284 lines
33 KiB
XML
284 lines
33 KiB
XML
<document id="AF9A12DC906E50902C1892435B8562DC" ID-DOI="10.1080/00222933.2019.1632952" ID-GBIF-Dataset="433c8642-c66c-4bdd-b5c8-d0174d063ce4" ID-ISSN="1464-5262" ID-Zenodo-Dep="3673335" ID-ZooBank="F8879B33-54DE-4EB2-92A8-FEA75B80CB22" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1582027448851" checkinUser="valdenar" docAuthor="Frolov, Andrey V., Akhmetova, Lilia A. & Vaz-de-Mello, Fernando Z." docDate="2019" docId="03D987CAFFDBA1139E20FCDFFC46EE81" docLanguage="en" docName="JNATHIST.53.17–18.1071–1077.pdf.imf" docOrigin="Journal of Natural History 53 (17)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Onorius sp. Frolov & Vaz-de-Mello 2015" docType="treatment" docVersion="9" lastPageNumber="1075" masterDocId="FFE0FFB2FFD9A1169C6BFFD2FFEAED14" masterDocTitle="Contribution to the knowledge of the Neotropical canopy dwelling genus Onorius (Coleopteraı Scarabaeidaeı Orphninae): another species from southern Ecuador and notes on the hind gut morphology in the Orphninae" masterLastPageNumber="1077" masterPageNumber="1071" pageNumber="1072" updateTime="1698718774306" updateUser="plazi">
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<mods:titleInfo id="261C8A67D900118CFEEC4A5ABC1CAE5A">
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<mods:title id="38239B46C2030015BE03787351441703">Contribution to the knowledge of the Neotropical canopy dwelling genus Onorius (Coleopteraı Scarabaeidaeı Orphninae): another species from southern Ecuador and notes on the hind gut morphology in the Orphninae</mods:title>
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<mods:namePart id="10FA3DCF1273963279914CE991393DEA">Frolov, Andrey V.</mods:namePart>
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<mods:roleTerm id="0592A9FE9E3EE0A2701D9BED81769DBC">Author</mods:roleTerm>
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<mods:namePart id="E3CEB9C30EF067D07BCC3AFD8F693FC5">Akhmetova, Lilia A.</mods:namePart>
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<mods:name id="7C237F7C224D49D0BDBB2CA7CA40BC09" type="personal">
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<mods:roleTerm id="FAE2BAE76B119CB0FFDDFB9A182476CF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BA2A192CCF3F7E7703DD41366EDC7D1F">Vaz-de-Mello, Fernando Z.</mods:namePart>
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<mods:typeOfResource id="175DF18F694DE853C09E85ABB00B5146">text</mods:typeOfResource>
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<mods:title id="81257C379A74D783475510CDF7FD0F30">Journal of Natural History</mods:title>
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<mods:part id="2AB6A39FE3353D06ADFB6673989A7CC5">
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<mods:date id="C60EF606B9AD251E791C8E9BAA4D9C56">2019</mods:date>
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<mods:number id="AA073D9670AB2452337AD62EC3EF1929">2019-06-25</mods:number>
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<mods:number id="3C21989644A7F9C08885B2147C944C61">53</mods:number>
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<mods:number id="67F8EFE5F9C48B799B8C6A7B1CE92A8B">17</mods:number>
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<mods:start id="6F3B465D8B11141D2D7F057CA8801911">1071</mods:start>
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<mods:classification id="0AA3B4055918EAF354116E0B138E3277">journal article</mods:classification>
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<mods:identifier id="ED2E43792608EE1D2A745D0CF127B9E1" type="DOI">10.1080/00222933.2019.1632952</mods:identifier>
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<mods:identifier id="C3DA50594E888F25697606088A473095" type="ISSN">1464-5262</mods:identifier>
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<mods:identifier id="F5E9688466391B56BB21CBF1494C83FD" type="Zenodo-Dep">3673335</mods:identifier>
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<treatment id="03D987CAFFDBA1139E20FCDFFC46EE81" ID-DOI="http://doi.org/10.5281/zenodo.3679693" ID-GBIF-Taxon="162161662" ID-Zenodo-Dep="3679693" LSID="urn:lsid:plazi:treatment:03D987CAFFDBA1139E20FCDFFC46EE81" httpUri="http://treatment.plazi.org/id/03D987CAFFDBA1139E20FCDFFC46EE81" lastPageId="5" lastPageNumber="1075" pageId="2" pageNumber="1072">
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<subSubSection id="C36A6557FFDBA1149E20FCDFFD32EE32" box="[587,728,781,806]" pageId="2" pageNumber="1072" type="nomenclature">
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<paragraph id="8BCF36DCFFDBA1149E20FCDFFD32EE32" blockId="2.[587,728,781,806]" box="[587,728,781,806]" pageId="2" pageNumber="1072">
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<heading id="D08781B0FFDBA1149E20FCDFFD32EE32" box="[587,728,781,806]" centered="true" fontSize="11" level="2" pageId="2" pageNumber="1072" reason="2">
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<taxonomicName id="4C704D5FFFDBA1149E20FCDFFD32EE32" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[587,728,781,806]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1072" phylum="Arthropoda" rank="species" species="sp.">
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<emphasis id="B904EACEFFDBA1149E20FCDFFD32EE32" bold="true" box="[587,728,781,806]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149E20FCDFFD41EE32" bold="true" box="[587,683,781,806]" italics="true" pageId="2" pageNumber="1072">Onorius</emphasis>
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sp.
