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<document id="C9356455C75AC2BED13AEE40C754D47F" ID-DOI="10.1163/18759866-20191416" ID-ISSN="1875-9866" ID-Zenodo-Dep="8343061" IM.bibliography_approvedBy="diego" IM.illustrations_approvedBy="diego" IM.materialsCitations_approvedBy="diego" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="diego" IM.taxonomicNames_approvedBy="diego" IM.treatmentCitations_approvedBy="diego" IM.treatments_approvedBy="diego" checkinTime="1694662606570" checkinUser="felipe" docAuthor="Hyman, Isabel T. &amp; Köhler, Frank" docDate="2019" docId="03F32A40FF8FB15851A59F57F504FE3A" docLanguage="en" docName="ContribZool.88.4.351-451.pdf" docOrigin="Contributions to Zoology 88 (4)" docSource="http://dx.doi.org/10.1163/18759866-20191416" docStyle="DocumentStyle:E1C3C5FD6CDDA08D62BFE928526826AF.2:ContribZool.2019-.journal_article" docStyleId="E1C3C5FD6CDDA08D62BFE928526826AF" docStyleName="ContribZool.2019-.journal_article" docStyleVersion="2" docTitle="Fastosarion tuljun Hyman &amp; Köhler 2019, sp. nov." docType="treatment" docVersion="4" lastPageNumber="438" masterDocId="FFCA5238FFD9B100533A9C10F477FFFB" masterDocTitle="Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)" masterLastPageNumber="451" masterPageNumber="351" pageNumber="437" updateTime="1695298548213" updateUser="diego" zenodo-license-document="CC-BY-4.0">
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<mods:title id="5034B3EF271D049DED63D58F6555148A">Phylogeny and systematic revision of the helicarionid semislugs of eastern Queensland (Stylommatophora, Helicarionidae)</mods:title>
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<mods:namePart id="AC84FFC3F33FEB7DFB41987543EC4B24">Hyman, Isabel T.</mods:namePart>
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<mods:namePart id="0B92061AABC3B4974BFAED431B003685">Köhler, Frank</mods:namePart>
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<mods:title id="50025C75614CB6772F3D49CD8DC615A6">Contributions to Zoology</mods:title>
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<mods:date id="C3796EE127458C4336D56D8CC6BF9B0E">2019</mods:date>
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<mods:number id="0C1C94CE2731050F80A1215E4F0F26D9">2019-10-03</mods:number>
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<mods:identifier id="7FFA8637E60EA13B8A7F7022AC312A48" type="DOI">10.1163/18759866-20191416</mods:identifier>
<mods:identifier id="485E165FB9028288ECD9D439155E9C1A" type="ISSN">1875-9866</mods:identifier>
<mods:identifier id="5062CDAF0A8D1A88FED5282DC2B695DA" type="Zenodo-Dep">8343061</mods:identifier>
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<treatment id="03F32A40FF8FB15851A59F57F504FE3A" LSID="urn:lsid:plazi:treatment:03F32A40FF8FB15851A59F57F504FE3A" httpUri="http://treatment.plazi.org/id/03F32A40FF8FB15851A59F57F504FE3A" lastPageId="88" lastPageNumber="438" pageId="86" pageNumber="437">
<subSubSection id="C340C8DDFF8FB15651A59F57F7BAFC9A" box="[671,973,839,866]" pageId="86" pageNumber="437" type="nomenclature">
<paragraph id="8BE59B56FF8FB15651A59F57F7BAFC9A" blockId="86.[671,973,839,866]" box="[671,973,839,866]" pageId="86" pageNumber="437">
<heading id="D0AD2C3AFF8FB15651A59F57F7BAFC9A" box="[671,973,839,866]" fontSize="11" level="2" pageId="86" pageNumber="437" reason="2">
<emphasis id="B92E4744FF8FB15651A59F57F7BAFC9A" bold="true" box="[671,973,839,866]" pageId="86" pageNumber="437">
<taxonomicName id="4C5AE0D5FF8FB15651A59F57F703FC99" ID-CoL="7RBS4" ID-ENA="2658028" authority="Hyman &amp; Köhler, 2019" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[671,884,839,866]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="species" species="tuljun" status="sp. nov.">
<emphasis id="B92E4744FF8FB15651A59F57F703FC99" bold="true" box="[671,884,839,866]" italics="true" pageId="86" pageNumber="437">Fastosarion tuljun</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21DFA3FFF8FB15650439F57F7BAFC9A" box="[889,973,839,865]" pageId="86" pageNumber="437" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C340C8DDFF8FB158519E9F7BF504FE3A" lastPageId="88" lastPageNumber="439" pageId="86" pageNumber="437" type="description">
<paragraph id="8BE59B56FF8FB156519E9F7BF73DFC7E" blockId="86.[676,842,875,901]" box="[676,842,875,901]" pageId="86" pageNumber="437">
<figureCitation id="136187D3FF8FB156519E9F7BF683FC7E" box="[676,756,875,901]" captionStart="FIGURE 18" captionStartId="66.[129,202,1582,1603]" captionTargetBox="[129,1187,188,1550]" captionTargetId="figure-8@66.[129,1187,188,1550]" captionTargetPageId="66" captionText="FIGURE 18 Distribution of Clade B species. █ = F.alyssa, ▼ = F.deensis, × = F.ephelis, ♢ = F.hannianus, ▲ = F. insularis, ◻ = F.katatonos, ∆ = F.longimentula, ● = F.mcdonaldi, ♠ = F.minerva, ◆ = F.papillosus, ♣ = F.pustulosus, ♥ = F.rowani, Ǫ = F.sarina, ▽ = F.tuljun." figureDoi="http://doi.org/10.5281/zenodo.8343097" httpUri="https://zenodo.org/record/8343097/files/figure.png" pageId="86" pageNumber="437">
Figs.
