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<document id="1DE7270AB4AD324DEB5CFC494169B737" ID="10.11646/zootaxa.4238.3.1SLASH10400" ID-CLB-Dataset="36345" ID-DOI="10.11646/zootaxa.4238.3.1" ID-GBIF-Dataset="030203e2-e6a1-4b59-bfa0-0689bf817b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="345756" ID-ZooBank="D123950C-DF2B-45EE-9966-813339A83491" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1488787785636" checkinUser="plazi" docAuthor="Krikken, J. &amp; Huijbregts, J." docDate="2017" docId="03E5BB4DE82AFFC2AAE4FC64FB88FC12" docLanguage="en" docName="zootaxa.4238.3.1SLASH10400.pdf" docOrigin="Zootaxa 4238 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Onthophagus bongkudai Krikken &amp; Huijbregts, new species" docType="treatment" docVersion="7" lastPageNumber="317" masterDocId="FFDCC335E824FFD2AA73FF98FFDAFFDA" masterDocTitle="Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="334" masterPageNumber="301" pageNumber="315" updateTime="1698437609924" updateUser="ExternalLinkService">
<mods:mods id="9FD4836592415B54C233E7972E15815C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="EE142F1B4C5E61667AF4F808CE454E2B">
<mods:title id="D5513DA8564862263D71E276D0352752">Sulawesi Onthophagus: seven new species in select groups (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
</mods:titleInfo>
<mods:name id="16693811162FF703904F8106E9526462" type="personal">
<mods:role id="D6B50F3D7BFBD4AEA0A9806942B019EB">
<mods:roleTerm id="E5DEF7957D923012F5AC8A84E94A2FE7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5C225B05A1CAC5C7103DA2337D65743F">Krikken, J.</mods:namePart>
</mods:name>
<mods:name id="C04EE593E98A2A7BA6FF6F06A6C51B4F" type="personal">
<mods:role id="D8D7303214E6BBD527426B2D258DA5C4">
<mods:roleTerm id="87B8B47F782E22B6E3BB40F2B1EE1026">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="951F83875334D8ECE208287B3F0585EF">Huijbregts, J.</mods:namePart>
</mods:name>
<mods:typeOfResource id="1903B0D6E3528DBC885EC1A69F988F03">text</mods:typeOfResource>
<mods:relatedItem id="482C825436E695C984ECB941FD76D3EB" type="host">
<mods:titleInfo id="C5483CCF1F54CC4B28C783FB0EC12224">
<mods:title id="3B0ED41F7CBE6C9C18709CBB76654F61">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F0A1DD44850B747BA44BFF45476FF7A0">
<mods:date id="3D9BE00495EA10AA538F0B593524D758">2017</mods:date>
<mods:detail id="3F7A8CC897DD6EAA2C8F97BD93F5B035" type="volume">
<mods:number id="F6646ACEF68F0FC60D5DBFC6F1AF3C24">4238</mods:number>
</mods:detail>
<mods:detail id="FB6649F8B8F362B9F82AE16AB59DAF9E" type="issue">
<mods:number id="56B022BB51F0FA928EE554E5EB61B170">3</mods:number>
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<mods:extent id="987815FE589932A11A00A27415B97B84" unit="page">
<mods:start id="3B74EDF8753D70A15E4F5DC2A8940D53">301</mods:start>
<mods:end id="8423D84576BD7F59367259F2940AF606">334</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="845310D4B205AD6A5C479C791C274181">journal article</mods:classification>
<mods:identifier id="A698DC64839175100F41502713A180CA" type="CLB-Dataset">36345</mods:identifier>
<mods:identifier id="29A380A5094E73044CADFD0F71E4BB8D" type="DOI">10.11646/zootaxa.4238.3.1</mods:identifier>
<mods:identifier id="3FAFDF40F1575BBE1F417538270F55AF" type="GBIF-Dataset">030203e2-e6a1-4b59-bfa0-0689bf817b08</mods:identifier>
<mods:identifier id="1A74F85B884B6FCA2381393E02C9993B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="084FE0560848A902BECA93507D08A208" type="Zenodo-Dep">345756</mods:identifier>
