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<document id="1B8570C8256FD06A38BE31DEBBA8BD4C" ID-CLB-Dataset="8531" ID-DOI="10.11646/zootaxa.4851.2.5" ID-GBIF-Dataset="7beefd33-c83a-4e05-82d1-7522b65c112c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4407593" ID-ZooBank="C0027037-9243-40E5-AAE1-22A9A272BD14" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609433360249" checkinUser="plazi" docAuthor="Park, Kyu-Tek &amp; Bae, Yang-Seop" docDate="2020" docId="03D987EDD948E0454BFEFB8D7BAAFE29" docLanguage="en" docName="zootaxa.4851.2.5.pdf" docOrigin="Zootaxa 4851 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torodora multispinata Park &amp; Bae 2020, sp. nov." docType="treatment" docVersion="8" lastPageNumber="316" masterDocId="FFE0FF95D940E04E4B69FFB37810FFDC" masterDocTitle="Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos with descriptions of five new species and two newly recorded species" masterLastPageNumber="318" masterPageNumber="305" pageNumber="313" updateTime="1698850697182" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="EDC2AB240E1F456A8D8EA9CF5EE0A798" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C8CD0F8E00D480972DA89F90E3E1C5F8">
<mods:title id="73497A82C3B3A602C75ADB1817252760">Genus Torodora Meyrick (Lepidoptera, Lecithoceridae, Torodorinae) in Laos with descriptions of five new species and two newly recorded species</mods:title>
</mods:titleInfo>
<mods:name id="2E4F9A9A6EBF3D32AC21CAF07892A4F4" type="personal">
<mods:role id="494CA8230051733EC2F4485770921F8C">
<mods:roleTerm id="63D0AD241FB61581F1CF367CEC7A089F">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="08BD6395CD8A357E98A55C9B41795E38">Park, Kyu-Tek</mods:namePart>
<mods:nameIdentifier id="75BBE5D55AB626C9132935302285C5F0" type="ORCID">0000-0001-9933-4497</mods:nameIdentifier>
<mods:affiliation id="30DD2B9080200FFFBA13CE9CCB033F1B">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497</mods:affiliation>
<mods:nameIdentifier id="85E193AADA64982087CA43D93D20F89B" type="email">ktpark02@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="237589D75E4451EB57A9C6299821330D" type="personal">
<mods:role id="D86AAD03938C66F8B235AAD71811BC3E">
<mods:roleTerm id="DFC3B9AF85A0ACB33BFF7277459A7D19">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="41FDE1D361F9BD7F2814493B39F2C78F">Bae, Yang-Seop</mods:namePart>
<mods:nameIdentifier id="91172A07AE38E8A182291250A0268E0D" type="ORCID">0000-0001-9933-4497</mods:nameIdentifier>
<mods:affiliation id="AC19D9F3D8A2CBB1BFE6AA4E4A327294">BioResource and Environmental Center, Division of Life Sciences, Incheon National University, Incheon, 22012 Korea ktpark 02 @ gmail. com; https: // orcid. org / 0000 - 0001 - 9933 - 4497 &amp; Division of Life Sciences, Incheon National University, Incheon, 22012 Korea</mods:affiliation>
<mods:nameIdentifier id="462E0C3EA72130B9CC05AF31C5A1A78D" type="email">ktpark02@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="A4A2F09AB2275C3DA923C67758B6F0D6">text</mods:typeOfResource>
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<mods:titleInfo id="2DA6FDFA9EA39E2313CF108D8407C89D">
<mods:title id="B88130FFE28A7117C1938A26033C5E21">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="425400016BD6DC1D2C8EB083CB15D032">
<mods:date id="140DEC58C6735177DED400E3869E2BB3">2020</mods:date>
<mods:detail id="C620B338C49B1AC9721A0C81178E3A22" type="pubDate">
