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<document id="DB0675A953E3AAD62137C9A28756FF78" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. &amp; Huijbregts, J." docDate="2017" docId="03E5BB4DE838FFCFAAE4FA59FEC9FC37" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus seseba Krikken &amp; Huijbregts, new species" docType="treatment" docVersion="7" lastPageNumber="330" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="329" updateTime="1698437609924" updateUser="ExternalLinkService">
<mods:mods id="E955A556E8A7C4CEBAD0B0353710204F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5AC987E2B76F3F7073500CC9CFEF1908">
<mods:title id="19F4EDA4BB1832801249893A17580E17">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="585C628DF8F6B99C6EE698D7A3357418" type="personal">
<mods:role id="2192696868D08ECCED55B8AF4DCA5F89">
<mods:roleTerm id="5BA9F114A393390821D15CBAE64B2729">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="57901A3BED6D715E30C63EFBFD204B3B">Krikken, J.</mods:namePart>
</mods:name>
<mods:name id="2FE0EB6246555A8723FA8111E4E4E113" type="personal">
<mods:role id="9939C04ACA44080CD4B8F3E6439677C8">
<mods:roleTerm id="733818F5D0A29B553B61E44ED99387E2">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5D9CEBE0E8A8EE2EECDFFEE27641055C">Huijbregts, J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="D12FC62806B75BB74EE9385D6C839857">text</mods:typeOfResource>
<mods:relatedItem id="C075D3B5F4E7B9BC940D2B443B615657" type="host">
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<mods:title id="4D0D84DAA2E05B4C1409FD6F04828401">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="5C7E705BDB83DE528A731FD22EAE0371">
<mods:date id="922C416F8D45ADDBE29EADF6A994AC57">2017</mods:date>
<mods:detail id="76BD60DEA990223B307E3B501400F053" type="volume">
<mods:number id="C80722FFA079D3714CF76FB325D885E5">4238</mods:number>
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<mods:number id="11335593421A3B95EA3AA5CEB7BAE6BA">3</mods:number>
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<mods:start id="1F9B807EA62670922E6AEEBBE4892B6F">301</mods:start>
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<mods:classification id="467E31D802CF089AB688292E211CAB3D">journal article</mods:classification>
<mods:identifier id="A11E67F865E9CE80709F2C31C23C68CB" type="CLB-Dataset">36345</mods:identifier>
<mods:identifier id="426D61AB93E857B32703D7FD204EA62F" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
<mods:identifier id="A030DDC32E5EAB6502CB5DA719FFF1FE" type="GBIF-Dataset">030203e2-e6a1-4b59-bfa0-0689bf817b08</mods:identifier>
<mods:identifier id="405261DE9F00BF7CB55A89F8086A5A62" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F75D3BDDF0AF7E8BCF701706F9DDC5DD" type="Zenodo-Dep">345756</mods:identifier>
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</mods:mods>
<treatment id="03E5BB4DE838FFCFAAE4FA59FEC9FC37" ID-DOI="http://doi.org/10.5281/zenodo.6030203" ID-GBIF-Taxon="127719079" ID-Zenodo-Dep="6030203" LSID="urn:lsid:plazi:treatment:03E5BB4DE838FFCFAAE4FA59FEC9FC37" httpUri="http://treatment.plazi.org/id/03E5BB4DE838FFCFAAE4FA59FEC9FC37" lastPageId="29" lastPageNumber="330" pageId="28" pageNumber="329">
<subSubSection id="C35659D0E838FFCEAAE4FA59FE58FA27" pageId="28" pageNumber="329" type="nomenclature">
<paragraph id="8BF30A5BE838FFCEAAE4FA59FC94FA01" blockId="28.[151,846,1473,1533]" box="[151,846,1473,1499]" pageId="28" pageNumber="329">
<heading id="D0BBBD37E838FFCEAAE4FA59FC94FA01" bold="true" box="[151,846,1473,1499]" fontSize="11" level="1" pageId="28" pageNumber="329" reason="1">
<emphasis id="B938D649E838FFCEAAE4FA59FC94FA01" bold="true" box="[151,846,1473,1499]" pageId="28" pageNumber="329">
<taxonomicName id="4C4C71D8E838FFCEAAE4FA59FD6BFA01" ID-CoL="7XP6R" authority="Krikken &amp; Huijbregts" authorityName="Krikken &amp; Huijbregts" box="[151,689,1473,1499]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="329" phylum="Arthropoda" rank="species" species="seseba" status="sp. nov.">
