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<document id="3FF1737F96E8FD8B8471B9D8B37A657E" ID-CLB-Dataset="84340" ID-DOI="10.11646/zootaxa.5159.2.3" ID-GBIF-Dataset="31b8b42c-af69-4e7b-8880-ecc214a692d6" ID-ISSN="1175-5326" ID-Zenodo-Dep="6779034" ID-ZooBank="0E55840F-FEB3-47AD-AFA3-202469567ACC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1656482715942" checkinUser="plazi" docAuthor="Martens, Jochen" docDate="2022" docId="03D287DBFFE6995AFBE2FBC7C592F97F" docLanguage="en" docName="zootaxa.5129.2.3.pdf" docOrigin="Zootaxa 5159 (2)" docSource="https://doi.org/10.11646/zootaxa.5159.2.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Filopalpus joschmidti Martens, 2022, sp. nov." docType="treatment" docVersion="5" lastPageNumber="229" masterDocId="FFEBFFA3FFE29952FB75FFF3C659FFE6" masterDocTitle="From the Ethiopian Bale Mountains hotspot-Filopalpinae subfam. nov., a new taxon of Laniatorean harvestmen based on external and genital morphology (Arachnida, Opiliones, Assamiidae)" masterLastPageNumber="244" masterPageNumber="221" pageNumber="225" updateTime="1699353417278" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="F7AB45BA2B4C93E96A7CB3A10B21FCC0">From the Ethiopian Bale Mountains hotspot-Filopalpinae subfam. nov., a new taxon of Laniatorean harvestmen based on external and genital morphology (Arachnida, Opiliones, Assamiidae)</mods:title>
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<mods:namePart id="2B367E6AB0D8DAAC1ADDA441C06F1089">Martens, Jochen</mods:namePart>
<mods:nameIdentifier id="0E61ACA5709C69553D862FD33464B2D5" type="ORCID">0000-0001-6863-8148</mods:nameIdentifier>
<mods:affiliation id="2B9222677840E3213B81E610CCE2DCA3">Johannes Gutenberg-Universität, Institut für Organismische und Molekulare Evolutionsbiologie (iomE), D- 55099 Mainz, Germany. Senckenberg Research Institute, Arachnology, D- 60325 Frankfurt am Main, Germany.</mods:affiliation>
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<treatment id="03D287DBFFE6995AFBE2FBC7C592F97F" ID-DOI="http://doi.org/10.5281/zenodo.6779050" ID-GBIF-Taxon="196368884" ID-Zenodo-Dep="6779050" LSID="urn:lsid:plazi:treatment:03D287DBFFE6995AFBE2FBC7C592F97F" httpUri="http://treatment.plazi.org/id/03D287DBFFE6995AFBE2FBC7C592F97F" lastPageId="8" lastPageNumber="229" pageId="4" pageNumber="225">
<subSubSection id="C3616546FFE69956FBE2FBC7C45AFBA9" box="[151,515,1076,1103]" pageId="4" pageNumber="225" type="nomenclature">
<paragraph id="8BC436CDFFE69956FBE2FBC7C45AFBA9" blockId="4.[151,515,1076,1139]" box="[151,515,1076,1103]" pageId="4" pageNumber="225">
<heading id="D08C81A1FFE69956FBE2FBC7C45AFBA9" bold="true" box="[151,515,1076,1103]" fontSize="11" level="1" pageId="4" pageNumber="225" reason="1">
<emphasis id="B90FEADFFFE69956FBE2FBC7C45AFBA9" bold="true" box="[151,515,1076,1103]" pageId="4" pageNumber="225">
<taxonomicName id="4C7B4D4EFFE69956FBE2FBC7C7C6FBA9" ID-CoL="9X5YT" authorityName="Martens" authorityYear="2022" box="[151,415,1076,1103]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="225" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFE69956FBE2FBC7C7C6FBA9" bold="true" box="[151,415,1076,1103]" italics="true" pageId="4" pageNumber="225">Filopalpus joschmidti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57A4FFE69956FAD3FBC6C45AFBA9" box="[422,515,1077,1103]" pageId="4" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3616546FFE69956FBE2FBABC740FB95" box="[151,281,1112,1139]" pageId="4" pageNumber="225" type="description">
<paragraph id="8BC436CDFFE69956FBE2FBABC740FB95" blockId="4.[151,515,1076,1139]" box="[151,281,1112,1139]" pageId="4" pageNumber="225">
(