<geoCoordinate id="EE6EFD91FF8FB15651E69F7BF683FC7E" box="[732,756,875,901]" degrees="18, 19" direction="west" orientation="longitude" pageId="86" pageNumber="437" precision="555" value="-18.19">18</geoCoordinate>
</figureCitation>
,
<figureCitation id="136187D3FF8FB15651C49F7BF750FC7E" box="[766,807,875,901]" captionStart="FIGURE 19" captionStartId="67.[98,171,1480,1501]" captionTargetBox="[98,1104,171,1448]" captionTargetId="figure-115@67.[98,1104,170,1448]" captionTargetPageId="67" captionText="FIGURE 19 Shells of Clade B species. A. Fastosarion alyssa (type locality of D.peterbrocki), NMV F230748. B. Fastosarion deensis, MO85834 (holotype). C. Fastosarion ephelis, MO85835 (holotype). D. Fastosarion hannianus, MO56684.E. Fastosarion insularis sp. nov., MO76354. F. Fastosarion katatonos, MO85837 (holotype). G.Fastosarion longimentula, MO65313.H. Fastosarion mcdonaldi, MO50434.I. Fastosarion minerva, MO78358. J.Fastosarion papillosus, MO78896 (Mt Superbus).K.Fastosarion papillosus, MO62397 (Kroombit Tops). L.Fastosarion pustulosus, MO78894 (Kalpowar SF).M. Fastosarion rowani, MO85839 (holotype). N. Fastosarion sarina, MO85840 (holotype).O. Fastosarion tujun (Denham Range), MO76081 (holotype). Scale bar 5 mm." figureDoi="http://doi.org/10.5281/zenodo.8343099" httpUri="https://zenodo.org/record/8343099/files/figure.png" pageId="86" pageNumber="437">19O</figureCitation>
,
<figureCitation id="136187D3FF8FB156500B9F7BF73DFC7E" box="[817,842,875,901]" captionStart="FIGURE 32" captionStartId="87.[98,171,1161,1182]" captionTargetBox="[98,1156,188,1129]" captionTargetId="figure-140@87.[98,1156,188,1129]" captionTargetPageId="87" captionText="FIGURE 32 Genitalia of F.tuljun, MO76081, Denham Range. A. Reproductive system. Arrow shows the junction of the penis and epiphallus.B. Penis interior. Scale bars 1 mm." figureDoi="http://doi.org/10.5281/zenodo.8343125" httpUri="https://zenodo.org/record/8343125/files/figure.png" pageId="86" pageNumber="437">32</figureCitation>
</paragraph>
<paragraph id="8BE59B56FF8FB15651A49FA3F761FC36" blockId="86.[670,1187,947,1152]" box="[670,790,947,973]" pageId="86" pageNumber="437">Etymology</paragraph>
<paragraph id="8BE59B56FF8FB156519E9FC7F754FB7B" blockId="86.[670,1187,947,1152]" pageId="86" pageNumber="437">
From
<taxonomicName id="4C5AE0D5FF8FB15651D39FC7F75DFC0A" authorityName="Hyman &amp; Köhler" authorityYear="2019" box="[745,810,983,1009]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="86" pageNumber="437" phylum="Mollusca" rank="species" species="tuljun">
<emphasis id="B92E4744FF8FB15651D39FC7F75DFC0A" box="[745,810,983,1009]" italics="true" pageId="86" pageNumber="437">tuljun</emphasis>
</taxonomicName>
(Wiri = hill or mountain), referring to the Tuljunburra (= mountain people) clan of the Wiri people whose traditional lands include the Denham Range; noun in apposition.