<mods:identifier id="EB9AA3D387BD7BBEAF1C4A363238DDEF" type="ZooBank">D123950C-DF2B-45EE-9966-813339A83491</mods:identifier>
</mods:mods>
<treatment id="03E5BB4DE82AFFC2AAE4FC64FB88FC12" ID-DOI="http://doi.org/10.5281/zenodo.6030184" ID-GBIF-Taxon="127719086" ID-Zenodo-Dep="6030184" LSID="urn:lsid:plazi:treatment:03E5BB4DE82AFFC2AAE4FC64FB88FC12" httpUri="http://treatment.plazi.org/id/03E5BB4DE82AFFC2AAE4FC64FB88FC12" lastPageId="16" lastPageNumber="317" pageId="14" pageNumber="315">
<subSubSection id="C35659D0E82AFFDCAAE4FC64FE44FBED" pageId="14" pageNumber="315" type="nomenclature">
<paragraph id="8BF30A5BE82AFFDCAAE4FC64FC5AFBCC" blockId="14.[151,896,1020,1079]" box="[151,896,1020,1046]" pageId="14" pageNumber="315">
<heading id="D0BBBD37E82AFFDCAAE4FC64FC5AFBCC" bold="true" box="[151,896,1020,1046]" fontSize="11" level="1" pageId="14" pageNumber="315" reason="1">
<emphasis id="B938D649E82AFFDCAAE4FC64FC5AFBCC" bold="true" box="[151,896,1020,1046]" pageId="14" pageNumber="315">
<taxonomicName id="4C4C71D8E82AFFDCAAE4FC64FD3EFBCC" ID-CoL="7XMHG" authority="Krikken &amp; Huijbregts" authorityName="Krikken &amp; Huijbregts" box="[151,740,1020,1046]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="315" phylum="Arthropoda" rank="species" species="bongkudai" status="sp. nov.">
<emphasis id="B938D649E82AFFDCAAE4FC64FE19FBCC" bold="true" box="[151,451,1020,1046]" italics="true" pageId="14" pageNumber="315">Onthophagus bongkudai</emphasis>
Krikken &amp; Huijbregts
</taxonomicName>
,
<taxonomicNameLabel id="A20B6B32E82AFFDCA882FC64FC5AFBCC" box="[753,896,1020,1046]" pageId="14" pageNumber="315" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FB86FE44FBED" blockId="14.[151,896,1020,1079]" box="[151,414,1054,1079]" pageId="14" pageNumber="315">
(
<figureCitation id="137716DEE82AFFDCAAECFB86FF37FBED" box="[159,237,1054,1079]" captionStart="FIGURES 1 – 8" captionStartId="5.[151,264,1919,1942]" captionTargetBox="[207,1396,233,1891]" captionTargetId="figure@5.[177,1413,195,1894]" captionTargetPageId="5" captionText="FIGURES 1 – 8. Sulawesi Onthophagus species, oblique habitus. 1, O. bisscrutator, holotype male from Toraut. 2, O. scrutator, male from Moramo. 3, O. bongkudai, holotype male from Moajat. 4, O. sangirensis, male from Toraut. 5, O. holosericus, male from Marena, 6, female from Marena. 7, O. sopu, holotype male from Sopu. 8, O. manguliensis, male from Mangoli. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345757/files/figure.png" pageId="14" pageNumber="315">Figs. 3</figureCitation>
,
<figureCitation id="137716DEE82AFFDCAA89FB86FE98FBED" box="[250,322,1054,1079]" captionStart="FIGURES 13 – 18" captionStartId="7.[151,264,1935,1958]" captionTargetBox="[162,1436,205,1903]" captionTargetId="figure@7.[151,1436,193,1915]" captionTargetPageId="7" captionText="FIGURES 13 – 18. Sulawesi Onthophagus species, dorsal forebody. 13, O. bisscrutator, holotype male from Toraut. 14, O. scrutator, male from Moramo. 15, O. bongkudai, holotype male from Moajat 16, paratype female. 17, O. sangirensis, male from Toraut. 18, O. sangirensis, female from Tangkoko. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345759/files/figure.png" pageId="14" pageNumber="315">15–16</figureCitation>
,