<figureCitation id="13402A48FFE69956FBEAFBABC748FB95" box="[159,273,1112,1139]" captionStart-0="FIGURES 16" captionStart-1="FIGURES 710" captionStart-2="FIGURES 1124" captionStartId-0="5.[151,264,1909,1933]" captionStartId-1="6.[151,264,1548,1572]" captionStartId-2="7.[151,264,1874,1898]" captionTargetBox-0="[243,1345,179,1882]" captionTargetBox-1="[167,1418,197,1517]" captionTargetBox-2="[208,1379,192,1838]" captionTargetId-0="figure-15@5.[243,1345,179,1882]" captionTargetId-1="figure-30@6.[151,1436,181,1524]" captionTargetId-2="figure-15@7.[195,1392,181,1850]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." captionText-1="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." captionText-2="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi-0="http://doi.org/10.5281/zenodo.6779036" figureDoi-1="http://doi.org/10.5281/zenodo.6779040" figureDoi-2="http://doi.org/10.5281/zenodo.6779044" httpUri-0="https://zenodo.org/record/6779036/files/figure.png" httpUri-1="https://zenodo.org/record/6779040/files/figure.png" httpUri-2="https://zenodo.org/record/6779044/files/figure.png" pageId="4" pageNumber="225">Figs 124</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3616546FFE69956FBE2FB52C7DDFAE4" pageId="4" pageNumber="225" type="materials_examined">
<paragraph id="8BC436CDFFE69956FBE2FB52C7DDFAE4" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<emphasis id="B90FEADFFFE69956FBE2FB52C713FB5C" bold="true" box="[151,330,1184,1211]" pageId="4" pageNumber="225">Type material:</emphasis>
<materialsCitation id="3B133C90FFE69956FA21FB53C527FB39" ID-GBIF-Occurrence="3820340309" collectingDate="2019-06-02" country="Ethiopia" county="Bale Mts." elevation="3630" latitude="7.029722" location="upper Salgen Valley" longLatPrecision="21" longitude="39.18389" municipality="Washia Forest" pageId="4" pageNumber="225" specimenCount="1" specimenCount-male="1" stateProvince="Oromiya" typeStatus="holotype">
<typeStatus id="54C0886FFFE69956FA21FB53C7E5FB5C" box="[340,444,1184,1210]" pageId="4" pageNumber="225" type="holotype">Holotype</typeStatus>
(CJM 9003),
<specimenCount id="9D7DFD44FFE69956F914FB52C421FB5D" box="[609,632,1185,1211]" pageId="4" pageNumber="225" type="male"></specimenCount>
,
<collectingCountry id="F36C765DFFE69956F9F2FB52C552FB5D" box="[647,779,1185,1211]" name="Ethiopia" pageId="4" pageNumber="225">ETHIOPIA</collectingCountry>
,
<collectingRegion id="49BFF82FFFE69956F86CFB53C528FB5C" box="[793,881,1184,1210]" country="Ethiopia" name="Oromiya" pageId="4" pageNumber="225">Oromia</collectingRegion>
,
<collectingCounty id="62A54E41FFE69956F80AFB53C5A8FB5D" box="[895,1009,1184,1211]" pageId="4" pageNumber="225">Bale Mts.</collectingCounty>
,
<collectingMunicipality id="6BA0ACB7FFE69956F88AFB53C2FAFB5D" box="[1023,1187,1184,1211]" pageId="4" pageNumber="225">Washia Forest</collectingMunicipality>
,
<location id="8EA46016FFE69956FFC4FB52C3C0FB5C" LSID="urn:lsid:plazi:treatment:03D287DBFFE6995AFBE2FBC7C592F97F:8EA46016FFE69956FFC4FB52C3C0FB5C" box="[1201,1433,1184,1211]" country="Ethiopia" county="Bale Mts." latitude="7.029722" longLatPrecision="21" longitude="39.18389" municipality="Washia Forest" name="upper Salgen Valley" pageId="4" pageNumber="225" stateProvince="Oromiya">upper Salgen Valley</location>
, above
<location id="8EA46016FFE69956FB94FB37C719FB38" LSID="urn:lsid:plazi:treatment:03D287DBFFE6995AFBE2FBC7C592F97F:8EA46016FFE69956FB94FB37C719FB38" box="[225,320,1220,1246]" country="Ethiopia" county="Bale Mts." latitude="7.029722" longLatPrecision="21" longitude="39.18389" municipality="Washia Forest" name="Sebsebe" pageId="4" pageNumber="225" stateProvince="Oromiya">Sebsebe</location>
,
<quantity id="4C839B28FFE69956FA39FB37C7FAFB39" box="[332,419,1220,1247]" metricMagnitude="3" metricUnit="m" metricValue="3.63" pageId="4" pageNumber="225" unit="m" value="3630.0">
<elevation id="0056D1FEFFE69956FA39FB37C7FAFB39" box="[332,419,1220,1247]" metricMagnitude="3" metricUnit="m" metricValue="3.63" pageId="4" pageNumber="225" unit="m" value="3630.0">3630 m</elevation>
</quantity>
, leg.
<date id="FFC5100DFFE69956FA95FB37C43BFB38" box="[480,610,1220,1246]" pageId="4" pageNumber="225" value="2019-06-02">
<collectingDate id="EF81E9E5FFE69956FA95FB37C43BFB38" box="[480,610,1220,1246]" pageId="4" pageNumber="225" value="2019-06-02">06.02.2019</collectingDate>
</date>
;
<geoCoordinate id="EE4F500AFFE69956F918FB36C4A0FB39" box="[621,761,1221,1247]" degrees="07" direction="north" minutes="01" orientation="latitude" pageId="4" pageNumber="225" precision="15" seconds="47" value="7.029722">07°0147”N</geoCoordinate>
,
<geoCoordinate id="EE4F500AFFE69956F870FB36C527FB39" box="[773,894,1221,1247]" degrees="39" direction="east" minutes="11" orientation="longitude" pageId="4" pageNumber="225" precision="15" seconds="2" value="39.18389">39°112”E</geoCoordinate>
</materialsCitation>
.
