368 lines
54 KiB
XML
368 lines
54 KiB
XML
<document id="6A170532A27B3629AD0E240847CF8BF6" ID-CLB-Dataset="22127" ID-DOI="10.5852/ejt.2013.70" ID-GBIF-Dataset="2b8d9c95-6a86-4b09-bf6e-06965827cf87" ID-ISSN="2118-9773" ID-Zenodo-Dep="3827701" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1589478096172" checkinUser="carolina" docAuthor="Lazányi, Eszter & Vagalinski, Boyan" docDate="2013" docId="03E5878D0477054CFD9E47935ECEB362" docLanguage="en" docName="ejt-2013-70.pdf.imf" docOrigin="European Journal of Taxonomy 70" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Enghophyllum Lazányi & Vagalinski 2013, gen. nov." docType="treatment" docUuid="15008961-CD77-4325-B694-13BA1F7E514E" docUuidSource="ZooBank" docVersion="7" lastPageNumber="4" masterDocId="FFDCFFF50476054FFFC343255D05B551" masterDocTitle="A new genus of the millipede tribe Brachyiulini (Diplopoda: Julida: Julidae) from the Aegean region" masterLastPageNumber="12" masterPageNumber="1" pageNumber="2" updateTime="1698839752143" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A2ECEB41D8D6A4875B03C1A5A082BDD0">A new genus of the millipede tribe Brachyiulini (Diplopoda: Julida: Julidae) from the Aegean region</mods:title>
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<mods:name id="432F387707815481E686BC179165F63C" type="personal">
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<mods:roleTerm id="498E34903B80FBB7B44B784D882A3A39">Author</mods:roleTerm>
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<mods:namePart id="A7D14E45AB187EEC54B97CC3622A062B">Lazányi, Eszter</mods:namePart>
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<mods:affiliation id="D4B357BC9F06731E1B58D279A94DDDDB">Corresponding author: Department of Zoology, Hungarian Natural History Museum, Baross u. 13, H- 1088 Budapest, Hungary. E-mail: pesca 12 @ gmail. com & urn: lsid: zoobank. org: author: 02 DB 48 F 1 - 624 C- 4435 - AF 85 - FA 87168 CD 85 A</mods:affiliation>
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<mods:namePart id="4064AF195A4F050233B54E96AB923700">Vagalinski, Boyan</mods:namePart>
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<mods:affiliation id="3F4AFE902F7DFB5651E9E5D6EDCE95B2">Faculty of Biology, Sofia University, 8 Dragan Tsankov Blvd., 1164 Sofia, Bulgaria. E-mail: boyan _ vagalinski @ excite. com & Institute of Biodiversity and Ecosystem Research, Bulgarian Academy of Sciences, 2 Yurii Gagarin Street, 1113, Sofia, Bulgaria. & urn: lsid: zoobank. org: author: 973 B 8725 - 039 E- 4 F 29 - 8 D 73 - 96 A 7 F 52 CF 934</mods:affiliation>
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<mods:typeOfResource id="B3E597ECDB318F95777413AD9F1D922A">text</mods:typeOfResource>
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<mods:title id="3AE4684D663153B0D15580D00CF21531">European Journal of Taxonomy</mods:title>
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<mods:part id="A864239C463014493092CF894C4B8DF9">
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<mods:date id="1B739C88723DFC8517F1FF124229DB22">2013</mods:date>
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<mods:number id="0B2BBCF16CA26B5D2035149BE15248BB">2013-12-17</mods:number>
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<mods:number id="19430F3858DFF577537E2BED0216323C">70</mods:number>
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<treatment id="03E5878D0477054CFD9E47935ECEB362" ID-DOI="http://doi.org/10.5281/zenodo.3846681" ID-GBIF-Taxon="164152705" ID-Zenodo-Dep="3846681" LSID="urn:lsid:zoobank.org:act:15008961-CD77-4325-B694-13BA1F7E514E" httpUri="http://treatment.plazi.org/id/03E5878D0477054CFD9E47935ECEB362" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2">
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<subSubSection id="C35665100477054EFD9E47935ED2B181" box="[605,983,1205,1232]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="8BF3369B0477054EFD9E47935ED2B181" blockId="1.[375,1213,1205,1268]" box="[605,983,1205,1232]" pageId="1" pageNumber="2">
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<heading id="D0BB81F70477054EFD9E47935ED2B181" box="[605,983,1205,1232]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
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Genus
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<taxonomicName id="4C4C4D180477054EFD6C47905E61B181" ID-CoL="7PD8Y" authority="Lazányi & Vagalinski, 2013" authorityName="Lazányi & Vagalinski" authorityYear="2013" box="[687,868,1205,1232]" class="Diplopoda" family="Julidae" genus="Enghophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="B938EA890477054EFD6C47905E61B181" bold="true" box="[687,868,1205,1232]" italics="true" pageId="1" pageNumber="2">Enghophyllum</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20B57F20477054EFCAF47935ED2B181" box="[876,983,1206,1232]" pageId="1" pageNumber="2" rank="genus">gen. nov.</taxonomicNameLabel>
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</heading>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C35665100477054EFEB447FC59B8B1A5" box="[375,1213,1241,1268]" pageId="1" pageNumber="2" type="description">
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<uri id="FFDD3A990477054EFEB447FC59B8B1A5" box="[375,1213,1241,1268]" pageId="1" pageNumber="2">
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urn:lsid:zoobank.org:act:
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35665100477054EFF7E463F5E1AB032" pageId="1" pageNumber="2" type="type_taxon">
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<paragraph id="8BF3369B0477054EFF7E463F5C5DB065" blockId="1.[189,344,1306,1332]" box="[189,344,1306,1332]" pageId="1" pageNumber="2">