<figureCitation id="137716DEE82AFFDCAB3DFB87FE4CFBED" box="[334,406,1055,1079]" captionStart="FIGURES 39 – 46" captionStartId="15.[151,264,1581,1604]" captionTargetBox="[151,1415,230,1560]" captionTargetId="figure@15.[151,1436,193,1560]" captionTargetPageId="15" captionText="FIGURES 39 – 46. Sulawesi Onthophagus body parts. 39, O. bongkudai, head full-face, holotype male from Moajat. 40, O. bongkudai, left elytron dorsal, holotype male from Moajat. 41, O. bongkudai, protibial upper side, holotype male from Moajat. 42, O. bongkudai, metatibial upperside, holotype male from Moajat. 43, O. bongkudai, aedeagus lateral, holotype male from Moajat. 44, O. bongkudai, head full-face, paratype female. 45, O. sangirensis, aedeagus lateral, male from Toraut. 46, O. holosericus, aedeagus lateral, male from Marena. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345763/files/figure.png" pageId="14" pageNumber="315">39–44</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E82AFFDCAAE4FBFEFEAAF9AC" pageId="14" pageNumber="315" type="materials_examined">
<paragraph id="8BF30A5BE82AFFDCAAE4FBFEFAC8FB78" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">
<materialsCitation id="3B240006E82AFFDCAAE4FBFEFAC8FB78" ID-GBIF-Occurrence="1438340110" collectingDate="1985-09-15" collectingDateMax="1985-09-16" collectingDateMin="1985-09-15" collectionCode="RMNH" collectorName="J. Huijbregts" country="Indonesia" elevation="1660" location="Mount Ambang Nature Reserve" municipality="Mount Moajat" pageId="14" pageNumber="315" specimenCount="1" typeStatus="holotype">
<emphasis id="B938D649E82AFFDCAAE4FBFEFE59FBA5" bold="true" box="[151,387,1126,1151]" pageId="14" pageNumber="315">Material examined.</emphasis>
<typeStatus id="54F7B4F9E82AFFDCABFEFBFEFE2FFBA5" box="[397,501,1126,1151]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED5D929EE82AFFDCA834FBFFFD45FBA5" box="[583,671,1127,1151]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
) from
<collectingCountry id="F35B4ACBE82AFFDCA882FBFEFCB8FBA5" box="[753,866,1126,1151]" name="Indonesia" pageId="14" pageNumber="315">Indonesia</collectingCountry>
:
<location id="8E935C80E82AFFDCA903FBFFFB3EFBA4" LSID="urn:lsid:plazi:treatment:03E5BB4DE82AFFC2AAE4FC64FB88FC12:8E935C80E82AFFDCA903FBFFFB3EFBA4" box="[880,1252,1126,1151]" country="Indonesia" municipality="Mount Moajat" name="Mount Ambang Nature Reserve" pageId="14" pageNumber="315">Mount Ambang Nature Reserve</location>
:
<collectingMunicipality id="6B979021E82AFFDCAE82FBFFFA42FBA5" box="[1265,1432,1126,1151]" pageId="14" pageNumber="315">Mount Moajat</collectingMunicipality>
,
<elevation id="0061ED68E82AFFDCAAE4FB12FF34FB78" box="[151,238,1162,1187]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="315" unit="m" value="1660.0">
<quantity id="4CB4A7BEE82AFFDCAAE4FB12FF34FB78" box="[151,238,1162,1187]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="315" unit="m" value="1660.0">1660 m</quantity>
</elevation>
,
<collectingDate id="EFB6D573E82AFFDCAA89FB12FE47FB79" box="[250,413,1162,1187]" pageId="14" pageNumber="315" value="1985-09-15" valueMax="1985-09-16" valueMin="1985-09-15">
<date id="FFF22C9BE82AFFDCAA89FB12FE47FB79" box="[250,413,1162,1187]" pageId="14" pageNumber="315" value="1985-09-15" valueMax="1985-09-16" valueMin="1985-09-15">15–16.ix.1985</date>
</collectingDate>
,
<collectorName id="26B96F8DE82AFFDCABDAFB13FDE1FB79" box="[425,571,1162,1187]" pageId="14" pageNumber="315">J. Huijbregts</collectorName>
, HH454A; multistratal evergreen forest,
<collectingMethod id="520D724CE82AFFDCAE76FB12FAD4FB78" box="[1029,1294,1162,1187]" pageId="14" pageNumber="315">human excrement traps</collectingMethod>
.
