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<mods:title id="868C78F9BB5FAED94D9065DD1100E6E0">Revisionary checklist of the Southern African Sesiini (Lepidoptera: Sesiidae) with description of new species</mods:title>
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<subSubSection id="C356657AFFD54410FF4FFD37FE57DDAD" box="[151,498,692,718]" pageId="26" pageNumber="27" type="nomenclature">
<paragraph id="8BF336F1FFD54410FF4FFD37FE57DDAD" blockId="26.[151,498,692,718]" box="[151,498,692,718]" pageId="26" pageNumber="27">
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<taxonomicName id="4C4C4D72FFD54410FF4FFD37FE57DDAD" ID-CoL="92Q9M" authority="Le Cerf, 1917" authorityName="Le Cerf" authorityYear="1917" box="[151,498,692,718]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFD54410FF4FFD37FE57DDAD" bold="true" box="[151,498,692,718]" pageId="26" pageNumber="27">
<emphasis id="B938EAE3FFD54410FF4FFD37FEE5DDAD" bold="true" box="[151,320,692,718]" italics="true" pageId="26" pageNumber="27">Vespanthedon</emphasis>
<bibRefCitation id="EFDD4B00FFD54410FE9FFD37FE57DDAD" author="Le Cerf" box="[327,498,692,718]" firstAuthor="Le Cerf" pageId="26" pageNumber="27" pagination="137 - 388" refId="ref29032" refString="Le Cerf, F. (1917) Contributions a l'etude des Aegeriidae. Description et Iconographie d'Especes et de Formes nouvelles ou peu connues. In: Oberthur, C. (Ed.), Etudes de Lepidopterologie Comparee, 14, 137 - 388, pls. 475 - 481." type="journal article" year="1917">Le Cerf, 1917</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFD54410FF4FFD78FF42DC0F" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="8BF336F1FFD54410FF4FFD78FC90DC73" blockId="26.[151,1436,762,876]" box="[151,821,762,784]" pageId="26" pageNumber="27">
<typeStatus id="54F78853FFD54410FF4FFD78FF6CDC73" box="[151,201,763,784]" pageId="26" pageNumber="27">Type</typeStatus>
species:
<taxonomicName id="4C4C4D72FFD54410FEFEFD79FD3ADC73" authority="Le Cerf, 1917" authorityName="Le Cerf" authorityYear="1917" box="[294,671,762,784]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cerceris">
<emphasis id="B938EAE3FFD54410FEFEFD79FDAFDC73" box="[294,522,762,784]" italics="true" pageId="26" pageNumber="27">Vespanthedon cerceris</emphasis>
<bibRefCitation id="EFDD4B00FFD54410FDC8FD78FD3ADC73" author="Le Cerf" box="[528,671,762,784]" firstAuthor="Le Cerf" pageId="26" pageNumber="27" pagination="137 - 388" refId="ref29032" refString="Le Cerf, F. (1917) Contributions a l'etude des Aegeriidae. Description et Iconographie d'Especes et de Formes nouvelles ou peu connues. In: Oberthur, C. (Ed.), Etudes de Lepidopterologie Comparee, 14, 137 - 388, pls. 475 - 481." type="journal article" year="1917">Le Cerf, 1917</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
<paragraph id="8BF336F1FFD54410FF4FFC9AFF42DC0F" blockId="26.[151,1436,762,876]" pageId="26" pageNumber="27">
Literature.