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<heading id="D0BB81F70477054EFF7E463F5C5DB065" bold="true" box="[189,344,1306,1332]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
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<emphasis id="B938EA890477054EFF7E463F5C5DB065" bold="true" box="[189,344,1306,1332]" pageId="1" pageNumber="2">
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<typeStatus id="54F788390477054EFF7E463F5DFFB065" box="[189,250,1306,1332]" pageId="1" pageNumber="2">Type</typeStatus>
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species
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF3369B0477054EFF7E466C5E1AB032" blockId="1.[189,799,1353,1379]" box="[189,799,1353,1379]" pageId="1" pageNumber="2">
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<taxonomicName id="4C4C4D180477054EFF7E466C5E1AB032" ID-CoL="7SS4R" authority="Verhoeff, 1901" authorityName="Verhoeff" authorityYear="1901" box="[189,799,1353,1379]" class="Diplopoda" family="Julidae" genus="Brachyiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="naxius" subGenus="Chromatoiulus">
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<emphasis id="B938EA890477054EFF7E466C5C4EB032" box="[189,331,1353,1379]" italics="true" pageId="1" pageNumber="2">Brachyiulus</emphasis>
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(
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<emphasis id="B938EA890477054EFE9F466C5F0EB032" box="[348,523,1353,1379]" italics="true" pageId="1" pageNumber="2">Chromatoiulus</emphasis>
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)
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<emphasis id="B938EA890477054EFDDF466F5F6CB032" box="[540,617,1354,1379]" italics="true" pageId="1" pageNumber="2">naxius</emphasis>
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<bibRefCitation id="EFDD4B6A0477054EFDAC466C5E1AB032" author="Verhoeff C." box="[623,799,1353,1379]" pageId="1" pageNumber="2" pagination="79 - 102" refId="ref5702" refString="Verhoeff C. 1901. Beitrage zur Kenntniss palaarktischer Myriopoden. XVII. Aufsatz: Diplopoden aus dem Mittelmeergebiet. Archiv fur Naturgeschichte 67 (1): 79 - 102." type="journal article" year="1901">Verhoeff, 1901</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35665100477054EFF7E46AF5F57B331" pageId="1" pageNumber="2" type="diagnosis">
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<paragraph id="8BF3369B0477054EFF7E46AF5C33B0F5" blockId="1.[189,310,1418,1444]" box="[189,310,1418,1444]" pageId="1" pageNumber="2">
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||
<heading id="D0BB81F70477054EFF7E46AF5C33B0F5" bold="true" box="[189,310,1418,1444]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
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<emphasis id="B938EA890477054EFF7E46AF5C33B0F5" bold="true" box="[189,310,1418,1444]" pageId="1" pageNumber="2">Diagnosis</emphasis>
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</heading>
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||
</paragraph>
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<paragraph id="8BF3369B0477054EFF7E469C5F57B331" blockId="1.[189,1399,1465,1632]" pageId="1" pageNumber="2">
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A genus of
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<taxonomicName id="4C4C4D180477054EFE89469C5CE5B082" authorityName="Verhoeff" authorityYear="1909" box="[330,480,1465,1491]" class="Diplopoda" family="Julidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Brachyiulini">Brachyiulini</taxonomicName>
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, differing from contribal genera by the following combination of characters: promere broad, shield-like,
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<emphasis id="B938EA890477054EFDCB46F85F57B0A7" box="[520,594,1501,1526]" italics="true" pageId="1" pageNumber="2">in situ</emphasis>
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protruding mostly posteriad, completely covering opisthomere and gonopodal sinus; transverse muscles and coxal apodemes of the promere fully reduced. Opisthomere with 3 well-differentiated processes: lateral (
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<emphasis id="B938EA890477054EFD2945075E04B36D" box="[746,769,1570,1596]" italics="true" pageId="1" pageNumber="2">lp</emphasis>
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), basal posterior (
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<emphasis id="B938EA890477054EFC2945075913B36D" box="[1002,1046,1570,1596]" italics="true" pageId="1" pageNumber="2">bpp</emphasis>
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) and apical posterior (
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<emphasis id="B938EA890477054EFAFA45065860B36D" box="[1337,1381,1571,1596]" italics="true" pageId="1" pageNumber="2">app</emphasis>
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). Solenomere rather simple, tubular.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35665100477054EFF7E45A25C96B247" pageId="1" pageNumber="2" type="etymology">
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<paragraph id="8BF3369B0477054EFF7E45A25C41B3F0" blockId="1.[189,324,1671,1697]" box="[189,324,1671,1697]" pageId="1" pageNumber="2">
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<heading id="D0BB81F70477054EFF7E45A25C41B3F0" bold="true" box="[189,324,1671,1697]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
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<emphasis id="B938EA890477054EFF7E45A25C41B3F0" bold="true" box="[189,324,1671,1697]" pageId="1" pageNumber="2">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF3369B0477054EFF7E45905C96B247" blockId="1.[189,1398,1717,1814]" pageId="1" pageNumber="2">