<emphasis id="B938D649E838FFCEAAE4FA59FE4BFA01" bold="true" box="[151,401,1473,1499]" italics="true" pageId="28" pageNumber="329">Onthophagus seseba</emphasis>
Krikken &amp; Huijbregts
</taxonomicName>
,
<taxonomicNameLabel id="A20B6B32E838FFCEA8CDFA59FC94FA01" box="[702,846,1473,1499]" pageId="28" pageNumber="329" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF30A5BE838FFCEAAE4FA7CFE58FA27" blockId="28.[151,846,1473,1533]" box="[151,386,1508,1533]" pageId="28" pageNumber="329">
(
<figureCitation id="137716DEE838FFCEAAECFA7CFF26FA26" box="[159,252,1508,1533]" captionStart="FIGURES 9 – 12" captionStartId="6.[151,264,1319,1342]" captionTargetBox="[172,1423,274,1297]" captionTargetId="figure@6.[151,1436,193,1298]" captionTargetPageId="6" captionText="FIGURES 9 – 12. Sulawesi Onthophagus species, oblique habitus. 9, O. begoniophilus, paratype male from Ambang. 10, O. annulopunctatus, paratype male from Mogogonipa. 11, O. hollowayi, holotype male from Dumoga. 12, O. seseba, holotype male from Batui (pygidium up). Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345758/files/figure.png" pageId="28" pageNumber="329">Figs. 12</figureCitation>
,
<figureCitation id="137716DEE838FFCEAB7BFA7CFEFCFA26" box="[264,294,1508,1532]" captionStart="FIGURES 25 – 30" captionStartId="9.[151,264,1925,1948]" captionTargetBox="[170,1436,198,1875]" captionTargetId="figure@9.[151,1436,193,1904]" captionTargetPageId="9" captionText="FIGURES 25 – 30. Sulawesi Onthophagus species, dorsal forebody. 25, O. sopu, holotype male from Sopu. 26, O. sopu, paratype male from Sopu. 27, O. manguliensis, male from Mangoli. 28, O. annulopunctatus, paratype male from Mogogonipa. 29, O. hollowayi, holotype male from Dumoga. 30, O. seseba, holotype male from Batui. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345761/files/figure.png" pageId="28" pageNumber="329">30</figureCitation>
,
<figureCitation id="137716DEE838FFCEAB41FA7CFEA0FA27" box="[306,378,1508,1533]" captionStart="FIGURES 68 – 81" captionStartId="30.[151,264,1760,1783]" captionTargetBox="[253,1344,205,1735]" captionTargetId="figure@30.[239,1344,191,1735]" captionTargetPageId="30" captionText="FIGURES 68 – 81. Sulawesi Onthophagus body parts. 68, O. seseba, head full-face, holotype male from Batui. 69, O. seseba, left elytron dorsal, holotype male from Batui. 70, O. seseba, protibial upper side, holotype male from Batui. 71, O. seseba, metatibial upperside, holotype male from Batui. 72, O. seseba, aedeagus lateral, holotype male from Batui. 73, O. seseba, head full-face, paratype female. 72, O. begoniophilus, aedeagus lateral, holotype from Ambang. 73, O. begoniophilus, head fullface, holotype from Ambang. 74, O. begoniophilus, head full-face, holotype from Ambang. 75, O. begoniophilus, left elytron dorsal, holotype from Ambang. 76, O. begoniophilus, head full-face, paratype male. 77, O. begoniophilus, left elytron caudal, paratype male. 78, O. begoniophilus, protibial upper side, paratype male. 79, O. begoniophilus, metatibial upperside, paratype male. 80, O. begoniophilus, aedeagus lateral, paratype male. 81, O. begoniophilus, head full-face, paratype female. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345766/files/figure.png" pageId="28" pageNumber="329">68–73</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E838FFCEAAE4F9B4FDE4F8C6" pageId="28" pageNumber="329" type="materials_examined">
<paragraph id="8BF30A5BE838FFCEAAE4F9B4FF0FF956" blockId="28.[151,1437,1580,2037]" pageId="28" pageNumber="329">
<materialsCitation id="3B240006E838FFCEAAE4F9B4FF0FF956" ID-GBIF-Occurrence="1438340102" collectingDate="1989-11-06" collectingDateMax="1989-11-09" collectingDateMin="1989-11-06" collectionCode="RMNH" collectorName="J. Krikken &amp; K. van der Blom" country="Indonesia" county="Banggai District" elevation="80" location="Estate" municipality="Batui" pageId="28" pageNumber="329" specimenCount="1" stateProvince="Central Sulawesi" typeStatus="holotype">
<emphasis id="B938D649E838FFCEAAE4F9B4FE58F99F" bold="true" box="[151,386,1580,1605]" pageId="28" pageNumber="329">Material examined.</emphasis>