<bibRefCitation id="EFDD4B00FFD54410FED2FC99FE6BDC4C" author="Le Cerf" box="[266,462,793,815]" firstAuthor="Le Cerf" page="329" pageId="26" pageNumber="27" pagination="137 - 388" refId="ref29032" refString="Le Cerf, F. (1917) Contributions a l'etude des Aegeriidae. Description et Iconographie d'Especes et de Formes nouvelles ou peu connues. In: Oberthur, C. (Ed.), Etudes de Lepidopterologie Comparee, 14, 137 - 388, pls. 475 - 481." type="journal article" year="1917">Le Cerf 1917: 329</bibRefCitation>
;
<bibRefCitation id="EFDD4B00FFD54410FE03FC99FD06DC4C" author="Hampson" box="[475,675,793,815]" firstAuthor="Hampson" page="77" pageId="26" pageNumber="27" pagination="46 - 119" refId="ref28526" refString="Hampson, G. F. (1919) A classification of the Aegeriadae of the Oriental and Ethiopian Regions. Novitates Zoologicae, 26 (1), 46 - 119." type="journal article" year="1919">Hampson 1919: 77</bibRefCitation>
;
<bibRefCitation id="EFDD4B00FFD54410FD68FC9AFC5ADC4C" author="Dalla Torre &amp; Strand" box="[688,1023,793,815]" firstAuthor="Dalla Torre" page="119" pageId="26" pageNumber="27" refId="ref27751" refString="Dalla Torre, K. W. &amp; Strand, E. (1925) Aegeriidae. Lepidopterorum Catalogus, 31. W. Junk, Berlin, 202 pp." type="book" year="1925">Dalla Torre &amp; Strand 1925: 119</bibRefCitation>
; Gaede 1929: 527;
<bibRefCitation id="EFDD4B00FFD54410FB16FC99FA32DC4C" author="Naumann" box="[1230,1431,793,815]" firstAuthor="Naumann" page="31" pageId="26" pageNumber="27" refId="ref29205" refString="Naumann, C. M. (1971) Untersuchungen zur Systematik und Phylogenese der holarktischen Sesiiden (Insecta, Lepidoptera). Bonner Zoologische Monographien, 1, 190 pp." type="book" year="1971">Naumann 1971: 31</bibRefCitation>
; Fletcher &amp; Nye 1982: 168;
<bibRefCitation id="EFDD4B00FFD54410FE39FCBBFC8ADC2D" author="Heppner &amp; Duckworth" box="[481,815,824,846]" firstAuthor="Heppner" page="42" pageId="26" pageNumber="27" pagination="1 - 144" refId="ref28716" refString="Heppner, J. B. &amp; Duckworth, W. D. (1981) Classification of the Superfamily Sesioidea (Lepidoptera, Ditrysia). Smithsonian Contributions to Zoology, 314, 1 - 144. http: // dx. doi. org / 10.5479 / si. 00810282.314" type="journal article" year="1981">Heppner &amp; Duckworth 1981: 42</bibRefCitation>
;
<bibRefCitation id="EFDD4B00FFD54410FCE3FCBBFB27DC2D" author="Vari, Kroon &amp; Kruger" box="[827,1154,824,846]" firstAuthor="Vari" page="67" pageId="26" pageNumber="27" refId="ref29547" refString="Vari, L., Kroon, D. M. &amp; Kruger, M. (2002) Classification and Checklist of the species of Lepidoptera recorded in Southern Afrika. Simple Solutions, Chatswood, Australia, 384 pp." type="book" year="2002">Vári, Kroon &amp; Krüger 2002: 67</bibRefCitation>
;
<bibRefCitation id="EFDD4B00FFD54410FB56FCBBFF46DC0F" author="Puhringer &amp; Kallies" firstAuthor="Puhringer" page="45" pageId="26" pageNumber="27" pagination="185" refId="ref29350" refString="Puhringer, F. &amp; Kallies, A. (2004) Provisional checklist of the Sesiidae of the world (Lepidoptera: Ditrysia). Mitteilungen der Entomologischen Arbeitsgemeinschaft Salzkammergut, 4, 185. Available from: http: // www. sesiidae. net / Checklst. htm (accessed 30 October 2013)" type="journal article" year="2004">Pühringer &amp; Kallies 2004: 45</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFD54410FF4FFC1AFDB5D93D" pageId="26" pageNumber="27" type="description">