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This genus is named in honor of Prof. Henrik Enghoff from the ZMUC, not only for his vast contribution to our knowledge of diplopods, but also for his continuous encouragement and help during our work with
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<taxonomicName id="4C4C4D180477054EFF3B45D95C8AB247" authorityName="Verhoeff" authorityYear="1909" box="[248,399,1788,1814]" class="Diplopoda" family="Julidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="tribe" tribe="Brachyiulini">Brachyiulini</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35665100477054CFF7E44185975B00B" lastPageId="3" lastPageNumber="4" pageId="1" pageNumber="2" type="description">
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<paragraph id="8BF3369B0477054EFF7E44185C4BB206" blockId="1.[189,334,1853,1879]" box="[189,334,1853,1879]" pageId="1" pageNumber="2">
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<heading id="D0BB81F70477054EFF7E44185C4BB206" bold="true" box="[189,334,1853,1879]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
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<emphasis id="B938EA890477054EFF7E44185C4BB206" bold="true" box="[189,334,1853,1879]" pageId="1" pageNumber="2">Description</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BF3369B0477054EFF7E44495FD1B2A1" blockId="1.[189,1399,1900,2032]" pageId="1" pageNumber="2">
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Two frontal, four supralabral and 14-18 labral setae. Antennomeres 4 and 5 subequal in length (
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<figureCitation id="13772A1E0477054EFACF4449586DB2D7" box="[1292,1384,1900,1926]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="1" pageNumber="2">Fig. 1A</figureCitation>
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). Gnathochilarium as on
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<figureCitation id="13772A1E0477054EFE0F44AA5F20B2F8" box="[460,549,1935,1961]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="1" pageNumber="2">Fig. 1B</figureCitation>
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. Metazona sparsely striated, as emphasised by
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<bibRefCitation id="EFDD4B6A0477054EFB8744AA5804B2F8" author="Verhoeff C." box="[1092,1281,1935,1961]" pageId="1" pageNumber="2" pagination="79 - 102" refId="ref5702" refString="Verhoeff C. 1901. Beitrage zur Kenntniss palaarktischer Myriopoden. XVII. Aufsatz: Diplopoden aus dem Mittelmeergebiet. Archiv fur Naturgeschichte 67 (1): 79 - 102." type="journal article" year="1901">
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Verhoeff (
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<date id="FFF2105B0477054EFB7A44AA59FDB2F8" box="[1209,1272,1935,1961]" pageId="1" pageNumber="2" value="1901">1901</date>
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||
)
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</bibRefCitation>
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; ozopores right on the suture (
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<figureCitation id="13772A1E0477054EFE6D44975F09B29D" box="[430,524,1970,1996]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="1" pageNumber="2">Fig. 1C</figureCitation>
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). Preanal process straight, anal valves sparsely pilose (
|
||
<figureCitation id="13772A1E0477054EFB6744975806B29D" box="[1188,1283,1970,1996]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="1" pageNumber="2">Fig. 1D</figureCitation>
|
||
); subanal scale triangular, with protruding tip (
|
||
<figureCitation id="13772A1E0477054EFDAF44F05FC3B2A1" box="[620,710,2005,2032]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="1" pageNumber="2">Fig. 1E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF3369B0474054DFF29422E5C30B474" blockId="2.[189,1399,267,505]" box="[234,309,267,293]" pageId="2" pageNumber="3">
|
||
<emphasis id="B938EA890474054DFF29422E5C30B474" bold="true" box="[234,309,267,293]" pageId="2" pageNumber="3">Males</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF3369B0474054CFF7E420B5E76B418" blockId="2.[189,1399,267,505]" lastBlockId="3.[189,1398,267,329]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
|
||
Mandibular stipes slightly protruding (
|
||
<figureCitation id="13772A1E0474054DFD63420B5FEDB419" box="[672,744,302,328]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
|
||
F-G). First leg-pair like simple, rounded, somewhat converging hooks; the two hooks converging in an obtuse angle (
|
||
<figureCitation id="13772A1E0474054DFC6E4274590CB43A" box="[941,1033,337,363]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="2" pageNumber="3">Fig. 1H</figureCitation>
|
||
). Second leg-pair (
|
||
<figureCitation id="13772A1E0474054DFB2742745837B43A" box="[1252,1330,337,363]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="2" pageNumber="3">
|
||
Fig.
|
||
<date id="FFF2105B0474054DFAD942745837B43A" box="[1306,1330,337,363]" pageId="2" pageNumber="3">1I</date>
|
||
</figureCitation>
|
||
) with two ventral pads: on postfemur (
|
||
<emphasis id="B938EA890474054DFD8642505F59B4DE" box="[581,604,373,399]" italics="true" pageId="2" pageNumber="3">pf</emphasis>
|
||
) and tibia (
|
||
<emphasis id="B938EA890474054DFD2942505FF7B4DF" box="[746,754,373,398]" italics="true" pageId="2" pageNumber="3">t</emphasis>
|
||
). Penis (
|
||
<figureCitation id="13772A1E0474054DFC9D42515EB5B4DE" box="[862,944,372,399]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="2" pageNumber="3">Fig. 1J</figureCitation>