</materialsCitation>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAB4FB36FD4FFB1D" blockId="14.[151,1437,1126,1979]" box="[199,661,1198,1223]" pageId="14" pageNumber="315">
The
<specimenCount id="9D4AC1D2E82AFFDCAA8AFB37FE52FB1D" box="[249,392,1199,1223]" pageId="14" pageNumber="315" type="generic" typeStatus="paratypes">90 paratypes</specimenCount>
are summarized below:
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FB4BFC91FB31" blockId="14.[151,1437,1126,1979]" box="[151,843,1235,1259]" pageId="14" pageNumber="315">
<collectingRegion id="4988C4B9E82AFFDCAAE4FB4BFE57FB31" box="[151,397,1235,1259]" country="Indonesia" name="Sulawesi Utara" pageId="14" pageNumber="315">
<collectionCode id="ED5D929EE82AFFDCAAE4FB4BFE57FB31" box="[151,397,1235,1259]" pageId="14" pageNumber="315">NORTH SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE82AFFDCABE4FB4BFC91FB31" box="[407,843,1235,1259]" pageId="14" pageNumber="315">DUMOGA BONE NATIONAL PARK</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FB6FFBF3FAD5" blockId="14.[151,1437,1126,1979]" box="[151,1065,1270,1295]" pageId="14" pageNumber="315">
- Zebra,
<date id="FFF22C9BE82AFFDCAB6EFB6EFE1BFAD5" box="[285,449,1270,1295]" pageId="14" pageNumber="315" value="1985-11-10" valueMax="1985-11-20" valueMin="1985-11-10">10–20.xi.1985</date>
, J. Van Stalle
<emphasis id="B938D649E82AFFDCA82DFB6FFD4DFAD4" box="[606,663,1270,1294]" italics="true" pageId="14" pageNumber="315">et al.</emphasis>
, upland site, dung trap,
<quantity id="4CB4A7BEE82AFFDCA9D6FB6FFC0AFAD5" box="[933,976,1270,1295]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="14" pageNumber="315" unit="in" value="2.0">2 in</quantity>
<collectionCode id="ED5D929EE82AFFDCA9A4FB6FFBF3FAD5" box="[983,1065,1271,1295]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="14" pageNumber="315">IRSNB</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FA83FCADFAE9" blockId="14.[151,1437,1126,1979]" box="[151,887,1307,1331]" pageId="14" pageNumber="315">
<collectingRegion id="4988C4B9E82AFFDCAAE4FA83FE57FAE9" box="[151,397,1307,1331]" country="Indonesia" name="Sulawesi Utara" pageId="14" pageNumber="315">
<collectionCode id="ED5D929EE82AFFDCAAE4FA83FE57FAE9" box="[151,397,1307,1331]" pageId="14" pageNumber="315">NORTH SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE82AFFDCABE4FA83FCADFAE9" box="[407,887,1307,1331]" pageId="14" pageNumber="315">MOUNT AMBANG NATURE RESERVE</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FAA7FE31FAA0" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">
- Mount Moajat,
<quantity id="4CB4A7BEE82AFFDCABFBFAA7FE32FA8C" box="[392,488,1343,1367]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="14" pageNumber="315" unit="m" value="1780.0">1780 m</quantity>
,
<date id="FFF22C9BE82AFFDCAB8EFAA6FD7AFA8D" box="[509,672,1342,1367]" pageId="14" pageNumber="315" value="1985-09-13" valueMax="1985-09-16" valueMin="1985-09-13">13–16.ix.1985</date>
, J. Huijbregts, hh451a, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE82AFFDCAB24FAFBFE4AFAA1" box="[343,400,1378,1403]" metricMagnitude="0" metricUnit="m" metricValue="1.8796" pageId="14" pageNumber="315" unit="in" value="74.0">74 in</quantity>
<collectionCode id="ED5D929EE82AFFDCABE4FAFBFE31FAA0" box="[407,491,1379,1402]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FA1FFE07FA18" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">
- Mount Moajat,
<quantity id="4CB4A7BEE82AFFDCABF4FA1FFE3DFA44" box="[391,487,1415,1439]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="14" pageNumber="315" unit="m" value="1780.0">1780 m</quantity>
,
<date id="FFF22C9BE82AFFDCAB8FFA1EFD7AFA45" box="[508,672,1414,1439]" pageId="14" pageNumber="315" value="1985-09-13" valueMax="1985-09-15" valueMin="1985-09-13">13–15.ix.1985</date>