<paragraph id="8BF336F1FFD54410FF4FFC1AFD4FDA79" blockId="26.[151,1436,921,1990]" pageId="26" pageNumber="27">
<emphasis id="B938EAE3FFD54410FF4FFC1AFEE1DCD1" bold="true" box="[151,324,921,946]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Redescription.</emphasis>
Medium sized with wingspan
<quantity id="4CB49B14FFD54410FD43FC1AFCB3DCD2" box="[667,790,921,946]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" metricValueMax="3.0" metricValueMin="2.6" pageId="26" pageNumber="27" unit="mm" value="28.0" valueMax="30.0" valueMin="26.0">2630 mm</quantity>
. Head with frons 1.5× the diameter of the eye; haustellum of medium length, weakly sclerotized; labial palpus upturned, reaching at least up to antenna base, second palpomere somewhat longer than other ones, first and second palpomere ventrally with long, hair-like scales; male antenna short, clavate, bipectinate, ciliate, rami short, distally tapering, lacking at tip; female antenna without pectination or ciliae. Thorax and abdomen slender, smooth scaled; abdominal segments 2 and 3 of the male somewhat constricted, forming a weak wasp-waist; anal tuft narrow, abdomen of the female spindle-shaped. Legs smooth scaled; hind leg as long as abdomen; all pairs of spurs with lateral ones half as long as mesal ones. Wings narrow, partially hyaline; forewing with costal area and costal parts of cell and apical area opaque; discal spots indistinct; that of hindwing very short and small. Wing venation (
<figureCitation id="13772A74FFD54410FC0DFB3AFB95DBB1" box="[981,1072,1209,1234]" captionStart="FIGURES 8491" captionStartId="38.[151,264,1763,1786]" captionTargetBox="[160,1428,202,1727]" captionTargetId="figure-70@38.[151,1436,193,1742]" captionTargetPageId="38" captionText="FIGURES 8491. Wing venations: (84) Cicinnoscelis longipes Holland, 1893; (85) Alonina rygchiiformis Walker, 1856; (86) Anaudia felderi Wallengren, 1863, syntype; (87) Austrosetia semirufa Felder &amp; Felder, 1874; (88) Felderiola candescens (Felder &amp; Felder, 1874); (89) Monopetalotaxis doleriformis (Walker, 1856); (90) Hagnogyna sanguicosta (Hampson, 1919); (91) Vespanthedon cerceris Le Cerf, 1917, holotype." figureDoi="http://doi.org/10.5281/zenodo.10099887" httpUri="https://zenodo.org/record/10099887/files/figure.png" pageId="26" pageNumber="27">Fig. 91</figureCitation>
) with forewing veins R1R3 approximated, R4 and R5 stalked, veins from common stalk of R4/R5 to CuA2 arise equally spaced from cross vein; hindwing with common stalk of M3 and CuA1.
</paragraph>
<paragraph id="8BF336F1FFD54410FF1FFAA6FCD3D959" blockId="26.[151,1436,921,1990]" pageId="26" pageNumber="27">
<emphasis id="B938EAE3FFD54410FF1FFAA6FE57DA5D" box="[199,498,1317,1342]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">
<emphasis id="B938EAE3FFD54410FF1FFAA6FED6DA5D" bold="true" box="[199,371,1317,1342]" pageId="26" pageNumber="27">Male genitalia</emphasis>
(
<figureCitation id="13772A74FFD54410FE5BFAA6FE43DA5D" box="[387,486,1317,1342]" captionStart="FIGURES 115116" captionStartId="45.[151,264,1784,1807]" captionTargetBox="[151,1424,205,1763]" captionTargetId="figure-33@45.[151,1436,193,1763]" captionTargetPageId="45" captionText="FIGURES 115116. Male genitalia: (115) Austrosetia semirufa Felder &amp; Felder, 1874; (116) Vespanthedon cerceris Le Cerf, 1917, holotype." figureDoi="http://doi.org/10.5281/zenodo.10099901" httpUri="https://zenodo.org/record/10099901/files/figure.png" pageId="26" pageNumber="27">Fig. 116</figureCitation>
).