<mods:number id="0B9211331BB1D8CC9F23D0635363AD96">2020-09-10</mods:number>
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<mods:detail id="D32B6CD404BCD697A93E0159CDD07983" type="volume">
<mods:number id="2B649B1AC93B95EC90EEA91348C044F7">4851</mods:number>
</mods:detail>
<mods:detail id="9CCDC6D7DE105A7EF4366B6A6F5F0742" type="issue">
<mods:number id="9DFEE21ECFB4A2262625CD4CF6DE4C34">2</mods:number>
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<mods:classification id="CE6DFE8A7D57210DA7E6FA3D6E3DF73F">journal article</mods:classification>
<mods:identifier id="60CA636943A5B43EDC8AE6AF0D4DE72A" type="CLB-Dataset">8531</mods:identifier>
<mods:identifier id="C2C0083995E73080E84C1EAC099B957B" type="DOI">10.11646/zootaxa.4851.2.5</mods:identifier>
<mods:identifier id="E028B99C8F8E61CDB2E3D8193E880299" type="GBIF-Dataset">7beefd33-c83a-4e05-82d1-7522b65c112c</mods:identifier>
<mods:identifier id="5EDCA949425A9E5B1381272C52D37166" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AD7A98884F84160F9CF7F52BCA288F04" type="Zenodo-Dep">4407593</mods:identifier>
<mods:identifier id="50D3B519F5E3E166747DD268187947F6" type="ZooBank">C0027037-9243-40E5-AAE1-22A9A272BD14</mods:identifier>
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<treatment id="03D987EDD948E0454BFEFB8D7BAAFE29" ID-DOI="http://doi.org/10.5281/zenodo.4407963" ID-GBIF-Taxon="177203854" ID-Zenodo-Dep="4407963" LSID="urn:lsid:plazi:treatment:03D987EDD948E0454BFEFB8D7BAAFE29" httpUri="http://treatment.plazi.org/id/03D987EDD948E0454BFEFB8D7BAAFE29" lastPageId="11" lastPageNumber="316" pageId="8" pageNumber="313">
<subSubSection id="C36A6570D948E0464BFEFB8D7A47FB84" box="[151,599,1086,1113]" pageId="8" pageNumber="313" type="nomenclature">
<paragraph id="8BCF36FBD948E0464BFEFB8D7A47FB84" blockId="8.[151,599,1086,1149]" box="[151,599,1086,1113]" pageId="8" pageNumber="313">
<heading id="D0878197D948E0464BFEFB8D7A47FB84" bold="true" box="[151,599,1086,1113]" fontSize="11" level="1" pageId="8" pageNumber="313" reason="1">
<emphasis id="B904EAE9D948E0464BFEFB8D7A47FB84" bold="true" box="[151,599,1086,1113]" pageId="8" pageNumber="313">
<taxonomicName id="4C704D78D948E0464BFEFB8D79FDFB84" ID-CoL="9FPX4" authority="Park" authorityName="Park &amp; Bae" authorityYear="2020" box="[151,493,1086,1113]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="multispinata" status="sp. nov.">
<emphasis id="B904EAE9D948E0464BFEFB8D79B6FB85" bold="true" box="[151,422,1086,1113]" italics="true" pageId="8" pageNumber="313">Torodora multispinata</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D948E0464A90FB8D7A47FB84" box="[505,599,1086,1112]" pageId="8" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D948E0464BFEFBD17933FBA1" box="[151,291,1122,1149]" pageId="8" pageNumber="313" type="description">
<paragraph id="8BCF36FBD948E0464BFEFBD17933FBA1" blockId="8.[151,599,1086,1149]" box="[151,291,1122,1149]" pageId="8" pageNumber="313">
(
<figureCitation id="134B2A7ED948E0464BF6FBD178FCFBA1" box="[159,236,1122,1149]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[161,1427,208,1839]" captionTargetId="figure-19@9.[151,1436,201,1853]" captionTargetPageId="9" captionText="FIGURE 6. Torodora multispinata Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, female genitalia, gen. slide no. CIS-8077, holotype, with close-up corpus bursae and signum; E, abdominal segment VIII and ostium bursae of paratype, gen. slide no. CIS-8100; F, ditto, close-up signum. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407610" httpUri="https://zenodo.org/record/4407610/files/figure.png" pageId="8" pageNumber="313">Figs. 6</figureCitation>