|
||
) significantly small, deeply hidden in penis sac, with two very short lobes and two rounded small lamellae running parallel, i.e., not diverging (
|
||
<emphasis id="B938EA890474054DFF04429E5DDEB484" box="[199,219,443,469]" italics="true" pageId="2" pageNumber="3">rl</emphasis>
|
||
and
|
||
<emphasis id="B938EA890474054DFED5429E5C23B484" box="[278,294,443,469]" italics="true" pageId="2" pageNumber="3">ll</emphasis>
|
||
for right and left lamella). Pleurotergum of the 7
|
||
<superScript id="7C399BD30474054DFCB1429C5E7AB496" attach="left" box="[882,895,441,455]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
|
||
body ring protruding like a simple shovel (broken in the unique
|
||
<typeStatus id="54F788390474054DFE7942FB5F24B4A9" box="[442,545,478,504]" pageId="2" pageNumber="3" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C4C4D180474054DFD8642FA5FBFB4A9" authority="Lazányi & Vagalinski, 2013" authorityName="Lazányi & Vagalinski" authorityYear="2013" box="[581,698,478,504]" class="Diplopoda" family="Julidae" genus="Enghophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sifnium" status="gen. et sp. nov.">
|
||
<emphasis id="B938EA890474054DFD8642FA5FBFB4A9" box="[581,698,478,504]" italics="true" pageId="2" pageNumber="3">E. sifnium</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57F20474054DFD0342FA5E6AB4A8" box="[704,879,479,505]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, thus investigated only in
|
||
<taxonomicName id="4C4C4D180474054DFB5942FA580AB4A9" baseAuthorityName="Verhoeff" baseAuthorityYear="1903" box="[1178,1295,479,504]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="naxium">
|
||
<emphasis id="B938EA890474054DFB5942FA580AB4A9" box="[1178,1295,479,504]" italics="true" pageId="2" pageNumber="3">E. naxium</emphasis>
|
||
</taxonomicName>
|
||
, see
|
||
<figureCitation id="13772A1E0474054CFA8B42FB5DE6B474" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">Fig. 1K</figureCitation>
|
||
). Walking legs (
|
||
<figureCitation id="13772A1E0475054CFE63422E5CDAB474" box="[416,479,267,293]" captionStart="Fig" captionStartId="2.[189,232,1758,1784]" captionTargetBox="[211,1393,605,1683]" captionTargetId="figure@2.[189,1399,591,1716]" captionTargetPageId="2" captionText="Fig. 1. Non-gonopod characters of Enghopyhllum gen. nov., males.A. Right antenna.B. Gnathochilarium. C. 8th and 9th body rings. D. Telson. E. Subanal scale. F. Mandibular stipital (“cheek”) lobe, lateral view. G. Mandibular stipital (“cheek”) lobe, ventro-lateral view. H. First leg-pair. I. Second leg-pair. J. Penis in the penis sac. K. Pleurotergum of the 7th body ring, ventral view. L. Anterior left leg on the 6th body ring. M. Anterior right leg on the 22nd body ring. – A-D, F, H-I, L-M: E. sifnium gen. et sp. nov., holotype. E: E. naxium, from Antiparos. G, J-K: E. naxium, from Mavri Islet." figureDoi="http://doi.org/10.5281/zenodo.3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
L-M) with two ventral pads: on postfemur (
|
||
<emphasis id="B938EA890475054CFC2342295EF2B477" box="[992,1015,268,294]" italics="true" pageId="3" pageNumber="4">pf</emphasis>
|
||
) and tibia (
|
||
<emphasis id="B938EA890475054CFBBE42295980B474" box="[1149,1157,268,293]" italics="true" pageId="3" pageNumber="4">t</emphasis>
|
||
). Tarsus of midbody legs slightly longer than tibia: tarsus / tibia = (1.05-1.1) / 1.
|
||
</paragraph>
|
||
<caption id="DF3366130474054DFF7E45FB5ED7B2F9" ID-DOI="http://doi.org/10.5281/zenodo.3827703" ID-Zenodo-Dep="3827703" httpUri="https://zenodo.org/record/3827703/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,232,1758,1784]" targetBox="[211,1393,605,1683]" targetPageId="2">
|
||
<paragraph id="8BF3369B0474054DFF7E45FB5ED7B2F9" blockId="2.[189,1399,1758,1961]" pageId="2" pageNumber="3">
|
||
<emphasis id="B938EA890474054DFF7E45FB5C02B3A9" bold="true" box="[189,263,1758,1784]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
|
||
Non-gonopod characters of
|
||
<taxonomicName id="4C4C4D180474054DFD8945FB5FF0B3A9" authority="Lazányi & Vagalinski, 2013" authorityName="Lazányi & Vagalinski" authorityYear="2013" box="[586,757,1758,1784]" class="Diplopoda" family="Julidae" genus="Enghopyhllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
||
<emphasis id="B938EA890474054DFD8945FB5FF0B3A9" box="[586,757,1758,1784]" italics="true" pageId="2" pageNumber="3">Enghopyhllum</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57F20474054DFD3945FA5E61B3A8" box="[762,868,1759,1785]" pageId="2" pageNumber="3" rank="genus">gen. nov.</taxonomicNameLabel>
|
||
, males.
|
||
<emphasis id="B938EA890474054DFC7F45FB5ED4B3A9" bold="true" box="[956,977,1758,1784]" pageId="2" pageNumber="3">A</emphasis>
|
||
. Right antenna.
|
||
<emphasis id="B938EA890474054DFB4A45FA5998B3A8" bold="true" box="[1161,1181,1759,1785]" pageId="2" pageNumber="3">B</emphasis>
|
||
. Gnathochilarium.
|
||
<emphasis id="B938EA890474054DFF7E44245DD7B24A" bold="true" box="[189,210,1793,1819]" pageId="2" pageNumber="3">C</emphasis>
|
||
. 8
|
||
<superScript id="7C399BD30474054DFF2D44255DFEB25F" attach="left" box="[238,251,1792,1806]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
|
||
and 9
|
||
<superScript id="7C399BD30474054DFE8244255C4BB25F" attach="left" box="[321,334,1792,1806]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
|
||
body rings.
|
||
<emphasis id="B938EA890474054DFE1D44275CF6B24D" bold="true" box="[478,499,1794,1820]" pageId="2" pageNumber="3">D</emphasis>
|
||
. Telson.
|
||
<emphasis id="B938EA890474054DFD9F44275F75B24D" bold="true" box="[604,624,1794,1820]" pageId="2" pageNumber="3">E</emphasis>
|
||
. Subanal scale.
