copulatrix with a swollen duct.
</paragraph>
</subSubSection>
</treatment>
</document>