<materialsCitation id="3B133C90FFE69956F8FFFB36C3C5FB38" ID-GBIF-Occurrence="3820340303" box="[906,1436,1220,1247]" collectingDate="2019-06-02" country="Ethiopia" county="Bale Mts." elevation="3630" latitude="7.029722" location="upper Salgen Valley" longLatPrecision="21" longitude="39.18389" municipality="Washia Forest" pageId="4" pageNumber="225" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Oromiya" typeStatus="paratype">
<typeStatus id="54C0886FFFE69956F8FFFB36C5AEFB39" box="[906,1015,1221,1247]" pageId="4" pageNumber="225" type="paratype">Paratypes</typeStatus>
(CJM 9002),
<specimenCount id="9D7DFD44FFE69956FFE0FB36C29BFB39" box="[1173,1218,1221,1247]" pageId="4" pageNumber="225" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D7DFD44FFE69956FFBBFB36C2A0FB39" box="[1230,1273,1221,1247]" pageId="4" pageNumber="225" type="female">2 ♀</specimenCount>
, same locality
</materialsCitation>
<materialsCitation id="3B133C90FFE69956FBE2FB1BC7D8FAE4" ID-GBIF-Occurrence="3820340301" box="[151,385,1256,1283]" collectingDate="2019-06-02" country="Ethiopia" county="Bale Mts." elevation="3630" latitude="7.029722" location="upper Salgen Valley" longLatPrecision="21" longitude="39.18389" municipality="Washia Forest" pageId="4" pageNumber="225" specimenCount="1" stateProvince="Oromiya" typeStatus="paratype">and data as holotype</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3616546FFE69956FBB2FAFFC71BFA50" pageId="4" pageNumber="225" type="diagnosis">
<paragraph id="8BC436CDFFE69956FBB2FAFFC71BFA50" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<emphasis id="B90FEADFFFE69956FBB2FAFFC71BFAC0" bold="true" box="[199,322,1292,1318]" pageId="4" pageNumber="225">Diagnosis.</emphasis>
Total body length 3.0
<quantity id="4C839B28FFE69956F935FAFEC4CCFAC1" box="[576,661,1293,1319]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="225" unit="mm" value="3.1">3.1 mm</quantity>
, dorsal side of body covered with a dense layer of polygonal tubercles, touching each other, male pedipalp four times longer than body length. Pedipalp Fe nearly twice as long as body, Pt slightly shorter. Penis parallel-sided truncus except last fifth, marked concavity on dorso-distal part of truncus, the ventral plate, i.e. distal part of penis slightly enlarged, glans tube surpassing distal rim of penis; strong armament of free tergites.
</paragraph>
</subSubSection>
<subSubSection id="C3616546FFE69956FBB2FA32C2F7FA18" pageId="4" pageNumber="225" type="discussion">
<paragraph id="8BC436CDFFE69956FBB2FA32C2F7FA18" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<emphasis id="B90FEADFFFE69956FBB2FA32C764FA3D" bold="true" box="[199,317,1473,1499]" pageId="4" pageNumber="225">Remarks:</emphasis>
<taxonomicName id="4C7B4D4EFFE69956FA32FA33C419FA3C" box="[327,576,1472,1498]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="4" pageNumber="225" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFE69956FA32FA33C419FA3C" box="[327,576,1472,1498]" italics="true" pageId="4" pageNumber="225">Filopalpus joschmidti</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFE69956F93CFA32C4FFFA3D" bold="true" box="[585,678,1473,1499]" pageId="4" pageNumber="225">
<taxonomicNameLabel id="A23C57A4FFE69956F93CFA32C4FFFA3D" box="[585,678,1473,1499]" pageId="4" pageNumber="225" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is the species bearing the longest male pedipalp of all congeners presently known; it slightly surpasses length of leg II and the length of the body by 5.1 times.
</paragraph>
</subSubSection>
<subSubSection id="C3616546FFE69956FBB2F9FAC769F98D" pageId="4" pageNumber="225" type="etymology">
<paragraph id="8BC436CDFFE69956FBB2F9FAC769F98D" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<emphasis id="B90FEADFFFE69956FBB2F9FAC70BF9C5" bold="true" box="[199,338,1545,1571]" pageId="4" pageNumber="225">Etymology:</emphasis>
It honors Joachim Schmidt, renowned entomologist working on ground beetles with focus on Himalayan fauna, especially their taxonomy, systematics, evolutionary and biogeographic affinities. Name in genitive case.
</paragraph>
</subSubSection>
<subSubSection id="C3616546FFE6995AFBB2F987C79FFA27" lastPageId="8" lastPageNumber="229" pageId="4" pageNumber="225" type="description">
<paragraph id="8BC436CDFFE69956FBB2F987C222F8A5" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<emphasis id="B90FEADFFFE69956FBB2F987C702F968" bold="true" box="[199,347,1652,1678]" pageId="4" pageNumber="225">Description:</emphasis>
<emphasis id="B90FEADFFFE69956FA14F986C42CF968" box="[353,629,1652,1679]" italics="true" pageId="4" pageNumber="225">MALE. Body, dorsal side</emphasis>
(
<figureCitation id="13402A48FFE69956F9F6F987C4BFF969" box="[643,742,1652,1679]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="4" pageNumber="225">Figs 13</figureCitation>
,
<figureCitation id="13402A48FFE69956F984F986C548F969" box="[753,785,1653,1679]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="4" pageNumber="225">10</figureCitation>
): Scutum densely covered with marked polygonal blackish tubercles, touching each other on Abd scutum but slightly separated from each other on prosoma, indicated by light patches between groups of tubercles, distal rim of prosoma equipped with a row of 11 (HT, 8 PA) pointed apophyses of various sizes, large ones on left and right third of distal rim; a single pair of para-median tubercles on the scutal areas IIV, on area V a row of long pointed tubercles; free tergites visible from above, each covered by a narrow row of slender, drawn-out pointed tubercles, longest in central third of tergites (
<figureCitation id="13402A48FFE69956F8ABF8DBC21AF8A5" box="[990,1091,1832,1859]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="4" pageNumber="225">Figs 13</figureCitation>
,
<figureCitation id="13402A48FFE69956FF3AF8DAC237F8A5" box="[1103,1134,1833,1859]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="4" pageNumber="225">10</figureCitation>
).