, J. Huijbregts, hh451b, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE82AFFDCAB24FA33FE58FA19" box="[343,386,1450,1475]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="14" pageNumber="315" unit="in" value="3.0">3 in</quantity>
<collectionCode id="ED5D929EE82AFFDCABFAFA33FE07FA18" box="[393,477,1451,1474]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4FA57FF36F9D0" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">
- Mount Moajat,
<quantity id="4CB4A7BEE82AFFDCAB08FA57FE0BFA3C" box="[379,465,1487,1511]" metricMagnitude="3" metricUnit="m" metricValue="1.78" pageId="14" pageNumber="315" unit="m" value="1780.0">1780 m</quantity>
,
<date id="FFF22C9BE82AFFDCABAEFA56FD5BFA3D" box="[477,641,1486,1511]" pageId="14" pageNumber="315" value="1985-09-13" valueMax="1985-09-16" valueMin="1985-09-13">13–16.ix.1985</date>
, J. Huijbregts, hh452, multistratal evergreen forest, hand collected,
<quantity id="4CB4A7BEE82AFFDCAF01FA57FA46FA3D" box="[1394,1436,1486,1511]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="315" unit="in" value="1.0">1 in</quantity>
<collectionCode id="ED5D929EE82AFFDCAAE4FA6BFF36F9D0" box="[151,236,1523,1546]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4F98FFD5BF9F5" blockId="14.[151,1437,1126,1979]" box="[151,641,1559,1583]" pageId="14" pageNumber="315">
<collectingRegion id="4988C4B9E82AFFDCAAE4F98FFE57F9F5" box="[151,397,1559,1583]" country="Indonesia" name="Sulawesi Utara" pageId="14" pageNumber="315">
<collectionCode id="ED5D929EE82AFFDCAAE4F98FFE57F9F5" box="[151,397,1559,1583]" pageId="14" pageNumber="315">NORTH SULAWESI</collectionCode>
</collectingRegion>
:
<collectionCode id="ED5D929EE82AFFDCABE4F98FFD5BF9F5" box="[407,641,1559,1583]" pageId="14" pageNumber="315">MOUNT AMBANG</collectionCode>
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAE4F9A3FEAAF9AC" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">
- Mt Moajat,
<quantity id="4CB4A7BEE82AFFDCAB23F9A2FE73F988" box="[336,425,1594,1619]" metricMagnitude="3" metricUnit="m" metricValue="1.66" pageId="14" pageNumber="315" unit="m" value="1660.0">1660 m</quantity>
,
<date id="FFF22C9BE82AFFDCABC4F9A2FD81F989" box="[439,603,1594,1619]" pageId="14" pageNumber="315" value="1985-09-15" valueMax="1985-09-16" valueMin="1985-09-15">15–16.ix.1985</date>
, J. Huijbregts, hh454a, multistratal evergreen forest, 2 human excrement traps,
<quantity id="4CB4A7BEE82AFFDCAAA8F9C7FECEF9AD" box="[219,276,1630,1655]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="14" pageNumber="315" unit="in" value="10.0">10 in</quantity>
<collectionCode id="ED5D929EE82AFFDCAB68F9C7FEAAF9AC" box="[283,368,1631,1654]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="14" pageNumber="315">RMNH</collectionCode>
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E82AFFC2AAB4F91AFD0FFC5A" lastPageId="16" lastPageNumber="317" pageId="14" pageNumber="315" type="description">
<paragraph id="8BF30A5BE82AFFDCAAB4F91AFBD1F939" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">
<emphasis id="B938D649E82AFFDCAAB4F91AFE83F941" bold="true" box="[199,345,1666,1691]" pageId="14" pageNumber="315">Description.</emphasis>
<typeStatus id="54F7B4F9E82AFFDCAB2CF91AFE10F941" box="[351,458,1666,1691]" pageId="14" pageNumber="315" type="holotype">Holotype</typeStatus>
, male. Body length approximately
<quantity id="4CB4A7BEE82AFFDCA920F91BFC76F940" box="[851,940,1667,1691]" metricMagnitude="-3" metricUnit="m" metricValue="7.8" pageId="14" pageNumber="315" unit="mm" value="7.8">7.8 mm</quantity>
. Head and pronotum dark-brown, sericeous; elytra submarmorate (scattered infuscated and yellow-brown patches), matt, largely due to microreticulation (
<date id="FFF22C9BE82AFFDCAF12F93FFA4AF965" box="[1377,1424,1703,1727]" pageId="14" pageNumber="315" value="1980-10">x80</date>
). Pilosity pale-yellow; dorsal surface with sparse, indistinct micro-stubbles only.