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36A6557FFDBA1149CCBFC94FBC2EE96" pageId="2" pageNumber="1072" type="materials_examined">
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<paragraph id="8BCF36DCFFDBA1149CCBFC94FE6AEE4B" blockId="2.[160,1064,838,898]" box="[160,384,838,863]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149CCBFC94FE6AEE4B" bold="true" box="[160,384,838,863]" italics="true" pageId="2" pageNumber="1072">Material examined</emphasis>
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</paragraph>
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<paragraph id="8BCF36DCFFDBA1149CCBFCBBFBC2EE96" blockId="2.[160,1064,838,898]" box="[160,1064,873,898]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149CCBFCBBFF17EE96" bold="true" box="[160,253,873,898]" italics="true" pageId="2" pageNumber="1072">Female.</emphasis>
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<emphasis id="B904EACEFFDBA1149D67FCB8FEFBEE96" box="[268,273,874,898]" italics="true" pageId="2" pageNumber="1072">‘</emphasis>
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<materialsCitation id="3B183C81FFDBA1149D79FCB8FCAEEE96" ID-GBIF-Occurrence="2565952200" box="[274,836,874,898]" collectingDate="2008-03" location="Paquisha Alto" pageId="2" pageNumber="1072" specimenCount="1" stateProvince="Zamora-Chinchipe">
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<collectingRegion id="49B4F83EFFDBA1149D79FCB8FE08EE96" box="[274,482,874,898]" country="Ecuador" name="Zamora-Chinchipe" pageId="2" pageNumber="1072">Zamora Chinchipe</collectingRegion>
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<location id="8EAF6007FFDBA1149D81FCB8FD6CEE96" LSID="urn:lsid:plazi:treatment:03D987CAFFDBA1139E20FCDFFC46EE81:8EAF6007FFDBA1149D81FCB8FD6CEE96" box="[490,646,874,898]" name="Paquisha Alto" pageId="2" pageNumber="1072" stateProvince="Zamora-Chinchipe">Paquisha Alto</location>
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<collectingDate id="EF8AE9F4FFDBA1149EE4FCB8FCFEEE96" box="[655,788,874,898]" pageId="2" pageNumber="1072" value="2008-03">
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<date id="FFCE101CFFDBA1149EE4FCB8FCFEEE96" box="[655,788,874,898]" pageId="2" pageNumber="1072" value="2008-03">Marzo 2008</date>
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</collectingDate>
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Col
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</materialsCitation>
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.
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<collectionCode id="ED61AE19FFDBA1149F24FCB8FC8AEE96" box="[847,864,874,898]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="1072" type="Museum">V</collectionCode>
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. Carvajal
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<emphasis id="B904EACEFFDBA1149FAEFCB8FC21EE96" box="[965,971,874,898]" italics="true" pageId="2" pageNumber="1072">’</emphasis>
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(
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<collectionCode id="ED61AE19FFDBA1149FB2FCB8FBF5EE96" box="[985,1055,874,898]" pageId="2" pageNumber="1072">CEMT</collectionCode>
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).
