353 lines
44 KiB
XML
353 lines
44 KiB
XML
<document id="CF42E115DC917ADA22AF5F47763087AB" ID-CLB-Dataset="28394" ID-DOI="10.11646/zootaxa.4565.2.5" ID-GBIF-Dataset="dc72f6e5-06b2-4846-89ed-cc8a213f054e" ID-ISSN="1175-5326" ID-Zenodo-Dep="2589895" ID-ZooBank="F79BF0D3-BD46-409E-8BEE-E57B24A024F7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1552287587612" checkinUser="plazi" docAuthor="Campos, Rogério & Mariano, Rodolfo" docDate="2019" docId="03D287AFFFCBFFA4FF1AE4F8FB89FC65" docLanguage="en" docName="zootaxa.4565.2.5.pdf" docOrigin="Zootaxa 4565 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thraulodes catoles Campos & Mariano 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="217" masterDocId="FFEBFFD7FFC9FFA0FF8DE425FFF4FFAD" masterDocTitle="New species of Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Northeastern Brazil" masterLastPageNumber="222" masterPageNumber="213" pageNumber="215" updateTime="1698701475782" updateUser="ExternalLinkService">
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<mods:title id="C7CB24A16C5698C8CA2154FBD765B9D2">New species of Thraulodes Ulmer, 1920 (Ephemeroptera: Leptophlebiidae: Atalophlebiinae) from Northeastern Brazil</mods:title>
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<mods:name id="8BBE6FEA3D2E228BD9968D4859C74FDE" type="personal">
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<mods:namePart id="811D841B9F5AFB7D2F7CF2A0FC8FDD65">Campos, Rogério</mods:namePart>
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<mods:namePart id="A03E5724F9C01798670B5B141C1FE422">Mariano, Rodolfo</mods:namePart>
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<mods:typeOfResource id="9138AABE33D0E246D042DA46B6118E9B">text</mods:typeOfResource>
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<mods:title id="426F8CDF08C8328CD3F573D44AD5B4D1">Zootaxa</mods:title>
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<mods:part id="8BD0FAB9BE90E49A055A1E611CA891F4">
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<mods:date id="FC310487A60FD6403854AB118F2362AC">2019</mods:date>
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<mods:number id="89044E0F265CE10D11C2D537F71BAE84">2019-03-08</mods:number>
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<mods:number id="E63CA9D87122DFDBD40FC069A47EC036">4565</mods:number>
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<mods:identifier id="46D25CB7177298003596EC6F95BCF2ED" type="DOI">10.11646/zootaxa.4565.2.5</mods:identifier>
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<treatment id="03D287AFFFCBFFA4FF1AE4F8FB89FC65" ID-DOI="http://doi.org/10.5281/zenodo.5925457" ID-GBIF-Taxon="154366918" ID-Zenodo-Dep="5925457" LSID="urn:lsid:plazi:treatment:03D287AFFFCBFFA4FF1AE4F8FB89FC65" httpUri="http://treatment.plazi.org/id/03D287AFFFCBFFA4FF1AE4F8FB89FC65" lastPageId="4" lastPageNumber="217" pageId="2" pageNumber="215">
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<subSubSection id="C3616532FFCBFFA2FF1AE4F8FE14FF5A" box="[151,480,221,247]" pageId="2" pageNumber="215" type="nomenclature">
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<paragraph id="8BC436B9FFCBFFA2FF1AE4F8FE14FF5A" blockId="2.[151,480,221,280]" box="[151,480,221,247]" pageId="2" pageNumber="215">
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<heading id="D08C81D5FFCBFFA2FF1AE4F8FE14FF5A" bold="true" box="[151,480,221,247]" fontSize="11" level="1" pageId="2" pageNumber="215" reason="1">
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<emphasis id="B90FEAABFFCBFFA2FF1AE4F8FE14FF5A" bold="true" box="[151,480,221,247]" pageId="2" pageNumber="215">
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<taxonomicName id="4C7B4D3AFFCBFFA2FF1AE4F8FE8FFF5A" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[151,379,221,247]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
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<emphasis id="B90FEAABFFCBFFA2FF1AE4F8FE8FFF5A" bold="true" box="[151,379,221,247]" italics="true" pageId="2" pageNumber="215">Thraulodes catoles</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23C57D0FFCBFFA2FE0EE4F8FE14FF5A" box="[387,480,221,247]" pageId="2" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616532FFCBFFA2FF1AE4DAFED4FEB5" box="[151,288,255,280]" pageId="2" pageNumber="215" type="description">
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<paragraph id="8BC436B9FFCBFFA2FF1AE4DAFED4FEB5" blockId="2.[151,480,221,280]" box="[151,288,255,280]" pageId="2" pageNumber="215">
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(
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<figureCitation id="13402A3CFFCBFFA2FF12E4DAFEE3FEB5" box="[159,279,255,280]" captionStart-0="FIGURES 2–7" captionStart-1="FIGURES 8–12" captionStartId-0="2.[151,264,1958,1981]" captionStartId-1="3.[151,264,1983,2006]" captionTargetBox-0="[161,1425,934,1917]" captionTargetBox-1="[183,1392,415,1910]" captionTargetId-0="figure@2.[161,1425,934,1917]" captionTargetId-1="figure@3.[161,1426,373,1962]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." captionText-1="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri-0="https://zenodo.org/record/2589899/files/figure.png" httpUri-1="https://zenodo.org/record/2589901/files/figure.png" pageId="2" pageNumber="215">Figs. 2–12</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616532FFCBFFA2FF1AE562FD2BFDB9" pageId="2" pageNumber="215" type="diagnosis">