</paragraph>
<paragraph id="8BF30A5BE82AFFDCAAB4F976FC62F860" blockId="14.[151,1437,1126,1979]" pageId="14" pageNumber="315">Clypeal border round; clypeus anteromedially, just behind border, with slightly deflexed, slender horn; clypeus posterior surface slightly concave, apex rounded off; clypeogenal transition at border continuous; genal border shortly round near eye; clypeofrontal ridge absent. Clypeal surface anteriorly slightly uneven, impunctate, shiny; remainder of head matt, also impunctate, with microreticulation; 2 low paramedian protrusions between anterior part of eyes. Vertex base medially slightly elevated. Eyes with approximately 11 facet rows across widest point; ratio interocular distance / single maximum eye width approximately 7.</paragraph>
<caption id="DF335AD3E82BFFDDAAE4F9B5FC07F964" httpUri="https://zenodo.org/record/345763/files/figure.png" pageId="15" pageNumber="316" targetBox="[151,1415,230,1560]" targetPageId="15">
<paragraph id="8BF30A5BE82BFFDDAAE4F9B5FC07F964" blockId="15.[151,1437,1581,1726]" pageId="15" pageNumber="316">
<emphasis id="B938D649E82BFFDDAAE4F9B5FE8CF999" bold="true" box="[151,342,1581,1604]" pageId="15" pageNumber="316">FIGURES 39–46.</emphasis>
Sulawesi
<taxonomicName id="4C4C71D8E82BFFDDABB1F9B5FD91F999" box="[450,587,1581,1603]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="genus">
<emphasis id="B938D649E82BFFDDABB1F9B5FD91F999" box="[450,587,1581,1603]" italics="true" pageId="15" pageNumber="316">Onthophagus</emphasis>
</taxonomicName>
body parts. 39,
<taxonomicName id="4C4C71D8E82BFFDDA884F9B6FC59F999" box="[759,899,1581,1603]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E82BFFDDA884F9B6FC59F999" box="[759,899,1581,1603]" italics="true" pageId="15" pageNumber="316">O. bongkudai</emphasis>
</taxonomicName>
, head full-face, holotype male from Moajat. 40,
<taxonomicName id="4C4C71D8E82BFFDDAFF0F9B6FED8F9B8" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E82BFFDDAFF0F9B6FED8F9B8" italics="true" pageId="15" pageNumber="316">O. bongkudai</emphasis>
</taxonomicName>
, left elytron dorsal, holotype male from Moajat. 41,
<taxonomicName id="4C4C71D8E82BFFDDA96BF9D5FC7BF9B8" box="[792,929,1612,1634]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E82BFFDDA96BF9D5FC7BF9B8" box="[792,929,1612,1634]" italics="true" pageId="15" pageNumber="316">O. bongkudai</emphasis>
</taxonomicName>
, protibial upper side, holotype male from Moajat. 42,
<taxonomicName id="4C4C71D8E82BFFDDAACDF9F3FE90F95B" box="[190,330,1643,1665]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E82BFFDDAACDF9F3FE90F95B" box="[190,330,1643,1665]" italics="true" pageId="15" pageNumber="316">O. bongkudai</emphasis>
</taxonomicName>
, metatibial upperside, holotype male from Moajat. 43,
<taxonomicName id="4C4C71D8E82BFFDDA9F6F9F3FBCBF95B" box="[901,1041,1643,1665]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E82BFFDDA9F6F9F3FBCBF95B" box="[901,1041,1643,1665]" italics="true" pageId="15" pageNumber="316">O. bongkudai</emphasis>
</taxonomicName>
, aedeagus lateral, holotype male from Moajat. 44,
<taxonomicName id="4C4C71D8E82BFFDDAB6BF912FE7CF945" box="[280,422,1673,1695]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E82BFFDDAB6BF912FE7CF945" box="[280,422,1673,1695]" italics="true" pageId="15" pageNumber="316">O. bongkudai</emphasis>
</taxonomicName>
, head full-face, paratype female. 45,
<taxonomicName id="4C4C71D8E82BFFDDA945F912FC10F945" box="[822,970,1674,1695]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="sangirensis">
<emphasis id="B938D649E82BFFDDA945F912FC10F945" box="[822,970,1674,1695]" italics="true" pageId="15" pageNumber="316">O. sangirensis</emphasis>
</taxonomicName>
, aedeagus lateral, male from Toraut. 46,
<taxonomicName id="4C4C71D8E82BFFDDAFF0F912FED6F964" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="316" phylum="Arthropoda" rank="species" species="holosericus">
<emphasis id="B938D649E82BFFDDAFF0F912FED6F964" italics="true" pageId="15" pageNumber="316">O. holosericus</emphasis>
</taxonomicName>
, aedeagus lateral, male from Marena. Scale lines approximately 1 mm.