</paragraph>
<paragraph id="8BE59B56FF8FB15651A498BEF705FB33" blockId="86.[670,1186,1198,1332]" box="[670,882,1198,1224]" pageId="86" pageNumber="437">Material examined</paragraph>
<paragraph id="8BE59B56FF8FB156519E98C2F7A1FACF" blockId="86.[670,1186,1198,1332]" pageId="86" pageNumber="437">
<materialsCitation id="3B32910BFF8FB156519E98C2F7A1FACF" collectingDate="2000-12-18" collectionCode="QM" collectorName="D. Cook. G. Monteith." location="Denham Range" pageId="86" pageNumber="437" specimenCode="MO76081" specimenCount="1" typeStatus="holotype">
<emphasis id="B92E4744FF8FB156519E98C2F772FB17" box="[676,773,1234,1260]" italics="true" pageId="86" pageNumber="437">
<typeStatus id="54E125F4FF8FB156519E98C2F772FB17" box="[676,773,1234,1260]" pageId="86" pageNumber="437">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="ED4B0393FF8FB156502D98C2F735FB17" box="[791,834,1234,1260]" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="86" pageNumber="437" type="Museum">QM</collectionCode>
<specimenCode id="DBFC332DFF8FB156507798C2F7C0FB17" box="[845,951,1234,1260]" collectionCode="QM" country="Australia" lsid="urn:lsid:biocol.org:col:35019" name="Queensland Museum" pageId="86" pageNumber="437" type="Museum">MO76081</specimenCode>
,
<location id="8E85CD8DFF8FB15650FC98C2F00DFB16" LSID="urn:lsid:plazi:treatment:03F32A40FF8FB15851A59F57F504FE3A:8E85CD8DFF8FB15650FC98C2F00DFB16" box="[966,1146,1234,1261]" name="Denham Range" pageId="86" pageNumber="437">Denham Range</location>
, ca
<locationDeviation id="9A578274FF8FB156519E98E6F7B1FAEB" box="[676,966,1270,1297]" location="Mt Robert" pageId="86" pageNumber="437">
<quantity id="4CA236B3FF8FB156519E98E6F6A9FAEB" box="[676,734,1270,1296]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="86" pageNumber="437" unit="km" value="2.0">2 km</quantity>
NNW of Mt Robert
</locationDeviation>
, MEQ. Coll.
<date id="FFE4BD96FF8FB156576D98E6F6ACFACF" pageId="86" pageNumber="437" value="2000-12-18">
<collectingDate id="EFA0447EFF8FB156576D98E6F6ACFACF" pageId="86" pageNumber="437" value="2000-12-18">18 Dec 2000</collectingDate>
</date>
,
<collectorName id="26AFFE80FF8FB15651DE990AF7A1FACF" box="[740,982,1306,1332]" pageId="86" pageNumber="437">D. Cook. G. Monteith.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="8BE59B56FF8FB15651A49972F77CFA87" blockId="86.[670,1187,1378,1763]" box="[670,779,1378,1404]" pageId="86" pageNumber="437">Diagnosis</paragraph>
<paragraph id="8BE59B56FF8FB156519E9996F007F967" blockId="86.[670,1187,1378,1763]" pageId="86" pageNumber="437">
<emphasis id="B92E4744FF8FB156519E9996F7E4FA5B" box="[676,915,1414,1440]" italics="true" pageId="86" pageNumber="437">External morphology</emphasis>
: Shell (fig. 19O) small (
<quantity id="4CA236B3FF8FB156519199BAF741FA3F" box="[683,822,1450,1476]" metricMagnitude="-3" metricUnit="m" metricValue="9.45" metricValueMax="10.2" metricValueMin="8.7" pageId="86" pageNumber="437" unit="mm" value="9.45" valueMax="10.2" valueMin="8.7">8.7–10.2 mm</quantity>
), golden, transparent, ear-shaped, very reduced, 2.6–3.0 whorls, flattened, thin, last whorl large. Body at least
<quantity id="4CA236B3FF8FB15650C599E1F03CF9F7" box="[1023,1099,1521,1548]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="86" pageNumber="437" unit="mm" value="13.0">13 mm</quantity>
, discoloured through immersion in pitfall trap; shell lappets and mantle lobes moderately large with some dark markings, smooth. Tail not keeled, slime network moderately strong.