A-F)
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D948E0464BFEFB197BE8FA88" pageId="8" pageNumber="313" type="materials_examined">
<paragraph id="8BCF36FBD948E0464BFEFB197BE8FA88" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BFEFB19795CFB18" bold="true" box="[151,332,1194,1220]" pageId="8" pageNumber="313">Type material.</emphasis>
<materialsCitation id="3B183CA6D948E0464A33FB197918FAD0" ID-GBIF-Occurrence="3028414313" collectingDate="2015-11-10" collectorName="YS Bae &amp; Bae, BS &amp; Park, SM &amp; Na, JW &amp; DJ Lee" country="Laos" elevation="470" latitude="18.395" location="Phou Khao Khouay National Protected Area" pageId="8" pageNumber="313" specimenCount="1" stateProvince="Bolikhamxai" typeStatus="holotype">
<typeStatus id="54CB8859D948E0464A33FB1979D7FB18" box="[346,455,1194,1220]" pageId="8" pageNumber="313" type="holotype">Holotype</typeStatus>
: female,
<collectingCountry id="F367766BD948E0464950FB197A96FB18" box="[569,646,1194,1220]" name="Laos" pageId="8" pageNumber="313">LAOS</collectingCountry>
:
<collectingRegion id="49B4F819D948E04649FEFB197B60FB19" box="[663,880,1194,1221]" country="Laos" name="Bolikhamxai" pageId="8" pageNumber="313">Bolikhamsai Prov.</collectingRegion>
,
<location id="8EAF6020D948E04648EBFB197D88FB18" LSID="urn:lsid:plazi:treatment:03D987EDD948E0454BFEFB8D7BAAFE29:8EAF6020D948E04648EBFB197D88FB18" box="[898,1432,1194,1220]" country="Laos" latitude="18.395" name="Phou Khao Khouay National Protected Area" pageId="8" pageNumber="313" stateProvince="Bolikhamxai">Phou Khao Khouay National Protected Area</location>
,
<geoCoordinate id="EE44503CD948E0464BFEFB7D7935FB34" box="[151,293,1230,1256]" degrees="18" direction="north" minutes="23" orientation="latitude" pageId="8" pageNumber="313" precision="15" seconds="42" value="18.395">18°23’42’’N</geoCoordinate>
<geoCoordinate id="EE44503CD948E0464A45FB7D79DCFB34" box="[300,460,1230,1256]" pageId="8" pageNumber="313">103°04’ 1’’8E</geoCoordinate>
,
<quantity id="4C889B1ED948E0464AB0FB7D7A33FB35" box="[473,547,1230,1257]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="313" unit="m" value="470.0">
<elevation id="005DD1C8D948E0464AB0FB7D7A33FB35" box="[473,547,1230,1257]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="8" pageNumber="313" unit="m" value="470.0">470 m</elevation>
</quantity>
,
<date id="FFCE103BD948E0464959FB7D7ABCFB35" box="[560,684,1230,1257]" pageId="8" pageNumber="313" value="2015-11-10">
<collectingDate id="EF8AE9D3D948E0464959FB7D7ABCFB35" box="[560,684,1230,1257]" pageId="8" pageNumber="313" value="2015-11-10">10 xi 2015</collectingDate>
</date>
, leg.
<collectorName id="2685532DD948E0464981FB7D7B52FB35" box="[744,834,1230,1257]" pageId="8" pageNumber="313">YS Bae</collectorName>
, BS
<collectorName id="2685532DD948E0464811FB7D7C2FFB35" box="[888,1087,1230,1257]" pageId="8" pageNumber="313">Park, SM Na, JW</collectorName>
Kim, &amp;
<collectorName id="2685532DD948E0464FCCFB7C7CE9FB35" box="[1189,1273,1231,1257]" pageId="8" pageNumber="313">DJ Lee</collectorName>
; gen. slide no. CIS-8087
</materialsCitation>
.