</paragraph>
</caption>
<paragraph id="8BF30A5BE82BFFDDAAB4F97EFB9CF854" blockId="15.[151,1436,1765,2006]" pageId="15" pageNumber="316">Pronotum strongly convex, discal midline not impressed; anterior section of lateral border distinctly sinuate, anterolateral corner slightly reflexed, angle approximately 80°; posterior section scarcely sinuate; pronotal base medially indistinctly, finely marginate; anterior declivity medially topped by slender, parallel-sided, anteriorly directed, slightly deflexed prong; anterior declivity impunctate, more shiny; rest of pronotal derm with sparse, vague, subocellate, fine punctures, mostly bearing fine suberect, short microstubble.</paragraph>
<paragraph id="8BF30A5BE82BFFC2AAB4F801FA87FF0F" blockId="15.[151,1436,1765,2006]" lastBlockId="16.[151,1437,151,968]" lastPageId="16" lastPageNumber="317" pageId="15" pageNumber="316">
Elytral striae discally well defined, with indistinct punctures, mostly separated by 3–6 times their diameter, scarcely crenulating interstriae; interstriae almost flat; interstriae 1–7 with sparse, microstubble-bearing, fine punctures and microreticulation (
<date id="FFF22C9BE834FFC2A864FF00FD9FFF6A" box="[535,581,152,176]" pageId="16" pageNumber="317" value="1980-10">x80</date>
), matt; interstria 8 with coarser, microseta-bearing punctures; derm more shiny; many punctures (especially on anterior section of interstriae) anteriorly with contiguous microgranule.
</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FF47FB0BFEBF" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">Antennal club brown. Metaventrite without distinct midline impression; with sparse, fine punctures only; lateral wings with abundant, seta-bearing, ocellate punctures. Visible abdominal ventrites 1–5 laterally with transverse row of seta-bearing, ocellate punctures; length of setae long, not appressed. Pygidium matt, yellow; darker medially with sparse, inconspicuous, seta-bearing, subocellate punctures; base marginate.</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FEE8FDF6FE2F" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">
Legs brown, femora with more yellow. Protibia with 3+(1) larger denticles, spur elongate-acuminate. Mesotibiae and metatibiae and spurs unmodified. Femoral undersides with sparse, seta-bearing punctures. Metatarsomere 1 nearly as long as metatarsomeres 2–5 combined; approximate proportions of spur and metatarsomeres 1–5: 12//
<date id="FFF22C9BE834FFC2ABC7FE44FE21FE2F" box="[436,507,476,501]" pageId="16" pageNumber="317" value="2014-03-06">14/6/3</date>
/2/5.
</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FE67FDCCFDC2" blockId="16.[151,1437,151,968]" box="[199,534,511,536]" pageId="16" pageNumber="317">
Aedeagus,
<figureCitation id="137716DEE834FFC2AB30FE67FE48FDC2" box="[323,402,511,536]" captionStart="FIGURES 39 – 46" captionStartId="15.[151,264,1581,1604]" captionTargetBox="[151,1415,230,1560]" captionTargetId="figure@15.[151,1436,193,1560]" captionTargetPageId="15" captionText="FIGURES 39 – 46. Sulawesi Onthophagus body parts. 39, O. bongkudai, head full-face, holotype male from Moajat. 40, O. bongkudai, left elytron dorsal, holotype male from Moajat. 41, O. bongkudai, protibial upper side, holotype male from Moajat. 42, O. bongkudai, metatibial upperside, holotype male from Moajat. 43, O. bongkudai, aedeagus lateral, holotype male from Moajat. 44, O. bongkudai, head full-face, paratype female. 45, O. sangirensis, aedeagus lateral, male from Toraut. 46, O. holosericus, aedeagus lateral, male from Marena. Scale lines approximately 1 mm." httpUri="https://zenodo.org/record/345763/files/figure.png" pageId="16" pageNumber="317">Fig. 43</figureCitation>
(
<typeStatus id="54F7B4F9E834FFC2ABD1FE67FDD0FDC2" box="[418,522,511,536]" pageId="16" pageNumber="317" type="holotype">holotype</typeStatus>
).
