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<document id="5B565A7341492251C967CC78CCD3E134" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFFA7AF0ED1DBDD6BFE09120A" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Degallieridium Vaz-De-Mello 2008, new genus" docType="treatment" docVersion="3" lastPageNumber="21" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="20" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="DDFC70B7ABA2014CAB44820FEE43C5E7">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
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<mods:namePart id="272FDCD7CDE116B26E35F1E91418BDC4">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date id="830CCA2F9857ACAC918D8FE161D75A22">2008</mods:date>
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<mods:number id="76757219905458BE7580ED68D212FD72">2008-12-05</mods:number>
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<treatment id="03D287CEFFA7AF0ED1DBDD6BFE09120A" ID-DOI="http://doi.org/10.5281/zenodo.5241989" ID-Zenodo-Dep="5241989" LSID="urn:lsid:plazi:treatment:03D287CEFFA7AF0ED1DBDD6BFE09120A" httpUri="http://treatment.plazi.org/id/03D287CEFFA7AF0ED1DBDD6BFE09120A" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<subSubSection id="C3616553FFA7AF09D1DBDD6BFE541161" box="[151,498,543,569]" pageId="19" pageNumber="20" type="nomenclature">
<paragraph id="8BC436D8FFA7AF09D1DBDD6BFE541161" blockId="19.[151,498,543,569]" box="[151,498,543,569]" pageId="19" pageNumber="20">
<heading id="D08C81B4FFA7AF09D1DBDD6BFE541161" bold="true" box="[151,498,543,569]" fontSize="11" level="1" pageId="19" pageNumber="20" reason="1">
<emphasis id="B90FEACAFFA7AF09D1DBDD6BFE541161" bold="true" box="[151,498,543,569]" pageId="19" pageNumber="20">
3.
<taxonomicName id="4C7B4D5BFFA7AF09D1F8DD6BFECC1161" ID-CoL="42GC" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[180,362,543,569]" class="Insecta" family="Scarabaeidae" genus="Degallieridium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEACAFFA7AF09D1F8DD6BFECC1161" bold="true" box="[180,362,543,569]" italics="true" pageId="19" pageNumber="20">Degallieridium</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57B1FFA7AF09D03DDD6BFE541161" box="[369,498,543,569]" pageId="19" pageNumber="20" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFA7AF09D1DBDD1EFBB4107C" pageId="19" pageNumber="20" type="diagnosis">
<paragraph id="8BC436D8FFA7AF09D1DBDD1EFBB4107C" blockId="19.[151,1436,618,2004]" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D1DBDD1EFEBF11DC" bold="true" box="[151,281,618,644]" pageId="19" pageNumber="20">Diagnosis:</emphasis>
Body short, oval, convex (
<figureCitation id="13402A5DFFA7AF09D313DD1EFD1011DC" box="[607,694,618,644]" captionStart="FIGURES 1522" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 1522. Dorsal views. 1516: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 2122: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="19" pageNumber="20">Fig. 18</figureCitation>
) and very small size (length 2.0 mm or less). Dorsum brown; dorsal setae thick, as wide as lateral elytral striae. Clypeus with two teeth, laterally rounded; eyes dorsally as long as twice their width (
<figureCitation id="13402A5DFFA7AF09D08BDDCEFDB9118C" box="[455,543,698,724]" captionStart="FIGURES 4754" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 4754. 4748: Degallieridium lilliputanum: 47. head; 48. parameres; 4951: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 5254. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="19" pageNumber="20">Fig. 47</figureCitation>
). Hypomeron without lateral longitudinal carina. Elytral interstriae with two rows of punctures, only one of them bearing setae. Pseudoepipleuron medially inflexed, but gradually narrowed posteriorly (
<figureCitation id="13402A5DFFA7AF09D03ADC7EFE6A107C" box="[374,460,778,804]" captionStart="FIGURES 511" captionStartId="54.[151,269,1929,1953]" captionTargetBox="[223,1320,197,1895]" captionTargetId="figure-60@54.[203,1372,197,1917]" captionTargetPageId="54" captionText="FIGURES 511. Left epipleuron and pseudoepipleuron, ventral view. 5. Trichillum externepunctatum; 6. Onoreidium cristatum; 7. Scatrichus bicarinatus; 8. Pedaridium hirsutum; 9. Silvinha unica; 10. Besourenga minutus; 11. Degallieridium lilliputanum. Scale = 1.0 mm." pageId="19" pageNumber="20">Fig. 11</figureCitation>
). Parameres strongly divergent apically (
<figureCitation id="13402A5DFFA7AF09D2E0DC7EFBA2107C" box="[940,1028,778,804]" captionStart="FIGURES 4754" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 4754. 4748: Degallieridium lilliputanum: 47. head; 48. parameres; 4951: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 5254. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="19" pageNumber="20">Fig. 48</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFA7AF09D18ADC46FF5C1534" pageId="19" pageNumber="20" type="description">
<paragraph id="8BC436D8FFA7AF09D18ADC46FB1716AC" blockId="19.[151,1436,618,2004]" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D18ADC46FEC61014" bold="true" box="[198,352,818,844]" pageId="19" pageNumber="20">Description:</emphasis>
Length 1.82.0 mm; body short (
<figureCitation id="13402A5DFFA7AF09D24CDC46FCFF1014" box="[768,857,818,844]" captionStart="FIGURES 1522" captionStartId="56.[151,269,1962,1986]" captionTargetBox="[207,1374,197,1940]" captionTargetId="figure-57@56.[207,1374,197,1940]" captionTargetPageId="56" captionText="FIGURES 1522. Dorsal views. 1516: Besourenga: 15. B. sp.; 16. B. horacioi; 17. Bradypodidium sp.; 18. Degallieridium lilliputanum; 19. Eutrichillum sp.; 20. Feeridium woodruffi; 2122: Genieridium: 21. G. bordoni; 22. G. cryptops." pageId="19" pageNumber="20">Fig. 18</figureCitation>
), oval, bright brown, with no traces of metallic sheen. Clypeus with two triangular teeth separated by V-shaped emargination, laterally simply rounded and continuous with genal border. Clypeofrontal and clypeogenal sutures indistinct. Eyes, in dorsal view, twice as long as wide; interocular width four times eye width (
<figureCitation id="13402A5DFFA7AF09D258DCDEFCCB109C" box="[788,877,938,964]" captionStart="FIGURES 4754" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 4754. 4748: Degallieridium lilliputanum: 47. head; 48. parameres; 4951: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 5254. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="19" pageNumber="20">Fig. 47</figureCitation>
). Pronotum lacking anterior or posterior beads; lateral callosity present, elongated and darker than remaining parts of pronotum; disc centrally with large setose punctures. Pronotum and hypomeron separated by simple complete carina, with its posterior two-thirds placed ventrally (anterior third lateral). Hypomeron laterally lacking longitudinal carina; mesoepimeron anteriorly lacking transverse carina. Metasternum with small glabrous punctures on the disc. Elytral interstriae with two rows of punctures on disc, inner one bearing setae; interstriae flat on disc, apically convex; striae apically clearly wider and deeper than on disc. Pseudoepipleura medially inflexed (posteriorly almost in a transverse plane in relation to anterior part), but without sharp angle and gradually narrowed posteriorly (
<figureCitation id="13402A5DFFA7AF09D420DBB6FF13165C" captionStart="FIGURES 511" captionStartId="54.[151,269,1929,1953]" captionTargetBox="[223,1320,197,1895]" captionTargetId="figure-60@54.[203,1372,197,1917]" captionTargetPageId="54" captionText="FIGURES 511. Left epipleuron and pseudoepipleuron, ventral view. 5. Trichillum externepunctatum; 6. Onoreidium cristatum; 7. Scatrichus bicarinatus; 8. Pedaridium hirsutum; 9. Silvinha unica; 10. Besourenga minutus; 11. Degallieridium lilliputanum. Scale = 1.0 mm." pageId="19" pageNumber="20">Fig. 11</figureCitation>
), completely glabrous. Protibiae laterally with three strong teeth distributed along apical seven-tenths, not denticulate basally, and ventrally lacking scale-like setae on teeth. Width of apices of mesotibiae and metatibiae about one-third length of tibia. Length of parameres shorter than one-half that of phallobase, right angled in relation to phallobase axis, apically flattened and strongly divergent with sharp lateral apical angles (
<figureCitation id="13402A5DFFA7AF09D420DA16FF1E16FC" captionStart="FIGURES 4754" captionStartId="60.[151,269,1940,1964]" captionTargetBox="[221,1365,194,1911]" captionTargetId="figure-53@60.[221,1369,194,1915]" captionTargetPageId="60" captionText="FIGURES 4754. 4748: Degallieridium lilliputanum: 47. head; 48. parameres; 4951: Eutrichillum hirsutum: 49. head; 50. parameres; 51. ♂ protarsus; 5254. Feeridium woodruffi: 52. head; 53. parameres; 54. ventral view of metalegs and abdomen." pageId="19" pageNumber="20">Fig. 48</figureCitation>
). Internal sac with a long helicoidal pseudoflagellum occupying most of phallobase length, and secondary pseudoflagellum flattened all along. Coxites small and well defined, glabrous. Spermatheca C-shaped, strongly narrowed apically, basally somewhat bulbous; spermathecal duct not sclerotized.
</paragraph>
<paragraph id="8BC436D8FFA7AF09D18AD976FF5C1534" blockId="19.[151,1436,618,2004]" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D18AD976FE631544" bold="true" box="[198,453,1538,1564]" pageId="19" pageNumber="20">Sexual dimorphism:</emphasis>
Males have proclaws strongly bent in the middle with last tarsomere hollowed to receive them, an apical mesal tooth in protibia directed downward, and less transverse pygidium compared to females.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFA7AF09D18AD90EFC4B15CC" box="[198,1005,1658,1684]" pageId="19" pageNumber="20" type="type_taxon">
<paragraph id="8BC436D8FFA7AF09D18AD90EFC4B15CC" blockId="19.[151,1436,618,2004]" box="[198,1005,1658,1684]" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D18AD90EFECC15CC" bold="true" box="[198,362,1658,1684]" pageId="19" pageNumber="20">
<typeStatus id="54C0887AFFA7AF09D18AD90EFEA515CC" box="[198,259,1658,1684]" pageId="19" pageNumber="20">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7B4D5BFFA7AF09D03ED90FFD1D15CC" ID-CoL="6CGLC" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[370,699,1659,1684]" class="Insecta" family="Scarabaeidae" genus="Degallieridium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="lilliputanum" status="sp. nov.">
<emphasis id="B90FEACAFFA7AF09D03ED90FFD1D15CC" box="[370,699,1659,1684]" italics="true" pageId="19" pageNumber="20">Degallieridium lilliputanum</emphasis>
</taxonomicName>
<emphasis id="B90FEACAFFA7AF09D38FD90EFCF515CC" bold="true" box="[707,851,1658,1684]" pageId="19" pageNumber="20">
<taxonomicNameLabel id="A23C57B1FFA7AF09D38FD90EFCF515CC" box="[707,851,1658,1684]" pageId="19" pageNumber="20" rank="species">new species</taxonomicNameLabel>
</emphasis>
(monotypy).
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFA7AF09D18AD9D6FDCA15BC" pageId="19" pageNumber="20" type="etymology">
<paragraph id="8BC436D8FFA7AF09D18AD9D6FDCA15BC" blockId="19.[151,1436,618,2004]" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D18AD9D6FEF115E4" bold="true" box="[198,343,1698,1724]" pageId="19" pageNumber="20">Etymology:</emphasis>
After Nicolas Degallier, friend, histeridologist and collector of part of the
<typeStatus id="54C0887AFFA7AF09D59DD9D6FAA515E4" box="[1233,1283,1698,1724]" pageId="19" pageNumber="20">type</typeStatus>
series of the species described below. Gender neuter.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFA7AF09D18AD986FC59146C" pageId="19" pageNumber="20" type="distribution">
<paragraph id="8BC436D8FFA7AF09D18AD986FC59146C" blockId="19.[151,1436,618,2004]" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D18AD986FECF1454" bold="true" box="[198,361,1778,1804]" pageId="19" pageNumber="20">Distribution:</emphasis>
Central
<collectingCountry id="F36C7648FFA7AF09D081D986FDB21454" box="[461,532,1778,1804]" name="Brazil" pageId="19" pageNumber="20">Brazil</collectingCountry>
(
<collectingRegion id="49BFF83AFFA7AF09D368D986FCA61454" box="[548,768,1778,1804]" country="Brazil" name="Minas Gerais" pageId="19" pageNumber="20">Minas Gerais state</collectingRegion>
and Distrito Federal) and
<collectingCountry id="F36C7648FFA7AF09D57FD986FB2A1454" box="[1075,1164,1778,1804]" name="Bolivia" pageId="19" pageNumber="20">Bolivia</collectingCountry>
, corresponding to scattered localities in Cerrado and Chaco provinces of the Chacoan subregion.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFA7AF0ED18AD836FE09120A" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20" type="discussion">
<paragraph id="8BC436D8FFA7AF0ED18AD836FC6D125A" blockId="19.[151,1436,618,2004]" lastBlockId="20.[151,1436,152,338]" lastPageId="20" lastPageNumber="21" pageId="19" pageNumber="20">
<emphasis id="B90FEACAFFA7AF09D18AD836FEE71404" bold="true" box="[198,321,1858,1884]" pageId="19" pageNumber="20">Remarks:</emphasis>
This genus appears closely related to
<taxonomicName id="4C7B4D5BFFA7AF09D27FD837FC0B1404" authorityName="Vaz-De-Mello" authorityYear="2008" box="[819,941,1859,1884]" class="Insecta" family="Scarabaeidae" genus="Feeridium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA7AF09D27FD837FC0B1404" box="[819,941,1859,1884]" italics="true" pageId="19" pageNumber="20">Feeridium</emphasis>
</taxonomicName>
, with which shares the absence of the hypomeral lateral longitudinal carina, pseudoepipleural longitudinal fold, and parameral and spermathecal general form. It is most easily distinguishable by size (
<taxonomicName id="4C7B4D5BFFA7AF09D26AD8E7FC0614F4" authorityName="Vaz-De-Mello" authorityYear="2008" box="[806,928,1939,1964]" class="Insecta" family="Scarabaeidae" genus="Feeridium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA7AF09D26AD8E7FC0614F4" box="[806,928,1939,1964]" italics="true" pageId="19" pageNumber="20">Feeridium</emphasis>
</taxonomicName>
is about five times longer) and body form (very elongated in
<taxonomicName id="4C7B4D5BFFA7AF09D038D8CFFE48148C" authorityName="Vaz-De-Mello" authorityYear="2008" box="[372,494,1979,2004]" class="Insecta" family="Scarabaeidae" genus="Feeridium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA7AF09D038D8CFFE48148C" box="[372,494,1979,2004]" italics="true" pageId="19" pageNumber="20">Feeridium</emphasis>
</taxonomicName>
), although many other differences do exist, for instance,
<taxonomicName id="4C7B4D5BFFA7AF09D5DCD8CFFAAC148C" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1168,1290,1979,2004]" class="Insecta" family="Scarabaeidae" genus="Feeridium" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA7AF09D5DCD8CFFAAC148C" box="[1168,1290,1979,2004]" italics="true" pageId="19" pageNumber="20">Feeridium</emphasis>
</taxonomicName>
lacks modification in male proclaws. Both genera belong to the clade that includes also
<emphasis id="B90FEACAFFA0AF0ED50CDFECFAC613E9" box="[1088,1376,152,177]" italics="true" pageId="20" pageNumber="21">
<taxonomicName id="4C7B4D5BFFA0AF0ED50CDFECFB1B13E9" box="[1088,1213,152,177]" class="Insecta" family="Scarabaeidae" genus="Trichillum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Trichillum</taxonomicName>
,
<taxonomicName id="4C7B4D5BFFA0AF0ED580DFECFAC613E9" authorityName="Martinez" authorityYear="1969" box="[1228,1376,152,177]" class="Insecta" family="Scarabaeidae" genus="Eutrichillum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">Eutrichillum</taxonomicName>
</emphasis>
, and
<taxonomicName id="4C7B4D5BFFA0AF0ED1DBDFB4FE841381" authorityName="Vaz-De-Mello" authorityYear="2008" box="[151,290,192,217]" class="Insecta" family="Scarabaeidae" genus="Besourenga" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA0AF0ED1DBDFB4FE841381" box="[151,290,192,217]" italics="true" pageId="20" pageNumber="21">Besourenga</emphasis>
</taxonomicName>
(see discussion above). A strong overall similarity exists between
<taxonomicName id="4C7B4D5BFFA0AF0ED57EDFB4FB431381" authorityName="Vaz-De-Mello" authorityYear="2008" box="[1074,1253,192,217]" class="Insecta" family="Scarabaeidae" genus="Degallieridium" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA0AF0ED57EDFB4FB431381" box="[1074,1253,192,217]" italics="true" pageId="20" pageNumber="21">Degallieridium</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7B4D5BFFA0AF0ED452DFB4FF651259" authorityName="Vaz-De-Mello" authorityYear="2008" class="Insecta" family="Scarabaeidae" genus="Leotrichillum" kingdom="Animalia" order="Coleoptera" pageId="20" pageNumber="21" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFFA0AF0ED452DFB4FF651259" italics="true" pageId="20" pageNumber="21">Leotrichillum</emphasis>
</taxonomicName>
, however relationships of the latter with other genera are obscure.
</paragraph>
<paragraph id="8BC436D8FFA0AF0ED18ADE7BFE09120A" blockId="20.[151,1436,152,338]" pageId="20" pageNumber="21">
<emphasis id="B90FEACAFFA0AF0ED18ADE7BFEC91271" bold="true" box="[198,367,271,297]" pageId="20" pageNumber="21">Composition:</emphasis>
Besides the
<typeStatus id="54C0887AFFA0AF0ED34BDE64FD9C1272" box="[519,570,272,298]" pageId="20" pageNumber="21">type</typeStatus>
species, the genus contains at least one new species currently in the process of being described.
</paragraph>
</subSubSection>
</treatment>
</document>