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</paragraph>
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</subSubSection>
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<subSection id="E2FF2F37FFDBA1139CCBFC61FC46EE81" lastPageId="5" lastPageNumber="1075" pageId="2" pageNumber="1072" type="multiple">
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<subSubSection id="C36A6557FFDBA1149CCBFC61FBD6E9C5" pageId="2" pageNumber="1072" type="diagnosis">
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<paragraph id="8BCF36DCFFDBA1149CCBFC61FECBEED9" blockId="2.[160,289,947,973]" box="[160,289,947,973]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149CCBFC61FECBEED9" bold="true" box="[160,289,947,973]" pageId="2" pageNumber="1072">Diagnosis</emphasis>
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</paragraph>
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<paragraph id="8BCF36DCFFDBA1149CCBFC3BFBD6E9C5" blockId="2.[160,1155,1000,1233]" pageId="2" pageNumber="1072">
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This species differs from
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<taxonomicName id="4C704D5FFFDBA1149DD3FC3BFDB5E915" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[440,607,1001,1025]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1072" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B904EACEFFDBA1149DD3FC3BFDB5E915" box="[440,607,1001,1025]" italics="true" pageId="2" pageNumber="1072">O. inexpectatus</emphasis>
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</taxonomicName>
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in the sculpture of the body (
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<figureCitation id="134B2A59FFDBA1149FDEFC3BFBCBE915" box="[949,1057,1001,1025]" captionStart="Figure 1" captionStartId="3.[160,225,1546,1569]" captionTargetBox="[165,1150,143,1512]" captionTargetId="figure@3.[163,1153,142,1514]" captionTargetPageId="3" captionText="Figure 1. Aegidiini. Onorius sp.¸ Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (a–c, f), O. inexpectatus (d, f), Aegidium onorei (e). Habitus in dorsal (a) and ventral (b) view, external female genitalia in ventral view (c–e), locality map (f; dot – O. inexpectatus, triangle – O. sp.)." figureDoi="http://doi.org/10.5281/zenodo.3673337" httpUri="https://zenodo.org/record/3673337/files/figure.png" pageId="2" pageNumber="1072">Figure 1a</figureCitation>
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): head is punctate with transversely elongate, sometimes adjoining punctures (punctures are round and relatively sparse in
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<taxonomicName id="4C704D5FFFDBA1149D99FBFCFD72E952" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[498,664,1070,1094]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1072" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B904EACEFFDBA1149D99FBFCFD72E952" box="[498,664,1070,1094]" italics="true" pageId="2" pageNumber="1072">O. inexpectatus</emphasis>
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</taxonomicName>
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); pronotum with coarser ocellate punctures (punctures are smaller and not ocellate in
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<taxonomicName id="4C704D5FFFDBA1149E12FB83FCF5E97D" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[633,799,1105,1129]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1072" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B904EACEFFDBA1149E12FB83FCF5E97D" box="[633,799,1105,1129]" italics="true" pageId="2" pageNumber="1072">O. inexpectatus</emphasis>
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</taxonomicName>
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); elytra with characteristic U- to V-shaped punctures each bearing a distinct yellowish seta posteriorly (punctures are round in
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<taxonomicName id="4C704D5FFFDBA1149D67FB44FE5FE9BA" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[268,437,1174,1198]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="1072" phylum="Arthropoda" rank="species" species="inexpectatus">
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<emphasis id="B904EACEFFDBA1149D67FB44FE5FE9BA" box="[268,437,1174,1198]" italics="true" pageId="2" pageNumber="1072">O. inexpectatus</emphasis>
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</taxonomicName>
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; setae in the punctures are smaller, barely visible at the same magnification); proximal ileum has two large, symmetrical diverticula (
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<figureCitation id="134B2A59FFDBA1149FA8FB6BFBDBE9C5" box="[963,1073,1209,1233]" captionStart="Figure 2" captionStartId="4.[160,225,1395,1418]" captionTargetBox="[166,1150,148,1360]" captionTargetId="figure@4.[163,1153,142,1363]" captionTargetPageId="4" captionText="Figure 2. Orphninae, hind gut. Onorius inexpectatus (a), O. sp., Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (b), Triodontus itremoi (c), T. nitidulus (d), Paraegidium howdeni (e), Aegidium asperatum (f), Paraegidium pokornyi (g). Ileum diverticula and protuberances in the Onorius are arrowed." figureDoi="http://doi.org/10.5281/zenodo.3673339" httpUri="https://zenodo.org/record/3673339/files/figure.png" pageId="2" pageNumber="1072">Figure 2b</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C36A6557FFDBA1139CCBFAD4FE99EF7D" lastPageId="5" lastPageNumber="1075" pageId="2" pageNumber="1072" type="description">
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<paragraph id="8BCF36DCFFDBA1149CCBFAD4FECCE80B" blockId="2.[160,1156,1286,1727]" box="[160,294,1286,1311]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149CCBFAD4FECCE80B" bold="true" box="[160,294,1286,1311]" italics="true" pageId="2" pageNumber="1072">Description</emphasis>
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</paragraph>
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<paragraph id="8BCF36DCFFDBA1149CCBFAFBFC22E871" blockId="2.[160,1156,1286,1727]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149CCBFAFBFE0FE856" bold="true" box="[160,485,1321,1346]" italics="true" pageId="2" pageNumber="1072">
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Female (
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<figureCitation id="134B2A59FFDBA1149D67FAFBFE94E856" box="[268,382,1321,1346]" captionStart="Figure 1" captionStartId="3.[160,225,1546,1569]" captionTargetBox="[165,1150,143,1512]" captionTargetId="figure@3.[163,1153,142,1514]" captionTargetPageId="3" captionText="Figure 1. Aegidiini. Onorius sp.¸ Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (a–c, f), O. inexpectatus (d, f), Aegidium onorei (e). Habitus in dorsal (a) and ventral (b) view, external female genitalia in ventral view (c–e), locality map (f; dot – O. inexpectatus, triangle – O. sp.)." figureDoi="http://doi.org/10.5281/zenodo.3673337" httpUri="https://zenodo.org/record/3673337/files/figure.png" pageId="2" pageNumber="1072">Figures 1</figureCitation>
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and
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<figureCitation id="134B2A59FFDBA1149DACFAFBFE3CE856" box="[455,470,1321,1346]" captionStart="Figure 2" captionStartId="4.[160,225,1395,1418]" captionTargetBox="[166,1150,148,1360]" captionTargetId="figure@4.[163,1153,142,1363]" captionTargetPageId="4" captionText="Figure 2. Orphninae, hind gut. Onorius inexpectatus (a), O. sp., Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (b), Triodontus itremoi (c), T. nitidulus (d), Paraegidium howdeni (e), Aegidium asperatum (f), Paraegidium pokornyi (g). Ileum diverticula and protuberances in the Onorius are arrowed." figureDoi="http://doi.org/10.5281/zenodo.3673339" httpUri="https://zenodo.org/record/3673339/files/figure.png" pageId="2" pageNumber="1072">2</figureCitation>
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).
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</emphasis>
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Body length 9.0 mm, width of elytra
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<quantity id="4C889B39FFDBA1149FDFFAF8FBFAE856" box="[948,1040,1322,1346]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="2" pageNumber="1072" unit="mm" value="4.6">4.6 mm</quantity>
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, width of pronotum
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||
<quantity id="4C889B39FFDBA1149D73FA9FFE85E871" box="[280,367,1357,1381]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="2" pageNumber="1072" unit="mm" value="3.5">3.5 mm</quantity>
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. Colour uniform brown with pronounced bronze tint.
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</paragraph>
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<paragraph id="8BCF36DCFFDBA1149CABFABDFDD5E8FB" blockId="2.[160,1156,1286,1727]" pageId="2" pageNumber="1072">
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||
<emphasis id="B904EACEFFDBA1149CABFABDFF13E893" box="[192,249,1391,1415]" italics="true" pageId="2" pageNumber="1072">Head</emphasis>
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densely punctate with transversely elongate punctures; some punctures fused near anterior margin of clypeus. Head without traces of medial horn or tubercle. Left mandible slightly protruding past clypeus in dorsal view; labrum not protruding past clypeus. Clypeus rounded anteriorly.
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</paragraph>
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<paragraph id="8BCF36DCFFDBA1149CABFA28FCCAEB6E" blockId="2.[160,1156,1286,1727]" pageId="2" pageNumber="1072">
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<emphasis id="B904EACEFFDBA1149CABFA28FEC0EB06" box="[192,298,1530,1554]" italics="true" pageId="2" pageNumber="1072">Pronotum</emphasis>
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1.5 times wider than long, widest at basal 0.3 of its length. Anterior margin without border, basal and lateral margins bordered. Disc with shallow impression anteriorly and near base, without tubercles or deep fossae. Surface punctate with round ocellate punctures separated by 1
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<emphasis id="B904EACEFFDBA1149E77F9B0FDC3EB6E" box="[540,553,1634,1658]" italics="true" pageId="2" pageNumber="1072">–</emphasis>
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3 puncture diameters.
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</paragraph>
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<paragraph id="8BCF36DCFFDBA1149CABF957FEE1EBAB" blockId="2.[160,1156,1286,1727]" pageId="2" pageNumber="1072">
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<taxonomicName id="4C704D5FFFDBA1149CABF957FEC0EB89" box="[192,298,1669,1693]" class="Trilobita" family="Styginidae" genus="Scutellum" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Corynexochida" pageId="2" pageNumber="1072" phylum="Arthropoda" rank="genus">
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<emphasis id="B904EACEFFDBA1149CABF957FEC0EB89" box="[192,298,1669,1693]" italics="true" pageId="2" pageNumber="1072">Scutellum</emphasis>
|
||
</taxonomicName>
|
||
triangular, narrowly rounded apically, about 1/15 length of elytra, coarsely punctate.
|
||
</paragraph>
|
||
<caption id="DF0F6654FFDAA1159CCBF9D8FD34EB48" ID-DOI="http://doi.org/10.5281/zenodo.3673337" ID-Zenodo-Dep="3673337" httpUri="https://zenodo.org/record/3673337/files/figure.png" pageId="3" pageNumber="1073" startId="3.[160,225,1546,1569]" targetBox="[165,1150,143,1512]" targetPageId="3">
|
||
<paragraph id="8BCF36DCFFDAA1159CCBF9D8FD34EB48" blockId="3.[160,1155,1546,1628]" pageId="3" pageNumber="1073">
|
||
<emphasis id="B904EACEFFDAA1159CCBF9D8FF11EB35" bold="true" box="[160,251,1546,1569]" pageId="3" pageNumber="1073">Figure 1.</emphasis>
|
||
<taxonomicName id="4C704D5FFFDAA1159D68F9D8FEB2EB35" authorityName="Paulian" authorityYear="1984" box="[259,344,1546,1569]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1073" phylum="Arthropoda" rank="tribe" tribe="Aegidiini">Aegidiini</taxonomicName>
|
||
.
|
||
<taxonomicName id="4C704D5FFFDAA1159D0BF9D8FE4CEB35" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[352,422,1546,1569]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1073" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904EACEFFDAA1159D0BF9D8FE4CEB35" box="[352,422,1546,1569]" italics="true" pageId="3" pageNumber="1073">Onorius</emphasis>
|
||
</taxonomicName>
|
||
sp.¸ Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (a–c, f),
|
||
<taxonomicName id="4C704D5FFFDAA1159F93F9D8FB68EB35" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[1016,1154,1546,1569]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1073" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B904EACEFFDAA1159F93F9D8FB68EB35" box="[1016,1154,1546,1569]" italics="true" pageId="3" pageNumber="1073">O. inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
(d, f),
|
||
<taxonomicName id="4C704D5FFFDAA1159CB8F9F5FE82EB2A" authorityName="Frolov, Akhmetova & Vaz-de-Mello" authorityYear="2017" box="[211,360,1575,1598]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1073" phylum="Arthropoda" rank="species" species="onorei">
|
||
<emphasis id="B904EACEFFDAA1159CB8F9F5FE82EB2A" box="[211,360,1575,1598]" italics="true" pageId="3" pageNumber="1073">Aegidium onorei</emphasis>
|
||
</taxonomicName>
|
||
(e). Habitus in dorsal (a) and ventral (b) view, external female genitalia in ventral view (c–e), locality map (f; dot –
|
||
<taxonomicName id="4C704D5FFFDAA1159DCEF997FDDBEB48" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[421,561,1605,1628]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="1073" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B904EACEFFDAA1159DCEF997FDDBEB48" box="[421,561,1605,1628]" italics="true" pageId="3" pageNumber="1073">O. inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
, triangle –
|
||
<emphasis id="B904EACEFFDAA1159EF5F997FD47EB48" box="[670,685,1605,1628]" italics="true" pageId="3" pageNumber="1073">O</emphasis>
|
||
. sp.).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0F6654FFDDA1129CCBFAA1FBF2E8D1" ID-DOI="http://doi.org/10.5281/zenodo.3673339" ID-Zenodo-Dep="3673339" httpUri="https://zenodo.org/record/3673339/files/figure.png" pageId="4" pageNumber="1074" startId="4.[160,225,1395,1418]" targetBox="[166,1150,148,1360]" targetPageId="4">
|
||
<paragraph id="8BCF36DCFFDDA1129CCBFAA1FBF2E8D1" blockId="4.[160,1157,1395,1477]" pageId="4" pageNumber="1074">
|
||
<emphasis id="B904EACEFFDDA1129CCBFAA1FF16E89E" bold="true" box="[160,252,1395,1418]" pageId="4" pageNumber="1074">Figure 2.</emphasis>
|
||
<taxonomicName id="4C704D5FFFDDA1129D6FFAA1FE86E89E" authorityName="Erichson" authorityYear="1847" box="[260,364,1395,1418]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="subFamily" subFamily="Orphninae">Orphninae</taxonomicName>
|
||
, hind gut.