</emphasis>
Tegumen rather slender, medio-distally somewhat laterally constricted; gnathos of unique, triangular, mitre-like shape; uncus short, basally narrow, ventro-lateral margins densely covered with multifurcate, scale-like, basad pointing setae; valva rectangular, margins almost straight, dorsal one somewhat convex, ventral one slightly concave, inner surface of valva distally and dorsally with numerous broad, multifurcate, scale-like setae and medio-ventrally with short, distally with ventral margin connected crista sacculi, which has distally a distinct tooth-like protrusion; juxta broad, very short protruded, manica with numerous sclerotized spines; saccus long, very narrow, apically rounded; phallus straight, narrow with short coecum penis; bulbous dorso-proximal part of vesica with numerous fine spines.
</paragraph>
<paragraph id="8BF336F1FFD54410FF1FF9C6FDB5D93D" blockId="26.[151,1436,921,1990]" box="[199,528,1605,1630]" pageId="26" pageNumber="27">
<emphasis id="B938EAE3FFD54410FF1FF9C6FE36D93D" bold="true" box="[199,403,1605,1630]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Female genitalia.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFD54410FF1FF9EAFAE4D81D" pageId="26" pageNumber="27" type="diagnosis">
<paragraph id="8BF336F1FFD54410FF1FF9EAFB80D9AA" blockId="26.[151,1436,921,1990]" pageId="26" pageNumber="27">
<emphasis id="B938EAE3FFD54410FF1FF9EAFEE4D9E1" bold="true" box="[199,321,1641,1666]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Diagnosis.</emphasis>
Putative synapomorphies are: (1) antenna very short, bipectinate, rami lacking at tip; (2) gnathos present, with unique, mitre-like shape; (3) setae of valva large, multifurcate, scale-like; (4) wings very narrow, hindwing tornus and anal veins somewhat reduced; (6) costal part of cell opaque.
</paragraph>
<paragraph id="8BF336F1FFD54410FF4FF956FAE4D81D" blockId="26.[151,1436,921,1990]" pageId="26" pageNumber="27">
Similar, scale-like setae of the valva are only known in
<taxonomicName id="4C4C4D72FFD54410FCD3F956FBEBD98D" authority="Hampson, 1910" authorityName="Hampson" authorityYear="1910" box="[779,1102,1749,1774]" class="Insecta" family="Sesiidae" genus="Sphecosesia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFD54410FCD3F956FC31D98D" box="[779,916,1749,1774]" italics="true" pageId="26" pageNumber="27">Sphecosesia</emphasis>
<bibRefCitation id="EFDD4B00FFD54410FC44F955FBEBD98D" author="Hampson" box="[924,1102,1749,1774]" firstAuthor="Hampson" pageId="26" pageNumber="27" pagination="145 - 160" refId="ref28465" refString="Hampson, G. F. (1910) Descriptions of new African Moths. The Annals and Magazine of Natural History (Eighth Series), 5 / 6, 430 - 496, 116 - 141, 145 - 160. [Sesiidae 149 - 156] http: // dx. doi. org / 10.1080 / 00222931008692834" type="journal article" year="1910">Hampson, 1910</bibRefCitation>
</taxonomicName>
from
<collectingRegion id="4988F813FFD54410FB4CF956FAA4D98D" box="[1172,1281,1749,1774]" country="Botswana" name="South East" pageId="26" pageNumber="27">Southeast</collectingRegion>
Asia (
<bibRefCitation id="EFDD4B00FFD54410FA93F956FE9FD871" author="Kallies &amp; Arita" firstAuthor="Kallies" pageId="26" pageNumber="27" pagination="65 - 95" refId="ref28884" refString="Kallies, A. &amp; Arita, Y. (2004) A survey of the clearwing moths of the tribe Sesiini (Lepidoptera, Sesiidae) from Vietnam and adjacent countries with a synopsis of the Oriental Sesiini fauna. Tinea, 18 (1), 65 - 95." type="journal article" year="2004">Kallies &amp; Arita 2004</bibRefCitation>