|
||
<emphasis id="B938EA890474054DFCE844275E38B24D" bold="true" box="[811,829,1794,1820]" pageId="2" pageNumber="3">F</emphasis>
|
||
. Mandibular stipital (“cheek”) lobe, lateral view.
|
||
<emphasis id="B938EA890474054DFF7E44005DD1B26E" bold="true" box="[189,212,1829,1855]" pageId="2" pageNumber="3">G</emphasis>
|
||
. Mandibular stipital (“cheek”) lobe, ventro-lateral view.
|
||
<emphasis id="B938EA890474054DFCAC44005E83B26E" bold="true" box="[879,902,1829,1855]" pageId="2" pageNumber="3">H</emphasis>
|
||
. First leg-pair.
|
||
<emphasis id="B938EA890474054DFBF944005940B26E" bold="true" box="[1082,1093,1829,1855]" pageId="2" pageNumber="3">I</emphasis>
|
||
. Second leg-pair.
|
||
<emphasis id="B938EA890474054DFAD94400582CB26E" bold="true" box="[1306,1321,1829,1855]" pageId="2" pageNumber="3">J</emphasis>
|
||
. Penis in the penis sac.
|
||
<emphasis id="B938EA890474054DFE41446D5C9CB233" bold="true" box="[386,409,1864,1890]" pageId="2" pageNumber="3">K</emphasis>
|
||
. Pleurotergum of the 7
|
||
<superScript id="7C399BD30474054DFD6944635FB2B205" attach="left" box="[682,695,1862,1876]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
|
||
body ring, ventral view.
|
||
<emphasis id="B938EA890474054DFC23446D5EF1B233" bold="true" box="[992,1012,1864,1890]" pageId="2" pageNumber="3">L</emphasis>
|
||
. Anterior left leg on the 6
|
||
<superScript id="7C399BD30474054DFAE444635831B205" attach="left" box="[1319,1332,1862,1876]" fontSize="6" pageId="2" pageNumber="3">th</superScript>
|
||
body ring.
|
||
<emphasis id="B938EA890474054DFF3A444E5C10B2D4" bold="true" box="[249,277,1899,1925]" pageId="2" pageNumber="3">M</emphasis>
|
||
. Anterior right leg on the 22
|
||
<superScript id="7C399BD30474054DFD99444C5F6EB226" attach="left" box="[602,619,1897,1911]" fontSize="6" pageId="2" pageNumber="3">nd</superScript>
|
||
body ring. – A-D, F, H-I, L-M:
|
||
<taxonomicName id="4C4C4D180474054DFC154449594EB2D4" authority="Lazányi & Vagalinski, 2013" authorityName="Lazányi & Vagalinski" authorityYear="2013" box="[982,1099,1899,1925]" class="Diplopoda" family="Julidae" genus="Enghophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="sifnium" status="gen. et sp. nov.">
|
||
<emphasis id="B938EA890474054DFC154449594EB2D4" box="[982,1099,1899,1925]" italics="true" pageId="2" pageNumber="3">E. sifnium</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57F20474054DFB92444959FBB2D7" box="[1105,1278,1900,1926]" pageId="2" pageNumber="3" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
, holotype. E:
|
||
<taxonomicName id="4C4C4D180474054DFF1D44AA5C50B2F9" baseAuthorityName="Verhoeff" baseAuthorityYear="1903" box="[222,341,1935,1960]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="naxium">
|
||
<emphasis id="B938EA890474054DFF1D44AA5C50B2F9" box="[222,341,1935,1960]" italics="true" pageId="2" pageNumber="3">E. naxium</emphasis>
|
||
</taxonomicName>
|
||
, from Antiparos. G, J-K:
|
||
<taxonomicName id="4C4C4D180474054DFD4044AA5FFCB2F9" baseAuthorityName="Verhoeff" baseAuthorityYear="1903" box="[643,761,1935,1960]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="naxium">
|
||
<emphasis id="B938EA890474054DFD4044AA5FFCB2F9" box="[643,761,1935,1960]" italics="true" pageId="2" pageNumber="3">E. naxium</emphasis>
|
||
</taxonomicName>
|
||
, from Mavri Islet.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BF3369B0475054CFF7E42555F7AB605" blockId="3.[189,1399,368,852]" pageId="3" pageNumber="4">
|
||
Gonopods noticeably big compared to body size, directed mostly posteriad. Promere bevelled in laterobasal corner to fit into the inner curve of the 7
|
||
<superScript id="7C399BD30475054CFD3242B45FFBB4CE" attach="left" box="[753,766,401,415]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
body ring; latero-basal corner of promere only with thin triangular lamella (
|
||
<emphasis id="B938EA890475054CFE1042935CE6B481" box="[467,483,438,464]" italics="true" pageId="3" pageNumber="4">tl</emphasis>
|
||
) (
|
||
<figureCitation id="13772A1E0475054CFE3E42935F46B480" box="[509,579,438,465]" captionStart="Fig" captionStartId="4.[189,232,1818,1844]" captionTargetBox="[214,1363,491,1766]" captionTargetId="figure@4.[195,1392,484,1778]" captionTargetPageId="4" captionText="Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827705" httpUri="https://zenodo.org/record/3827705/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
D-E). Promeres attached to each other in their meso-basal corner with a chitinous bridge. A short, oblique ridge (
|
||
<emphasis id="B938EA890475054CFD7E42FF5FCDB4A2" box="[701,712,474,499]" italics="true" pageId="3" pageNumber="4">r</emphasis>
|
||
) on posterior surface of promere (
|
||
<figureCitation id="13772A1E0475054CFBAA42FC59C9B4A5" box="[1129,1228,473,500]" captionStart="Fig" captionStartId="4.[189,232,1818,1844]" captionTargetBox="[214,1363,491,1766]" captionTargetId="figure@4.[195,1392,484,1778]" captionTargetPageId="4" captionText="Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827705" httpUri="https://zenodo.org/record/3827705/files/figure.png" pageId="3" pageNumber="4">Figs 2A</figureCitation>