<materialsCitation id="3B183CA6D948E0464A7DFB407C52FAEC" ID-GBIF-Occurrence="3028414311" collectingDate="2019-07-26" collectorName="YS Bae &amp; Bae, MJ &amp; Qi, DJ &amp; Lee, TG &amp; Lee, YB &amp; Cha, JH &amp; CM Jang" country="Laos" elevation="479" latitude="18.45639" location="Phou Khao Khouay National Protected Area" pageId="8" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Bolikhamxai" typeStatus="paratype">
<typeStatus id="54CB8859D948E0464A7DFB407995FAD1" box="[276,389,1267,1293]" pageId="8" pageNumber="313" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D76FD72D948E0464AF8FB4179A4FAD0" box="[401,436,1266,1292]" pageId="8" pageNumber="313" type="female">1♀</specimenCount>
, same locality as holotype,
<geoCoordinate id="EE44503CD948E046499DFB417B91FAD0" box="[756,897,1266,1292]" degrees="18" direction="north" minutes="27" orientation="latitude" pageId="8" pageNumber="313" precision="15" seconds="23" value="18.45639">18°27’23’’N</geoCoordinate>
<geoCoordinate id="EE44503CD948E04648E3FB417C34FAD1" box="[906,1060,1266,1293]" pageId="8" pageNumber="313">103°03’0’’5E</geoCoordinate>
, alt.
<quantity id="4C889B1ED948E0464F34FB417CB8FAD1" box="[1117,1192,1266,1293]" metricMagnitude="2" metricUnit="m" metricValue="4.79" pageId="8" pageNumber="313" unit="m" value="479.0">
<elevation id="005DD1C8D948E0464F34FB417CB8FAD1" box="[1117,1192,1266,1293]" metricMagnitude="2" metricUnit="m" metricValue="4.79" pageId="8" pageNumber="313" unit="m" value="479.0">479 m</elevation>
</quantity>
,
<date id="FFCE103BD948E0464FDCFB417D2BFAD0" box="[1205,1339,1266,1292]" pageId="8" pageNumber="313" value="2019-07-26">
<collectingDate id="EF8AE9D3D948E0464FDCFB417D2BFAD0" box="[1205,1339,1266,1292]" pageId="8" pageNumber="313" value="2019-07-26">26 vii 2019</collectingDate>
</date>
, leg.
<collectorName id="2685532DD948E0464E11FB4178D5FAED" pageId="8" pageNumber="313">YS Bae</collectorName>
, MJ
<collectorName id="2685532DD948E0464B92FAA57A7EFAED" box="[251,622,1302,1329]" pageId="8" pageNumber="313">Qi, DJ Lee, TG Lee, YB Cha, JH</collectorName>
Ko, &amp;
<collectorName id="2685532DD948E04649A8FAA57B3BFAED" box="[705,811,1302,1329]" pageId="8" pageNumber="313">CM Jang</collectorName>
; gen. slide no. CIS-8094
</materialsCitation>
;
<materialsCitation id="3B183CA6D948E0464F27FAA57BE4FA88" ID-GBIF-Occurrence="3028414306" collectorName="J. Heppner" country="Cambodia" location="Kep Prov." pageId="8" pageNumber="313" specimenCount="1" specimenCount-female="1" stateProvince="Kep" typeStatus="paratype">
<specimenCount id="9D76FD72D948E0464F27FAA57C60FAEC" box="[1102,1136,1302,1328]" pageId="8" pageNumber="313" type="female">1♀</specimenCount>
,
<collectingCountry id="F367766BD948E0464F17FAA57D0BFAEC" box="[1150,1307,1302,1328]" name="Cambodia" pageId="8" pageNumber="313">CAMBODIA</collectingCountry>
:
<collectingRegion id="49B4F819D948E0464E4FFAA47D89FAED" box="[1318,1433,1303,1329]" country="Cambodia" name="Kep" pageId="8" pageNumber="313">Kep Prov.</collectingRegion>
,
<collectingRegion id="49B4F819D948E0464BFEFA8878D8FA89" box="[151,200,1339,1365]" country="Cambodia" name="Kep" pageId="8" pageNumber="313">Kep</collectingRegion>
, Spring Vall. Lodge, 4-6 v 201, leg.
<collectorName id="2685532DD948E0464936FA887ACCFA89" box="[607,732,1339,1365]" pageId="8" pageNumber="313">J. Heppner</collectorName>
; gen. slide no. CIS-8100
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D948E0464BAEFAED7A36F9D4" pageId="8" pageNumber="313" type="diagnosis">
<paragraph id="8BCF36FBD948E0464BAEFAED7A36F9D4" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BAEFAED7952FAA4" bold="true" box="[199,322,1374,1400]" pageId="8" pageNumber="313">Diagnosis.</emphasis>
This new species is superficially similar to the preceding new species,
<taxonomicName id="4C704D78D948E0464F0AFAEC7CE6FAA4" authority="Park &amp; Bae, 2020" authorityName="Park &amp; Bae" authorityYear="2020" box="[1123,1270,1374,1400]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="313" phylum="Arthropoda" rank="species" species="cordisigna" status="sp. nov.">
<emphasis id="B904EAE9D948E0464F0AFAEC7CE6FAA4" box="[1123,1270,1374,1400]" italics="true" pageId="8" pageNumber="313">T. cordisigna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2375792D948E0464F94FAEC7D46FAA5" box="[1277,1366,1375,1401]" pageId="8" pageNumber="313" rank="species">sp. nov.</taxonomicNameLabel>
, but it can be distinguished by the darker forewing ground color having a more distinct wedge-shaped costal patch, fringe with no yellowish–white basal line nor white scales apically; and the forewing venation with R
<subScript id="17F434BED948E0464FBBFA0A7CCAFA1B" attach="left" box="[1234,1242,1465,1479]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
stalked for basal 1/3. The female genitalia can be distinguished from any known species of the genus by having two zones with numerous spines in the corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D948E0454BAEF9A17968FE55" lastPageId="11" lastPageNumber="316" pageId="8" pageNumber="313" type="description">
<paragraph id="8BCF36FBD948E0464BAEF9A17D01F800" blockId="8.[151,1437,1194,2013]" pageId="8" pageNumber="313">
<emphasis id="B904EAE9D948E0464BAEF9A17949F9F0" bold="true" box="[199,345,1554,1580]" pageId="8" pageNumber="313">Description.</emphasis>
Female (
<figureCitation id="134B2A7ED948E0464AABF9A17A00F9F1" box="[450,528,1554,1581]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[161,1427,208,1839]" captionTargetId="figure-19@9.[151,1436,201,1853]" captionTargetPageId="9" captionText="FIGURE 6. Torodora multispinata Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, female genitalia, gen. slide no. CIS-8077, holotype, with close-up corpus bursae and signum; E, abdominal segment VIII and ostium bursae of paratype, gen. slide no. CIS-8100; F, ditto, close-up signum. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407610" httpUri="https://zenodo.org/record/4407610/files/figure.png" pageId="8" pageNumber="313">Figs. 6</figureCitation>
A-C). Wingspan 14.0 mm.
<emphasis id="B904EAE9D948E0464828F9A17B6EF9F0" box="[833,894,1554,1580]" italics="true" pageId="8" pageNumber="313">Head</emphasis>
: Dark yellowish brown dorsally, with yellowish white scales laterally. Antenna slightly longer than forewing; basal segment dilated distally, dark brown dorsally and ventrally, yellowish white on anterior and posterior surface; flagellum yellowish white, with dark-brown annulations. Second segment of labial palpus thickened, arched, dark yellowish brown in basal 3/4 and yellowish white in distal 1/4 with distinct dark-brown ring preapically; yellowish white on inner surface; 3
<superScript id="7C059BB3D948E0464F0DF9117C62F96C" attach="left" box="[1124,1138,1698,1712]" fontSize="6" pageId="8" pageNumber="313">rd</superScript>
segment, slender, strongly upturned, longer than 2
<superScript id="7C059BB3D948E0464ACCF97579A5F908" attach="left" box="[421,437,1734,1748]" fontSize="6" pageId="8" pageNumber="313">nd</superScript>
segment, yellowish white, dark brown ventrally.
<emphasis id="B904EAE9D948E046489BF9757C5CF93C" box="[1010,1100,1734,1760]" italics="true" pageId="8" pageNumber="313">Thorax:</emphasis>
Tegula and thorax yellowish white dorsally. Hind tibia with yellowish-white rough scales above and creamy-white scales at apex. Forewing ground color dark yellowish brown evenly, with a small wedge-shaped, yellowish-white costal patch before 3/4 of costa; apex obtuse; termen concave beyond apex; fringe concolorous with ground color, without white basal line; venation with R
<subScript id="17F434BED948E0464A26F8DA7947F8AB" attach="left" box="[335,343,1897,1911]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
arising from near middle; distance between origins of R
<subScript id="17F434BED948E046488DF8DA7BFCF8AB" attach="left" box="[996,1004,1897,1911]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
and R
<subScript id="17F434BED948E0464F53F8DA7C52F8AB" attach="left" box="[1082,1090,1897,1911]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
less than 1/2 the length of R
<subScript id="17F434BED948E0464EFDF8DA7D8CF8AB" attach="left" box="[1428,1436,1897,1911]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
and R
<subScript id="17F434BED948E0464BB3F83E78F2F847" attach="left" box="[218,226,1933,1947]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
; R
<subScript id="17F434BED948E0464A6DF83E791CF847" attach="left" box="[260,268,1933,1947]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
and R
<subScript id="17F434BED948E0464A3AF83E794BF847" attach="left" box="[339,347,1933,1947]" fontSize="6" pageId="8" pageNumber="313">4</subScript>
stalked for basal 1/3; R
<subScript id="17F434BED948E046490CF83E7A7DF847" attach="left" box="[613,621,1933,1947]" fontSize="6" pageId="8" pageNumber="313">4</subScript>
and R
<subScript id="17F434BED948E04649DDF83F7AACF846" attach="left" box="[692,700,1932,1946]" fontSize="6" pageId="8" pageNumber="313">5</subScript>
stalked beyond distal 1/4; M
<subScript id="17F434BED948E0464F6BF83E7C1AF847" attach="left" box="[1026,1034,1933,1947]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
much nearer to M
<subScript id="17F434BED948E0464FB2F83E7CF3F847" attach="left" box="[1243,1251,1933,1947]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
; CuA
<subScript id="17F434BED948E0464E4EF83E7D3FF847" attach="both" box="[1319,1327,1933,1947]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
and CuA
<subScript id="17F434BED948E0464EFDF83E7D8CF847" attach="left" box="[1428,1436,1933,1947]" fontSize="6" pageId="8" pageNumber="313">2</subScript>
stalked for basal half; cell weakly closed. Hind wing pale brownish gray, broader than forewing; venation with M
<subScript id="17F434BED948E0464EFDF8027D8CF863" attach="left" box="[1428,1436,1969,1983]" fontSize="6" pageId="8" pageNumber="313">3</subScript>
and CuA
<subScript id="17F434BED948E0464B95F8667914F83F" attach="left" box="[252,260,2005,2019]" fontSize="6" pageId="8" pageNumber="313">1</subScript>
short-stalked; cell closed; apex acute; termen sinuate; fringe concolorous with ground color.
</paragraph>
<caption id="DF0F6673D949E0474BFEF8E67CFAF868" ID-DOI="http://doi.org/10.5281/zenodo.4407610" ID-Zenodo-Dep="4407610" httpUri="https://zenodo.org/record/4407610/files/figure.png" pageId="9" pageNumber="314" startId="9.[151,250,1877,1901]" targetBox="[161,1427,208,1839]" targetPageId="9">
<paragraph id="8BCF36FBD949E0474BFEF8E67CFAF868" blockId="9.[151,1436,1877,1973]" pageId="9" pageNumber="314">
<emphasis id="B904EAE9D949E0474BFEF8E67903F8B1" bold="true" box="[151,275,1877,1901]" pageId="9" pageNumber="314">FIGURE 6.</emphasis>
<taxonomicName id="4C704D78D949E0474A70F8E67A23F8B1" authority="Park" authorityName="Park &amp; Bae" authorityYear="2020" box="[281,563,1877,1901]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="9" pageNumber="314" phylum="Arthropoda" rank="species" species="multispinata" status="sp. nov.">
<emphasis id="B904EAE9D949E0474A70F8E679ECF8B1" box="[281,508,1877,1901]" italics="true" pageId="9" pageNumber="314">Torodora multispinata</emphasis>
Park
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D949E0474954F8E67A80F8B0" box="[573,656,1877,1901]" pageId="9" pageNumber="314" rank="species">
<emphasis id="B904EAE9D949E0474954F8E67A98F8B1" bold="true" box="[573,648,1877,1901]" pageId="9" pageNumber="314">sp. nov</emphasis>
.
</taxonomicNameLabel>
: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, female genitalia, gen. slide no. CIS-8077, holotype, with close-up corpus bursae and signum; E, abdominal segment VIII and ostium bursae of paratype, gen. slide no. CIS-8100; F, ditto, close-up signum. Scale bar for genitalia and aedeagus: 1.0 mm.
</paragraph>
</caption>
<caption id="DF0F6673D94AE0444BFEF8937A76F87F" ID-DOI="http://doi.org/10.5281/zenodo.4407612" ID-Zenodo-Dep="4407612" httpUri="https://zenodo.org/record/4407612/files/figure.png" pageId="10" pageNumber="315" startId="10.[151,250,1824,1848]" targetBox="[159,1429,207,1785]" targetPageId="10">
<paragraph id="8BCF36FBD94AE0444BFEF8937A76F87F" blockId="10.[151,1437,1824,1956]" pageId="10" pageNumber="315">
<emphasis id="B904EAE9D94AE0444BFEF8937903F8E4" bold="true" box="[151,275,1824,1848]" pageId="10" pageNumber="315">FIGURE 7.</emphasis>
<taxonomicName id="4C704D78D94AE0444A70F8937A9EF8E4" authority="(Meyrick, 1911)" authorityName="Park &amp; Bae" authorityYear="2020" baseAuthorityName="Meyrick" baseAuthorityYear="1911" box="[281,654,1824,1848]" class="Insecta" family="Lecithoceridae" genus="Torodora" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="315" phylum="Arthropoda" rank="species" species="metasaris" status="comb. nov.">
<emphasis id="B904EAE9D94AE0444A70F89379F0F8E4" box="[281,480,1824,1848]" italics="true" pageId="10" pageNumber="315">Torodora metasaris</emphasis>
(
<bibRefCitation id="EFE14B0AD94AE0444A84F8937A96F8E4" author="Meyrick, E." box="[493,646,1824,1848]" pageId="10" pageNumber="315" pagination="706 - 736" refId="ref6096" refString="Meyrick, E. (1911) Descriptions of Indian Microlepidoptera. Journal of the Bombay Natural History Society, 20, 706 - 736." type="journal article" year="1911">Meyrick, 1911</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A2375792D94AE04449F1F8937B1EF8EB" box="[664,782,1824,1848]" pageId="10" pageNumber="315" rank="species">
<emphasis id="B904EAE9D94AE04449F1F8937B16F8E4" bold="true" box="[664,774,1824,1848]" pageId="10" pageNumber="315">comb. nov</emphasis>
.
</taxonomicNameLabel>
: A, adult; B, lateral view of a paratype, with expansible brush of long orange-white hairs; C, head with labial palpus; D. venation of hindwing; E, male genitalia, gen. slide no. CIS-8075, holotype; F, ditto, aedeagus; G, uncus-saccus of a paratype, gen. slide CIS-8074; H, ditto, aedeagus; I; abdominal segments VI-VIII. Scale bar for genitalia and aedeagus: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BCF36FBD94BE0454BAEFF247AF6FEB9" blockId="11.[151,1437,150,501]" pageId="11" pageNumber="316">
<emphasis id="B904EAE9D94BE0454BAEFF247996FF6D" box="[199,390,151,177]" italics="true" pageId="11" pageNumber="316">Female genitalia</emphasis>
(
<figureCitation id="134B2A7ED94BE0454AFDFF2479F0FF6E" box="[404,480,151,178]" captionStart="FIGURE 6" captionStartId="9.[151,250,1877,1901]" captionTargetBox="[161,1427,208,1839]" captionTargetId="figure-19@9.[151,1436,201,1853]" captionTargetPageId="9" captionText="FIGURE 6. Torodora multispinata Park, sp. nov.: A, holotype; B, labial palpus in lateral view; C, hind tibia; D, female genitalia, gen. slide no. CIS-8077, holotype, with close-up corpus bursae and signum; E, abdominal segment VIII and ostium bursae of paratype, gen. slide no. CIS-8100; F, ditto, close-up signum. Scale bar for genitalia and aedeagus: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4407610" httpUri="https://zenodo.org/record/4407610/files/figure.png" pageId="11" pageNumber="316">Figs. 6</figureCitation>
D-F): Apophyses anteriores about 3/5 the length of apophyses posteriores. Abdominal sternite VIII deeply emarginated in U-shape medially; lateral corner on anterior margin sharply produced laterally. Antrum membranous, funnel-shaped. Ductus bursae narrow, shorter than the length of corpus bursae; ductus seminalis arising from near middle. Corpus bursae extremely large, ovate, with two series of unique spine-like scales posteriorly; signum heart-shaped, with strong spines along heavily sclerotized posterior margin; numerous conic spines on surface’ deeply incised on anterior margin.
</paragraph>
<paragraph id="8BCF36FBD94BE0454BAEFEDC7968FE55" blockId="11.[151,1437,150,501]" box="[199,376,367,393]" pageId="11" pageNumber="316">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C36A6570D94BE0454BAEFE217B06FE71" box="[199,790,402,430]" pageId="11" pageNumber="316" type="distribution">
<paragraph id="8BCF36FBD94BE0454BAEFE217B06FE71" blockId="11.[151,1437,150,501]" box="[199,790,402,430]" pageId="11" pageNumber="316">
<emphasis id="B904EAE9D94BE0454BAEFE217971FE70" bold="true" box="[199,353,402,428]" pageId="11" pageNumber="316">Distribution.</emphasis>
<collectingCountry id="F367766BD94BE0454A0EFE20798DFE71" box="[359,413,403,429]" name="Laos" pageId="11" pageNumber="316">Laos</collectingCountry>
(
<collectingRegion id="49B4F819D94BE0454AC4FE207A51FE72" box="[429,577,403,430]" country="Laos" name="Bolikhamxai" pageId="11" pageNumber="316">Bolikhamsai</collectingRegion>
),
<collectingCountry id="F367766BD94BE045493AFE207AD6FE71" box="[595,710,403,429]" name="Cambodia" pageId="11" pageNumber="316">Cambodia</collectingCountry>
(
<collectingRegion id="49B4F819D94BE04549BCFE207B19FE71" box="[725,777,403,429]" country="Cambodia" name="Kep" pageId="11" pageNumber="316">Kep</collectingRegion>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6570D94BE0454BAEFE047BAAFE29" pageId="11" pageNumber="316" type="etymology">
<paragraph id="8BCF36FBD94BE0454BAEFE047BAAFE29" blockId="11.[151,1437,150,501]" pageId="11" pageNumber="316">
<emphasis id="B904EAE9D94BE0454BAEFE04795EFE0D" bold="true" box="[199,334,439,465]" pageId="11" pageNumber="316">Etymology.</emphasis>
The species name is derived from the Latin,
<emphasis id="B904EAE9D94BE0454822FE047B86FE0D" box="[843,918,439,465]" italics="true" pageId="11" pageNumber="316">multus</emphasis>
(= abundant) and
<emphasis id="B904EAE9D94BE0454F0CFE047CB2FE0C" box="[1125,1186,439,464]" italics="true" pageId="11" pageNumber="316">spina</emphasis>
(= thorn), referring to numerous thorn-like spines in the corpus bursae of the female genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>
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