<typeStatus id="54F7B4F9E838FFCEABF9F9B4FE28F99F" box="[394,498,1580,1605]" pageId="28" pageNumber="329" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED5D929EE838FFCEA831F9B4FD40F99E" box="[578,666,1580,1604]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="329">RMNH</collectionCode>
) from
<collectingCountry id="F35B4ACBE838FFCEA89AF9B4FC81F99F" box="[745,859,1580,1605]" name="Indonesia" pageId="28" pageNumber="329">Indonesia</collectingCountry>
:
<collectingRegion id="4988C4B9E838FFCEA915F9B4FBF1F99F" box="[870,1067,1580,1605]" country="Indonesia" name="Sulawesi Tengah" pageId="28" pageNumber="329">Central Sulawesi</collectingRegion>
:
<collectingCounty id="629272D7E838FFCEAE44F9B4FB2CF99F" box="[1079,1270,1580,1605]" pageId="28" pageNumber="329">Banggai District</collectingCounty>
:
<collectingMunicipality id="6B979021E838FFCEAF72F9B4FA98F99F" box="[1281,1346,1580,1605]" pageId="28" pageNumber="329">Batui</collectingMunicipality>
:
<taxonomicName id="4C4C71D8E838FFCEAF3DF9B4FA46F99F" box="[1358,1436,1580,1605]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">Seseba</taxonomicName>
<location id="8E935C80E838FFCEAAE4F9C8FF04F9BD" LSID="urn:lsid:plazi:treatment:03E5BB4DE838FFCFAAE4FA59FEC9FC37:8E935C80E838FFCEAAE4F9C8FF04F9BD" box="[151,222,1616,1639]" country="Indonesia" county="Banggai District" municipality="Batui" name="Estate" pageId="28" pageNumber="329" stateProvince="Central Sulawesi">Estate</location>
,
<elevation id="0061ED68E838FFCEAA98F9C8FEFDF9BD" box="[235,295,1616,1640]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="329" unit="m" value="80.0">
<quantity id="4CB4A7BEE838FFCEAA98F9C8FEFDF9BD" box="[235,295,1616,1640]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="329" unit="m" value="80.0">80 m</quantity>
</elevation>
,
<collectingDate id="EFB6D573E838FFCEAB47F9D7FE67F9B2" box="[308,445,1615,1640]" pageId="28" pageNumber="329" value="1989-11-06" valueMax="1989-11-09" valueMin="1989-11-06">
<date id="FFF22C9BE838FFCEAB47F9D7FE67F9B2" box="[308,445,1615,1640]" pageId="28" pageNumber="329" value="1989-11-06" valueMax="1989-11-09" valueMin="1989-11-06">6–9.xi.1989</date>
</collectingDate>
,
<collectorName id="26B96F8DE838FFCEABB9F9C8FD9AF9B2" box="[458,576,1615,1640]" pageId="28" pageNumber="329">J. Krikken</collectorName>
&amp;
<collectorName id="26B96F8DE838FFCEA815F9C8FCFFF9B2" box="[614,805,1615,1640]" pageId="28" pageNumber="329">K. van der Blom</collectorName>
sw30a; multistratal evergreen forest,
<collectingMethod id="520D724CE838FFCEAEA0F9D7FF0BF956" pageId="28" pageNumber="329">human excrement traps</collectingMethod>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE838FFCEAAB4F90FFD4FF96A" blockId="28.[151,1437,1580,2037]" box="[199,661,1687,1712]" pageId="28" pageNumber="329">The 15 paratypes are summarized below:</paragraph>
<paragraph id="8BF30A5BE838FFCEAAE4F924FD65F90E" blockId="28.[151,1437,1580,2037]" box="[151,703,1724,1748]" pageId="28" pageNumber="329">
<materialsCitation id="3B240006E838FFCEAAE4F924FD65F90E" ID-GBIF-Occurrence="1438340113" box="[151,703,1724,1748]" location="CENTRAL SULAWESI" pageId="28" pageNumber="329" specimenCount="1" stateProvince="CENTRAL SULAWESI">
<collectingRegion id="4988C4B9E838FFCEAAE4F924FE75F90E" box="[151,431,1724,1748]" country="Indonesia" name="Sulawesi Tengah" pageId="28" pageNumber="329">CENTRAL SULAWESI</collectingRegion>
: BANGGAI DISTRICT
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE838FFCEAAE4F978FDE4F8C6" blockId="28.[151,1437,1580,2037]" pageId="28" pageNumber="329">
- Batui:
<taxonomicName id="4C4C71D8E838FFCEAB66F947FEB9F922" box="[277,355,1759,1784]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="331" phylum="Arthropoda" rank="species" species="seseba">Seseba</taxonomicName>
Estate,
<quantity id="4CB4A7BEE838FFCEABB7F978FDD8F92D" box="[452,514,1760,1784]" metricMagnitude="1" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="329" unit="m" value="80.0">80 m</quantity>
,
<date id="FFF22C9BE838FFCEA862F947FD40F922" box="[529,666,1759,1784]" pageId="28" pageNumber="329" value="1989-11-06" valueMax="1989-11-09" valueMin="1989-11-06">6–9.xi.1989</date>
, Krikken &amp; Van der Blom, sw30a, multistratal evergreen forest, 4 human excrement traps,
<quantity id="4CB4A7BEE838FFCEABD9F89CFE39F8C7" box="[426,483,1796,1821]" metricMagnitude="-1" metricUnit="m" metricValue="4.064" pageId="28" pageNumber="329" unit="in" value="16.0">16 in</quantity>
<collectionCode id="ED5D929EE838FFCEAB99F89DFDE4F8C6" box="[490,574,1797,1820]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="28" pageNumber="329">RMNH</collectionCode>
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E838FFCFAAB4F8BFFB65FC7F" lastPageId="29" lastPageNumber="330" pageId="28" pageNumber="329" type="description">
<paragraph id="8BF30A5BE838FFCEAAB4F8BFFAB8F8BE" blockId="28.[151,1437,1580,2037]" pageId="28" pageNumber="329">
<emphasis id="B938D649E838FFCEAAB4F8BFFE83F89A" bold="true" box="[199,345,1831,1856]" pageId="28" pageNumber="329">Description.</emphasis>
<typeStatus id="54F7B4F9E838FFCEAB1BF8BFFE09F89A" box="[360,467,1831,1856]" pageId="28" pageNumber="329" type="holotype">Holotype</typeStatus>
, male. Body length approximately
<quantity id="4CB4A7BEE838FFCEA9FBF8B0FC30F8E5" box="[904,1002,1832,1856]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="28" pageNumber="329" unit="mm" value="3.8">3.8 mm</quantity>
. Largely black, with orange elytral markings, dorsal side matt. Pilosity abundant, pale yellow, dorsal side with inconspicuous short (white) setae.
</paragraph>
<paragraph id="8BF30A5BE838FFCEAAB4F8F7FD8AF82E" blockId="28.[151,1437,1580,2037]" pageId="28" pageNumber="329">Clypeal border round-truncate, clypeogenal transition continuous; clypeofrontal transition with fine arcuate ridge just reaching clypeogenal sutures on either side; head densely punctate throughout, punctures distinct, with pale microstubble. Eyes with approximately 6 facet rows across widest point; ratio interocular distance / single maximum eye width approximately 10.</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FF00FB5BFEC7" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">Pronotum evenly moderately convex, discal midline not impressed; anterior section of lateral border widely round, posterior section slightly sinuate to obtuse posterolateral angle; pronotal base indistinctly, finely marginate. Pronotum with disc densely, superficially but densely annulate-punctate, punctures with short seta (length mainly twice punctural diameter); punctures on lateral declivities smaller, deeper, more irregular.</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FEBFFE5DFE77" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">Elytra bicoloured; striae well defined, shallow, with distinct punctures; mostly separated by 2–4 times their diameter, distinctly crenulate on interstriae; discal interstriae flat, with irregular rows of seta-bearing microgranulae (setae fine, short), derm generally microreticulate; surface of lateral interstriae more uneven, almost rugulatepunctate, more shiny.</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FE2FFE8BFDBA" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">Antennal club yellowish brown. Metaventrite shiny brown, without distinct midline impression, anterior lobe with slight median ridge in front; most of discal surface with abundant fine punctation; lateral wings with coarser, more-or-less annulate punctation, with longer setae. Abdominal ventrites with transverse row of (more-or-less annulate) punctures, setae appressed caudad. Pygidium evenly convex, abundantly punctate-setose; base with transverse ridge.</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FDF4FC16FD17" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">
Protibia with 3+(1) larger external denticles; spur curved-acuminate. Metatibial spur long, tapering. Femoral undersides all with (more-or-less scattered) seta-bearing punctures. Metatarsomere 1 shorter than 2–5 combined; approximate proportions of spur and metatarsomeres 1–5: 11//
<date id="FFF22C9BE839FFCFA921FD2CFC46FD17" box="[850,924,692,717]" pageId="29" pageNumber="330" value="2013-04-06">13/6/4</date>
/3/5.
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FD4FFDCCFD2A" blockId="29.[151,1437,151,1005]" box="[199,534,727,752]" pageId="29" pageNumber="330">
Aedeagus,
<figureCitation id="137716DEE839FFCFAB30FD4FFE48FD2A" box="[323,402,727,752]" captionStart="FIGURES 68 – 81" captionStartId="30.[151,264,1760,1783]" captionTargetBox="[253,1344,205,1735]" captionTargetId="figure@30.[239,1344,191,1735]" captionTargetPageId="30" captionText="FIGURES 68 – 81. Sulawesi Onthophagus body parts. 68, O. seseba, head full-face, holotype male from Batui. 69, O. seseba, left elytron dorsal, holotype male from Batui. 70, O. seseba, protibial upper side, holotype male from Batui. 71, O. seseba, metatibial upperside, holotype male from Batui. 72, O. seseba, aedeagus lateral, holotype male from Batui. 73, O. seseba, head full-face, paratype female. 72, O. begoniophilus, aedeagus lateral, holotype from Ambang. 73, O. begoniophilus, head fullface, holotype from Ambang. 74, O. begoniophilus, head full-face, holotype from Ambang. 75, O. begoniophilus, left elytron dorsal, holotype from Ambang. 76, O. begoniophilus, head full-face, paratype male. 77, O. begoniophilus, left elytron caudal, paratype male. 78, O. begoniophilus, protibial upper side, paratype male. 79, O. begoniophilus, metatibial upperside, paratype male. 80, O. begoniophilus, aedeagus lateral, paratype male. 81, O. begoniophilus, head full-face, paratype female. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345766/files/figure.png" pageId="29" pageNumber="330">Fig. 72</figureCitation>
(
<typeStatus id="54F7B4F9E839FFCFABD1FD4FFDD0FD2A" box="[418,522,727,752]" pageId="29" pageNumber="330" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FD65FD66FCE2" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">Measurements in mm. Maximum width of head 1.1. Median length of pronotum 1.2, maximum width 1.9. Sutural length of elytra 1.6, maximum width 2.1.</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FCDCFF23FC5A" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">
<emphasis id="B938D649E839FFCFAAB4FCDCFDB7FC87" bold="true" box="[199,621,836,861]" pageId="29" pageNumber="330">Variation and sexual dimorphism.</emphasis>
Body length
<quantity id="4CB4A7BEE839FFCFA960FCDCFC57FC86" box="[787,909,836,861]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="4.5" metricValueMin="3.0" pageId="29" pageNumber="330" unit="mm" value="3.75" valueMax="4.5" valueMin="3.0">3–4.5 mm</quantity>
. Microsculpture, pilosity, and colour slightly variable.
</paragraph>
<paragraph id="8BF30A5BE839FFCFAAB4FC14FB65FC7F" blockId="29.[151,1437,151,1005]" box="[199,1215,908,933]" pageId="29" pageNumber="330">
<emphasis id="B938D649E839FFCFAAB4FC14FEFCFC7E" box="[199,294,908,932]" italics="true" pageId="29" pageNumber="330">Females</emphasis>
with distinct, low clypeofrontal ridge, sexual dimorphism otherwise insignificant.
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E839FFCFAAB4FC37FEC9FC37" pageId="29" pageNumber="330" type="etymology">
<paragraph id="8BF30A5BE839FFCFAAB4FC37FEC9FC37" blockId="29.[151,1437,151,1005]" pageId="29" pageNumber="330">
<emphasis id="B938D649E839FFCFAAB4FC37FE94FC12" bold="true" box="[199,334,943,968]" pageId="29" pageNumber="330">Etymology.</emphasis>
Named for the
<typeStatus id="54F7B4F9E839FFCFAB8DFC28FDF4FC12" box="[510,558,944,968]" pageId="29" pageNumber="330">type</typeStatus>
locality in eastern
<collectingRegion id="4988C4B9E839FFCFA977FC37FC1CFC12" box="[772,966,943,968]" country="Indonesia" name="Sulawesi Tengah" pageId="29" pageNumber="330">Central Sulawesi</collectingRegion>
. This name should be treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>
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