</paragraph>
<paragraph id="8BC436CDFFE69956FBB2F8BFC416F86C" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<emphasis id="B90FEADFFFE69956FBB2F8BFC734F880" box="[199,365,1868,1894]" italics="true" pageId="4" pageNumber="225">Colour pattern</emphasis>
(
<figureCitation id="13402A48FFE69956FA0FF8BFC7A1F881" box="[378,504,1868,1895]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="4" pageNumber="225">Figs 13, 5</figureCitation>
): Most tubercles brownish to blackish, small light yellow parts on prosoma, blackish on rim of Abd scutum and free tergites.
</paragraph>
<paragraph id="8BC436CDFFE69956FBB2F867C2F1F810" blockId="4.[151,1437,1184,2039]" pageId="4" pageNumber="225">
<taxonomicName id="4C7B4D4EFFE69956FBB2F867C722F848" box="[199,379,1940,1966]" class="Agaricomycetes" family="Paxillaceae" genus="Tuber" kingdom="Fungi" order="Boletales" pageId="4" pageNumber="225" phylum="Basidiomycota" rank="species" species="oculorum">
<emphasis id="B90FEADFFFE69956FBB2F867C722F848" box="[199,379,1940,1966]" italics="true" pageId="4" pageNumber="225">Tuber oculorum</emphasis>
</taxonomicName>
(
<figureCitation id="13402A48FFE69956FAF9F867C7ABF849" box="[396,498,1940,1967]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="4" pageNumber="225">Figs 3, 5</figureCitation>
,
<figureCitation id="13402A48FFE69956FA8AF866C446F849" box="[511,543,1941,1967]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="4" pageNumber="225">10</figureCitation>
): Small, front part less than half of its length apart from distal rim of prosomal scutum, equipped with a number of long pointed tubercles, longest on anterior and posterior rim of Tu oc, pointing upwards, few small tubercles around eye, longest reaching and slightly surpassing distal rim.
</paragraph>
<caption id="DF046645FFE79957FBE2F886C31BF81F" ID-DOI="http://doi.org/10.5281/zenodo.6779036" ID-Zenodo-Dep="6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="5" pageNumber="226" startId="5.[151,264,1909,1933]" targetBox="[243,1345,179,1882]" targetPageId="5">
<paragraph id="8BC436CDFFE79957FBE2F886C31BF81F" blockId="5.[151,1437,1909,2041]" pageId="5" pageNumber="226">
<emphasis id="B90FEADFFFE79957FBE2F886C76AF86B" bold="true" box="[151,307,1909,1933]" pageId="5" pageNumber="226">FIGURES 16</emphasis>
.
<taxonomicName id="4C7B4D4EFFE79957FA4AF886C445F86B" box="[319,540,1909,1933]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="5" pageNumber="226" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFE79957FA4AF886C445F86B" box="[319,540,1909,1933]" italics="true" pageId="5" pageNumber="226">Filopalpus joschmidti</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFE79957F955F886C429F86B" bold="true" box="[544,624,1909,1933]" pageId="5" pageNumber="226">
<taxonomicNameLabel id="A23C57A4FFE79957F955F886C429F86B" box="[544,624,1909,1933]" pageId="5" pageNumber="226" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top. A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26).
</paragraph>
</caption>
<caption id="DF046645FFE49954FBE2F9FFC419F9AE" ID-DOI="http://doi.org/10.5281/zenodo.6779040" ID-Zenodo-Dep="6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="6" pageNumber="227" startId="6.[151,264,1548,1572]" targetBox="[167,1418,197,1517]" targetPageId="6">
<paragraph id="8BC436CDFFE49954FBE2F9FFC419F9AE" blockId="6.[151,1436,1548,1609]" pageId="6" pageNumber="227">
<emphasis id="B90FEADFFFE49954FBE2F9FFC719F9C2" bold="true" box="[151,320,1548,1572]" pageId="6" pageNumber="227">FIGURES 710</emphasis>
.
<taxonomicName id="4C7B4D4EFFE49954FA39F9FFC470F9C2" box="[332,553,1548,1572]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="6" pageNumber="227" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFE49954FA39F9FFC470F9C2" box="[332,553,1548,1572]" italics="true" pageId="6" pageNumber="227">Filopalpus joschmidti</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFE49954F95BF9FEC427F9C3" bold="true" box="[558,638,1549,1573]" pageId="6" pageNumber="227">
<taxonomicNameLabel id="A23C57A4FFE49954F95BF9FEC427F9C3" box="[558,638,1549,1573]" pageId="6" pageNumber="227" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm.
</paragraph>
</caption>
<paragraph id="8BC436CDFFE49954FBB2F982C402F911" blockId="6.[151,1437,1649,2036]" pageId="6" pageNumber="227">
<emphasis id="B90FEADFFFE49954FBB2F982C7CAF96D" box="[199,403,1649,1675]" italics="true" pageId="6" pageNumber="227">Body, ventral side</emphasis>
(
<figureCitation id="13402A48FFE49954FAD1F982C7B2F96D" box="[420,491,1649,1675]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="6" pageNumber="227">Fig. 9</figureCitation>
): Cx irregularly covered all over with large pointed tubercles of various forms (not markedly polygonal), most of them with minute seta on top. Op gen light yellowish in front, the five distal sternites dark brownish each, with a row of tubercles, longest and massive on lateral sides, the large sternite and Op anale light with low tubercles densely packed.
</paragraph>
<paragraph id="8BC436CDFFE49954FBB2F8F1C2D3F8D9" blockId="6.[151,1437,1649,2036]" pageId="6" pageNumber="227">
<emphasis id="B90FEADFFFE49954FBB2F8F1C6A5F8FD" box="[199,252,1794,1819]" italics="true" pageId="6" pageNumber="227">Legs</emphasis>
: All articles light brownish to blackish. Tr smooth except for few minute tubercles, the other articles irregularly covered with low tubercles, all pointed with minute seta (except articles of Ta).
</paragraph>
<paragraph id="8BC436CDFFE49954FBB2F8BAC5E9F815" blockId="6.[151,1437,1649,2036]" pageId="6" pageNumber="227">
<emphasis id="B90FEADFFFE49954FBB2F8BAC775F885" box="[199,300,1865,1891]" italics="true" pageId="6" pageNumber="227">Pedipalp</emphasis>
(
<figureCitation id="13402A48FFE49954FA4EF8BAC7C6F885" box="[315,415,1865,1891]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="6" pageNumber="227">Figs 12</figureCitation>
,
<figureCitation id="13402A48FFE49954FADEF8BAC790F885" box="[427,457,1865,1891]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="6" pageNumber="227">10</figureCitation>
,
<figureCitation id="13402A48FFE49954FAA1F8BAC446F885" box="[468,543,1865,1891]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="6" pageNumber="227">2123</figureCitation>
): Extremely long, thin and filiform, all articles slenderer than those of legs IIV, Fe and Pt of nearly equal length (Pt slightly shorter), four times longer than body, Fe and Pt slightly broadened close to distal end (
<figureCitation id="13402A48FFE49954FA5BF862C7D8F84D" box="[302,385,1937,1963]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="6" pageNumber="227">Fig. 22</figureCitation>
), Ti extremely short, parallel-sided, Ta shortest article, similar to Ti but fusiform (
<figureCitation id="13402A48FFE49954FE7BF862C3C8F84D" box="[1294,1425,1937,1963]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="6" pageNumber="227">Figs 2223</figureCitation>
). All palpal articles devoid of any strong and elevated Tu, few fine light setae on all articles, coarse surface on Ti and Ta. One light pseudo-articulation each on Fe distally and on Pt basally.
</paragraph>
<caption id="DF046645FFE59955FBE2F8A1C45EF81C" ID-DOI="http://doi.org/10.5281/zenodo.6779044" ID-Zenodo-Dep="6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="7" pageNumber="228" startId="7.[151,264,1874,1898]" targetBox="[208,1379,192,1838]" targetPageId="7">
<paragraph id="8BC436CDFFE59955FBE2F8A1C45EF81C" blockId="7.[151,1437,1874,2043]" pageId="7" pageNumber="228">
<emphasis id="B90FEADFFFE59955FBE2F8A1C70BF88D" bold="true" box="[151,338,1874,1899]" pageId="7" pageNumber="228">FIGURES 1124.</emphasis>
<taxonomicName id="4C7B4D4EFFE59955FA22F8A0C46DF88D" box="[343,564,1875,1899]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="7" pageNumber="228" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFE59955FA22F8A0C46DF88D" box="[343,564,1875,1899]" italics="true" pageId="7" pageNumber="228">Filopalpus joschmidti</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFE59955F94FF8A0C4D0F88D" bold="true" box="[570,649,1875,1899]" pageId="7" pageNumber="228">
<taxonomicNameLabel id="A23C57A4FFE59955F94FF8A0C4D0F88D" box="[570,649,1875,1899]" pageId="7" pageNumber="228" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm.
</paragraph>
</caption>
<paragraph id="8BC436CDFFEA995AFBB2FF64C51FFEFB" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FF64C76CFF57" box="[199,309,151,177]" italics="true" pageId="8" pageNumber="229">Chelicera</emphasis>
(
<figureCitation id="13402A48FFEA995AFA30FF64C790FF57" box="[325,457,151,177]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="8" pageNumber="229">Figs 1617</figureCitation>
): Basal article slender, dorsally strongly concave at mid portion, indistinctly on ventral side, few strong tubercles dorso- and ventro-distally with minute seta on top, 2
<superScript id="7C0E9B85FFEA995AF885FF49C259FF2E" attach="left" box="[1008,1024,186,200]" fontSize="6" pageId="8" pageNumber="229">nd</superScript>
article moderately inflated, fusiform, single strong setae frontally and distally close to insertion 3
<superScript id="7C0E9B85FFEA995AF85EFF2DC560FF0A" attach="left" box="[811,825,222,236]" fontSize="6" pageId="8" pageNumber="229">rd</superScript>
article, a strong elongated cone-shaped apophysis near mid-half on pro-lateral side close to frontal side of 2
<superScript id="7C0E9B85FFEA995AF994FEF1C4A8FEF6" attach="left" box="[737,753,258,272]" fontSize="6" pageId="8" pageNumber="229">nd</superScript>
article.