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<paragraph id="8BC436B9FFCBFFA2FF1AE562FD2BFDB9" blockId="2.[151,1437,327,856]" pageId="2" pageNumber="215">
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<emphasis id="B90FEAABFFCBFFA2FF1AE562FEE7FECD" bold="true" box="[151,275,327,352]" pageId="2" pageNumber="215">Diagnosis:</emphasis>
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<taxonomicName id="4C7B4D3AFFCBFFA2FE91E562FE07FECD" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[284,499,327,352]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
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<emphasis id="B90FEAABFFCBFFA2FE91E562FE07FECD" box="[284,499,327,352]" italics="true" pageId="2" pageNumber="215">Thraulodes catoles</emphasis>
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</taxonomicName>
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<emphasis id="B90FEAABFFCBFFA2FE71E562FDA3FECD" bold="true" box="[508,599,327,352]" pageId="2" pageNumber="215">
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<taxonomicNameLabel id="A23C57D0FFCBFFA2FE71E562FDA3FECD" box="[508,599,327,352]" pageId="2" pageNumber="215" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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imago can be separated from all other species by the following character combinations: 1) forewings with three crossveins basal to bulla (
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<figureCitation id="13402A3CFFCBFFA2FCFCE549FC42FE29" box="[881,950,364,389]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="2" pageNumber="215">Fig. 8</figureCitation>
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); 2) costal and subcostal areas of forewing translucent; 3) a reddish band on apex of forelegs femora (
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<figureCitation id="13402A3CFFCBFFA2FCA8E5AAFC9CFE05" box="[805,872,399,424]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="2" pageNumber="215">Fig. 3</figureCitation>
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); 4) abdominal color pattern reddish brown with a pair of triangular midway spots and two large submedian spots (
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<figureCitation id="13402A3CFFCBFFA2FCF8E591FC2DFE60" box="[885,985,436,461]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="2" pageNumber="215">Fig. 5–7</figureCitation>
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); 5) styliger plate with dorsal extension digitiform (
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<figureCitation id="13402A3CFFCBFFA2FE9AE5F2FE9EFE5D" box="[279,362,471,496]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="2" pageNumber="215">Fig. 10</figureCitation>
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); 6) penes with apicolateral area forming an “ear-like” projection, long spines on penes, lapel on inner margin and lateral pouch present (
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<figureCitation id="13402A3CFFCBFFA2FDD8E5D9FD26FDB9" box="[597,722,508,533]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="2" pageNumber="215">Fig. 11–12</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616532FFCBFFA4FF4AE63AFD70FE7D" lastPageId="4" lastPageNumber="217" pageId="2" pageNumber="215" type="description">
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<paragraph id="8BC436B9FFCBFFA2FF4AE63AFA72FD9A" blockId="2.[151,1437,327,856]" box="[199,1414,543,568]" pageId="2" pageNumber="215">
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<emphasis id="B90FEAABFFCBFFA2FF4AE63AFEAFFD95" bold="true" box="[199,347,543,568]" pageId="2" pageNumber="215">Male imago.</emphasis>
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Body length:
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<quantity id="4C839B5CFFCBFFA2FE77E63AFD54FD9A" box="[506,672,543,568]" metricMagnitude="-3" metricUnit="m" metricValue="6.35" metricValueMax="6.62" metricValueMin="6.08" pageId="2" pageNumber="215" unit="mm" value="6.35" valueMax="6.62" valueMin="6.08">6.08–6.62 mm</quantity>
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; forewing length:
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<quantity id="4C839B5CFFCBFFA2FCE1E63AFBE6FD9A" box="[876,1042,543,568]" metricMagnitude="-3" metricUnit="m" metricValue="6.785" metricValueMax="7.03" metricValueMin="6.54" pageId="2" pageNumber="215" unit="mm" value="6.785" valueMax="7.03" valueMin="6.54">6.54–7.03 mm</quantity>
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; hind wing length: 1.0–
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<quantity id="4C839B5CFFCBFFA2FA96E605FA76FD9A" box="[1307,1410,544,568]" metricMagnitude="-3" metricUnit="m" metricValue="1.34" pageId="2" pageNumber="215" unit="mm" value="1.34">1.34 mm</quantity>
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.