|
||
<taxonomicName id="4C704D5FFFDDA1129DB2FAA1FD74E89E" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[473,670,1395,1418]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B904EACEFFDDA1129DB2FAA1FD74E89E" box="[473,670,1395,1418]" italics="true" pageId="4" pageNumber="1074">Onorius inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
(a),
|
||
<emphasis id="B904EACEFFDDA1129EA0FAA1FD30E89E" box="[715,730,1395,1418]" italics="true" pageId="4" pageNumber="1074">O</emphasis>
|
||
. sp., Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (b),
|
||
<taxonomicName id="4C704D5FFFDDA1129D53FA42FE0EE8B3" box="[312,484,1424,1447]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="species" species="itremoi">
|
||
<emphasis id="B904EACEFFDDA1129D53FA42FE0EE8B3" box="[312,484,1424,1447]" italics="true" pageId="4" pageNumber="1074">Triodontus itremoi</emphasis>
|
||
</taxonomicName>
|
||
(c),
|
||
<taxonomicName id="4C704D5FFFDDA1129E64FA42FD9DE8B3" baseAuthorityName="Guérin-Méneville" baseAuthorityYear="1829" box="[527,631,1424,1447]" class="Insecta" family="Orphnidae" genus="Triodontus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="species" species="nitidulus">
|
||
<emphasis id="B904EACEFFDDA1129E64FA42FD9DE8B3" box="[527,631,1424,1447]" italics="true" pageId="4" pageNumber="1074">T. nitidulus</emphasis>
|
||
</taxonomicName>
|
||
(d),
|
||
<taxonomicName id="4C704D5FFFDDA1129ECFFA42FC99E8B3" authorityName="Frolov, Akhmetova & Vaz-de-Mello" authorityYear="2017" box="[676,883,1424,1447]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="species" species="howdeni">
|
||
<emphasis id="B904EACEFFDDA1129ECFFA42FC99E8B3" box="[676,883,1424,1447]" italics="true" pageId="4" pageNumber="1074">Paraegidium howdeni</emphasis>
|
||
</taxonomicName>
|
||
(e),
|
||
<taxonomicName id="4C704D5FFFDDA1129FF4FA42FB89E8B3" authorityName="Preudhomme de Borre" authorityYear="1886" box="[927,1123,1424,1447]" class="Insecta" family="Hybosoridae" genus="Aegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="species" species="asperatum">
|
||
<emphasis id="B904EACEFFDDA1129FF4FA42FB89E8B3" box="[927,1123,1424,1447]" italics="true" pageId="4" pageNumber="1074">Aegidium asperatum</emphasis>
|
||
</taxonomicName>
|
||
(f),
|
||
<taxonomicName id="4C704D5FFFDDA1129CCBFA7CFE85E8D1" authorityName="Frolov, Akhmetova & Vaz-de-Mello" authorityYear="2017" box="[160,367,1454,1477]" class="Insecta" family="Hybosoridae" genus="Paraegidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="species" species="pokornyi">
|
||
<emphasis id="B904EACEFFDDA1129CCBFA7CFE85E8D1" box="[160,367,1454,1477]" italics="true" pageId="4" pageNumber="1074">Paraegidium pokornyi</emphasis>
|
||
</taxonomicName>
|
||
(g). Ileum diverticula and protuberances in the
|
||
<taxonomicName id="4C704D5FFFDDA1129F20FA7CFC79E8D1" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[843,915,1454,1477]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="1074" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904EACEFFDDA1129F20FA7CFC79E8D1" box="[843,915,1454,1477]" italics="true" pageId="4" pageNumber="1074">Onorius</emphasis>
|
||
</taxonomicName>
|
||
are arrowed.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCF36DCFFDDA1129CABFA27FEE0EB61" blockId="4.[160,1155,1525,1687]" pageId="4" pageNumber="1074">
|
||
<emphasis id="B904EACEFFDDA1129CABFA27FF10EB19" box="[192,250,1525,1549]" italics="true" pageId="4" pageNumber="1074">Elytra</emphasis>
|
||
