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<mods:title id="61156F6F0B8756332328CCB619AEBB7C">Thrinchostoma Saussure, little known bee genus from China, with description of a new species (Hymenoptera: Halictidae)</mods:title>
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<mods:namePart id="F612F24D0580730A726F0402DAEC9FA7">Zeqing Niu</mods:namePart>
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<mods:namePart id="ECCF1ADD93D25FD4544314152A2E7FE5">Feng Liu</mods:namePart>
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Key to Chinese known species of 
<taxonomicName id="4C704D03FFAB8702FE64FD4EFCC9FD4B" ID-CoL="7WC3" box="[475,801,707,729]" class="Insecta" family="Halictidae" genus="Thrinchostoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="genus">
<emphasis id="B904EA92FFAB8702FE64FD4EFD9BFD4B" bold="true" box="[475,627,707,729]" italics="true" pageId="2" pageNumber="209">Thrinchostoma</emphasis>
(
<emphasis id="B904EA92FFAB8702FD3EFD4EFCF1FD4B" bold="true" box="[641,793,707,729]" italics="true" pageId="2" pageNumber="209">Thrinchostoma</emphasis>
)
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.
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<paragraph id="8BCF3680FFAB8702FF37FD7DFA41FC97" blockId="2.[136,1459,752,1126]" box="[136,1449,752,774]" pageId="2" pageNumber="209">1. Flagellum 11-segmented, male ...........................................................................................................................................................2</paragraph>
<paragraph id="8BCF3680FFAB8702FF07FC9DFA41FCB7" blockId="2.[136,1459,752,1126]" box="[184,1449,784,806]" pageId="2" pageNumber="209">Flagellum 10-segmented, female .......................................................................................................................................................3</paragraph>
<paragraph id="8BCF3680FFAB8702FF37FCBDFA42FCF7" blockId="2.[136,1459,752,1126]" pageId="2" pageNumber="209">
2. Malar area nearly as long as the width of mandible base (
<figureCitation id="134B2A05FFAB8702FD55FCBDFCCEFCD4" box="[746,806,816,838]" captionStart="Figures 1 – 6" captionStartId="1.[155,227,1494,1516]" captionTargetBox="[189,1399,259,1461]" captionTargetId="figure@1.[189,1399,259,1461]" captionTargetPageId="1" captionText="Figures 1 – 6. Thrinchostoma (T.) sladeni Cockerell, 1913, male. 1. Habitus, lateral view. 2. Head, frontal view. 3. Head, lateral view. 4. Metasoma, dorsal view, showing T 1 – 2. 5. Metasoma, ventral view. 6. Hind leg, lateral view, showing the trochanter, femur and tibia. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399741/files/figure.png" pageId="2" pageNumber="209">Fig. 3</figureCitation>
); trochanter of hind leg ventrally with angular projection (
<figureCitation id="134B2A05FFAB8702FADDFCBDFA77FCD4" box="[1378,1439,816,838]" captionStart="Figures 1 – 6" captionStartId="1.[155,227,1494,1516]" captionTargetBox="[189,1399,259,1461]" captionTargetId="figure@1.[189,1399,259,1461]" captionTargetPageId="1" captionText="Figures 1 – 6. Thrinchostoma (T.) sladeni Cockerell, 1913, male. 1. Habitus, lateral view. 2. Head, frontal view. 3. Head, lateral view. 4. Metasoma, dorsal view, showing T 1 – 2. 5. Metasoma, ventral view. 6. Hind leg, lateral view, showing the trochanter, femur and tibia. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399741/files/figure.png" pageId="2" pageNumber="209">Fig. 6</figureCitation>
); second marginal crossvein reaching the radial nervure (
<figureCitation id="134B2A05FFAB8702FD74FCDDFCECFCF7" box="[715,772,848,870]" captionStart="Figures 7 – 10" captionStartId="2.[156,228,2017,2039]" captionTargetBox="[189,1399,1171,1971]" captionTargetId="figure@2.[189,1399,1171,1972]" captionTargetPageId="2" captionText="Figures 7 – 10. Thrinchostoma (T.) sladeni Cockerell, 1913, male. 7. Fore wing, dorsal view, showing second submarginal crossvein and the hair spot. 8. Genitalia, dorsal view. 9. Genitalia, lateral view. 10. S 7 – 8, ventral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399742/files/figure.png" pageId="2" pageNumber="209">Fig. 7</figureCitation>
) ...................................... 