</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FDBDFD66FDBA" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">Measurements in mm. Maximum width of head 2.2. Median length of pronotum 2.4, maximum width 3.8. Sutural length of elytra 3.4, maximum width 4.2.</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FDF4FA56FD72" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">
<emphasis id="B938D649E834FFC2AAB4FDF4FD0EFD5F" bold="true" box="[199,724,620,645]" pageId="16" pageNumber="317">Variation, sexual dimorphism, distribution.</emphasis>
Body length 4.8–8.0 mm. Colour and microsculpture slightly variant (elytral pattern marmorate, but variably strongly defined, and also variant in amount of light versus dark).
</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FD2CFD18FCCE" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">
<emphasis id="B938D649E834FFC2AAB4FD2CFEC6FD16" box="[199,284,692,716]" italics="true" pageId="16" pageNumber="317">Female</emphasis>
with clypeus entirely densely rugulate; clypeal horn absent; clypeofrontal ridge curvilinear, medially distinct; frons between anterior part of eyes with two distinct paramedian protrusions. Pronotal prong tapering but greatly reduced. Pygidium more strongly convex.
</paragraph>
<paragraph id="8BF30A5BE834FFC2AAB4FCB8FD0FFC5A" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">
Note that this species was exclusively collected in two montane locations in
<collectingRegion id="4988C4B9E834FFC2AE53FC87FB0AFCE2" box="[1056,1232,799,824]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="317">North Sulawesi</collectingRegion>
, one appearing on most maps as Mount Moajat, the other being more to the west, indicated as Zebra (this will be Ice Station Zebra during Project Wallace,
<geoCoordinate id="EE786C9CE834FFC2ABD6FCFFFE29FC5A" box="[421,499,871,896]" direction="north" orientation="latitude" pageId="16" pageNumber="317" precision="925" value="0.43333334">0°26'N</geoCoordinate>
<geoCoordinate id="EE786C9CE834FFC2AB89FCF0FDBEFC5A" box="[506,612,872,896]" direction="east" orientation="longitude" pageId="16" pageNumber="317" precision="925" value="123.816666">123°49'E</geoCoordinate>
,
<quantity id="4CB4A7BEE834FFC2A81CFCFFFD13FC5A" box="[623,713,871,896]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="16" pageNumber="317" unit="m" value="1520.0">1520 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C35659D0E834FFC2AAB4FC14FB88FC12" pageId="16" pageNumber="317" type="etymology">
<paragraph id="8BF30A5BE834FFC2AAB4FC14FB88FC12" blockId="16.[151,1437,151,968]" pageId="16" pageNumber="317">
<emphasis id="B938D649E834FFC2AAB4FC14FE94FC7F" bold="true" box="[199,334,908,933]" pageId="16" pageNumber="317">Etymology.</emphasis>
Named after a village near the
<typeStatus id="54F7B4F9E834FFC2A8B8FC15FD21FC7F" box="[715,763,909,933]" pageId="16" pageNumber="317">type</typeStatus>
locality in
<collectingRegion id="4988C4B9E834FFC2A9F4FC14FBE6FC7F" box="[903,1084,908,933]" country="Indonesia" name="Sulawesi Utara" pageId="16" pageNumber="317">North Sulawesi</collectingRegion>
. The name
<taxonomicName id="4C4C71D8E834FFC2AEBAFC14FAE5FC7E" box="[1225,1343,908,932]" class="Insecta" family="Scarabaeidae" genus="Onthophagus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="317" phylum="Arthropoda" rank="species" species="bongkudai">
<emphasis id="B938D649E834FFC2AEBAFC14FAE5FC7E" box="[1225,1343,908,932]" italics="true" pageId="16" pageNumber="317">bongkudai</emphasis>
</taxonomicName>
is non- Latinized noun in apposition, and the village location can be found via Google Earth.
</paragraph>
</subSubSection>
</treatment>
</document>
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