|
||
, D-E, 4A, C). Flagellum emerging from a well-developed sinus (
|
||
<emphasis id="B938EA890475054CFCC942D85E23B746" box="[778,806,509,535]" italics="true" pageId="3" pageNumber="4">fsi</emphasis>
|
||
) (
|
||
<figureCitation id="13772A1E0475054CFCFC42D85E81B746" box="[831,900,509,535]" captionStart="Fig" captionStartId="4.[189,232,1818,1844]" captionTargetBox="[214,1363,491,1766]" captionTargetId="figure@4.[195,1392,484,1778]" captionTargetPageId="4" captionText="Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827705" httpUri="https://zenodo.org/record/3827705/files/figure.png" pageId="3" pageNumber="4">Fig. 2</figureCitation>
|
||
D-E). At the level of this sinus a deepening on anterior surface of the promere (
|
||
<emphasis id="B938EA890475054CFDAE41055F79B76B" box="[621,636,544,570]" italics="true" pageId="3" pageNumber="4">d</emphasis>
|
||
) (
|
||
<figureCitation id="13772A1E0475054CFD5B41055FFDB76B" box="[664,760,544,570]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[203,1385,607,1806]" captionTargetId="figure@7.[151,1514,593,1819]" captionTargetPageId="7" captionText="Fig. 4. Gonopods of Enghophyllum naxium, male from Mavri Islet. A. Gonopods, posterior view. B. Gonopods, antero-lateral view. C. Right gonopods, mesal view. D. Left opisthomere, lateral view. – app = apical posterior process; bpp = basal posterior process; d = deepening; fc = flagellum channel; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; sc = sperm canal; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827709" httpUri="https://zenodo.org/record/3827709/files/figure.png" pageId="3" pageNumber="4">Fig. 4B</figureCitation>
|
||
). Opisthomere entirely concealed under the promere, protruding postero-ventrad, the two promeres tightly closing the ventral opening on 7
|
||
<superScript id="7C399BD30475054CFB71416459BAB71E" attach="left" box="[1202,1215,577,591]" fontSize="6" pageId="3" pageNumber="4">th</superScript>
|
||
ring like a pair of shutters. Opisthomere not embedded in the groove formed by the oblique ridge on the promere’s posterior surface (unlike in
|
||
<taxonomicName id="4C4C4D180475054CFDCB41AF5FACB7F5" authorityName="Verhoeff" authorityYear="1894" box="[520,681,650,676]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938EA890475054CFDCB41AF5FACB7F5" box="[520,681,650,676]" italics="true" pageId="3" pageNumber="4">Megaphyllum</emphasis>
|
||
</taxonomicName>
|
||
), but “riding” on the ridge (
|
||
<figureCitation id="13772A1E0475054CFC3841AC595BB7F5" box="[1019,1118,649,676]" captionStart="Fig" captionStartId="4.[189,232,1818,1844]" captionTargetBox="[214,1363,491,1766]" captionTargetId="figure@4.[195,1392,484,1778]" captionTargetPageId="4" captionText="Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827705" httpUri="https://zenodo.org/record/3827705/files/figure.png" pageId="3" pageNumber="4">Figs 2A</figureCitation>
|
||
,
|
||
<figureCitation id="13772A1E0475054CFBAF41AF5996B7F5" box="[1132,1171,650,676]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[203,1385,607,1806]" captionTargetId="figure@7.[151,1514,593,1819]" captionTargetPageId="7" captionText="Fig. 4. Gonopods of Enghophyllum naxium, male from Mavri Islet. A. Gonopods, posterior view. B. Gonopods, antero-lateral view. C. Right gonopods, mesal view. D. Left opisthomere, lateral view. – app = apical posterior process; bpp = basal posterior process; d = deepening; fc = flagellum channel; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; sc = sperm canal; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827709" httpUri="https://zenodo.org/record/3827709/files/figure.png" pageId="3" pageNumber="4">4C</figureCitation>
|
||
): posterior basis of opisthomere with a characteristic riffle (
|
||
<emphasis id="B938EA890475054CFD4E418B5FA4B796" box="[653,673,686,711]" italics="true" pageId="3" pageNumber="4">ri</emphasis>
|
||
) formed by chitinous lamellae, where the ridge of the promere is positioned (
|
||
<figureCitation id="13772A1E0475054CFEA141F55CEDB7BB" box="[354,488,720,746]" captionStart="Fig" captionStartId="4.[189,232,1818,1844]" captionTargetBox="[214,1363,491,1766]" captionTargetId="figure@4.[195,1392,484,1778]" captionTargetPageId="4" captionText="Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827705" httpUri="https://zenodo.org/record/3827705/files/figure.png" pageId="3" pageNumber="4">Figs 2A, G</figureCitation>
|
||
,
|
||
<figureCitation id="13772A1E0475054CFE3741F55F06B7BB" box="[500,515,720,746]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[203,1385,607,1806]" captionTargetId="figure@7.[151,1514,593,1819]" captionTargetPageId="7" captionText="Fig. 4. Gonopods of Enghophyllum naxium, male from Mavri Islet. A. Gonopods, posterior view. B. Gonopods, antero-lateral view. C. Right gonopods, mesal view. D. Left opisthomere, lateral view. – app = apical posterior process; bpp = basal posterior process; d = deepening; fc = flagellum channel; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; sc = sperm canal; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827709" httpUri="https://zenodo.org/record/3827709/files/figure.png" pageId="3" pageNumber="4">4</figureCitation>
|
||
C-D). Opisthomere with a well-developed lateral process (
|
||
<emphasis id="B938EA890475054CFB7041F559CFB7BB" box="[1203,1226,720,746]" italics="true" pageId="3" pageNumber="4">lp</emphasis>
|
||
) and an apical and a basal posterior process (
|
||
<emphasis id="B938EA890475054CFDE341D15F49B65C" box="[544,588,756,781]" italics="true" pageId="3" pageNumber="4">app</emphasis>
|
||
and
|
||
<emphasis id="B938EA890475054CFD4541D65FB7B65C" box="[646,690,755,781]" italics="true" pageId="3" pageNumber="4">bpp</emphasis>
|
||
). Solenomere (
|
||
<emphasis id="B938EA890475054CFCAB41D15E76B65C" box="[872,883,756,781]" italics="true" pageId="3" pageNumber="4">s</emphasis>
|
||
) simple, with no processes but basally with a micro-spinose pillow (
|
||
<emphasis id="B938EA890475054CFE1D40325F08B661" box="[478,525,791,816]" italics="true" pageId="3" pageNumber="4">msp</emphasis>
|
||
) (
|
||
<figureCitation id="13772A1E0475054CFDE440335F74B660" box="[551,625,790,817]" captionStart="Fig" captionStartId="4.[189,232,1818,1844]" captionTargetBox="[214,1363,491,1766]" captionTargetId="figure@4.[195,1392,484,1778]" captionTargetPageId="4" captionText="Fig. 2. Gonopods of Enghophyllum sifnium gen. et sp. nov., holotype ♂. A. Left gonopods, mesal view. B. Leftgonopods,antero-lateralview.C.Leftgonopods,meso-posteriorview.D.Leftpromere,meso-posterior view. E. Left promere, postero-mesal view. F. Right opisthomere, lateral view. G. Right opisthomere, mesal view. – app = apical posterior process; bpp = basal posterior process; fc = flagellum channel; fl = flagellum; fsi = flagellum sinus; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827705" httpUri="https://zenodo.org/record/3827705/files/figure.png" pageId="3" pageNumber="4">Figs 2</figureCitation>
|
||
A-C, F-G, 4A-D). Flagellum channel and sperm canal (
|
||
<emphasis id="B938EA890475054CFAC74032581CB660" box="[1284,1305,791,817]" italics="true" pageId="3" pageNumber="4">fc</emphasis>
|
||
and
|
||
<emphasis id="B938EA890475054CFA9740325869B661" box="[1364,1388,791,816]" italics="true" pageId="3" pageNumber="4">sc</emphasis>
|
||
) each with an apical opening (
|
||
<figureCitation id="13772A1E0475054CFDD6401C5F77B605" box="[533,626,825,852]" captionStart="Fig" captionStartId="7.[189,232,1858,1884]" captionTargetBox="[203,1385,607,1806]" captionTargetId="figure@7.[151,1514,593,1819]" captionTargetPageId="7" captionText="Fig. 4. Gonopods of Enghophyllum naxium, male from Mavri Islet. A. Gonopods, posterior view. B. Gonopods, antero-lateral view. C. Right gonopods, mesal view. D. Left opisthomere, lateral view. – app = apical posterior process; bpp = basal posterior process; d = deepening; fc = flagellum channel; lp = lateral process; msp = micro-spinose pillow; P = promere; r = ridge; ri = riffle; s = solenomere; sc = sperm canal; tl = triangular lamella. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827709" httpUri="https://zenodo.org/record/3827709/files/figure.png" pageId="3" pageNumber="4">Fig. 4C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BF3369B0475054CFF29405E5C54B6C4" blockId="3.[189,1399,891,1058]" box="[234,337,891,917]" pageId="3" pageNumber="4">
|
||
<emphasis id="B938EA890475054CFF29405E5C54B6C4" bold="true" box="[234,337,891,917]" pageId="3" pageNumber="4">Females</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BF3369B0475054CFF7E40BB5CC5B173" blockId="3.[189,1399,891,1058]" pageId="3" pageNumber="4">
|
||
The only available female of
|
||
<taxonomicName id="4C4C4D180475054CFDE140BA5F9FB6E9" authority="Lazányi & Vagalinski, 2013" authorityName="Lazányi & Vagalinski" authorityYear="2013" box="[546,666,926,952]" class="Diplopoda" family="Julidae" genus="Enghophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sifnium" status="gen. et sp. nov.">
|
||
<emphasis id="B938EA890475054CFDE140BA5F9FB6E9" box="[546,666,926,952]" italics="true" pageId="3" pageNumber="4">E. sifnium</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57F20475054CFD6640BA5E59B6E8" box="[677,860,927,953]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
is in stadium IX (with 8 rows of ocelli) and proved to be subadult, but females of
|
||
<taxonomicName id="4C4C4D180475054CFD4540E75FFAB68A" baseAuthorityName="Verhoeff" baseAuthorityYear="1903" box="[646,767,962,987]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="naxium">
|
||
<emphasis id="B938EA890475054CFD4540E75FFAB68A" box="[646,767,962,987]" italics="true" pageId="3" pageNumber="4">E. naxium</emphasis>
|
||
</taxonomicName>
|
||
in stadia VIII-IX (with 7-8 rows of ocelli) had fully developed vulvae, which we used here for the description of female sexual characters. First two legpairs slightly swollen.