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</paragraph>
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<paragraph id="8BC436B9FFCBFFA2FF4AE661FBBEFDF0" blockId="2.[151,1437,327,856]" box="[199,1098,580,605]" pageId="2" pageNumber="215">
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General coloration (
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<figureCitation id="13402A3CFFCBFFA2FE2AE661FE19FDF1" box="[423,493,580,605]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="2" pageNumber="215">Fig. 2</figureCitation>
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): reddish translucent with black spots. Wings hyaline.
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</paragraph>
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<paragraph id="8BC436B9FFCBFFA2FF4AE642FC96FD08" blockId="2.[151,1437,327,856]" pageId="2" pageNumber="215">Head: dark reddish with black spots. Scape and pedicel dark brown, flagellum pale. Ocelli white surrounded by a black ring. Upper portion of eyes orange, lower portion black.</paragraph>
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<paragraph id="8BC436B9FFCBFFA2FF4AE68AFF41FCF5" blockId="2.[151,1437,327,856]" pageId="2" pageNumber="215">
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Thorax: brown with black spots. Pronotum with median line and hind margin dark brown. Mesonotum with anteronotal protuberance chestnut, washed red on lateral margin; median longitudinal suture brown, latero and parapsidial sutures yellow-chestnut. Metanotum with hind margin of metascutelum reddish. Pleurae chestnut with black spots and some red areas. Prosternum white; mesosternum washed in brown and furcasternum brown (
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<figureCitation id="13402A3CFFCBFFA2FAE2E739FF5CFCF5" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="2" pageNumber="215">Fig. 4</figureCitation>
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).
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</paragraph>
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<caption id="DF046631FFCBFFA2FF1AE383FE0DF876" httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="2" pageNumber="215" startId="2.[151,264,1958,1981]" targetBox="[161,1425,934,1917]" targetPageId="2">
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||
<paragraph id="8BC436B9FFCBFFA2FF1AE383FE0DF876" blockId="2.[151,1436,1958,2011]" pageId="2" pageNumber="215">
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||
<emphasis id="B90FEAABFFCBFFA2FF1AE383FEC0F811" bold="true" box="[151,308,1958,1981]" pageId="2" pageNumber="215">FIGURES 2–7</emphasis>
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.
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<taxonomicName id="4C7B4D3AFFCBFFA2FECFE382FDF0F810" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[322,516,1959,1981]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="2" pageNumber="215" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
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<emphasis id="B90FEAABFFCBFFA2FECFE382FDF0F810" box="[322,516,1959,1981]" italics="true" pageId="2" pageNumber="215">Thraulodes catoles</emphasis>
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</taxonomicName>
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||
<taxonomicNameLabel id="A23C57D0FFCBFFA2FD86E382FDA8F810" box="[523,604,1959,1981]" pageId="2" pageNumber="215" rank="species">
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||
<emphasis id="B90FEAABFFCBFFA2FD86E382FDD7F811" bold="true" box="[523,547,1959,1980]" pageId="2" pageNumber="215">sp</emphasis>
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||
.
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<emphasis id="B90FEAABFFCBFFA2FDBDE382FDA2F811" bold="true" box="[560,598,1959,1980]" pageId="2" pageNumber="215">nov</emphasis>
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.
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</taxonomicNameLabel>
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, male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation.
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</paragraph>
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</caption>
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<paragraph id="8BC436B9FFCAFFA3FF4AE4B2FB4AFF55" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="216">
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Wings (
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<figureCitation id="13402A3CFFCAFFA3FE93E4B2FE78FF1D" box="[286,396,151,176]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="3" pageNumber="216">Figs. 8–9</figureCitation>
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): membrane of the forewing hyaline with a costal brace dark brown (
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<figureCitation id="13402A3CFFCAFFA3FB1EE4B2FB2CFF1D" box="[1171,1240,151,176]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="3" pageNumber="216">Fig. 8</figureCitation>
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). Forewings with three cross veins basal to bullae and 7–10 cross veins distal to bullae. Hind wings hyaline with a brown stain extending on basal portion of subcostal to base of radial sector veins and 7 cross veins (
|
||
<figureCitation id="13402A3CFFCAFFA3FBE6E4FAFB44FF55" box="[1131,1200,223,248]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="3" pageNumber="216">Fig. 9</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436B9FFCAFFA3FF4AE520FC27FEED" blockId="3.[151,1436,151,320]" pageId="3" pageNumber="216">
|
||
Legs (
|
||
<figureCitation id="13402A3CFFCAFFA3FE80E521FEA7FEB1" box="[269,339,260,285]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="3" pageNumber="216">Fig. 3</figureCitation>
|
||
): all legs translucent yellow; fore and middle femora with red and brown bands on 2/3 of femur; femora of hind legs red with a brown band on apex. Claws yellow-brown.
|
||
</paragraph>
|
||
<caption id="DF046631FFCAFFA3FF1AE39AFAA8F858" httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="3" pageNumber="216" startId="3.[151,264,1983,2006]" targetBox="[183,1392,415,1910]" targetPageId="3">
|
||
<paragraph id="8BC436B9FFCAFFA3FF1AE39AFAA8F858" blockId="3.[151,1436,1983,2037]" pageId="3" pageNumber="216">
|
||
<emphasis id="B90FEAABFFCAFFA3FF1AE39AFECBF878" bold="true" box="[151,319,1983,2006]" pageId="3" pageNumber="216">FIGURES 8–12</emphasis>
|
||
.
|
||
<taxonomicName id="4C7B4D3AFFCAFFA3FEC1E3E5FDF8F87B" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[332,524,1984,2006]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="216" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
|
||
<emphasis id="B90FEAABFFCAFFA3FEC1E3E5FDF8F87B" box="[332,524,1984,2006]" italics="true" pageId="3" pageNumber="216">Thraulodes catoles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23C57D0FFCAFFA3FD9FE3E5FD96F87B" box="[530,610,1984,2006]" pageId="3" pageNumber="216" rank="species">
|
||
<emphasis id="B90FEAABFFCAFFA3FD9FE3E5FDDEF878" bold="true" box="[530,554,1984,2005]" pageId="3" pageNumber="216">sp</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCAFFA3FDBBE3E5FDA8F878" bold="true" box="[566,604,1984,2005]" pageId="3" pageNumber="216">nov</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC436B9FFCDFFA4FF4AE4B2FA7EFEB0" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="217">
|
||
Abdomen (
|
||
<figureCitation id="13402A3CFFCDFFA4FEC8E4B2FE5CFF1D" box="[325,424,151,176]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="4" pageNumber="217">Fig. 5–7</figureCitation>
|
||
): terga entirely red, with a pair of large midway black spots and a pair of submedian black spots on terga I–VII (
|
||
<figureCitation id="13402A3CFFCDFFA4FE06E499FE0DFF78" box="[395,505,188,213]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="4" pageNumber="217">Figs. 5–6</figureCitation>
|
||
); terga VIII–X red opaque with diffuse black spots, with some minor intraspecific variations (
|
||
<figureCitation id="13402A3CFFCDFFA4FE99E4FAFEACFF55" box="[276,344,223,248]" captionStart="FIGURES 2–7" captionStartId="2.[151,264,1958,1981]" captionTargetBox="[161,1425,934,1917]" captionTargetId="figure@2.[161,1425,934,1917]" captionTargetPageId="2" captionText="FIGURES 2–7. Thraulodes catoles sp. nov., male imago: 2, body lateral view. 3, foreleg. 4, thoracic sternae. 5–7, abdominal spot intraspecific pattern variation." httpUri="https://zenodo.org/record/2589899/files/figure.png" pageId="4" pageNumber="217">Fig. 7</figureCitation>
|
||
). Sternum I entirely washed brown with two medial black spots, sterna II–IX translucent red with a pair of midway and submedian small red spots. Caudal filaments white, black annulations on base of segments.
|
||
</paragraph>
|
||
<paragraph id="8BC436B9FFCDFFA4FF4AE502FD8EFE01" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="217">
|
||
Genitalia (
|
||
<figureCitation id="13402A3CFFCDFFA4FEB3E502FE3EFEED" box="[318,458,295,320]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="4" pageNumber="217">Figs. 10–12</figureCitation>
|
||
): styliger plate yellow translucent, with a short extension dorsal finger-like projection (
|
||
<figureCitation id="13402A3CFFCDFFA4FF12E569FF03FEC9" box="[159,247,332,357]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="4" pageNumber="217">Fig. 10</figureCitation>
|
||
); forceps yellow translucent. Penes yellow and completely divided, apicolateral area with an ear-like projection, external margin forming lateral pouch and a lapel on inner margin; spines of penes long and narrow, 1.9x the width of penes base (
|
||
<figureCitation id="13402A3CFFCDFFA4FE68E5B1FD99FE01" box="[485,621,404,429]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="4" pageNumber="217">Figs. 11–12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC436B9FFCDFFA4FF4AE592FD70FE7D" blockId="4.[151,1437,151,968]" box="[199,644,439,464]" pageId="4" pageNumber="217">
|
||
<emphasis id="B90FEAABFFCDFFA4FF4AE592FDFBFE7D" bold="true" box="[199,527,439,464]" pageId="4" pageNumber="217">Female imago and nymphs.</emphasis>
|
||
Unknown
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3616532FFCDFFA4FF4AE5F9FCDBFDCD" pageId="4" pageNumber="217" type="materials_examined">
|
||
<paragraph id="8BC436B9FFCDFFA4FF4AE5F9FCDBFDCD" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="217">
|
||
<emphasis id="B90FEAABFFCDFFA4FF4AE5F9FE45FE58" bold="true" box="[199,433,476,501]" pageId="4" pageNumber="217">
|
||
<collectingCounty id="62A54E35FFCDFFA4FF4AE5F9FEDBFE58" box="[199,303,476,501]" pageId="4" pageNumber="217">Material</collectingCounty>
|
||
examined.
|
||
</emphasis>
|
||
<materialsCitation id="3B133CE4FFCDFFA4FE37E5F9FCD6FDBA" ID-GBIF-Occurrence="2012823393" collectingDate="2013-11-01" collectionCode="UV" collectorName="de Catoles" country="Brazil" county="State" elevation="1183" location="Cachoeira Samambaia" municipality="Distrito de Catoles" pageId="4" pageNumber="217" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
|
||
<typeStatus id="54C0881BFFCDFFA4FE37E5F9FDD2FE58" box="[442,550,476,501]" pageId="4" pageNumber="217" type="holotype">Holotype</typeStatus>
|
||
.
|
||
<collectingCountry id="F36C7629FFCDFFA4FDBCE5F9FD6DFE59" box="[561,665,476,500]" name="Brazil" pageId="4" pageNumber="217">BRAZIL</collectingCountry>
|
||
,
|
||
<emphasis id="B90FEAABFFCDFFA4FD28E5F9FCDBFE58" bold="true" box="[677,815,476,501]" pageId="4" pageNumber="217">
|
||
<collectingRegion id="49BFF85BFFCDFFA4FD28E5F9FD1FFE58" box="[677,747,476,501]" country="Brazil" name="Bahia" pageId="4" pageNumber="217">Bahia</collectingRegion>
|
||
<collectingCounty id="62A54E35FFCDFFA4FD79E5F9FCDBFE58" box="[756,815,476,501]" pageId="4" pageNumber="217">State</collectingCounty>
|
||
</emphasis>
|
||
,
|
||
<collectingCounty id="62A54E35FFCDFFA4FCB3E5F9FC79FE58" box="[830,909,476,501]" pageId="4" pageNumber="217">Abaíra</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6BA0ACC3FFCDFFA4FC17E5F9FB86FE58" box="[922,1138,476,501]" pageId="4" pageNumber="217">
|
||
Distrito
|
||
<collectorName id="268E536FFFCDFFA4FC75E5F9FB86FE58" box="[1016,1138,476,501]" pageId="4" pageNumber="217">de Catolés</collectorName>
|
||
</collectingMunicipality>
|
||
:
|
||
<location id="8EA46062FFCDFFA4FBF3E5F9FA8FFE58" LSID="urn:lsid:plazi:treatment:03D287AFFFCBFFA4FF1AE4F8FB89FC65:8EA46062FFCDFFA4FBF3E5F9FA8FFE58" box="[1150,1403,476,501]" country="Brazil" county="State" municipality="Distrito de Catoles" name="Cachoeira Samambaia" pageId="4" pageNumber="217" stateProvince="Bahia">Cachoeira Samambaia</location>
|
||
(S 13°18ʹ21.6ʺ; W 41°53ʹ16ʺ;
|
||
<elevation id="0056D18AFFCDFFA4FE45E625FDAEFDB5" box="[456,602,511,536]" metricMagnitude="3" metricUnit="m" metricValue="1.183" pageId="4" pageNumber="217" unit="m" value="1183.0">
|
||
<quantity id="4C839B5CFFCDFFA4FE45E625FDE9FDBA" box="[456,541,512,536]" metricMagnitude="3" metricUnit="m" metricValue="1.183" pageId="4" pageNumber="217" unit="m" value="1183.0">1183 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
),
|
||
<date id="FFC51079FFCDFFA4FDFFE5DAFD18FDB5" box="[626,748,511,536]" pageId="4" pageNumber="217" value="2013-11-01">
|
||
<collectingDate id="EF81E991FFCDFFA4FDFFE5DAFD18FDB5" box="[626,748,511,536]" pageId="4" pageNumber="217" value="2013-11-01">01.xi.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectionCode id="ED6AAE7CFFCDFFA4FD74E625FCD6FDBA" box="[761,802,512,535]" country="Colombia" name="Departamento de Biologia de la Universidad del Valle" pageId="4" pageNumber="217">UV</collectionCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B133CE4FFCDFFA4FCA7E5DAFE35FD91" ID-GBIF-Occurrence="2012823401" collectionCode="MZUSP" collectorName="Calor AR & Dias ES & Campos, R" country="Brazil" county="State" pageId="4" pageNumber="217" specimenCount="1" stateProvince="Bahia" typeStatus="holotype">
|
||
and white light onto white sheet,
|
||
<collectorName id="268E536FFFCDFFA4FB28E5DAFAE2FDB5" box="[1189,1302,511,536]" pageId="4" pageNumber="217">Calor AR</collectorName>
|
||
,
|
||
<collectorName id="268E536FFFCDFFA4FAAFE5DAFA8BFDB5" box="[1314,1407,511,536]" pageId="4" pageNumber="217">Dias ES</collectorName>
|
||
&
|
||
<collectorName id="268E536FFFCDFFA4FF1AE601FEE1FD91" box="[151,277,548,572]" pageId="4" pageNumber="217">Campos, R</collectorName>
|
||
<emphasis id="B90FEAABFFCDFFA4FE90E601FECBFD90" box="[285,319,548,573]" italics="true" pageId="4" pageNumber="217">leg</emphasis>
|
||
. (
|
||
<collectionCode id="ED6AAE7CFFCDFFA4FEDBE601FE41FD91" box="[342,437,548,572]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="4" pageNumber="217">MZUSP</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
<materialsCitation id="3B133CE4FFCDFFA4FE42E600FCDFFDCD" ID-GBIF-Occurrence="2012823398" collectingDate="2013-10-31" collectionCode="UFVB, UFBA" collectorName="Calor AR & Dias ES & Campos, R" country="Brazil" county="State" location="Grota river" pageId="4" pageNumber="217" specimenCount="2" specimenCount-male="2" stateProvince="Bahia" typeStatus="paratype">
|
||
<typeStatus id="54C0881BFFCDFFA4FE42E600FDB4FD90" box="[463,576,549,573]" pageId="4" pageNumber="217" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
<specimenCount id="9D7DFD30FFCDFFA4FDC6E601FD83FD91" box="[587,631,548,572]" pageId="4" pageNumber="217" type="male">1 ♂</specimenCount>
|
||
imago (
|
||
<collectionCode id="ED6AAE7CFFCDFFA4FD5BE601FCD2FD91" box="[726,806,548,572]" country="Brazil" httpUri="http://grbio.org/cool/3i20-rx1w" name="Vicosa, Universidade Federal de Vicosa, Museum of Entomology" pageId="4" pageNumber="217">UFVB</collectionCode>
|
||
), same data as holotype;
|
||
<specimenCount id="9D7DFD30FFCDFFA4FBCAE601FB86FD91" box="[1095,1138,548,572]" pageId="4" pageNumber="217" type="male">1 ♂</specimenCount>
|
||
imago,
|
||
<location id="8EA46062FFCDFFA4FB5DE601FABAFD90" LSID="urn:lsid:plazi:treatment:03D287AFFFCBFFA4FF1AE4F8FB89FC65:8EA46062FFCDFFA4FB5DE601FABAFD90" box="[1232,1358,548,573]" country="Brazil" county="State" name="Grota river" pageId="4" pageNumber="217" stateProvince="Bahia">Grota river</location>
|
||
(S 13° 17ʹ10ʺ; W 41° 52ʹ08.3ʺ),
|
||
<date id="FFC51079FFCDFFA4FE3DE66DFDD6FDCD" box="[432,546,584,608]" pageId="4" pageNumber="217" value="2013-10-31">
|
||
<collectingDate id="EF81E991FFCDFFA4FE3DE66DFDD6FDCD" box="[432,546,584,608]" pageId="4" pageNumber="217" value="2013-10-31">31.x.2013</collectingDate>
|
||
</date>
|
||
,
|
||
<collectingMethod id="523A4EAEFFCDFFA4FDA3E662FD30FDF2" box="[558,708,583,608]" pageId="4" pageNumber="217">light pan trap</collectingMethod>
|
||
(
|
||
<collectionCode id="ED6AAE7CFFCDFFA4FD5FE66DFCD7FDCD" box="[722,803,584,608]" pageId="4" pageNumber="217">UFBA</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3616532FFCDFFA4FF4AE649FA87FD28" box="[199,1395,620,645]" pageId="4" pageNumber="217" type="etymology">
|
||
<paragraph id="8BC436B9FFCDFFA4FF4AE649FA87FD28" blockId="4.[151,1437,151,968]" box="[199,1395,620,645]" pageId="4" pageNumber="217">
|
||
<emphasis id="B90FEAABFFCDFFA4FF4AE649FEBAFD28" bold="true" box="[199,334,620,645]" pageId="4" pageNumber="217">Etymology.</emphasis>
|
||
This species is named as noun in apposition in reference to the collection site, Catolés district.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3616532FFCDFFA4FF4AE6AAFB89FC65" pageId="4" pageNumber="217" type="discussion">
|
||
<paragraph id="8BC436B9FFCDFFA4FF4AE6AAFB89FC65" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="217">
|
||
<emphasis id="B90FEAABFFCDFFA4FF4AE6AAFEB8FD05" bold="true" box="[199,332,655,680]" pageId="4" pageNumber="217">Discussion.</emphasis>
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FED9E6AAFDDEFD05" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[340,554,655,680]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
|
||
<emphasis id="B90FEAABFFCDFFA4FED9E6AAFDDEFD05" box="[340,554,655,680]" italics="true" pageId="4" pageNumber="217">Thraulodes catoles</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAABFFCDFFA4FDBEE6AAFD79FD05" bold="true" box="[563,653,655,680]" pageId="4" pageNumber="217">
|
||
<taxonomicNameLabel id="A23C57D0FFCDFFA4FDBEE6AAFD79FD05" box="[563,653,655,680]" pageId="4" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as well as
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FC81E6B4FBE8FD05" authority="Dominguez" authorityName="Dominguez" box="[780,1052,656,680]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="consortis">
|
||
<emphasis id="B90FEAABFFCDFFA4FC81E6B4FCEDFD05" box="[780,793,657,680]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FCA5E6B4FC7BFD05" box="[808,911,657,680]" italics="true" pageId="4" pageNumber="217">consortis</emphasis>
|
||
Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FBA5E6B4FA88FD05" authority="Traver & Edmunds" authorityName="Traver & Edmunds" authorityYear="1967" box="[1064,1404,655,680]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="regulus">
|
||
<emphasis id="B90FEAABFFCDFFA4FBA5E6B4FBC1FD05" box="[1064,1077,657,680]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FBC9E6AAFB62FD05" box="[1092,1174,655,680]" italics="true" pageId="4" pageNumber="217">regulus</emphasis>
|
||
Traver & Edmunds
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FA0AE6B4FE65FD61" authority="Dominguez" authorityName="Dominguez" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="liminaris">
|
||
<emphasis id="B90FEAABFFCDFFA4FA0AE6B4FA60FD05" box="[1415,1428,657,680]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FF1AE691FF08FD60" box="[151,252,692,717]" italics="true" pageId="4" pageNumber="217">liminaris</emphasis>
|
||
Domínguez
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FE2EE690FCD8FD60" authority="Mariano & Froehlich" authorityName="Mariano & Froehlich" authorityYear="2011" box="[419,812,692,717]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="amanda">
|
||
<emphasis id="B90FEAABFFCDFFA4FE2EE690FE44FD61" box="[419,432,693,716]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FE48E691FDEBFD60" box="[453,543,692,717]" italics="true" pageId="4" pageNumber="217">amanda</emphasis>
|
||
Mariano & Froehlich
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FCFCE690FAC3FD60" authority="Mariano & Froehlich" authorityName="Mariano & Froehlich" authorityYear="2011" box="[881,1335,692,717]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="xavantinensis">
|
||
<emphasis id="B90FEAABFFCDFFA4FCFCE690FC8AFD61" box="[881,894,693,716]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FC19E690FBD8FD61" box="[916,1068,693,716]" italics="true" pageId="4" pageNumber="217">xavantinensis</emphasis>
|
||
Mariano & Froehlich
|
||
</taxonomicName>
|
||
has the abdominal segments entirely pigmented, however
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FD45E6FCFCC6FD5D" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[712,818,727,752]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
|
||
<emphasis id="B90FEAABFFCDFFA4FD45E6FCFD22FD5D" box="[712,726,729,752]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FD6EE6F2FCC6FD5D" box="[739,818,727,752]" italics="true" pageId="4" pageNumber="217">catoles</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAABFFCDFFA4FCB4E6F2FC66FD5D" bold="true" box="[825,914,727,752]" pageId="4" pageNumber="217">
|
||
<taxonomicNameLabel id="A23C57D0FFCDFFA4FCB4E6F2FC66FD5D" box="[825,914,727,752]" pageId="4" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has reddish abdominal segments with a pair of large midway black spots and a pair of submedian black spots while the other species have the color pattern varying from dark brown to orange-brown with different placement of black spots.
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FBB7E73AFAFBFC95" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[1082,1295,799,824]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
|
||
<emphasis id="B90FEAABFFCDFFA4FBB7E73AFAFBFC95" box="[1082,1295,799,824]" italics="true" pageId="4" pageNumber="217">Thraulodes catoles</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAABFFCDFFA4FA9BE73AFA84FC95" bold="true" box="[1302,1392,799,824]" pageId="4" pageNumber="217">
|
||
<taxonomicNameLabel id="A23C57D0FFCDFFA4FA9BE73AFA84FC95" box="[1302,1392,799,824]" pageId="4" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
has a digitiform dorsal extension on styliger plate and general shape of penes similar to
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FBFCE760FADCFCF1" authorityName="Mariano & Froehlich" authorityYear="2011" box="[1137,1320,837,860]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="xavantinensis">
|
||
<emphasis id="B90FEAABFFCDFFA4FBFCE760FB8AFCF1" box="[1137,1150,837,860]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FB1DE760FADCFCF1" box="[1168,1320,837,860]" italics="true" pageId="4" pageNumber="217">xavantinensis</emphasis>
|
||
</taxonomicName>
|
||
, although differs by the shape and size of styliger plate and by the lapel on inner margin of penes. Furthermore the recurved folds parallel to the inner margin of the penes as in
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FD57E7A8FC7BFC09" authorityName="Mariano & Froehlich" authorityYear="2011" box="[730,911,909,932]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="xavantinensis">
|
||
<emphasis id="B90FEAABFFCDFFA4FD57E7A8FD13FC09" box="[730,743,909,932]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FD7BE7A8FC7BFC09" box="[758,911,909,932]" italics="true" pageId="4" pageNumber="217">xavantinensis</emphasis>
|
||
</taxonomicName>
|
||
is absent in
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FB91E7A8FB73FC08" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[1052,1159,908,933]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
|
||
<emphasis id="B90FEAABFFCDFFA4FB91E7A8FB73FC08" box="[1052,1159,908,933]" italics="true" pageId="4" pageNumber="217">T. catoles</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAABFFCDFFA4FB03E7A9FB1CFC08" bold="true" box="[1166,1256,908,933]" pageId="4" pageNumber="217">
|
||
<taxonomicNameLabel id="A23C57D0FFCDFFA4FB03E7A9FB1CFC08" box="[1166,1256,908,933]" pageId="4" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
An asymmetric variation in lateral pouch has been observed in specimens of
|
||
<taxonomicName id="4C7B4D3AFFCDFFA4FCCCE794FC58FC65" authority="Campos & Mariano, 2019" authorityName="Campos & Mariano" authorityYear="2019" box="[833,940,943,968]" class="Insecta" family="Leptophlebiidae" genus="Thraulodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Ephemeroptera" pageId="4" pageNumber="217" phylum="Arthropoda" rank="species" species="catoles" status="sp. nov.">
|
||
<emphasis id="B90FEAABFFCDFFA4FCCCE794FCBBFC65" box="[833,847,945,968]" italics="true" pageId="4" pageNumber="217">T</emphasis>
|
||
.
|
||
<emphasis id="B90FEAABFFCDFFA4FCD0E78AFC58FC65" box="[861,940,943,968]" italics="true" pageId="4" pageNumber="217">catoles</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90FEAABFFCDFFA4FC3EE78AFBF9FC65" bold="true" box="[947,1037,943,968]" pageId="4" pageNumber="217">
|
||
<taxonomicNameLabel id="A23C57D0FFCDFFA4FC3EE78AFBF9FC65" box="[947,1037,943,968]" pageId="4" pageNumber="217" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13402A3CFFCDFFA4FB91E78AFB9BFC65" box="[1052,1135,943,968]" captionStart="FIGURES 8–12" captionStartId="3.[151,264,1983,2006]" captionTargetBox="[183,1392,415,1910]" captionTargetId="figure@3.[161,1426,373,1962]" captionTargetPageId="3" captionText="FIGURES 8–12. Thraulodes catoles sp. nov., male imago: 8, forewing. 9, hind wing.10, styliger plate (ventral view). 11, penes in detail (ventral view). 12, asymmetric variations of lateral pouch in penes indicated by the black arrows (ventral view)." httpUri="https://zenodo.org/record/2589901/files/figure.png" pageId="4" pageNumber="217">Fig. 12</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |