treatments-xml/data/03/E5/87/03E58783FFB8FFE546BAF9E8FEEFFEA2.xml
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<document id="208DE10400EC98C2D848AB9E9D907BDE" ID-CLB-Dataset="7435" ID-DOI="10.11646/zootaxa.4937.1.1" ID-GBIF-Dataset="712f8615-91da-4442-99f9-1e0a6a211cb5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4559438" ID-ZooBank="A1DDB75B-27E5-463A-8FA6-975B89163A50" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614161496698" checkinUser="plazi" docAuthor="Sanborn, Allen F." docDate="2021" docId="03E58783FFB8FFE546BAF9E8FEEFFEA2" docLanguage="en" docName="zootaxa.4937.1.1.pdf" docOrigin="Zootaxa 4937 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Nelbroma betrokaensis Sanborn 2021, n. sp." docType="treatment" docVersion="10" lastPageNumber="42" masterDocId="FFDCFFFBFF9EFFCC462DFFFBFFCAFFBF" masterDocTitle="The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species" masterLastPageNumber="79" masterPageNumber="1" pageNumber="39" updateTime="1698931083349" updateUser="ExternalLinkService">
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<mods:title id="D3BCC8F92145F38EE8071B62418361E5">The cicadas (Hemiptera: Cicadoidea: Cicadidae) of Madagascar including a new tribe, five new genera, twelve new species, four new species synonymies, five revised species status, ten new combinations, new tribal assignments for four genera, one new subtribe synonymy, a checklist and key to the species</mods:title>
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<mods:namePart id="4B7152A9C9F130ED5EDE8DCF14939E91">Sanborn, Allen F.</mods:namePart>
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<mods:title id="1FCBE20B9B2AE5F82360E481B27796E6">Zootaxa</mods:title>
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<mods:date id="A1EF901D56382971C87744D76F3C0DE1">2021</mods:date>
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<treatment id="03E58783FFB8FFE546BAF9E8FEEFFEA2" ID-DOI="http://doi.org/10.5281/zenodo.4673279" ID-GBIF-Taxon="180885759" ID-Zenodo-Dep="4673279" LSID="urn:lsid:plazi:treatment:03E58783FFB8FFE546BAF9E8FEEFFEA2" httpUri="http://treatment.plazi.org/id/03E58783FFB8FFE546BAF9E8FEEFFEA2" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39">
<subSubSection id="C356651EFFB8FFEA46BAF9E8FE3AF992" box="[151,496,1555,1581]" pageId="38" pageNumber="39" type="nomenclature">
<paragraph id="8BF33695FFB8FFEA46BAF9E8FE3AF992" blockId="38.[151,496,1555,1617]" box="[151,496,1555,1581]" pageId="38" pageNumber="39">
<heading id="D0BB81F9FFB8FFEA46BAF9E8FE3AF992" box="[151,496,1555,1581]" fontSize="11" level="2" pageId="38" pageNumber="39" reason="5">
<taxonomicName id="4C4C4D16FFB8FFEA46BAF9E8FE6CF992" ID-CoL="8KGR5" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[151,422,1555,1581]" class="Insecta" family="Cicadidae" genus="Nelbroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="38" pageNumber="39" phylum="Arthropoda" rank="species" species="betrokaensis" status="sp. nov.">
<emphasis id="B938EA87FFB8FFEA46BAF9E8FE6CF992" box="[151,422,1555,1581]" italics="true" pageId="38" pageNumber="39">Nelbroma betrokaensis</emphasis>
</taxonomicName>
<emphasis id="B938EA87FFB8FFEA4780F9E8FE3AF992" bold="true" box="[429,496,1555,1581]" pageId="38" pageNumber="39">
<taxonomicNameLabel id="A20B57FCFFB8FFEA4780F9E8FE3AF992" box="[429,496,1555,1581]" pageId="38" pageNumber="39" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C356651EFFB8FFE546BAF9CCFEEFFEA2" lastPageId="41" lastPageNumber="42" pageId="38" pageNumber="39" type="description">
<paragraph id="8BF33695FFB8FFEA46BAF9CCFF21F9EE" blockId="38.[151,496,1555,1617]" box="[151,235,1590,1617]" pageId="38" pageNumber="39">
(
<figureCitation id="13772A10FFB8FFEA468DF9CCFF28F9EE" box="[160,226,1590,1617]" captionStart="FIG" captionStartId="39.[151,196,1944,1968]" captionTargetBox="[151,1435,417,1921]" captionTargetId="figure-121@39.[151,1435,417,1921]" captionTargetPageId="39" captionText="FIG. 5. Nelbroma betrokaensis n. gen., n. sp.: A, Holotype male habitus; B, holotype male dorsum; C, holotype male timbal; D, holotype male operculum; E, holotype male lateral view of genitalia; F, holotype male posterior view of genitalia. Scale bar: A, 2 cm; B, 2 mm; C—F, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4559454" httpUri="https://zenodo.org/record/4559454/files/figure.png" pageId="38" pageNumber="39">Fig. 5</figureCitation>
)
</paragraph>
<paragraph id="8BF33695FFB8FFEA46BAF984FED0F95E" blockId="38.[151,1436,1662,2013]" pageId="38" pageNumber="39">
Type material.—
<materialsCitation id="3B243CC8FFB8FFEA477BF984FED0F95E" ID-GBIF-Occurrence="3069571308" collectingDate="1994-12-04" collectingDateMax="1994-12-09" collectingDateMin="1994-12-04" collectionCode="MTEC, CASC" collectorName="M. A. Ivie &amp; D. A. Pollack" country="Madagascar" elevation="825" latitude="-23.163334" location="Betroka" longLatPrecision="21" longitude="45.968613" pageId="38" pageNumber="39" specimenCount="1" typeStatus="holotype">
<typeStatus id="54F78837FFB8FFEA477BF984FE0BF926" box="[342,449,1663,1689]" pageId="38" pageNumber="39" type="holotype">Holotype</typeStatus>
. “
<collectingCountry id="F35B7605FFB8FFEA47F9F984FD69F926" box="[468,675,1663,1689]" name="Madagascar" pageId="38" pageNumber="39">MADAGASCAR</collectingCountry>
: Tulear Pr. /
<quantity id="4CB49B70FFB8FFEA451DF984FCBAF926" box="[816,880,1663,1689]" metricMagnitude="4" metricUnit="m" metricValue="1.8" pageId="38" pageNumber="39" unit="km" value="18.0">18km</quantity>
NNW
<location id="8E93604EFFB8FFEA45EDF984FBD6F926" LSID="urn:lsid:plazi:treatment:03E58783FFB8FFE546BAF9E8FEEFFEA2:8E93604EFFB8FFEA45EDF984FBD6F926" box="[960,1052,1663,1689]" country="Madagascar" latitude="-23.163334" longLatPrecision="21" longitude="45.968613" name="Betroka" pageId="38" pageNumber="39">Betroka</location>
,
<quantity id="4CB49B70FFB8FFEA420AF985FBADF926" box="[1063,1127,1662,1689]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="38" pageNumber="39" unit="m" value="825.0">
<elevation id="0061D1A6FFB8FFEA420AF985FBADF926" box="[1063,1127,1662,1689]" metricMagnitude="2" metricUnit="m" metricValue="8.25" pageId="38" pageNumber="39" unit="m" value="825.0">825m</elevation>
</quantity>
/
<geoCoordinate id="EE785052FFB8FFEA4250F984FADFF926" box="[1149,1301,1663,1689]" degrees="23" direction="south" minutes="09" orientation="latitude" pageId="38" pageNumber="39" precision="15" seconds="48" value="-23.163334">23° 09 48” S</geoCoordinate>
<geoCoordinate id="EE785052FFB8FFEA4336F985FF62F902" degrees="45" direction="east" minutes="58" orientation="longitude" pageId="38" pageNumber="39" precision="15" seconds="07" value="45.968613">45° 58 07” E</geoCoordinate>
/
<date id="FFF21055FFB8FFEA46ECF958FE48F902" box="[193,386,1699,1725]" pageId="38" pageNumber="39" value="1994-12-04" valueMax="1994-12-09" valueMin="1994-12-04">
<collectingDate id="EFB6E9BDFFB8FFEA46ECF958FE48F902" box="[193,386,1699,1725]" pageId="38" pageNumber="39" value="1994-12-04" valueMax="1994-12-09" valueMin="1994-12-04">0409DEC 1994</collectingDate>
</date>
, F.I.T. /
<collectorName id="26B95343FFB8FFEA47CCF958FD9BF902" box="[481,593,1699,1725]" pageId="38" pageNumber="39">M.A. Ivie</collectorName>
&amp;
<collectorName id="26B95343FFB8FFEA445AF958FCC7F902" box="[631,781,1699,1725]" pageId="38" pageNumber="39">D.A. Pollack</collectorName>
” male (Michael Ivie collection,
<collectionCode id="ED5DAE50FFB8FFEA4251F958FB00F902" box="[1148,1226,1699,1725]" country="USA" httpUri="http://grbio.org/cool/zm6z-4gpy" name="Montana State Entomology Collection" pageId="38" pageNumber="39" type="Museum">MTEC</collectionCode>
to be deposited in the
<collectionCode id="ED5DAE50FFB8FFEA46EDF93CFEC4F95E" box="[192,270,1735,1761]" pageId="38" pageNumber="39">CASC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BF33695FFB8FFEA46EAF910FF29F895" blockId="38.[151,1436,1662,2013]" pageId="38" pageNumber="39">
Etymology.—The name is a combination of
<emphasis id="B938EA87FFB8FFEA44FBF910FCE0F8BA" box="[726,810,1771,1797]" italics="true" pageId="38" pageNumber="39">betroka</emphasis>
for the
<typeStatus id="54F78837FFB8FFEA45B2F910FC05F8BA" box="[927,975,1771,1797]" pageId="38" pageNumber="39">type</typeStatus>
locality and
<emphasis id="B938EA87FFB8FFEA4242F910FB7FF8BB" box="[1135,1205,1771,1796]" italics="true" pageId="38" pageNumber="39">ensis</emphasis>
(L. suffix denoting place).
</paragraph>
<paragraph id="8BF33695FFB8FFEA46EAF8C8FE80F8F2" blockId="38.[151,1436,1662,2013]" box="[199,330,1843,1869]" pageId="38" pageNumber="39">Description</paragraph>
<paragraph id="8BF33695FFB8FFEA46EAF8ACFABCF8CE" blockId="38.[151,1436,1662,2013]" box="[199,1398,1879,1905]" pageId="38" pageNumber="39">Head and thorax tawny marked with piceous and castaneous, abdomen testaceous marked with castaneous.</paragraph>
<paragraph id="8BF33695FFB8FFEA46EAF880FECEF82A" blockId="38.[151,1436,1662,2013]" box="[199,260,1915,1941]" pageId="38" pageNumber="39">
<emphasis id="B938EA87FFB8FFEA46EAF880FECEF82A" box="[199,260,1915,1941]" italics="true" pageId="38" pageNumber="39">Head</emphasis>
</paragraph>
<paragraph id="8BF33695FFB8FFEB46EAF864FF1DFE36" blockId="38.[151,1436,1662,2013]" lastBlockId="39.[151,1437,151,393]" lastPageId="39" lastPageNumber="40" pageId="38" pageNumber="39">Head wider than mesonotum, castaneous except ground color supra-antennal plates, anteromedial vertex parallel to frontoclypeal suture curving to anterior cranial depression, anterolateral corner of frons, and epicranial suture posterior to median ocellus. Dorsal head covered with sparse, short silvery, longer posterior to eye. Ocelli red, eyes ochraceous but cells have retracted from eye surface so they appear hollow. Ventral head ground color with castaneous eye margin and castaneous spot in posterior lorum, short white pile on lorum and gena, denser than on dorsal head. Postclypeus ground color with 12 castaneous transverse ridges, short white pile radiating in lateral transverse grooves and posterior margin. Anteclypeus castaneous covered with white pile. Mentum castaneous, labium castaneous with piceous tip, rostrum radiating short golden pile, reaching to posterior coxae. Antennal segments ground color.</paragraph>
<caption id="DF33661DFFB9FFEB46BAF863FE15F847" ID-DOI="http://doi.org/10.5281/zenodo.4559454" ID-Zenodo-Dep="4559454" httpUri="https://zenodo.org/record/4559454/files/figure.png" pageId="39" pageNumber="40" startId="39.[151,196,1944,1968]" targetBox="[151,1435,417,1921]" targetPageId="39">
<paragraph id="8BF33695FFB9FFEB46BAF863FE15F847" blockId="39.[151,1437,1944,2041]" pageId="39" pageNumber="40">
<emphasis id="B938EA87FFB9FFEB46BAF863FF2EF80E" bold="true" box="[151,228,1944,1969]" pageId="39" pageNumber="40">FIG. 5.</emphasis>
<taxonomicName id="4C4C4D16FFB9FFEB46C1F863FE1CF80F" authority="Sanborn, 2021" authorityName="Sanborn" authorityYear="2021" box="[236,470,1944,1968]" class="Insecta" family="Cicadidae" genus="Nelbroma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="39" pageNumber="40" phylum="Arthropoda" rank="species" species="betrokaensis" status="n. gen., n. sp.">
<emphasis id="B938EA87FFB9FFEB46C1F863FE1CF80F" box="[236,470,1944,1968]" italics="true" pageId="39" pageNumber="40">Nelbroma betrokaensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20B57FCFFB9FFEB47F0F862FDA7F80E" box="[477,621,1945,1969]" pageId="39" pageNumber="40" rank="species">
<emphasis id="B938EA87FFB9FFEB47F0F862FDECF80E" bold="true" box="[477,550,1945,1969]" pageId="39" pageNumber="40">n. gen.</emphasis>
,
<emphasis id="B938EA87FFB9FFEB441EF862FDA7F80E" bold="true" box="[563,621,1945,1969]" pageId="39" pageNumber="40">n. sp.</emphasis>
</taxonomicNameLabel>
: A, Holotype male habitus; B, holotype male dorsum; C, holotype male timbal; D, holotype male operculum; E, holotype male lateral view of genitalia; F, holotype male posterior view of genitalia. Scale bar: A, 2 cm; B, 2 mm; C—F, 1 mm.
</paragraph>
</caption>
<paragraph id="8BF33695FFB6FFE446EAFF6CFED2FF0E" blockId="40.[151,1437,151,2013]" box="[199,280,151,177]" pageId="40" pageNumber="41">
<emphasis id="B938EA87FFB6FFE446EAFF6CFED2FF0E" box="[199,280,151,177]" italics="true" pageId="40" pageNumber="41">Thorax</emphasis>
</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFF40FBC9FE4A" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">Pronotum ground color with castaneous longitudinal fascia along midline expanding laterally along ambient fissure not reaching level of posterior lateral fissure, castaneous within lateral fissure and lateral ambient fissure surrounding lateral disk, some irregular castaneous markings on disk between paramedial and lateral fissures and anterior lateral disk. Pronotal collar castaneous with ground color posterior margin between lateral angles. Pronotum with sparse short silvery pile. Mesonotum castaneous with piceous sigillae and scutal depressions, ochraceous lateral cruciform elevation. Metanotum ochraceous with castaneous spot in middle wing groove. Dorsum with short silvery pile on anterolateral margin, between anterior arms of cruciform elevation, between anterior and posterior arms of cruciform elevation, on posterior mesonotum, in wing groove, and on posterior metanotum. Ventral thoracic segments ground color laterally, castaneous medially, covered with white pile.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFE04FEC1FDA7" blockId="40.[151,1437,151,2013]" box="[199,267,511,536]" pageId="40" pageNumber="41">
<emphasis id="B938EA87FFB6FFE446EAFE04FEC1FDA7" box="[199,267,511,536]" italics="true" pageId="40" pageNumber="41">Wings</emphasis>
</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFDD8FE2FFD4E" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">Fore wings and hind wings opaque medially, hyaline fore wings with eight apical cells, hind wings damaged distally but five apical cells clearly present in remaining section attached to underside of fore wing. Venation ground color proximally, becoming castaneous distally, dark castaneous fascia along base of radius &amp; subcostal vein in proximal basal cell. Basal cell hyaline. Pterostigma present, not reaching radial crossvein. Basal membrane of fore wing gray. Hind wing venation similar to fore wing. Anal cell 3, anal cell 2 along anal veins 2 and 3, and anal cell 1 along anal vein 2 light gray.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFD00FF36FCAB" blockId="40.[151,1437,151,2013]" box="[199,252,763,788]" pageId="40" pageNumber="41">
<emphasis id="B938EA87FFB6FFE446EAFD00FF36FCAB" box="[199,252,763,788]" italics="true" pageId="40" pageNumber="41">Legs</emphasis>
</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFCE4FE3DFC52" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">Legs castaneous, ground color on distomedial coxae and distal femora. Fore femora with primary spine longest, angled to femoral axis, secondary spine upright with curved tip, tertiary spine smallest about parallel to secondary spine, spines dark castaneous with lighter tip. Tarsi castaneous except ground color proximal half of hind tarsi, tibial spurs and comb dark castaneous. Pretarsal claws castaneous. Meracanthus broadly triangular with curved lateral margin, ochraceous with castaneous base, not reaching anterior margin of medial operculum. Legs with short white pile, radiating long golden pile.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFC0CFE8EFBAE" blockId="40.[151,1437,151,2013]" box="[199,324,1015,1041]" pageId="40" pageNumber="41">
<emphasis id="B938EA87FFB6FFE446EAFC0CFE8EFBAE" box="[199,324,1015,1041]" italics="true" pageId="40" pageNumber="41">Operculum</emphasis>
</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFBE0FE75FB7A" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">Male operculum ochraceous with castaneous base, lateral margin straight, posterolateral margin curved, posterior margin straight at approximate right angle to lateral margin, rounded posteromedial margin connecting concave anteromedial margin, not reaching to middle of hind coxa medially so medial margins well separated, reaching to anterior sternite II medially but not covering tympanal cavity posteriorly or laterally, covered with white pile at base, radiating long golden pile.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFB34FEFBFB56" blockId="40.[151,1437,151,2013]" box="[199,305,1231,1257]" pageId="40" pageNumber="41">
<emphasis id="B938EA87FFB6FFE446EAFB34FEFBFB56" box="[199,305,1231,1257]" italics="true" pageId="40" pageNumber="41">Abdomen</emphasis>
</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFB08FA93FA5A" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">Abdominal tergites dark castaneous except ground color lateral tergite 2, dorsolateral tergites 35, posterior margins of tergites 38 and dorsolateral tergite 8, tergites covered with sparse short silvery pile, white pile on dorsum of tergites 1, 2, and anterior of tergite 3. Timbal cover absent, timbal completely exposed, posterior timbal strongly angled mediad, timbal with nine ribs. Male sternites castaneous except ground color posterolateral margin of sternite I, all of sternite II except castaneous posterior margin and auditory capsule, and posterolateral corners of sternite VII. Epipleurites castaneous with ground color posterior margin. Sternites and epipleurites radiating long silvery pile. Sternite VIII light castaneous with ground color anterolateral margin, radiating long silvery pile.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAFA14FEFBF9B6" blockId="40.[151,1437,151,2013]" box="[199,305,1519,1545]" pageId="40" pageNumber="41">
<emphasis id="B938EA87FFB6FFE446EAFA14FEFBF9B6" box="[199,305,1519,1545]" italics="true" pageId="40" pageNumber="41">Genitalia</emphasis>
</paragraph>
<paragraph id="8BF33695FFB6FFE446EAF9E8FD00F896" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">Male pygofer castaneous with ground color dorsal beak margin, distal shoulder undeveloped, dorsal beak pointed, broadly triangular, pygofer with short silvery pile dorsally and laterally. Upper pygofer lobe small, flattened, angled mediad, radiating short silvery pile, superimposed on basal pygofer lobe, pygofer basal lobes extending about half length of pygofer, adpressed to lateral pygofer margin, with dark castaneous, semicircular mediad extensions radiating short silvery pile in middle and on distal end. Anal styles dark castaneous radiating short silvery pile, anal tube ground color. Uncus absent, claspers castaneous with darker base, laterally flattened, curving mediad to meet along midline, tapering to transverse apex, surrounding aedeagus distally. Aedeagus small, tubular, light castaneous, restrained by claspers, lacking conjunctival claws.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAF8C8FE60F8F2" blockId="40.[151,1437,151,2013]" box="[199,426,1843,1869]" pageId="40" pageNumber="41">Female is unknown.</paragraph>
<paragraph id="8BF33695FFB6FFE446EAF8ACFE0CF806" blockId="40.[151,1437,151,2013]" pageId="40" pageNumber="41">
Measurements (mm).—N =
<specimenCount id="9D4AFD1CFFB6FFE4442EF8ACFD98F8CE" box="[515,594,1879,1905]" pageId="40" pageNumber="41" type="male">1 male</specimenCount>
. Length of body: 8.60; length of fore wing: 11.75; width of fore wing: 3.85; length of head: 2.25; width of head including eyes: 3.65; width of pronotum including suprahumeral plates: 3.40; width of mesonotum: 3.00.
</paragraph>
<paragraph id="8BF33695FFB6FFE546EAF838FA52FF6A" blockId="40.[151,1437,151,2013]" lastBlockId="41.[151,1436,151,285]" lastPageId="41" lastPageNumber="42" pageId="40" pageNumber="41">
Diagnosis.—This new species is the smallest known species in
<collectingCountry id="F35B7605FFB6FFE445A2F838FBD2F862" box="[911,1048,1987,2013]" name="Madagascar" pageId="40" pageNumber="41">Madagascar</collectingCountry>
. The measurements can be used to distinguish it from the remaining species. It can also be distinguished from the other two small new species by the ground color supra-antennal plates and the large claspers that extend out from under the uncus in this new species.
</paragraph>
<paragraph id="8BF33695FFB7FFE546EAFF24FEEFFEA2" blockId="41.[151,1436,151,285]" pageId="41" pageNumber="42">
Distribution.—The species is only known from the
<typeStatus id="54F78837FFB7FFE5453DFF24FC8AFF46" box="[784,832,223,249]" pageId="41" pageNumber="42">type</typeStatus>
specimen collected in Tulear Province, southwestern
<collectingCountry id="F35B7605FFB7FFE546BAFEF8FEEAFEA2" box="[151,288,259,285]" name="Madagascar" pageId="41" pageNumber="42">Madagascar</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>