convex, with marked humeral and apical humps, without ridges or other elevated areas.Maximum width approximately at the middle. Striae indistinct. Elytra punctate with U- or V-shaped punctures; each puncture bears a short, yellow seta posteriorly. Base of elytra not bordered.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDDA1129CABF9ADFE2BEB83" blockId="4.[160,1155,1525,1687]" box="[192,449,1663,1687]" pageId="4" pageNumber="1074">
|
||
<emphasis id="B904EACEFFDDA1129CABF9ADFEE8EB83" box="[192,258,1663,1687]" italics="true" pageId="4" pageNumber="1074">Wings</emphasis>
|
||
fully developed.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDCA1139CABFF5CFC3FEC1A" blockId="5.[160,1156,142,617]" pageId="5" pageNumber="1075">
|
||
<emphasis id="B904EACEFFDCA1139CABFF5CFEF4EDB2" box="[192,286,142,166]" italics="true" pageId="5" pageNumber="1075">Protibiae</emphasis>
|
||
with 3 outer teeth, without smaller inner tooth. Lateral margin basad of outer teeth not crenulate. Apex and internal margin of tibia with a few slender setae. Protarsi well developed, about 2/3 length of protibiae. Claws 1/3 length of tarsomere 5. Tarsomere 5 as long as tarsomeres 2
|
||
<emphasis id="B904EACEFFDCA1139DEEFF24FE78EC1A" box="[389,402,246,270]" italics="true" pageId="5" pageNumber="1075">–</emphasis>
|
||
4 combined. Tarsomere 1 as long as tarsomeres 2
|
||
<emphasis id="B904EACEFFDCA1139FDFFF24FC2BEC1A" box="[948,961,246,270]" italics="true" pageId="5" pageNumber="1075">–</emphasis>
|
||
4.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDCA1139CABFECBFDACECCA" blockId="5.[160,1156,142,617]" pageId="5" pageNumber="1075">
|
||
<emphasis id="B904EACEFFDCA1139CABFECBFE74EC25" box="[192,414,281,305]" italics="true" pageId="5" pageNumber="1075">Hind and middle legs</emphasis>
|
||
similar in shape; metafemora and metatibiae about 1.2 times longer than mesofemora and mesotibiae. Femora sparsely punctate with elongate punctures. Tibiae somewhat triangular, with 2 apical spurs. Upper tibial spur as long as tarsomeres 1
|
||
<emphasis id="B904EACEFFDCA1139CC6FE53FF50EC8D" box="[173,186,385,409]" italics="true" pageId="5" pageNumber="1075">–</emphasis>
|
||
3; lower spur as long as or slightly longer than tarsomere 1. Claws 1/3 length of tarsomere 5. Tarsomere 5 relatively robust, as wide as the other tarsomeres, as long as tarsomeres 2
|
||
<emphasis id="B904EACEFFDCA1139803FE71FB9FECAF" box="[1128,1141,419,443]" italics="true" pageId="5" pageNumber="1075">–</emphasis>
|
||
4 combined and as long as tarsomere 1.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDCA1139CABFE3BFE31EF30" blockId="5.[160,1156,142,617]" pageId="5" pageNumber="1075">
|
||
<emphasis id="B904EACEFFDCA1139CABFE3BFE73EF15" box="[192,409,489,513]" italics="true" pageId="5" pageNumber="1075">Abdominal sternites</emphasis>
|
||
punctate with elongate punctures. Sternite 8 about 2 times wider than sternites 4
|
||
<emphasis id="B904EACEFFDCA1139D39FDD9FEB5EF37" box="[338,351,523,547]" italics="true" pageId="5" pageNumber="1075">–</emphasis>
|
||
7 medially.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDCA1139CABFDFCFC24EF52" blockId="5.[160,1156,142,617]" box="[192,974,558,582]" pageId="5" pageNumber="1075">
|
||
<taxonomicName id="4C704D5FFFDCA1139CABFDFCFECFEF52" box="[192,293,558,582]" family="Trichomycteridae" genus="Pygidium" higherTaxonomySource="CoL" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="1075" phylum="Chordata" rank="genus">
|
||
<emphasis id="B904EACEFFDCA1139CABFDFCFECFEF52" box="[192,293,558,582]" italics="true" pageId="5" pageNumber="1075">Pygidium</emphasis>
|
||
</taxonomicName>
|
||
transverse, irregularly punctate, partly hidden under elytra.
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDCA1139CABFD83FE99EF7D" blockId="5.[160,1156,142,617]" box="[192,371,593,617]" pageId="5" pageNumber="1075">
|
||
<emphasis id="B904EACEFFDCA1139CABFD83FF10EF7D" bold="true" box="[192,250,593,617]" pageId="5" pageNumber="1075">Male</emphasis>
|
||
unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36A6557FFDCA1139CCBFD58FC46EE81" pageId="5" pageNumber="1075" type="distribution">
|
||
<paragraph id="8BCF36DCFFDCA1139CCBFD58FE29EFB7" blockId="5.[160,1155,650,917]" box="[160,451,650,675]" pageId="5" pageNumber="1075">
|
||
<emphasis id="B904EACEFFDCA1139CCBFD58FE29EFB7" bold="true" box="[160,451,650,675]" italics="true" pageId="5" pageNumber="1075">Distribution and habitat</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BCF36DCFFDCA1139CCBFD7FFC46EE81" blockId="5.[160,1155,650,917]" pageId="5" pageNumber="1075">
|
||
The new species is known from only one locality, Alto Marzo in
|
||
<collectingRegion id="49B4F83EFFDCA1139FDBFD7FFEEEEFFC" country="Ecuador" name="Zamora-Chinchipe" pageId="5" pageNumber="1075">Zamora-Chinchipe province</collectingRegion>
|
||
of
|
||
<collectingCountry id="F367764CFFDCA1139D5FFD02FE7BEFFC" box="[308,401,720,744]" name="Ecuador" pageId="5" pageNumber="1075">Ecuador</collectingCountry>
|
||
(
|
||
<figureCitation id="134B2A59FFDCA1139DCDFD02FDF8EFFC" box="[422,530,720,744]" captionStart="Figure 1" captionStartId="3.[160,225,1546,1569]" captionTargetBox="[165,1150,143,1512]" captionTargetId="figure@3.[163,1153,142,1514]" captionTargetPageId="3" captionText="Figure 1. Aegidiini. Onorius sp.¸ Ecuador, Zamora-Chinchipe, Paquisha, Alto Marzo (a–c, f), O. inexpectatus (d, f), Aegidium onorei (e). Habitus in dorsal (a) and ventral (b) view, external female genitalia in ventral view (c–e), locality map (f; dot – O. inexpectatus, triangle – O. sp.)." figureDoi="http://doi.org/10.5281/zenodo.3673337" httpUri="https://zenodo.org/record/3673337/files/figure.png" pageId="5" pageNumber="1075">Figure 1f</figureCitation>
|
||
), a place on the western side of the Cordillera del Cóndor, an elevated (over
|
||
<quantity id="4C889B39FFDCA1139DB6FD21FDC6EE1F" box="[477,556,755,779]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="1075" unit="m" value="2000.0">2000m</quantity>
|
||
) enclave in the Amazonian hylaea that appears to present high endemicity. The exact habitat and collecting circumstances of the specimen are unknown. Considering the characters of the genus and complete absence of the soil contamination, normally found in all
|
||
<taxonomicName id="4C704D5FFFDCA1139EDCFC89FCF6EE67" authorityName="Paulian" authorityYear="1984" box="[695,796,859,883]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1075" phylum="Arthropoda" rank="tribe" tribe="Aegidiini">Aegidiini</taxonomicName>
|
||
taxa except
|
||
<taxonomicName id="4C704D5FFFDCA1139FDCFC89FBE7EE67" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[951,1037,859,883]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1075" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B904EACEFFDCA1139FDCFC89FBE7EE67" box="[951,1037,859,883]" italics="true" pageId="5" pageNumber="1075">Onorius</emphasis>
|
||
</taxonomicName>
|
||
, we think that the new species is a canopy dweller similar to
|
||
<taxonomicName id="4C704D5FFFDCA1139E92FCAFFC4FEE81" authorityName="Frolov & Vaz-de-Mello" authorityYear="2015" box="[761,933,893,917]" class="Insecta" family="Orphnidae" genus="Onorius" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="1075" phylum="Arthropoda" rank="species" species="inexpectatus">
|
||
<emphasis id="B904EACEFFDCA1139E92FCAFFC4FEE81" box="[761,933,893,917]" italics="true" pageId="5" pageNumber="1075">O. inexpectatus</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</subSection>
|
||
</treatment>
|
||
</document> |