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FED4C247FE37" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FED4C7CCFEA7" box="[199,405,295,321]" italics="true" pageId="8" pageNumber="229">Penis morphology</emphasis>
(
<figureCitation id="13402A48FFEA995AFAD3FED4C473FEA7" box="[422,554,294,321]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="8" pageNumber="229">Figs 1115</figureCitation>
): Truncus penis slender (la and do/ve views) from the small slightly enlarged base continuously enlarged towards distal end, but more markedly in distal forth, markedly broadened and rounded shortly before and along the movable glans (do/ve view), disto-frontal rim of truncus slightly invaginated. Equipment of penis with minute setae inconspicuous and slightly asymmetrical (
<figureCitation id="13402A48FFEA995AF8D1FE60C270FE4B" box="[932,1065,402,429]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="8" pageNumber="229">Figs 1315</figureCitation>
): two on or close to distal rim of truncus, three or five on lateral sides, all in the enlarged glans section of truncus.
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FE29C457FDDB" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FE29C702FE12" bold="true" box="[199,347,474,500]" pageId="8" pageNumber="229">Description:</emphasis>
<emphasis id="B90FEADFFFEA995AFA10FE28C789FE13" box="[357,464,475,501]" italics="true" pageId="8" pageNumber="229">FEMALE</emphasis>
. Characters of external morphology generally like in male including coloration and equipment of body with tubercles, leg length and leg armament, but apophyses on rear end of opisthosoma and on free tergites shorter (
<figureCitation id="13402A48FFEA995AFA0AFDD0C791FDDB" box="[383,456,547,573]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="8" pageNumber="229">Figs 4</figureCitation>
,
<figureCitation id="13402A48FFEA995AFAA1FDD0C458FDDB" box="[468,513,547,573]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="8" pageNumber="229">78</figureCitation>
).
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FDB4C4E1FD63" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FDB4C76CFD87" box="[199,309,583,609]" italics="true" pageId="8" pageNumber="229">Chelicera</emphasis>
(
<figureCitation id="13402A48FFEA995AFA30FDB4C791FD87" box="[325,456,583,609]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="8" pageNumber="229">Figs 1819</figureCitation>
): Generally as in male with similar armament, ve concavity of 1
<superScript id="7C0E9B85FFEA995AFFEDFDB4C2FAFDB3" attach="left" box="[1176,1187,583,597]" fontSize="6" pageId="8" pageNumber="229">st</superScript>
article even less than in male, no cone-shaped apophysis on 2
<superScript id="7C0E9B85FFEA995AF926FD99C43AFD9E" attach="left" box="[595,611,618,632]" fontSize="6" pageId="8" pageNumber="229">nd</superScript>
article.
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FD7CC5A4FC67" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FD7CC775FD4F" box="[199,300,655,681]" italics="true" pageId="8" pageNumber="229">Pedipalp</emphasis>
(
<figureCitation id="13402A48FFEA995AFA4EFD7CC7F8FD4F" box="[315,417,655,681]" captionStart="FIGURES 710" captionStartId="6.[151,264,1548,1572]" captionTargetBox="[167,1418,197,1517]" captionTargetId="figure-30@6.[151,1436,181,1524]" captionTargetPageId="6" captionText="FIGURES 710. Filopalpus joschmidti sp. nov. 78. Female paratype; 910. Male holotype. Body dorsal view (8), lateral view (7, 10) and ventral view (9). Scale: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.6779040" httpUri="https://zenodo.org/record/6779040/files/figure.png" pageId="8" pageNumber="229">Figs 78</figureCitation>
,
<figureCitation id="13402A48FFEA995AFAD8FD7CC797FD4F" box="[429,462,655,681]" captionStart="FIGURES 1124" captionStartId="7.[151,264,1874,1898]" captionTargetBox="[208,1379,192,1838]" captionTargetId="figure-15@7.[195,1392,181,1850]" captionTargetPageId="7" captionText="FIGURES 1124. Filopalpus joschmidti sp. nov. Male holotype (1117; 2123) and female paratype (1820; 24). 1112. Penis dorsal and lateral views. 1315. Glans dorsal, ventral and lateral views. 1619. Chelicera pro-lateral and retro-lateral views. 20. Pedipalpus retro-lateral view. 21. Pedipalpal femur-patella joint, retro-lateral view male. 22. Pedipalp retro-lateral view. 23. Pedipalpal tibia and tarsus. 24. Ovipositor distal part. Scales: 1112: 0.3 mm, 1319: 0.1 mm, 20: 0.5 mm, 21: 0.13 mm, 22: 0.25 mm, 23: 0.5 mm, 24: 0.05 mm." figureDoi="http://doi.org/10.5281/zenodo.6779044" httpUri="https://zenodo.org/record/6779044/files/figure.png" pageId="8" pageNumber="229">20</figureCitation>
): Very different from male, extremely short (total length slightly less than body length, Ti and Ta of about equal length, Pt slightly larger). Tr ventrally enlarged, one ve apophysis with strong seta on top; Fe smooth, nearly parallel-sided, slightly enlarged distally, with few scattered setae on dorsal side, ventrally one tubercle each on near base and amidst the article; Pt slightly curved to ventral side and enlarged distally, few scattered minute setae; Ti as long as Pt, equipped with five elevated tubercle of different sizes ventrally, the central one with a large immovable setae on top; Ta with three low seta-topped apophyses ventrally and few small additional setae interspersed, those also on do side; claw strong, slightly shorter than Ta.
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FC78C489FC43" blockId="8.[151,1437,151,1689]" box="[199,720,907,933]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FC78C72FFC43" bold="true" box="[199,374,907,933]" pageId="8" pageNumber="229">Measurements</emphasis>
(male, female in parentheses):
</paragraph>
<paragraph id="8BC436CDFFEA995AFB82FC5CC6A7FBF7" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFB82FC5CC7DDFC2F" box="[247,388,943,969]" italics="true" pageId="8" pageNumber="229">Body length</emphasis>
: Including pointed tubercles on frontal and posterior rim of scutum; male
<typeStatus id="54C0886FFFEA995AFE65FC5CC32CFC2F" box="[1296,1397,943,969]" pageId="8" pageNumber="229" type="holotype">holotype</typeStatus>
, in parentheses without apophysis, but including free tergites: 3.0 (2.7),
<typeStatus id="54C0886FFFEA995AF8FBFC20C5B7FC0B" box="[910,1006,979,1005]" pageId="8" pageNumber="229" type="paratype">paratype</typeStatus>
male 3.1 (2.8);
<typeStatus id="54C0886FFFEA995AFFE9FC20C2A5FC0B" box="[1180,1276,979,1005]" pageId="8" pageNumber="229" type="paratype">paratype</typeStatus>
females twice 3.1 (2.8).
</paragraph>
<paragraph id="8BC436CDFFEA995AFB82FBE8C314FBD3" blockId="8.[151,1437,151,1689]" box="[247,1357,1050,1077]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFB82FBE8C762FBD2" box="[247,315,1051,1076]" italics="true" pageId="8" pageNumber="229">Leg II</emphasis>
: Tr 0.3 (0.3), Fe 2.2 (1.8), Pt 0.5 (0.5), Ti 1.6 (1.4), Mt 1.5 (1.4), Ta 1.4 (1.2); total: 7.5 (6.6).
</paragraph>
<paragraph id="8BC436CDFFEA995AFB82FBCCC4A5FB9B" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFB82FBCCC705FBBF" box="[247,348,1087,1113]" italics="true" pageId="8" pageNumber="229">Pedipalp</emphasis>
: Tr 0.3 (0.2), Fe 5.4 (0.75), Pt 5.2 (0.45), Ti 0.9 (0.4), Ta 0.6 (0.4), claw 0.3 (0.3); total: 12.7 (2.5). Male pedipalp 5.08 times longer than female pedipalp.
</paragraph>
<paragraph id="8BC436CDFFEA995AFB82FB74C3D0FB47" blockId="8.[151,1437,151,1689]" box="[247,1417,1158,1185]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFB82FB74C4D9FB46" box="[247,640,1159,1185]" italics="true" pageId="8" pageNumber="229">Number of tarsomeres on legs IIV</emphasis>
(right and left leg): I 4/4 (4/4), II 7/7 (6/7), III 5/5 (5/5), IV 6/6 (5/6).
</paragraph>
<paragraph id="8BC436CDFFEA995AFB82FB58C7E2FB23" blockId="8.[151,1437,151,1689]" box="[247,443,1195,1221]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFB82FB58C7DBFB23" box="[247,386,1195,1221]" italics="true" pageId="8" pageNumber="229">Penis length</emphasis>
: 1.2.
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FB3DC74BFAEB" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FB3DC718FB0E" bold="true" box="[199,321,1230,1256]" pageId="8" pageNumber="229">Variation:</emphasis>
Size and number of apophyses on scutal area V and on free tergites may differ between individuals (
<figureCitation id="13402A48FFEA995AFBEAFB00C75DFAEB" box="[159,260,1267,1293]" captionStart="FIGURES 16" captionStartId="5.[151,264,1909,1933]" captionTargetBox="[243,1345,179,1882]" captionTargetId="figure-15@5.[243,1345,179,1882]" captionTargetPageId="5" captionText="FIGURES 16. Filopalpus joschmidti sp. nov. Body dorsal view. 13, 5. Male; 4, 6 female. 12. Left pedipalp stretched out and surpassing length of leg II; 1. and 23. represent different males (note different equipment of apophyses of rear end of opisthosoma). 4. Female with short pedipalp; 56. Distal part of prosoma; note cover of hedgehog-like tubercles with seta on top.A and b in Figs 1 and 2 indicate the pedipalp (a) and leg II (b). Photographs by Jean Severin (1) and Joachim Schmidt (26)." figureDoi="http://doi.org/10.5281/zenodo.6779036" httpUri="https://zenodo.org/record/6779036/files/figure.png" pageId="8" pageNumber="229">Figs 12</figureCitation>
).
</paragraph>
<paragraph id="8BC436CDFFEA995AFBB2FAE5C79FFA27" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FAE5C72BFAD6" bold="true" box="[199,370,1302,1328]" pageId="8" pageNumber="229">Relationships:</emphasis>
<taxonomicName id="4C7B4D4EFFEA995AFA0FFAE4C453FAD7" box="[378,522,1303,1329]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFEA995AFA0FFAE4C453FAD7" box="[378,522,1303,1329]" italics="true" pageId="8" pageNumber="229">F. joschmidti</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFEA995AF964FAE4C432FAD7" bold="true" box="[529,619,1303,1329]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="A23C57A4FFEA995AF964FAE4C432FAD7" box="[529,619,1303,1329]" pageId="8" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C7B4D4EFFEA995AF849FAE4C59CFAD7" box="[828,965,1303,1329]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="kakaensis" status="sp. nov.">
<emphasis id="B90FEADFFFEA995AF849FAE4C59CFAD7" box="[828,965,1303,1329]" italics="true" pageId="8" pageNumber="229">F. kakaensis</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFEA995AF8B9FAE4C27FFAD7" bold="true" box="[972,1062,1303,1329]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="A23C57A4FFEA995AF8B9FAE4C27FFAD7" box="[972,1062,1303,1329]" pageId="8" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, especially in its male pedipalpal proportions, also the lack of elevated Tu on all pedipalp articles.
