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<document id="F7BE2F60DD6F96434C8BDA356A62FEC5" ID-DOI="10.11646/zootaxa.3986.5.2" ID-GBIF-Dataset="9c0709d6-a583-4253-b9b6-bcf5e5d3e8f9" ID-ISSN="1175-5326" ID-Zenodo-Dep="244398" ID-ZooBank="E348A984-7A2C-46D7-A2AD-2B0E89BF95AE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461264957410" checkinUser="plazi" docAuthor="Maldaner, Maria E., Nunes, Rafael V. &amp; Vaz-De-Mello, Fernando Z." docDate="2015" docId="03D987CF547AFFB07DBBFE38FF04FC06" docLanguage="en" docName="zt03986p560.pdf" docOrigin="Zootaxa 3986 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Dichotomius (Luederwaldtinia) alvarengai Maldaner, Nunes &amp; Vaz-De-Mello, 2015, new species" docType="treatment" docVersion="8" lastPageNumber="556" masterDocId="FFE0FFB7547CFFB77D2CFFACFFDEFFEA" masterDocTitle="Taxonomic revision of the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae)" masterLastPageNumber="560" masterPageNumber="549" pageNumber="555" updateTime="1698610292206" updateUser="plazi">
<mods:mods id="10F7ECC9B6807E7AC8AA8FAACFFD830A" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="353400413BA378B67AD49E166443D2E2">Taxonomic revision of the Dichotomius speciosus (Waterhouse, 1891) species group (Coleoptera: Scarabaeidae: Scarabaeinae)</mods:title>
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<mods:name id="3E5B8748663DB7395453A0C8EB8611C4" type="personal">
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<mods:namePart id="9A061A0E9B6287E4DF91204A3D0BBC9D">Maldaner, Maria E.</mods:namePart>
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<mods:namePart id="2665129B50B15AD4E75F03B4F04EADE7">Nunes, Rafael V.</mods:namePart>
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<mods:namePart id="EE5B29EC53D0E47637CB334B14386A43">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:title id="A8DEFECD05767813929F0675D4B82D3B">Zootaxa</mods:title>
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<mods:part id="1406CEE1F24AC36EDE5296EDD5CDDB0D">
<mods:date id="D5A0F656D7C5B19D64B4ED538B036914">2015</mods:date>
<mods:detail id="EAB8B24FB616C450B81881D00498EA64" type="volume">
<mods:number id="F633C5FB344FBA3ED8F0885096987872">3986</mods:number>
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<mods:identifier id="6631F5A62054DC5FF4100D3882BAA201" type="DOI">10.11646/zootaxa.3986.5.2</mods:identifier>
<mods:identifier id="EA8A45AAEDE83F99F60EF19F33997817" type="GBIF-Dataset">9c0709d6-a583-4253-b9b6-bcf5e5d3e8f9</mods:identifier>
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<treatment id="03D987CF547AFFB07DBBFE38FF04FC06" ID-DOI="http://doi.org/10.5281/zenodo.6109950" ID-GBIF-Taxon="119637727" ID-Zenodo-Dep="6109950" LSID="urn:lsid:plazi:treatment:03D987CF547AFFB07DBBFE38FF04FC06" httpUri="http://treatment.plazi.org/id/03D987CF547AFFB07DBBFE38FF04FC06" lastPageId="7" lastPageNumber="556" pageId="6" pageNumber="555">
<subSubSection id="C36A6552547AFFB17DBBFE38FC45FDB5" pageId="6" pageNumber="555" type="nomenclature">
<paragraph id="8BCF36D9547AFFB17DBBFE38FCE7FE44" blockId="6.[151,825,404,463]" box="[151,825,404,430]" pageId="6" pageNumber="555">
<heading id="D08781B5547AFFB17DBBFE38FCE7FE44" bold="true" box="[151,825,404,430]" fontSize="11" level="1" pageId="6" pageNumber="555" reason="1">