</paragraph>
<paragraph id="8BE59B56FF8FB15751F89AB5F746FA65" blockId="86.[670,1187,1378,1763]" lastBlockId="87.[646,1157,1377,1438]" lastPageId="87" lastPageNumber="438" pageId="86" pageNumber="437">
<emphasis id="B92E4744FF8FB15651F89AB5F7F1F944" box="[706,902,1701,1727]" italics="true" pageId="86" pageNumber="437">Genital anatomy:</emphasis>
Genitalia (fig. 32) with moderately short vagina; bursa copulatrix short, duct distinct, bursa oval. Vagina, free oviduct and base of bursa copulatrix folded into foot. Penis cylindrical, internally with chevron-shaped diagonal ridges distally, a single ridged longitudinal pilaster medially to proximally, proximal portion of penial wall sculptured with transverse ridges; 70% of penis contained in penial tunica. Epiphallus approx. equal to penis length; entering penis through a short, conical verge; epiphallus arms equal in length; epiphallic caecum of moderate length; flagellum of moderate length, slender.
</paragraph>
<caption id="DF25CBDEFF8EB15753589899F681FB40" ID-DOI="http://doi.org/10.5281/zenodo.8343125" ID-Zenodo-Dep="8343125" httpUri="https://zenodo.org/record/8343125/files/figure.png" pageId="87" pageNumber="438" startId="87.[98,171,1161,1182]" targetBox="[98,1156,188,1129]" targetPageId="87" targetType="figure">
<paragraph id="8BE59B56FF8EB15753589899F681FB40" blockId="87.[98,1144,1160,1212]" pageId="87" pageNumber="438">
FIGURE 32 Genitalia of
<taxonomicName id="4C5AE0D5FF8EB15752769899F5B7FB65" authority=", MO" authorityName="MO" box="[332,448,1160,1182]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="87" pageNumber="438" phylum="Mollusca" rank="species" species="tuljun">
<emphasis id="B92E4744FF8EB15752769899F5E3FB66" box="[332,404,1160,1182]" italics="true" pageId="87" pageNumber="438">F.tuljun</emphasis>
, MO
</taxonomicName>
76081, Denham Range. A. Reproductive system. Arrow shows the junction of the penis and epiphallus. B. Penis interior. Scale bars 1 mm.
</paragraph>
</caption>
<paragraph id="8BE59B56FF8EB15751BA99DCF695FA1D" blockId="87.[640,1157,1484,1762]" box="[640,738,1484,1510]" pageId="87" pageNumber="438">Remarks</paragraph>
<paragraph id="8BE59B56FF8EB15851BC99E0F504FE3A" blockId="87.[640,1157,1484,1762]" lastBlockId="88.[129,640,171,449]" lastPageId="88" lastPageNumber="439" pageId="87" pageNumber="438">
This species (identified above as ‘
<taxonomicName id="4C5AE0D5FF8EB15750C799E0F6D9F9D5" class="Gastropoda" family="Helicarionidae" kingdom="Animalia" order="Stylommatophora" pageId="87" pageNumber="438" phylum="Mollusca" rank="family">Helicarionidae</taxonomicName>
sp. Denham Range’) is only known from a single live specimen found in rainforest on the Denham Range, west of Mackay, and several shells from the surrounding region (Mt Bassett, Mt Charlton, Eungella NP) (fig. 18). In the phylogenetic analysis it grouped with
<taxonomicName id="4C5AE0D5FF81B15853BB9CBCF54EFF3E" box="[129,313,171,197]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="88" pageNumber="439" phylum="Mollusca" rank="species" species="longimentula">
<emphasis id="B92E4744FF81B15853BB9CBCF54EFF3E" box="[129,313,171,197]" italics="true" pageId="88" pageNumber="439">F. longimentula</emphasis>
</taxonomicName>
and
<taxonomicName id="4C5AE0D5FF81B15852B59CBCF58CFF3E" box="[399,507,172,197]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="88" pageNumber="439" phylum="Mollusca" rank="species" species="sarina">
<emphasis id="B92E4744FF81B15852B59CBCF58CFF3E" box="[399,507,172,197]" italics="true" pageId="88" pageNumber="439">F. sarina</emphasis>
</taxonomicName>
. However, morpho-anatomically it is more similar to
<taxonomicName id="4C5AE0D5FF81B15853BB9CE4F567FEF6" box="[129,272,243,269]" class="Gastropoda" family="Helicarionidae" genus="Fastosarion" kingdom="Animalia" order="Stylommatophora" pageId="88" pageNumber="439" phylum="Mollusca" rank="species" species="mcdonaldi">
<emphasis id="B92E4744FF81B15853BB9CE4F567FEF6" box="[129,272,243,269]" italics="true" pageId="88" pageNumber="439">F. mcdonaldi</emphasis>
</taxonomicName>
, both in its small, flattened, reduced shell and in its genitalia (in particular, the presence of a penial verge and the equal length arms of the epipihallus). It can be distinguished from all other Qld semislugs by its unique penis interior.
</paragraph>
</subSubSection>
</treatment>
</document>
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