<taxonomicName id="4C7B4D4EFFEA995AF8F6FAC8C257FAB3" box="[899,1038,1339,1365]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="kakaensis" status="sp. nov.">
<emphasis id="B90FEADFFFEA995AF8F6FAC8C257FAB3" box="[899,1038,1339,1365]" italics="true" pageId="8" pageNumber="229">F. kakaensis</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFEA995AFF62FAC8C22BFAB3" bold="true" box="[1047,1138,1339,1365]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="A23C57A4FFEA995AFF62FAC8C22BFAB3" box="[1047,1138,1339,1365]" pageId="8" pageNumber="229" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is smaller, has a different genital morphology (cavity on penis dorso-distal) and all minute apophyses on ventral and dorsal side of body are distally rounded, stump. The collecting localities of both filopaplid species are elevationally separated by about slightly more than
<quantity id="4C839B28FFEA995AFA1FFA54C79BFA27" box="[362,450,1447,1473]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="229" unit="m" value="1000.0">1000 m</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C3616546FFEA995AFBB2FA39C592F97F" pageId="8" pageNumber="229" type="distribution">
<paragraph id="8BC436CDFFEA995AFBB2FA39C592F97F" blockId="8.[151,1437,151,1689]" pageId="8" pageNumber="229">
<emphasis id="B90FEADFFFEA995AFBB2FA39C7ADFA02" bold="true" box="[199,500,1482,1509]" pageId="8" pageNumber="229">Distribution and habitat:</emphasis>
Up to present
<taxonomicName id="4C7B4D4EFFEA995AF9EFFA38C573FA03" box="[666,810,1483,1509]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="joschmidti" status="sp. nov.">
<emphasis id="B90FEADFFFEA995AF9EFFA38C573FA03" box="[666,810,1483,1509]" italics="true" pageId="8" pageNumber="229">F. joschmidti</emphasis>
</taxonomicName>
<emphasis id="B90FEADFFFEA995AF844FA38C528FA03" bold="true" box="[817,881,1483,1509]" pageId="8" pageNumber="229">
<taxonomicNameLabel id="A23C57A4FFEA995AF844FA38C528FA03" box="[817,881,1483,1509]" pageId="8" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
is only known from the northern face of the Bale Mts between Adaba and Dinsho at
<quantity id="4C839B28FFEA995AF969FA1CC42AF9EF" box="[540,627,1519,1545]" metricMagnitude="3" metricUnit="m" metricValue="3.63" pageId="8" pageNumber="229" unit="m" value="3630.0">3630 m</quantity>
. The specimens have been found under stones on a brook side shadowed by single
<taxonomicName id="4C7B4D4EFFEA995AFA73F9E0C7BDF9CB" box="[262,484,1555,1581]" class="Magnoliopsida" family="Rosaceae" genus="Hagenia" kingdom="Plantae" order="Rosales" pageId="8" pageNumber="229" phylum="Tracheophyta" rank="species" species="abyssinica">
<emphasis id="B90FEADFFFEA995AFA73F9E0C7BDF9CB" box="[262,484,1555,1581]" italics="true" pageId="8" pageNumber="229">Hagenia abyssinica</emphasis>
</taxonomicName>
trees and
<taxonomicName id="4C7B4D4EFFEA995AF92FF9E0C4A0F9CB" box="[602,761,1555,1581]" class="Arachnida" family="Salticidae" genus="Erica" kingdom="Animalia" order="Araneae" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="arborea">
<emphasis id="B90FEADFFFEA995AF92FF9E0C4A0F9CB" box="[602,761,1555,1581]" italics="true" pageId="8" pageNumber="229">Erica arborea</emphasis>
</taxonomicName>
shrubs within a canyon on a steep south-exposed mountain slope. The mountainous area beyond this canyon is widely deforested. Remains of the Sebsebe Washia Forest, which is famous in
<collectingCountry id="F36C765DFFEA995AFA1AF9A8C788F993" box="[367,465,1627,1653]" name="Ethiopia" pageId="8" pageNumber="229">Ethiopia</collectingCountry>
, existed in 2019 only within the lower parts of the main valley up to an altitude of about
<quantity id="4C839B28FFEA995AFBE2F98CC6B7F97F" box="[151,238,1663,1689]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="8" pageNumber="229" unit="m" value="3200.0">3200 m</quantity>
, much lower than the localities of the present
<taxonomicName id="4C7B4D4EFFEA995AF987F98CC592F97F" box="[754,971,1663,1689]" class="Arachnida" family="Assamiidae" genus="Filopalpus" kingdom="Animalia" order="Opiliones" pageId="8" pageNumber="229" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B90FEADFFFEA995AF987F98CC532F97F" box="[754,875,1663,1689]" italics="true" pageId="8" pageNumber="229">Filopalpus</emphasis>
species.
</taxonomicName>
</paragraph>
</subSubSection>
</treatment>
</document>