<emphasis id="B904EACB547AFFB17DBBFE38FCE7FE44" bold="true" box="[151,825,404,430]" pageId="6" pageNumber="555">
<emphasis id="B904EACB547AFFB17DBBFE38FD7CFE44" bold="true" box="[151,674,404,430]" italics="true" pageId="6" pageNumber="555">
<taxonomicName id="4C704D5A547AFFB17DBBFE38FD42FE44" ID-CoL="6D3R6" box="[151,668,404,430]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="555" phylum="Arthropoda" rank="species" species="alvarengai" status="sp. nov." subGenus="Luederwaldtinia">Dichotomius (Luederwaldtinia) alvarengai</taxonomicName>
,
</emphasis>
<taxonomicNameLabel id="A23757B0547AFFB17F85FE38FCE7FE44" box="[681,825,404,430]" pageId="6" pageNumber="555" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF36D9547AFFB17DBBFE1AFF32FE25" blockId="6.[151,825,404,463]" box="[151,236,438,463]" pageId="6" pageNumber="555">
<emphasis id="B904EACB547AFFB17DBBFE1AFF7EFE25" bold="true" box="[151,160,438,463]" pageId="6" pageNumber="555">(</emphasis>
<figureCitation id="134B2A5C547AFFB17D8CFE1AFF3DFE25" box="[160,227,438,463]" captionStart="FIGURE 4" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[164,1430,776,1836]" captionTargetId="figure@6.[152,1435,767,1836]" captionTargetPageId="6" captionText="FIGURE 4. (A) dorsal habitus of a male Dichotomius alvarengai n. sp. (B) Dorsal habitus of a female D. alvarengai n. sp. (C) Ventral view of a male paramera. (D) Lateral view of a male paramera." httpUri="https://zenodo.org/record/244402/files/figure.png" pageId="6" pageNumber="555">Fig. 4</figureCitation>
)
</paragraph>
<paragraph id="8BCF36D9547AFFB17DBBFE52FC45FDB5" blockId="6.[151,1437,510,715]" pageId="6" pageNumber="555">
<emphasis id="B904EACB547AFFB17DBBFE52FDD9FDFD" bold="true" box="[151,519,510,535]" pageId="6" pageNumber="555">
Specimens studied.
<typeStatus id="54CB887B547AFFB17CBDFE52FDDCFDFD" box="[401,514,510,535]" pageId="6" pageNumber="555" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂
<collectingCountry id="F3677649547AFFB17F17FE53FD7DFDFD" box="[571,675,511,535]" name="Brazil" pageId="6" pageNumber="555">BRASIL</collectingCountry>
:
<emphasis id="B904EACB547AFFB17F99FE52FCEDFDFD" bold="true" box="[693,819,510,535]" pageId="6" pageNumber="555">São Paulo</emphasis>
: São José do Barreiro. Serra da Bocaina.
<quantity id="4C889B3C547AFFB1786BFE52FA47FDFD" box="[1351,1433,510,535]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="555" unit="m" value="1500.0">1500m</quantity>
.
<date id="FFCE1019547AFFB17DBBFD8EFED2FDD1" box="[151,268,546,571]" pageId="6" pageNumber="555" value="1965-11-04">4.XI.1965</date>
. F. M. Oliveira. [DZUP].
<emphasis id="B904EACB547AFFB17F11FD8EFD60FDD1" bold="true" box="[573,702,546,571]" pageId="6" pageNumber="555">
<typeStatus id="54CB887B547AFFB17F11FD8EFD67FDD1" box="[573,697,546,571]" pageId="6" pageNumber="555" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<collectingCountry id="F3677649547AFFB17FE4FD8FFCEEFDD1" box="[712,816,547,571]" name="Brazil" pageId="6" pageNumber="555">BRASIL</collectingCountry>
:
<emphasis id="B904EACB547AFFB17E12FD8EFC66FDD1" bold="true" box="[830,952,546,571]" pageId="6" pageNumber="555">São Paulo</emphasis>
: São José do Barreiro. Serra da Bocaina.
<quantity id="4C889B3C547AFFB17DBBFDEAFF37FDB5" box="[151,233,582,607]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="6" pageNumber="555" unit="m" value="1500.0">1500m</quantity>
.