|
||
</paragraph>
|
||
<paragraph id="8BF3369B0475054CFF7E476C5975B00B" blockId="3.[189,1399,1097,1370]" pageId="3" pageNumber="4">
|
||
Vulva (
|
||
<figureCitation id="13772A1E0475054CFED0476C5C5FB132" box="[275,346,1097,1123]" captionStart="Fig" captionStartId="6.[189,232,1909,1935]" captionTargetBox="[403,1186,568,1855]" captionTargetId="figure@6.[365,1253,544,1902]" captionTargetPageId="6" captionText="Fig. 3. Right vulva of Enghophyllum naxium, female from Antiparos Island, anterior view. – at = apodematic tube; ca = central ampulla; ct = connecting tube; da = distal ampulla; lv = lateral valve; op = operculum; si = sinus. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.3827707" httpUri="https://zenodo.org/record/3827707/files/figure.png" pageId="3" pageNumber="4">Fig. 3</figureCitation>
|
||
): Subcylindric in shape, the mesal half shorter, the opening large, oval, apical. Operculum shorter than bursa, with around 20-25 setae. Mesal and lateral sclerites with 2-3 setae each. Apodematic tube not opening apically into a longitudinal median cleft as in, e.g.,
|
||
<taxonomicName id="4C4C4D180475054CFC3547B55992B1FB" authorityName="Verhoeff" authorityYear="1894" box="[1014,1175,1168,1194]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B938EA890475054CFC3547B55992B1FB" box="[1014,1175,1168,1194]" italics="true" pageId="3" pageNumber="4">Megaphyllum</emphasis>
|
||
</taxonomicName>
|
||
species, but into a wide sinus (
|
||
<emphasis id="B938EA890475054CFE8E47915C64B19C" box="[333,353,1204,1229]" italics="true" pageId="3" pageNumber="4">si</emphasis>
|
||
). Around this sinus a thin wall, formed by the apical part of the bursa. These walls with small dot-like pores; most setae (around 20/side) emerging from this apical region. Apodematic tube (
|
||
<emphasis id="B938EA890475054CFA9547F25868B1A1" box="[1366,1389,1239,1264]" italics="true" pageId="3" pageNumber="4">at</emphasis>
|
||
) ending in two receptaculi seminis or ampullae. Central ampulla (
|
||
<emphasis id="B938EA890475054CFC7747DF5ED5B042" box="[948,976,1274,1299]" italics="true" pageId="3" pageNumber="4">ca</emphasis>
|
||
) drop-like. Distal, globular ampulla (
|
||
<emphasis id="B938EA890475054CFF0446395DE1B067" box="[199,228,1308,1334]" italics="true" pageId="3" pageNumber="4">da</emphasis>
|
||
) joining the apodematic tube through a twisted connecting tube (
|
||
<emphasis id="B938EA890475054CFC1446385EE9B067" box="[983,1004,1309,1334]" italics="true" pageId="3" pageNumber="4">ct</emphasis>
|
||
). Connecting tube very long, thus the distal ampulla hanging out of the bursa; easily broken off during preparation.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35665100475054CFF7E46A45ECEB362" pageId="3" pageNumber="4" type="discussion">
|
||
<paragraph id="8BF3369B0475054CFF7E46A45C2AB0CA" blockId="3.[189,303,1409,1435]" box="[189,303,1409,1435]" pageId="3" pageNumber="4">
|
||
<heading id="D0BB81F70475054CFF7E46A45C2AB0CA" bold="true" box="[189,303,1409,1435]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="B938EA890475054CFF7E46A45C2AB0CA" bold="true" box="[189,303,1409,1435]" pageId="3" pageNumber="4">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BF3369B0475054CFF7E46955ECEB362" blockId="3.[189,1399,1456,1588]" pageId="3" pageNumber="4">
|
||
The everted penes of the adult
|
||
<taxonomicName id="4C4C4D180475054CFDF546955FB5B098" baseAuthorityName="Verhoeff" baseAuthorityYear="1903" box="[566,688,1456,1481]" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="naxium">
|
||
<emphasis id="B938EA890475054CFDF546955FB5B098" box="[566,688,1456,1481]" italics="true" pageId="3" pageNumber="4">E. naxium</emphasis>
|
||
</taxonomicName>
|
||
male from Antiparos was quite soft, amorphous and huge, contrary to the stout, minute penes, deeply hidden under the 2
|
||
<superScript id="7C399BD30475054CFC7F46F45EC8B08E" attach="left" box="[956,973,1489,1503]" fontSize="6" pageId="3" pageNumber="4">nd</superScript>
|
||
leg-pair coxae, observed in the
|
||
<taxonomicName id="4C4C4D180475054CFA9E46F15C16B341" baseAuthorityName="Verhoeff" baseAuthorityYear="1903" class="Diplopoda" family="Julidae" genus="Megaphyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="naxium">
|
||
<emphasis id="B938EA890475054CFA9E46F15C16B341" italics="true" pageId="3" pageNumber="4">E. naxium</emphasis>
|
||
</taxonomicName>
|
||
male from Mavri Islet and the
|
||
<taxonomicName id="4C4C4D180475054CFD4F46D25E01B341" authority="Lazányi & Vagalinski, 2013" authorityName="Lazányi & Vagalinski" authorityYear="2013" box="[652,772,1526,1552]" class="Diplopoda" family="Julidae" genus="Enghophyllum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="sifnium" status="gen. et sp. nov.">
|
||
<emphasis id="B938EA890475054CFD4F46D25E01B341" box="[652,772,1526,1552]" italics="true" pageId="3" pageNumber="4">E. sifnium</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A20B57F20475054CFCCD46D25EC6B340" box="[782,963,1527,1553]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
male from Sifnos. It is possible that penes vary in size between copulating and non-copulating periods.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |