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<document ID-DOI="http://doi.org/10.5281/zenodo.6073575" ID-GBIF-Dataset="ba81d59c-9df9-4bd9-acdb-8bf938086141" ID-GBIF-Taxon="126093296" ID-Zenodo-Dep="6073575" checkinTime="1483605343215" checkinUser="plazi" docAuthor="Sigfrid Ingrisch, Klaus Riede &amp; George Beccalon" docDate="2016" docId="03D987D4A63DFF960F47F9E880522EB6" docLanguage="en" docName="jOrthopResearch.25.2.67.pdf" docOrigin="Journal of Orthoptera Research 25 (2)" docStyle="DocumentStyle{}" docTitle="Eulophophyllum Hebard 1922" docType="treatment" docVersion="5" lastPageNumber="68" masterDocId="FFE0FFACA63FFF940E1EFFB686582D00" masterDocTitle="The Pink Katydids of Sabah (Orthoptera: Tettigoniidae: Phaneropterinae: Eulophophyllum) with Description of Two New Species" masterLastPageNumber="74" masterPageNumber="67" pageId="2" pageNumber="68" updateTime="1644825361482" updateUser="ExternalLinkService">
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<mods:title>The Pink Katydids of Sabah (Orthoptera: Tettigoniidae: Phaneropterinae: Eulophophyllum) with Description of Two New Species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Sigfrid Ingrisch</mods:namePart>
</mods:name>
<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Klaus Riede</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>George Beccalon</mods:namePart>
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<mods:title>Journal of Orthoptera Research</mods:title>
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<mods:part>
<mods:date>2016</mods:date>
<mods:detail type="volume">
<mods:number>25</mods:number>
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<mods:detail type="issue">
<mods:number>2</mods:number>
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<mods:start>67</mods:start>
<mods:end>74</mods:end>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1665/034.025.0205</mods:identifier>
<mods:identifier type="GBIF-Dataset">ba81d59c-9df9-4bd9-acdb-8bf938086141</mods:identifier>
<mods:identifier type="Zenodo-Dep">230806</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6073575" ID-GBIF-Taxon="126093296" ID-Zenodo-Dep="6073575" LSID="urn:lsid:plazi:treatment:03D987D4A63DFF960F47F9E880522EB6" httpUri="http://treatment.plazi.org/id/03D987D4A63DFF960F47F9E880522EB6" lastPageNumber="68" pageId="2" pageNumber="68">
<subSubSection box="[345,661,1630,1652]" pageId="2" pageNumber="68" type="nomenclature">
<paragraph blockId="2.[242,764,1630,1682]" box="[345,661,1630,1652]" pageId="2" pageNumber="68">
<heading bold="true" box="[345,661,1630,1652]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="68" reason="4">
<taxonomicName authority="Hebard, 1922" authorityName="Hebard" authorityYear="1922" box="[345,661,1630,1652]" class="Insecta" family="Phaneropteridae" genus="Eulophophyllum" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="genus">
<emphasis bold="true" box="[345,661,1630,1652]" pageId="2" pageNumber="68">
<emphasis bold="true" box="[345,508,1630,1652]" italics="true" pageId="2" pageNumber="68">Eulophophyllum</emphasis>
Hebard, 1922
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection box="[242,764,1660,1682]" pageId="2" pageNumber="68" type="reference_group">
<paragraph blockId="2.[242,764,1630,1682]" box="[242,764,1660,1682]" pageId="2" pageNumber="68">
<title box="[242,323,1660,1682]" jopScore="0.075617634" pageId="2" pageNumber="68">Hebard</title>
.
<year box="[332,388,1660,1681]" pageId="2" pageNumber="68">1922</year>
.
<journalOrPublisher box="[398,686,1660,1682]" jopScore="1.132351" pageId="2" pageNumber="68">Proc. Acad. Nat. Sci. Philad.</journalOrPublisher>
<part box="[691,720,1660,1682]" pageId="2" pageNumber="68" type="volume">74</part>
:
<pagination box="[723,764,1660,1682]" pageId="2" pageNumber="68" type="pagination">160</pagination>
</paragraph>
</subSubSection>
<subSubSection box="[225,780,1719,1741]" pageId="2" pageNumber="68" type="nomenclature">
<paragraph blockId="2.[163,843,1719,2034]" box="[225,780,1719,1741]" pageId="2" pageNumber="68">urn:lsid:Orthoptera.speciesfile.org:TaxonName:12499</paragraph>
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<subSubSection pageId="2" pageNumber="68" type="diagnosis">
<paragraph blockId="2.[163,843,1719,2034]" pageId="2" pageNumber="68">
<emphasis box="[163,288,1778,1799]" italics="true" pageId="2" pageNumber="68">Diagnosis.—</emphasis>
The genus has a unique tegminal venation as discussed above. Apart from this it differs from
<emphasis box="[557,656,1806,1828]" italics="true" pageId="2" pageNumber="68">Leptoderes</emphasis>
by a more robust and in frontal view shorter head and by a rather short and stout pronotum, instead of an elongate pronotum with a laterally concave disc. The tibial tympana are conchate internally and are open on the external side.
</paragraph>
<paragraph blockId="2.[163,843,1719,2034]" lastBlockId="2.[866,1546,224,422]" pageId="2" pageNumber="68">
Apart from the tegminal venation and the modified hind tibiae, the two new species are notable for their color polymorphism comprising two color variants, (1) a variant with striking pink and red-brown body color, with wings and legs contrasting with the pastel green tegminal veins and other marks of the same color and (2) a uniform green variant so far only observed in males. Color pattern of
<taxonomicName box="[970,1115,312,334]" class="Insecta" family="Phaneropteridae" genus="Eulophophyllum" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="species" species="thaumasium">
<emphasis box="[970,1115,312,334]" italics="true" pageId="2" pageNumber="68">E. thaumasium</emphasis>
</taxonomicName>
have been described as ochreous buff with yellowish pattern (Hebard 1922), but this description was based on a dead female specimen whose color would have altered after death.
</paragraph>
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<subSubSection pageId="2" pageNumber="68" type="key">
<paragraph blockId="2.[1131,1281,458,480]" box="[1131,1281,458,480]" pageId="2" pageNumber="68">
<heading bold="true" box="[1131,1281,458,480]" centered="true" fontSize="9" level="2" pageId="2" pageNumber="68" reason="2">
<emphasis bold="true" box="[1131,1281,458,480]" pageId="2" pageNumber="68">Key to species</emphasis>
</heading>
</paragraph>
<paragraph blockId="2.[866,1546,518,951]" pageId="2" pageNumber="68">
<materialsCitation ID-GBIF-Occurrence="1423614528" pageId="2" pageNumber="68">
1. Hind tibia simple. Female tegmen with six dorsal branches of MA plus R. Labuan
<collectingCountry box="[1069,1133,547,569]" name="Iceland" pageId="2" pageNumber="68">Island</collectingCountry>
.........
<taxonomicName authority="Hebardi 1922" authorityName="Hebardi" authorityYear="1922" box="[1250,1545,546,569]" class="Insecta" family="Phaneropteridae" genus="Eulophophyllum" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="species" species="thaumasium">
<emphasis box="[1250,1397,546,569]" italics="true" pageId="2" pageNumber="68">E. thaumasium</emphasis>
Hebard, 1922
</taxonomicName>
</materialsCitation>
</paragraph>
<paragraph blockId="2.[866,1546,518,951]" box="[866,1546,576,598]" pageId="2" pageNumber="68"> Dorsal angles of hind tibia forming widened lobes........ 2.</paragraph>
<paragraph blockId="2.[866,1546,518,951]" pageId="2" pageNumber="68">
2. Tegmen with five dorsal branches of MA plus R. Dorsal area of tegmen behind R in female semi-oval, in male strongly raised, almost semi-circular with concave anterior area along stridulatory area of dorsal field in lateral view (
<figureCitation box="[1236,1293,694,716]" captionStart="Fig. 1" captionStartId="3.[182,217,1381,1403]" captionTargetBox="[324,1453,228,1336]" captionTargetId="figure@3.[299,1471,210,1371]" captionTargetPageId="3" captionText="Fig. 1. A-Fı Comparison of venation of left (Aı F) or right tegmina (B-E) between Dysmorpha, Leptoderes and Eulophophyllum species (only main veins and stronger cross veinsı most drawings without dorsal field): Aı male of Dysmorpha sp. (Myanmarı Tenasserim); Bı female of Leptoderes sp. (Thailandı Takı Mae Salid); Cı male (holotype) and Dı female of E. lobulatum sp. n. (Sabahı Kinabalu); Eı fe- male (holotype) and Fı male of E. kirki sp. n. G-Hı Left male cercus of E. lobulatum sp. n. (Gı latero-dorsal view) and E. kirki sp. n. (Hı dorso-lateral view). Left tegmina (Aı F) mirror inverted to ease comparison. Abbreviations for veins: C Costaı Ma Media anteriorı Mp Media posteriorı R Radiusı Rs Radius sectorı Sc Subcosta. Not to scale. All after photographs. Stippled line in F indicates part of tegmen hidden by hind tibia." httpUri="https://zenodo.org/record/230808/files/figure.png" pageId="2" pageNumber="68">Fig. 1</figureCitation>
C-D). Male cercus thick at base, narrowing towards and gradually transforming into apical third; moderately curved with spinule at tip
<figureCitation box="[1317,1369,752,775]" captionStart="Fig. 1" captionStartId="3.[182,217,1381,1403]" captionTargetBox="[324,1453,228,1336]" captionTargetId="figure@3.[299,1471,210,1371]" captionTargetPageId="3" captionText="Fig. 1. A-Fı Comparison of venation of left (Aı F) or right tegmina (B-E) between Dysmorpha, Leptoderes and Eulophophyllum species (only main veins and stronger cross veinsı most drawings without dorsal field): Aı male of Dysmorpha sp. (Myanmarı Tenasserim); Bı female of Leptoderes sp. (Thailandı Takı Mae Salid); Cı male (holotype) and Dı female of E. lobulatum sp. n. (Sabahı Kinabalu); Eı fe- male (holotype) and Fı male of E. kirki sp. n. G-Hı Left male cercus of E. lobulatum sp. n. (Gı latero-dorsal view) and E. kirki sp. n. (Hı dorso-lateral view). Left tegmina (Aı F) mirror inverted to ease comparison. Abbreviations for veins: C Costaı Ma Media anteriorı Mp Media posteriorı R Radiusı Rs Radius sectorı Sc Subcosta. Not to scale. All after photographs. Stippled line in F indicates part of tegmen hidden by hind tibia." httpUri="https://zenodo.org/record/230808/files/figure.png" pageId="2" pageNumber="68">Fig. 1</figureCitation>
G). Mt. Kinabalu........................................
<taxonomicName box="[1361,1483,781,803]" class="Insecta" family="Phaneropteridae" genus="Eulophophyllum" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="species" species="lobulatum" status="sp. n.">
<emphasis box="[1361,1483,781,803]" italics="true" pageId="2" pageNumber="68">E. lobulatum</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1490,1546,782,803]" pageId="2" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
<paragraph blockId="2.[866,1546,518,951]" pageId="2" pageNumber="68">
Tegmen with seven to eight dorsal branches of MA plus R. Dorsal area of female tegmen behind R nearly semi-oval in shape, in male slightly higher (
<figureCitation box="[1021,1071,870,892]" captionStart="Fig. 1" captionStartId="3.[182,217,1381,1403]" captionTargetBox="[324,1453,228,1336]" captionTargetId="figure@3.[299,1471,210,1371]" captionTargetPageId="3" captionText="Fig. 1. A-Fı Comparison of venation of left (Aı F) or right tegmina (B-E) between Dysmorpha, Leptoderes and Eulophophyllum species (only main veins and stronger cross veinsı most drawings without dorsal field): Aı male of Dysmorpha sp. (Myanmarı Tenasserim); Bı female of Leptoderes sp. (Thailandı Takı Mae Salid); Cı male (holotype) and Dı female of E. lobulatum sp. n. (Sabahı Kinabalu); Eı fe- male (holotype) and Fı male of E. kirki sp. n. G-Hı Left male cercus of E. lobulatum sp. n. (Gı latero-dorsal view) and E. kirki sp. n. (Hı dorso-lateral view). Left tegmina (Aı F) mirror inverted to ease comparison. Abbreviations for veins: C Costaı Ma Media anteriorı Mp Media posteriorı R Radiusı Rs Radius sectorı Sc Subcosta. Not to scale. All after photographs. Stippled line in F indicates part of tegmen hidden by hind tibia." httpUri="https://zenodo.org/record/230808/files/figure.png" pageId="2" pageNumber="68">Fig.1</figureCitation>
E-F).Male cercus moderately thickened at base, narrowingtowardsapicalthird;apicalthirdmarkedlythinner,slightly curved
<figureCitation box="[941,994,928,951]" captionStart="Fig. 1" captionStartId="3.[182,217,1381,1403]" captionTargetBox="[324,1453,228,1336]" captionTargetId="figure@3.[299,1471,210,1371]" captionTargetPageId="3" captionText="Fig. 1. A-Fı Comparison of venation of left (Aı F) or right tegmina (B-E) between Dysmorpha, Leptoderes and Eulophophyllum species (only main veins and stronger cross veinsı most drawings without dorsal field): Aı male of Dysmorpha sp. (Myanmarı Tenasserim); Bı female of Leptoderes sp. (Thailandı Takı Mae Salid); Cı male (holotype) and Dı female of E. lobulatum sp. n. (Sabahı Kinabalu); Eı fe- male (holotype) and Fı male of E. kirki sp. n. G-Hı Left male cercus of E. lobulatum sp. n. (Gı latero-dorsal view) and E. kirki sp. n. (Hı dorso-lateral view). Left tegmina (Aı F) mirror inverted to ease comparison. Abbreviations for veins: C Costaı Ma Media anteriorı Mp Media posteriorı R Radiusı Rs Radius sectorı Sc Subcosta. Not to scale. All after photographs. Stippled line in F indicates part of tegmen hidden by hind tibia." httpUri="https://zenodo.org/record/230808/files/figure.png" pageId="2" pageNumber="68">Fig. 1</figureCitation>
H). Danum Valley...................
<taxonomicName box="[1414,1484,928,950]" class="Insecta" family="Phaneropteridae" genus="Eulophophyllum" kingdom="Animalia" order="Orthoptera" pageId="2" pageNumber="68" phylum="Arthropoda" rank="species" species="kirki" status="sp. n.">
<emphasis box="[1414,1484,928,950]" italics="true" pageId="2" pageNumber="68">E. kirki</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1490,1546,929,950]" pageId="2" pageNumber="68" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
</treatment>
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