<date id="FFCE1019547AFFB17DDFFDEAFEB6FDB5" box="[243,360,582,607]" pageId="6" pageNumber="555" value="1965-11-04">4.XI.1965</date>
. F. M. Oliveira. [
<specimenCount id="9D76FD50547AFFB17F0AFDEBFD87FDB5" box="[550,601,583,607]" pageId="6" pageNumber="555" type="generic">17♂</specimenCount>
8 ♀ DZUP;
<specimenCount id="9D76FD50547AFFB17FC4FDEAFCD5FDB4" box="[744,779,582,607]" pageId="6" pageNumber="555" type="generic">6♂</specimenCount>
4♀ CEMT].
</paragraph>
</subSubSection>
<subSubSection id="C36A6552547AFFB17DEBFDC6FC59FD21" pageId="6" pageNumber="555" type="diagnosis">
<paragraph id="8BCF36D9547AFFB17DEBFDC6FC59FD21" blockId="6.[151,1437,510,715]" pageId="6" pageNumber="555">
<emphasis id="B904EACB547AFFB17DEBFDC6FE9FFD69" bold="true" box="[199,321,618,643]" pageId="6" pageNumber="555">Diagnosis.</emphasis>
Within the group,
<taxonomicName id="4C704D5A547AFFB17F3FFDC7FD73FD69" box="[531,685,619,643]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="555" phylum="Arthropoda" rank="species" species="alvarengai" status="sp. nov.">
<emphasis id="B904EACB547AFFB17F3FFDC7FD73FD69" box="[531,685,619,643]" italics="true" pageId="6" pageNumber="555">D. alvarengai</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23757B0547AFFB17F98FDC6FD2DFD69" box="[692,755,618,643]" pageId="6" pageNumber="555" rank="species">
<emphasis id="B904EACB547AFFB17F98FDC6FD2DFD69" bold="true" box="[692,755,618,643]" pageId="6" pageNumber="555">n. sp.</emphasis>
</taxonomicNameLabel>
is unique in having the following combination of characters: body surface green to blue; males with cephalic carina straight, bearing one tubercle at each end, pronotum bearing a centrally projected lobe which is feebly bifurcated longitudinally.
</paragraph>
</subSubSection>
<caption id="DF0F6651547AFFB17DBBF8EEFCB8F89D" httpUri="https://zenodo.org/record/244402/files/figure.png" pageId="6" pageNumber="555" targetBox="[164,1430,776,1836]" targetPageId="6">
<paragraph id="8BCF36D9547AFFB17DBBF8EEFCB8F89D" blockId="6.[151,1436,1857,1911]" pageId="6" pageNumber="555">
<emphasis id="B904EACB547AFFB17DBBF8EEFEE2F8B2" bold="true" box="[151,316,1857,1880]" pageId="6" pageNumber="555">FIGURE 4. (A)</emphasis>
dorsal habitus of a male
<taxonomicName id="4C704D5A547AFFB17F14F8EEFCF2F8B2" box="[568,812,1858,1880]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="555" phylum="Arthropoda" rank="species" species="alvarengai" status="sp. nov.">
<emphasis id="B904EACB547AFFB17F14F8EEFCF2F8B2" box="[568,812,1858,1880]" italics="true" pageId="6" pageNumber="555">Dichotomius alvarengai</emphasis>
</taxonomicName>
<emphasis id="B904EACB547AFFB17E1EF8EEFC4DF8B2" bold="true" box="[818,915,1857,1880]" pageId="6" pageNumber="555">
<taxonomicNameLabel id="A23757B0547AFFB17E1EF8EEFCB5F8BD" box="[818,875,1858,1879]" pageId="6" pageNumber="555" rank="species">n. sp.</taxonomicNameLabel>
(B)
</emphasis>
Dorsal habitus of a female
<taxonomicName id="4C704D5A547AFFB17985F8EFFAEAF8B2" box="[1193,1332,1858,1880]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="555" phylum="Arthropoda" rank="species" species="alvarengai" status="sp. nov.">
<emphasis id="B904EACB547AFFB17985F8EFFAEAF8B2" box="[1193,1332,1858,1880]" italics="true" pageId="6" pageNumber="555">D. alvarengai</emphasis>
</taxonomicName>
<emphasis id="B904EACB547AFFB17816F8EEFA42F8B2" bold="true" box="[1338,1436,1857,1880]" pageId="6" pageNumber="555">
<taxonomicNameLabel id="A23757B0547AFFB17816F8EEFAADF8BD" box="[1338,1395,1858,1879]" pageId="6" pageNumber="555" rank="species">n. sp.</taxonomicNameLabel>
(C)
</emphasis>
Ventral view of a male paramera.
<emphasis id="B904EACB547AFFB17CC1F8CCFDCFF89D" bold="true" box="[493,529,1888,1911]" pageId="6" pageNumber="555">(D)</emphasis>
Lateral view of a male paramera.
</paragraph>
</caption>
<subSubSection id="C36A6552547AFFB07DEBF80FFD04FD6E" lastPageId="7" lastPageNumber="556" pageId="6" pageNumber="555" type="materials_examined">
<paragraph id="8BCF36D9547AFFB07DEBF80FFD04FD6E" blockId="6.[151,1436,1955,2016]" lastBlockId="7.[151,1437,151,1004]" lastPageId="7" lastPageNumber="556" pageId="6" pageNumber="555">
<emphasis id="B904EACB547AFFB17DEBF80FFEE5F856" bold="true" box="[199,315,1955,1980]" pageId="6" pageNumber="555">
<typeStatus id="54CB887B547AFFB17DEBF80FFEE8F856" box="[199,310,1955,1980]" pageId="6" pageNumber="555" type="holotype">Holotype</typeStatus>
.
</emphasis>
Male: Length:
<quantity id="4C889B3C547AFFB17CDCF808FDE5F856" box="[496,571,1956,1980]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="555" unit="mm" value="14.0">14mm</quantity>
. Maximum width (pronotum):
<quantity id="4C889B3C547AFFB17E8EF808FC3EF856" box="[930,992,1956,1980]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="6" pageNumber="555" unit="mm" value="9.0">9mm</quantity>
. Surface metallic, bluish-green.
<emphasis id="B904EACB547AFFB1787FF80FFA42F856" bold="true" box="[1363,1436,1955,1980]" pageId="6" pageNumber="555">Head:</emphasis>
Clypeo-genal junction rounded. Cephalic carina straight, bearing one horn on each end. Clypeal surface covered by ocellate punctures.
<emphasis id="B904EACB547BFFB07C43FF3BFE2FFF5A" bold="true" box="[367,497,151,176]" pageId="7" pageNumber="556">Pronotum:</emphasis>
Bearing an anterior central lobe which is weakly bifurcated longitudinally with two weak knobs. Pronotal disc, posterior and lateral borders with ocellate punctures evenly spaced. Anterior angles rounded.
<emphasis id="B904EACB547BFFB07C2DFF73FE47FF12" bold="true" box="[257,409,223,248]" pageId="7" pageNumber="556">Hypomeron:</emphasis>
with dense lateral setae similar to that on metasternum and femora, some of setae dorsally visible.
<emphasis id="B904EACB547BFFB07DDCFEA8FE42FEF7" bold="true" box="[240,412,260,285]" pageId="7" pageNumber="556">Mesosternum:</emphasis>
Covered by ocellate punctures with dense setae.
<emphasis id="B904EACB547BFFB07EEAFEA8FBAFFEF7" bold="true" box="[966,1137,260,285]" pageId="7" pageNumber="556">Metasternum:</emphasis>
Bearing dense setae at the sides and near the borders of the anterior lobe. Setae the same kind and length as those found on hypomeron, femora, and mesepisternum. Mesometasternal suture distinct.
<emphasis id="B904EACB547BFFB07E60FEE0FC7CFE8F" bold="true" box="[844,930,332,357]" pageId="7" pageNumber="556">Elytra:</emphasis>
Striae deeply impressed and bearing ocellate punctures spaced one and a half times their diameter. Interstriae bearing fine punctures (
<date id="FFCE1019547BFFB07947FEDCFB47FE62" box="[1131,1177,368,392]" pageId="7" pageNumber="556" value="1930-10">30x</date>
).
<emphasis id="B904EACB547BFFB07980FEC3FB33FE62" bold="true" box="[1196,1261,367,392]" pageId="7" pageNumber="556">Legs:</emphasis>
Ventral surface of fore femur with strong ocellate punctures all over surface. Middle and hind femora with few setose points. Anterior and posterior borders of all femora bearing dense dark brown setae, the same
<typeStatus id="54CB887B547BFFB0794EFE14FB4CFE3A" box="[1122,1170,440,464]" pageId="7" pageNumber="556">type</typeStatus>
as those on hypomeron and metasternum. Middle tibiae with green and blue sheen.
<emphasis id="B904EACB547BFFB07E14FE70FC68FE1F" bold="true" box="[824,950,476,501]" pageId="7" pageNumber="556">Abdomen:</emphasis>
6th ventrite strongly shortened medially in relation to the other ventrites. setae restricted to the sides of the abdominal ventrites. Ventrites with ocellate punctures along anterior margin.
<emphasis id="B904EACB547BFFB07F25FD88FD5BFDD7" bold="true" box="[521,645,548,573]" pageId="7" pageNumber="556">Pygidium:</emphasis>
As long as wide, covered by fine punctures (
<date id="FFCE1019547BFFB07950FD88FB77FDD6" box="[1148,1193,548,572]" pageId="7" pageNumber="556" value="1930-10">30x</date>
) equally spaced, with weak purple/violet sheen.
<emphasis id="B904EACB547BFFB07CE9FDEBFD99FD8A" bold="true" box="[453,583,583,608]" pageId="7" pageNumber="556">Paramera:</emphasis>
apex, in dorsal view rounded and slightly curved inward; in lateral view, apex with abrupt end, almost truncated (
<figureCitation id="134B2A5C547BFFB07F76FDC0FD43FD6E" box="[602,669,620,645]" captionStart="FIGURE 4" captionStartId="6.[151,250,1858,1880]" captionTargetBox="[164,1430,776,1836]" captionTargetId="figure@6.[152,1435,767,1836]" captionTargetPageId="6" captionText="FIGURE 4. (A) dorsal habitus of a male Dichotomius alvarengai n. sp. (B) Dorsal habitus of a female D. alvarengai n. sp. (C) Ventral view of a male paramera. (D) Lateral view of a male paramera." httpUri="https://zenodo.org/record/244402/files/figure.png" pageId="7" pageNumber="556">Fig. 4</figureCitation>
C, D)
</paragraph>
</subSubSection>
<subSubSection id="C36A6552547BFFB07DEBFD23FC55FCD2" pageId="7" pageNumber="556" type="description">
<paragraph id="8BCF36D9547BFFB07DEBFD23FC55FCD2" blockId="7.[151,1437,151,1004]" pageId="7" pageNumber="556">
<emphasis id="B904EACB547BFFB07DEBFD23FEE5FD42" bold="true" box="[199,315,655,680]" pageId="7" pageNumber="556">Variation.</emphasis>
Total length varying from
<quantity id="4C889B3C547BFFB07F41FD3CFD23FD4D" box="[621,765,655,680]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="7" pageNumber="556" unit="mm" value="13.5" valueMax="16.0" valueMin="11.0">11 to 16 mm</quantity>
. Small males may have cephalic carina and pronotal central lobe reduced. Females differ from males in the following respects:
<emphasis id="B904EACB547BFFB07E85FD18FC2CFD27" bold="true" box="[937,1010,692,717]" pageId="7" pageNumber="556">Head:</emphasis>
Clypeo-frontal carina lower than on males, straight and lacking tubercles at each end, sometimes almost lacking.
<emphasis id="B904EACB547BFFB07928FD7BFB58FD1A" bold="true" box="[1028,1158,727,752]" pageId="7" pageNumber="556">Pronotum:</emphasis>
Simply convex, lacking anterior lobed projection or strong declivities.
<emphasis id="B904EACB547BFFB07FF9FD50FCC8FCFF" bold="true" box="[725,790,764,789]" pageId="7" pageNumber="556">Legs:</emphasis>
Fore calcar straight, not curved downward apically.
<emphasis id="B904EACB547BFFB07DBBFCB3FECBFCD2" bold="true" box="[151,277,799,824]" pageId="7" pageNumber="556">Abdomen:</emphasis>
Sixth ventrite with the same medial width of the others.
</paragraph>
</subSubSection>
<subSubSection id="C36A6552547BFFB07DEBFCE8FE1BFC6A" pageId="7" pageNumber="556" type="distribution">
<paragraph id="8BCF36D9547BFFB07DEBFCE8FE1BFC6A" blockId="7.[151,1437,151,1004]" pageId="7" pageNumber="556">
<emphasis id="B904EACB547BFFB07DEBFCE8FEBEFCB7" bold="true" box="[199,352,836,861]" pageId="7" pageNumber="556">Distribution.</emphasis>
Apparently restricted to the
<taxonomicName id="4C704D5A547BFFB07F8AFCE8FC63FCB7" authority="Mountain Range" authorityName="Mountain Range" box="[678,957,836,861]" class="Arachnida" family="Gonyleptidae" genus="Bocaina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="552" phylum="Arthropoda" rank="genus">Bocaina mountain range</taxonomicName>
, known only from the
<typeStatus id="54CB887B547BFFB079ECFCE9FB2EFCB7" box="[1216,1264,837,861]" pageId="7" pageNumber="556">type</typeStatus>
locality in São Paulo state,
<collectingCountry id="F3677649547BFFB07C33FCCBFEBDFC6A" box="[287,355,871,896]" name="Brazil" pageId="7" pageNumber="556">Brazil</collectingCountry>
(
<figureCitation id="134B2A5C547BFFB07C5FFCCBFE69FC6A" box="[371,439,871,896]" captionStart="FIGURE 7" captionStartId="10.[151,250,1679,1701]" captionTargetBox="[155,1420,691,1655]" captionTargetId="figure@10.[151,1435,685,1667]" captionTargetPageId="10" captionText="FIGURE 7. distribution map of the Dichotomius speciosus (Waterhouse) species group, except for D. malyi, with uncertain locality. (A) Distribution of D. speciosus, D. opalescens (Felsche) and D. alvarengai n. sp. (B) detailed map of the distribution of D. speciosus and D. alvarengai n. sp. showing Serra da Mantiqueira and Serra da Boqueira Ranges. (C) Detailed distribution of D. opalescens showing the Serra Geral range." httpUri="https://zenodo.org/record/244405/files/figure.png" pageId="7" pageNumber="556">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A6552547BFFB07DEBFC20FF04FC06" pageId="7" pageNumber="556" type="etymology">
<paragraph id="8BCF36D9547BFFB07DEBFC20FF04FC06" blockId="7.[151,1437,151,1004]" pageId="7" pageNumber="556">
<emphasis id="B904EACB547BFFB07DEBFC20FE90FC4F" bold="true" box="[199,334,908,933]" pageId="7" pageNumber="556">Etymology.</emphasis>
Named after Moacyr Alvarenga (
<date id="FFCE1019547BFFB07FFFFC20FC88FC4F" box="[723,854,908,933]" pageId="7" pageNumber="556" value="1915" valueMax="2010">1915–2010</date>
), one of the great Brazilian Coleopterists, who has six genera and more than one hundred species named after him and from whose collection the
<typeStatus id="54CB887B547BFFB0799EFC1CFB3CFC22" box="[1202,1250,944,968]" pageId="7" pageNumber="556">type</typeStatus>
series originally came.
</paragraph>
</subSubSection>
</treatment>
</document>
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