<taxonomicName id="4C704D03FFAB8702FC47FCDDFA42FCF7" ID-CoL="56LYF" authority="Cockerell, 1913 Malar" authorityName="Cockerell, 1913 Malar" box="[1016,1450,848,870]" class="Insecta" family="Halictidae" genus="Thrinchostoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="sladeni" subGenus="Thrinchostoma">
<emphasis id="B904EA92FFAB8702FC47FCDDFA42FCF7" bold="true" box="[1016,1450,848,870]" pageId="2" pageNumber="209">
<emphasis id="B904EA92FFAB8702FC47FCDDFB78FCF4" bold="true" box="[1016,1168,848,870]" italics="true" pageId="2" pageNumber="209">Thrinchostoma</emphasis>
(
<emphasis id="B904EA92FFAB8702FB21FCDDFB47FCF7" bold="true" box="[1182,1199,848,869]" italics="true" pageId="2" pageNumber="209">T.</emphasis>
) 
<emphasis id="B904EA92FFAB8702FB00FCDDFAEDFCF4" bold="true" box="[1215,1285,848,870]" italics="true" pageId="2" pageNumber="209">sladeni</emphasis>
<bibRefCitation id="EFE14B71FFAB8702FAB4FCDDFA42FCF7" author="Cockerell, T. D. A." box="[1291,1450,848,870]" journalOrPublisher="Canadian Entomologist" pageId="2" pageNumber="209" pagination="35 - 36" part="45" refString="Cockerell, T. D. A. 1913. The bee genus Thrinchostoma in Asia. Canadian Entomologist, 45: 35 - 36." title="The bee genus Thrinchostoma in Asia" type="journal article" year="1913">Cockerell, 1913</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BCF3680FFAB8702FF07FCFDFA41FC57" blockId="2.[136,1459,752,1126]" pageId="2" pageNumber="209">
Malar area longer than the width of mandible base (
<figureCitation id="134B2A05FFAB8702FD02FCFDFCEEFC14" box="[701,774,880,902]" captionStart="Figures 17 – 22" captionStartId="4.[155,227,1983,2005]" captionTargetBox="[189,1399,749,1951]" captionTargetId="figure@4.[189,1399,749,1951]" captionTargetPageId="4" captionText="Figures 17 – 22. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., male. 17. Habitus, lateral view. 18. Head, frontal view. 19. Head, lateral view. 20. Metasoma, dorsal view, showing T 1 – 2. 21. Metasoma, ventral view. 22. Hind leg, lateral view, showing the trochanter, femur and tibia. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399744/files/figure.png" pageId="2" pageNumber="209">Fig. 19</figureCitation>
); trochanter of hind leg ventrally normal, without angular projection (
<figureCitation id="134B2A05FFAB8702FF00FC1DFEEFFC34" box="[191,263,912,934]" captionStart="Figures 17 – 22" captionStartId="4.[155,227,1983,2005]" captionTargetBox="[189,1399,749,1951]" captionTargetId="figure@4.[189,1399,749,1951]" captionTargetPageId="4" captionText="Figures 17 – 22. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., male. 17. Habitus, lateral view. 18. Head, frontal view. 19. Head, lateral view. 20. Metasoma, dorsal view, showing T 1 – 2. 21. Metasoma, ventral view. 22. Hind leg, lateral view, showing the trochanter, femur and tibia. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399744/files/figure.png" pageId="2" pageNumber="209">Fig. 22</figureCitation>
); second marginal crossvein not reaching the radial nervure (
<figureCitation id="134B2A05FFAB8702FCEEFC1DFC7FFC34" box="[849,919,912,934]" captionStart="Figures 23 – 26" captionStartId="5.[156,228,2012,2034]" captionTargetBox="[189,1399,1165,1965]" captionTargetId="figure@5.[189,1399,1165,1966]" captionTargetPageId="5" captionText="Figures 23 – 26. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., male. 23. Fore wing, dorsal view, showing second submarginal crossvein and the hair spot. 24. Genitalia, dorsal view. 25. Genitalia, lateral view. 26. S 7 – 8, ventral view. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399745/files/figure.png" pageId="2" pageNumber="209">Fig. 23</figureCitation>
) ..................................................................................... ....................................................................................................................... 
<emphasis id="B904EA92FFAB8702FC23FC22FA41FC57" bold="true" box="[924,1449,943,966]" pageId="2" pageNumber="209">
<taxonomicName id="4C704D03FFAB8702FC23FC22FABAFC57" authority="Niu &amp; Zhu" authorityName="Niu &amp; Zhu" box="[924,1362,943,966]" class="Insecta" family="Halictidae" genus="Thrinchostoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="yunnanense" status="sp. nov." subGenus="Thrinchostoma">
<emphasis id="B904EA92FFAB8702FC23FC22FBDCFC57" bold="true" box="[924,1076,943,965]" italics="true" pageId="2" pageNumber="209">Thrinchostoma</emphasis>
(
<emphasis id="B904EA92FFAB8702FBFDFC3DFBBBFC57" bold="true" box="[1090,1107,944,965]" italics="true" pageId="2" pageNumber="209">T.</emphasis>
) 
<emphasis id="B904EA92FFAB8702FBDCFC3DFB33FC57" bold="true" box="[1123,1243,944,965]" italics="true" pageId="2" pageNumber="209">yunnanense</emphasis>
Niu &amp; Zhu
</taxonomicName>
, 
<taxonomicNameLabel id="A23757E9FFAB8702FAE3FC3DFA41FC57" box="[1372,1449,944,965]" pageId="2" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BCF3680FFAB8702FF37FC5DFA42FBB7" blockId="2.[136,1459,752,1126]" pageId="2" pageNumber="209">
3. Malar area about one-third as long as the width of mandible base (
<figureCitation id="134B2A05FFAB8702FC8AFC5DFC94FC74" box="[821,892,976,998]" captionStart="Figures 11 – 16" captionStartId="3.[156,228,1976,1998]" captionTargetBox="[190,1400,741,1943]" captionTargetId="figure@3.[190,1400,741,1943]" captionTargetPageId="3" captionText="Figures 11 – 16. Thrinchostoma (T.) sladeni Cockerell, 1913, female. 11. Habitus, lateral view. 12. Head, frontal view. 13. Head, lateral view. 14. Fore wing, dorsal view, showing second submarginal crossvein angulated medially. 15. Metasoma, dorsal view, showing T 1 – 2. 16. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399743/files/figure.png" pageId="2" pageNumber="209">Fig. 13</figureCitation>
); second marginal crossvein angulated medially (
<figureCitation id="134B2A05FFAB8702FAE7FC5DFA48FC74" box="[1368,1440,976,998]" captionStart="Figures 11 – 16" captionStartId="3.[156,228,1976,1998]" captionTargetBox="[190,1400,741,1943]" captionTargetId="figure@3.[190,1400,741,1943]" captionTargetPageId="3" captionText="Figures 11 – 16. Thrinchostoma (T.) sladeni Cockerell, 1913, female. 11. Habitus, lateral view. 12. Head, frontal view. 13. Head, lateral view. 14. Fore wing, dorsal view, showing second submarginal crossvein angulated medially. 15. Metasoma, dorsal view, showing T 1 – 2. 16. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399743/files/figure.png" pageId="2" pageNumber="209">Fig. 14</figureCitation>
); apical one-third part of clypeus yellowish-brown, other part of clypeus black (
<figureCitation id="134B2A05FFAB8702FC18FC7DFC07FB94" box="[935,1007,1008,1030]" captionStart="Figures 11 – 16" captionStartId="3.[156,228,1976,1998]" captionTargetBox="[190,1400,741,1943]" captionTargetId="figure@3.[190,1400,741,1943]" captionTargetPageId="3" captionText="Figures 11 – 16. Thrinchostoma (T.) sladeni Cockerell, 1913, female. 11. Habitus, lateral view. 12. Head, frontal view. 13. Head, lateral view. 14. Fore wing, dorsal view, showing second submarginal crossvein angulated medially. 15. Metasoma, dorsal view, showing T 1 – 2. 16. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399743/files/figure.png" pageId="2" pageNumber="209">Fig. 12</figureCitation>
); T2 with broad orange apical margin (
<figureCitation id="134B2A05FFAB8702FAD9FC7DFA44FB94" box="[1382,1452,1008,1030]" captionStart="Figures 11 – 16" captionStartId="3.[156,228,1976,1998]" captionTargetBox="[190,1400,741,1943]" captionTargetId="figure@3.[190,1400,741,1943]" captionTargetPageId="3" captionText="Figures 11 – 16. Thrinchostoma (T.) sladeni Cockerell, 1913, female. 11. Habitus, lateral view. 12. Head, frontal view. 13. Head, lateral view. 14. Fore wing, dorsal view, showing second submarginal crossvein angulated medially. 15. Metasoma, dorsal view, showing T 1 – 2. 16. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399743/files/figure.png" pageId="2" pageNumber="209">Fig. 15</figureCitation>
) ....................................................................................................................................... 
<taxonomicName id="4C704D03FFAB8702FC47FB82FA42FBB7" ID-CoL="56LYF" authority="Cockerell, 1913 Malar" authorityName="Cockerell, 1913 Malar" box="[1016,1450,1039,1062]" class="Insecta" family="Halictidae" genus="Thrinchostoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="sladeni" subGenus="Thrinchostoma">
<emphasis id="B904EA92FFAB8702FC47FB82FA42FBB7" bold="true" box="[1016,1450,1039,1062]" pageId="2" pageNumber="209">
<emphasis id="B904EA92FFAB8702FC47FB82FB78FBB7" bold="true" box="[1016,1168,1039,1061]" italics="true" pageId="2" pageNumber="209">Thrinchostoma</emphasis>
(
<emphasis id="B904EA92FFAB8702FB21FB9DFB47FBB7" bold="true" box="[1182,1199,1040,1061]" italics="true" pageId="2" pageNumber="209">T.</emphasis>
) 
<emphasis id="B904EA92FFAB8702FB00FB82FAEDFBB7" bold="true" box="[1215,1285,1039,1061]" italics="true" pageId="2" pageNumber="209">sladeni</emphasis>
<bibRefCitation id="EFE14B71FFAB8702FAB4FB9DFA42FBB7" author="Cockerell, T. D. A." box="[1291,1450,1040,1062]" journalOrPublisher="Canadian Entomologist" pageId="2" pageNumber="209" pagination="35 - 36" part="45" refString="Cockerell, T. D. A. 1913. The bee genus Thrinchostoma in Asia. Canadian Entomologist, 45: 35 - 36." title="The bee genus Thrinchostoma in Asia" type="journal article" year="1913">Cockerell, 1913</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BCF3680FFAB8702FF07FBBDFA41FBF7" blockId="2.[136,1459,752,1126]" pageId="2" pageNumber="209">
Malar area longer than the width of mandible base (
<figureCitation id="134B2A05FFAB8702FD0DFBBDFD12FBD4" box="[690,762,1072,1094]" captionStart="Figures 27 – 32" captionStartId="6.[155,227,1494,1516]" captionTargetBox="[189,1399,259,1461]" captionTargetId="figure@6.[189,1399,259,1461]" captionTargetPageId="6" captionText="Figures 27 – 32. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., female. 27. Habitus, lateral view. 28. Head, frontal view. 29. Head, lateral view. 30. Fore wing, dorsal view, showing second submarginal crossvein stright medially. 31 Metasoma, dorsal view, showing T 1 – 2. 32. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399746/files/figure.png" pageId="2" pageNumber="209">Fig. 29</figureCitation>
); second marginal crossvein straight medially (
<figureCitation id="134B2A05FFAB8702FB7AFBBDFAE6FBD4" box="[1221,1294,1072,1094]" captionStart="Figures 27 – 32" captionStartId="6.[155,227,1494,1516]" captionTargetBox="[189,1399,259,1461]" captionTargetId="figure@6.[189,1399,259,1461]" captionTargetPageId="6" captionText="Figures 27 – 32. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., female. 27. Habitus, lateral view. 28. Head, frontal view. 29. Head, lateral view. 30. Fore wing, dorsal view, showing second submarginal crossvein stright medially. 31 Metasoma, dorsal view, showing T 1 – 2. 32. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399746/files/figure.png" pageId="2" pageNumber="209">Fig. 30</figureCitation>
); whole clypeus dust yellow (
<figureCitation id="134B2A05FFAB8702FE8AFBDDFE95FBF4" box="[309,381,1104,1126]" captionStart="Figures 27 – 32" captionStartId="6.[155,227,1494,1516]" captionTargetBox="[189,1399,259,1461]" captionTargetId="figure@6.[189,1399,259,1461]" captionTargetPageId="6" captionText="Figures 27 – 32. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., female. 27. Habitus, lateral view. 28. Head, frontal view. 29. Head, lateral view. 30. Fore wing, dorsal view, showing second submarginal crossvein stright medially. 31 Metasoma, dorsal view, showing T 1 – 2. 32. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399746/files/figure.png" pageId="2" pageNumber="209">Fig. 28</figureCitation>
); T2 without broad orange apical margin (
<figureCitation id="134B2A05FFAB8702FCADFBDDFCB0FBF4" box="[786,856,1104,1126]" captionStart="Figures 27 – 32" captionStartId="6.[155,227,1494,1516]" captionTargetBox="[189,1399,259,1461]" captionTargetId="figure@6.[189,1399,259,1461]" captionTargetPageId="6" captionText="Figures 27 – 32. Thrinchostoma (T.) yunnanense Niu &amp; Zhu, sp. nov., female. 27. Habitus, lateral view. 28. Head, frontal view. 29. Head, lateral view. 30. Fore wing, dorsal view, showing second submarginal crossvein stright medially. 31 Metasoma, dorsal view, showing T 1 – 2. 32. Hind tibia, showing the inner tibial spur. Scale bars = 1 mm." httpUri="https://zenodo.org/record/399746/files/figure.png" pageId="2" pageNumber="209">Fig. 31</figureCitation>
)......... 
<emphasis id="B904EA92FFAB8702FC23FBC2FA41FBF7" bold="true" box="[924,1449,1103,1126]" pageId="2" pageNumber="209">
<taxonomicName id="4C704D03FFAB8702FC23FBC2FABAFBF7" authority="Niu &amp; Zhu" authorityName="Niu &amp; Zhu" box="[924,1362,1103,1126]" class="Insecta" family="Halictidae" genus="Thrinchostoma" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="209" phylum="Arthropoda" rank="species" species="yunnanense" status="sp. nov." subGenus="Thrinchostoma">
<emphasis id="B904EA92FFAB8702FC23FBC2FBDCFBF7" bold="true" box="[924,1076,1103,1125]" italics="true" pageId="2" pageNumber="209">Thrinchostoma</emphasis>
(
<emphasis id="B904EA92FFAB8702FBFDFBDDFBBBFBF7" bold="true" box="[1090,1107,1104,1125]" italics="true" pageId="2" pageNumber="209">T.</emphasis>
) 
<emphasis id="B904EA92FFAB8702FBDDFBDDFB32FBF7" bold="true" box="[1122,1242,1104,1125]" italics="true" pageId="2" pageNumber="209">yunnanense</emphasis>
Niu &amp; Zhu
</taxonomicName>
, 
<taxonomicNameLabel id="A23757E9FFAB8702FAE3FBDDFA41FBF7" box="[1372,1449,1104,1125]" pageId="2" pageNumber="209" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>