). Species of this genus have also narrow wings and short antennas, but differ distinctly by the strongly wasp-waisted abdomen, the rough and short scales of the first and the smooth scales of the second palpomere rather than long, hair-like scales in
<taxonomicName id="4C4C4D72FFD54410FD69F8C2FCE8D839" authorityName="Le Cerf" authorityYear="1917" box="[689,845,1857,1882]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFD54410FD69F8C2FCE8D839" box="[689,845,1857,1882]" italics="true" pageId="26" pageNumber="27">Vespanthedon</emphasis>
</taxonomicName>
. The male genitalia of
<taxonomicName id="4C4C4D72FFD54410FB86F8C2FB42D839" authorityName="Hampson" authorityYear="1910" box="[1118,1255,1857,1882]" class="Insecta" family="Sesiidae" genus="Sphecosesia" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFD54410FB86F8C2FB42D839" box="[1118,1255,1857,1882]" italics="true" pageId="26" pageNumber="27">Sphecosesia</emphasis>
</taxonomicName>
have the uncus lobes very large (small in
<taxonomicName id="4C4C4D72FFD54410FE62F8E6FDF3D81D" authorityName="Le Cerf" authorityYear="1917" box="[442,598,1893,1918]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFD54410FE62F8E6FDF3D81D" box="[442,598,1893,1918]" italics="true" pageId="26" pageNumber="27">Vespanthedon</emphasis>
</taxonomicName>
) and lack the crista sacculi of the valva (present in
<taxonomicName id="4C4C4D72FFD54410FB4DF8E6FA94D81D" authorityName="Le Cerf" authorityYear="1917" box="[1173,1329,1893,1918]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="genus">
<emphasis id="B938EAE3FFD54410FB4DF8E6FA94D81D" box="[1173,1329,1893,1918]" italics="true" pageId="26" pageNumber="27">Vespanthedon</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C356657AFFD54410FF1FF80AFE22D8A5" pageId="26" pageNumber="27" type="distribution">
<paragraph id="8BF336F1FFD54410FF1FF80AFE22D8A5" blockId="26.[151,1436,921,1990]" pageId="26" pageNumber="27">
<emphasis id="B938EAE3FFD54410FF1FF80AFEC5D8C1" bold="true" box="[199,352,1929,1954]" inLineHeading="true" pageId="26" pageNumber="27" reason="1">Distribution.</emphasis>
Insufficiently known;
<taxonomicName id="4C4C4D72FFD54410FDA8F80AFCD5D8C2" authorityName="Le Cerf" authorityYear="1917" box="[624,880,1929,1954]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="cerceris">
<emphasis id="B938EAE3FFD54410FDA8F80AFCD5D8C2" box="[624,880,1929,1954]" italics="true" pageId="26" pageNumber="27">Vespanthedon cerceris</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4C4D72FFD54410FC6AF809FBFFD8C1" baseAuthorityName="Hampson" baseAuthorityYear="1919" box="[946,1114,1929,1954]" class="Insecta" family="Sesiidae" genus="Vespanthedon" kingdom="Animalia" order="Lepidoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="chalciphora">
<emphasis id="B938EAE3FFD54410FC6AF809FBFFD8C1" box="[946,1114,1929,1954]" italics="true" pageId="26" pageNumber="27">V. chalciphora</emphasis>
</taxonomicName>
occur in
<collectingCountry id="F35B7661FFD54410FB0BF80AFAC2D8C1" box="[1235,1383,1929,1954]" name="Mozambique" pageId="26" pageNumber="27">Mozambique</collectingCountry>
and
<collectingCountry id="F35B7661FFD54410FF4FF82EFF4AD8A5" box="[151,239,1965,1990]" name="Malawi" pageId="26" pageNumber="27">Malawi</collectingCountry>
, respectively.
</paragraph>
</subSubSection>
</treatment>
</document>