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<document id="DB4043D6FC36C7EE1A1707A101451216" ID-DOI="10.11646/zootaxa.3670.2.9" ID-GBIF-Dataset="492d3ca4-105c-4c92-8a7d-6cb180d00b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="223576" ID-ZooBank="33F668E4-95BC-4203-B892-80365B4F3B89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633857078" checkinUser="plazi" docAuthor="Filippini, Valentina, Micó, Estefania &amp; Galante, Eduardo" docDate="2013" docId="03D28781FFFAFFDC5AA6606AFB68F823" docLanguage="en" docName="zt03670p273.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala pseudoeucoma Filippini, Mico" docType="treatment" docVersion="8" lastPageNumber="269" masterDocId="FFEBFFF9FFF7FFD25A31622DFFA8FFB7" masterDocTitle="Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" masterLastPageNumber="273" masterPageNumber="255" pageNumber="268" updateTime="1698334918277" updateUser="plazi">
<mods:mods id="A0514B727B942A32D2335F21DA31787D" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8E270B2A85B05FA555B8C7BF1738EE0E">
<mods:title id="8B54B8B06B2BA382C4AD1B213830AC81">Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name id="D93D7E4EBFE1994E388CB28F5B529CDF" type="personal">
<mods:role id="8FA31A466358184379A0595862F5A10C">
<mods:roleTerm id="CA694F22E8119B2FFBA3AAF3C1D754E9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B13B8F92FC94B2FEC989F3FD8EBDE76">Filippini, Valentina</mods:namePart>
</mods:name>
<mods:name id="00A1BF2DC8DA33DD8A951DAB56F7A5A6" type="personal">
<mods:role id="1F325E0B0E278CD96749E11AEE6F5E10">
<mods:roleTerm id="33367782C6C069773C38E15A2464B489">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="243731D7923066F5D7C43781924A2E1D">Micó, Estefania</mods:namePart>
</mods:name>
<mods:name id="B71EF9F640238685DC6BBC318BB96A08" type="personal">
<mods:role id="7AB678BF8AB79372494686D234F0441D">
<mods:roleTerm id="1BB3E05A1D0CAC879E20D844535BB3E6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F00ACCD1B6468B77128A9A5EE18DAA15">Galante, Eduardo</mods:namePart>
</mods:name>
<mods:typeOfResource id="ED31B570750B3F02FA0BCAB288010422">text</mods:typeOfResource>
<mods:relatedItem id="E21061761C912B65965436664793CC81" type="host">
<mods:titleInfo id="C7EFF04C694B828287E21E50E4ECE58B">
<mods:title id="980251B00D36FBA3257186635086CC58">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="19072F74A55091CCE208234F8D36EA41">
<mods:date id="6CEE0FEA2DB312AF7F61CD31DE2C16F6">2013</mods:date>
<mods:detail id="9D9C7F865345C972EBE44D99D511B0B4" type="volume">
<mods:number id="7227450821542FB3A932C9E9D4891414">3670</mods:number>
</mods:detail>
<mods:detail id="9A5E3A3E6B91D9795109368577A546DD" type="issue">
<mods:number id="5108723CC9F8045F17D8ED013FFB36BA">2</mods:number>
</mods:detail>
<mods:extent id="DEEE9D29F266B9B489F205B29C7FD38A" unit="page">
<mods:start id="DF0844EC18B25F181D4D0DD0DEDEDC13">255</mods:start>
<mods:end id="3EBDAA182098F4ED4C1070F2AAFB86EF">273</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="44FBE012DD254059A520795158D0C837">journal article</mods:classification>
<mods:identifier id="37493497CB981909608B07EEE0120AF2" type="DOI">10.11646/zootaxa.3670.2.9</mods:identifier>
<mods:identifier id="259626753846FE0BE7A22E974C8552F2" type="GBIF-Dataset">492d3ca4-105c-4c92-8a7d-6cb180d00b08</mods:identifier>
<mods:identifier id="E00E403C60F017361FFB03F99D3FC4A6" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="B1DC963971E8D15D33812E8A4AF773A3" type="Zenodo-Dep">223576</mods:identifier>
<mods:identifier id="441DAA16C77F8D429D9FA6BD6FAEF628" type="ZooBank">33F668E4-95BC-4203-B892-80365B4F3B89</mods:identifier>
</mods:mods>
<treatment id="03D28781FFFAFFDC5AA6606AFB68F823" ID-DOI="http://doi.org/10.5281/zenodo.6159039" ID-GBIF-Taxon="119570128" ID-Zenodo-Dep="6159039" LSID="urn:lsid:plazi:treatment:03D28781FFFAFFDC5AA6606AFB68F823" httpUri="http://treatment.plazi.org/id/03D28781FFFAFFDC5AA6606AFB68F823" lastPageId="14" lastPageNumber="269" pageId="13" pageNumber="268">
<subSubSection id="C361651CFFFAFFDF5AA6606AFF70FD35" pageId="13" pageNumber="268" type="nomenclature">
<paragraph id="8BC43697FFFAFFDF5AA6606AFC18FDD6" blockId="13.[151,944,583,642]" box="[151,944,583,609]" pageId="13" pageNumber="268">
<heading id="D08C81FBFFFAFFDF5AA6606AFC18FDD6" bold="true" box="[151,944,583,609]" fontSize="11" level="1" pageId="13" pageNumber="268" reason="1">
<emphasis id="B90FEA85FFFAFFDF5AA6606AFC18FDD6" bold="true" box="[151,944,583,609]" pageId="13" pageNumber="268">
<taxonomicName id="4C7B4D14FFFAFFDF5AA6606AFD2AFDD6" authority="Filippini, Mico" authorityName="Filippini, Mico" box="[151,642,583,609]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="268" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFFAFFDF5AA6606AFE68FDD6" bold="true" box="[151,448,583,609]" italics="true" pageId="13" pageNumber="268">Anomala pseudoeucoma</emphasis>
<collectingCountry id="F36C7607FFFAFFDF5BF6606AFD9CFDD6" box="[455,564,583,609]" name="Philippines" pageId="13" pageNumber="268">Filippini</collectingCountry>
, Micó
</taxonomicName>
, &amp; Galante, new species
</emphasis>
</heading>
</paragraph>
<paragraph id="8BC43697FFFAFFDF5AA66047FF70FD35" blockId="13.[151,944,583,642]" box="[151,216,618,642]" pageId="13" pageNumber="268">
<figureCitation id="13402A12FFFAFFDF5AA66047FF70FD35" box="[151,216,618,642]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1929,1953]" captionTargetBox="[193,1406,206,1890]" captionTargetId="figure@4.[184,1420,194,1919]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Habitus images. 2: Anomala eucoma (paralectotype, Las Mercedes, Guatemala). 3: A. flavaeucoma (female paratype, Estación Hitoy Cerere, Costa Rica). 4. A. megaparamera (holotype). 5: A. pseudoeucoma (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/223578/files/figure.png" pageId="13" pageNumber="268">Fig. 5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFFAFFDF5AA6609FFB46F90D" pageId="13" pageNumber="268" type="materials_examined">
<paragraph id="8BC43697FFFAFFDF5AA6609FFDA3FCA5" blockId="13.[151,1437,690,1975]" pageId="13" pageNumber="268">
<emphasis id="B90FEA85FFFAFFDF5AA6609FFE2CFD7C" bold="true" box="[151,388,690,715]" pageId="13" pageNumber="268">Material examined.</emphasis>
<typeStatus id="54C08835FFFAFFDF5BBF609FFE53FD7D" box="[398,507,690,714]" pageId="13" pageNumber="268" type="holotype">Holotype</typeStatus>
: ɞ &quot;Est. Hitoy Cerere,
<quantity id="4C839B72FFFAFFDF5921609FFCFAFD7D" box="[784,850,690,714]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="268" unit="m" value="100.0">100m</quantity>
, R. Cerere, Res. Biol. Hitoy Cerere, Prov. Limón,
<collectingCountry id="F36C7607FFFAFFDF5AA660FBFEF8FD59" box="[151,336,726,750]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
, R. Guzman.
<date id="FFC51057FFFAFFDF581360FBFC9EFD59" box="[546,822,726,750]" pageId="13" pageNumber="268" value="1992-04-12" valueMax="1992-04-28" valueMin="1992-04-12">12 – 28 Abr 1992</date>
. L_N_184600_643400 #1137 /
<collectingCountry id="F36C7607FFFAFFDF5ED460FBFA34FD59" box="[1253,1436,726,750]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000393407&quot; (
<collectionCode id="ED6AAE52FFFAFFDF5B9A60D7FE57FCA5" box="[427,511,762,786]" pageId="13" pageNumber="268">INBIO</collectionCode>
).
</paragraph>
<paragraph id="8BC43697FFFAFFDF5AF66132FB46F90D" blockId="13.[151,1437,690,1975]" pageId="13" pageNumber="268">
<typeStatus id="54C08835FFFAFFDF5AF66132FE9CFC80" box="[199,308,799,823]" pageId="13" pageNumber="268" type="paratype">Paratypes</typeStatus>
(20): 1ɞ &quot;Alajuela,
<collectingCountry id="F36C7607FFFAFFDF58106133FD37FC81" box="[545,671,798,822]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
, San Ramon, Rio S. Lorencito,
<quantity id="4C839B72FFFAFFDF5E3A6133FBE3FC81" box="[1035,1099,798,822]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="13" pageNumber="268" unit="m" value="800.0">800m</quantity>
<date id="FFC51057FFFAFFDF5E656133FB57FC81" box="[1108,1279,798,822]" pageId="13" pageNumber="268" value="1986-10-03">3 octubre 1986</date>
Col: A. Solis Blanco. /
<collectingCountry id="F36C7607FFFAFFDF5B3C616FFE1DFCED" box="[269,437,834,858]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI002517836&quot;; 1Ƥ &quot;Alajuela,
<collectingCountry id="F36C7607FFFAFFDF59B7616FFBA1FCED" box="[902,1033,834,858]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
. Reserva de San Ramón. Río San Lorencito,
<specimenCount id="9D7DFD1EFFFAFFDF5B24614BFEF5FCC9" box="[277,349,870,894]" pageId="13" pageNumber="268" type="generic">
<quantity id="4C839B72FFFAFFDF5B24614BFEF1FCC9" box="[277,345,870,894]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="268" unit="m" value="850.0">850m</quantity>
.
</specimenCount>
<date id="FFC51057FFFAFFDF5B56614BFE5DFCC9" box="[359,501,870,894]" pageId="13" pageNumber="268" value="1987-04-01">1 abril 1987</date>
Angel Solís /
<collectingCountry id="F36C7607FFFAFFDF5895614BFCE3FCC9" box="[676,843,870,894]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI002517838&quot;; 1Ƥ &quot;Alajuela,
<collectingCountry id="F36C7607FFFAFFDF5F26614BFA30FCC9" box="[1303,1432,870,894]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
. Reserva de San Ramón. Río San Lorencito,
<specimenCount id="9D7DFD1EFFFAFFDF58CD61A7FCEBFC15" box="[764,835,906,930]" pageId="13" pageNumber="268" type="generic">
<quantity id="4C839B72FFFAFFDF58CD61A7FC97FC15" box="[764,831,906,930]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="268" unit="m" value="850.0">850m</quantity>
.
</specimenCount>
<date id="FFC51057FFFAFFDF596B61A7FBACFC15" box="[858,1028,906,930]" pageId="13" pageNumber="268" value="1987-04-01">1 abril 1987</date>
Angel Solís /
<collectingCountry id="F36C7607FFFAFFDF5ED661A7FA34FC15" box="[1255,1436,906,930]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI002517839&quot;; 1Ƥ &quot;Alajuela,
<collectingCountry id="F36C7607FFFAFFDF587F6183FD64FC71" box="[590,716,942,966]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
. Reserva de San Ramón. Río San Lorencito,
<specimenCount id="9D7DFD1EFFFAFFDF5EF76183FAA5FC71" box="[1222,1293,942,966]" pageId="13" pageNumber="268" type="generic">
<quantity id="4C839B72FFFAFFDF5EF76183FAA1FC71" box="[1222,1289,942,966]" metricMagnitude="2" metricUnit="m" metricValue="8.5" pageId="13" pageNumber="268" unit="m" value="850.0">850m</quantity>
.
</specimenCount>
<date id="FFC51057FFFAFFDF5F256183FA34FC71" box="[1300,1436,942,966]" pageId="13" pageNumber="268" value="1987-04-01">1 abril 1987</date>
Angel Solís /
<collectingCountry id="F36C7607FFFAFFDF5B0361FFFE7EFC5D" box="[306,470,978,1002]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI002517840&quot;; 2ɞ &quot;Albergue Heliconias, Prov. Alajuela,
<collectingCountry id="F36C7607FFFAFFDF5EFC61FFFAE4FC5D" box="[1229,1356,978,1002]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
.
<date id="FFC51057FFFAFFDF5F6661FFFF67FBB9" pageId="13" pageNumber="268" value="2007-11-11">11/11/ 2007</date>
Leg. D. Gutiérrez&quot;
<collectionCode id="ED6AAE52FFFAFFDF5B8761DBFDAAFBB9" box="[438,514,1014,1038]" pageId="13" pageNumber="268">CEUA</collectionCode>
00105078 and
<collectionCode id="ED6AAE52FFFAFFDF589C61DBFD53FBB9" box="[685,763,1014,1038]" pageId="13" pageNumber="268">CEUA</collectionCode>
00105079; 1Ƥ &quot;Albergue Heliconias, Prov. Alajuela,
<collectingCountry id="F36C7607FFFAFFDF5F6D61DBFF66FB85" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
.
<date id="FFC51057FFFAFFDF5AE86637FEF0FB85" box="[217,344,1050,1074]" pageId="13" pageNumber="268" value="2008-01-31">31/01/2008</date>
Leg. D. Gutiérrez /
<collectionCode id="ED6AAE52FFFAFFDF580F6637FD26FB85" box="[574,654,1050,1074]" pageId="13" pageNumber="268">CEUA</collectionCode>
00105081&quot;; 1ɞ &quot;Albergue Heliconias, Prov. Alajuela,
<collectingCountry id="F36C7607FFFAFFDF5EDE6637FAC5FB85" box="[1263,1389,1050,1074]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
.
<date id="FFC51057FFFAFFDF5F496637FF5BFBE1" pageId="13" pageNumber="268" value="2007-09-14">14/ 09/2007</date>
Leg. D. Gutiérrez /
<collectionCode id="ED6AAE52FFFAFFDF5BD26613FD9AFBE1" box="[483,562,1086,1110]" pageId="13" pageNumber="268">CEUA</collectionCode>
00105082&quot;; 1ɞ &quot;Cerro Tortuguero,
<quantity id="4C839B72FFFAFFDF59FA6613FB81FBE1" box="[971,1065,1086,1110]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="13" pageNumber="268" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, P. N. Tortuguero, Prov. Limon,
<collectingCountry id="F36C7607FFFAFFDF5AA6664FFE96FBCD" box="[151,318,1122,1146]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
, R. Delgado,
<date id="FFC51057FFFAFFDF5BEB664FFD61FBCD" box="[474,713,1122,1146]" pageId="13" pageNumber="268" value="1992-01-26" valueMax="1992-02-14" valueMin="1992-01-26">26 Ene–14 Feb 1992</date>
, L- N 285000_588000 /
<collectingCountry id="F36C7607FFFAFFDF59D3664FFB2EFBCD" box="[994,1158,1122,1146]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000334543&quot;; 1ɞ &quot;Cerro Tortuguero,
<quantity id="4C839B72FFFAFFDF5B9D66ABFDA2FB29" box="[428,522,1158,1182]" metricMagnitude="1" metricUnit="m" metricValue="6.0" metricValueMax="12.0" metricValueMin="0.0" pageId="13" pageNumber="268" unit="m" value="60.0" valueMax="120.0" valueMin="0.0">0–120m</quantity>
, P. N. Tortuguero, Prov. Limon,
<collectingCountry id="F36C7607FFFAFFDF59A566ABFB97FB29" box="[916,1087,1158,1182]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
, R. Delgado,
<date id="FFC51057FFFAFFDF5ED666ABFAF8FB29" box="[1255,1360,1158,1182]" pageId="13" pageNumber="268" value="1991-09">Set 1991</date>
, L- N 285000_588000 /
<collectingCountry id="F36C7607FFFAFFDF5B536687FDADFB75" box="[354,517,1194,1218]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000479636&quot;; 1ɞ &quot;Cerro Tortuguero, P. N. Tortuguero,
<quantity id="4C839B72FFFAFFDF5EDA6687FAFFFB75" box="[1259,1367,1194,1218]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="13" pageNumber="268" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0 – 100m</quantity>
, Prov. Limon,
<collectingCountry id="F36C7607FFFAFFDF5AC066E3FE33FB51" box="[241,411,1230,1254]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
. J. Solano,
<date id="FFC51057FFFAFFDF582E66E2FD3FFB51" box="[543,663,1230,1255]" pageId="13" pageNumber="268" value="1990-05">May 1990</date>
. L- N 285000_588000 /
<collectingCountry id="F36C7607FFFAFFDF598566E3FBF1FB51" box="[948,1113,1230,1254]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000242771&quot;; 1Ƥ &quot;Cerro Tortuguero, P. N. Tortuguero, Prov. Limon,
<collectingCountry id="F36C7607FFFAFFDF592A66DFFC64FABD" box="[795,972,1266,1290]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFFAFFDF59D066DFFB92FABD" box="[993,1082,1266,1290]" pageId="13" pageNumber="268" type="generic">
<quantity id="4C839B72FFFAFFDF59D066DFFB9DFABD" box="[993,1077,1266,1290]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="268" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
<date id="FFC51057FFFAFFDF5E7B66DEFB6FFABD" box="[1098,1223,1266,1290]" pageId="13" pageNumber="268" value="1989-11">Nov 1989</date>
. J. Solano, L N 285000_588000 /
<collectingCountry id="F36C7607FFFAFFDF5B56673BFDA5FA99" box="[359,525,1302,1326]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000055767&quot;; 1ɞ &quot;Cerro Tortuguero, P. N. Tortuguero, Prov. Limon,
<collectingCountry id="F36C7607FFFAFFDF5AA66717FEE9FAE5" box="[151,321,1338,1362]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFFAFFDF5B7F6717FE37FAE5" box="[334,415,1338,1362]" pageId="13" pageNumber="268" type="generic">
<quantity id="4C839B72FFFAFFDF5B7F6717FE33FAE5" box="[334,411,1338,1362]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="268" unit="m" value="100.0">100 m</quantity>
.
</specimenCount>
<date id="FFC51057FFFAFFDF5B986716FDB6FAE5" box="[425,542,1338,1362]" pageId="13" pageNumber="268" value="1989-11">Nov 1989</date>
. J. Solano, L N 285000_588000 /
<collectingCountry id="F36C7607FFFAFFDF59806717FBF0FAE5" box="[945,1112,1338,1362]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000141229&quot;; 1ɞ &quot;Est. La Casona,
<quantity id="4C839B72FFFAFFDF5B5D6773FE61FAC1" box="[364,457,1374,1398]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="13" pageNumber="268" unit="m" value="1520.0">1520 m</quantity>
, Res. Biol. Monteverde, Prov. Punt.
<collectingCountry id="F36C7607FFFAFFDF59B86773FB9FFAC1" box="[905,1079,1374,1398]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
, N. Obando,
<date id="FFC51057FFFAFFDF5EEF6773FAE6FAC1" box="[1246,1358,1374,1398]" pageId="13" pageNumber="268" value="1991-10">Oct 1991</date>
, L- N 253250_4497001 /
<collectingCountry id="F36C7607FFFAFFDF5B4567AFFDB2FA2D" box="[372,538,1410,1434]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000632624&quot;; 1ɞ &quot;Est. La Casona,
<quantity id="4C839B72FFFAFFDF5E1C67AFFB2EFA2D" box="[1069,1158,1410,1434]" metricMagnitude="3" metricUnit="m" metricValue="1.52" pageId="13" pageNumber="268" unit="m" value="1520.0">1520 m</quantity>
, Res. Biol. Monteverde, Prov. Punt.
<collectingCountry id="F36C7607FFFAFFDF5B2D678BFE6CFA09" box="[284,452,1446,1470]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
, N. Obando,
<date id="FFC51057FFFAFFDF586D678BFD6FFA09" box="[604,711,1446,1470]" pageId="13" pageNumber="268" value="1991-10">Oct 1991</date>
, L- N 253250_449700 /
<collectingCountry id="F36C7607FFFAFFDF59D3678BFB2EFA09" box="[994,1158,1446,1470]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000632689&quot;; 1ɞ &quot;Fca. Cafrosa, Est. Las Mellizas, P. N. Amistad, Puntarenas Pr.
<collectingCountry id="F36C7607FFFAFFDF59A367E7FB91FA55" box="[914,1081,1482,1506]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
,
<quantity id="4C839B72FFFAFFDF5E7467E7FB36FA55" box="[1093,1182,1482,1506]" metricMagnitude="3" metricUnit="m" metricValue="1.3" pageId="13" pageNumber="268" unit="m" value="1300.0">1300 m</quantity>
.
<date id="FFC51057FFFAFFDF5E9B67E7FA30FA55" box="[1194,1432,1482,1507]" pageId="13" pageNumber="268" value="1989-08-20" valueMax="1989-09-04" valueMin="1989-08-20">20 Aug – 4 Set 1989</date>
, M. Ramirez &amp; G. Mora, L S 316100_596100 /
<collectingCountry id="F36C7607FFFAFFDF58FA67C3FCDBF9B1" box="[715,883,1518,1542]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI000093803&quot;; 1Ƥ &quot;
<collectingCountry id="F36C7607FFFAFFDF5EFE67C3FB52F9B1" box="[1231,1274,1518,1542]" name="Iceland" pageId="13" pageNumber="268">Isla</collectingCountry>
Bonita, Prov. Alajuela,
<collectingCountry id="F36C7607FFFAFFDF5B37643FFE2FF99D" box="[262,391,1554,1578]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
.
<date id="FFC51057FFFAFFDF5BA5643FFDBCF99D" box="[404,532,1554,1578]" pageId="13" pageNumber="268" value="2007-07-12">12/07/2007</date>
Leg. M. Moraga /
<collectionCode id="ED6AAE52FFFAFFDF58C6643FFCEEF99D" box="[759,838,1554,1578]" pageId="13" pageNumber="268">CEUA</collectionCode>
00105080&quot;; 1ɞ &quot;Limon,
<collectingCountry id="F36C7607FFFAFFDF5E52643FFB48F99D" box="[1123,1248,1554,1578]" name="Costa Rica" pageId="13" pageNumber="268">Costa Rica</collectingCountry>
Reserva Hitoy– Cerere Rio Cerere 100 msnm
<date id="FFC51057FFFAFFDF5BDA641BFD3DF9F9" box="[491,661,1590,1614]" pageId="13" pageNumber="268" value="1987-03-24">24 marzo 1987</date>
Angel Solis /
<collectingCountry id="F36C7607FFFAFFDF5908641BFC75F9F9" box="[825,989,1590,1614]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI002517319&quot;; 1ɞ &quot;San Luis, Monteverde, Prov. Punta.,
<collectingCountry id="F36C7607FFFAFFDF5BF96477FDD9F9C5" box="[456,625,1626,1650]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
,
<quantity id="4C839B72FFFAFFDF584F6477FD70F9C5" box="[638,728,1626,1650]" metricMagnitude="3" metricUnit="m" metricValue="1.04" pageId="13" pageNumber="268" unit="m" value="1040.0">1040 m</quantity>
.
<date id="FFC51057FFFAFFDF58D46476FCFBF9C5" box="[741,851,1626,1650]" pageId="13" pageNumber="268" value="1993-11">nov 1993</date>
. Z. Fuentes L- N
<date id="FFC51057FFFAFFDF5E126477FBDEF9C5" box="[1059,1142,1626,1650]" pageId="13" pageNumber="268" value="50-08-25">250850</date>
_449250 #2443 /
<collectingCountry id="F36C7607FFFAFFDF5F736477FF72F921" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI001937988&quot;; 1ɞ &quot;R.B. Hitoy Cerere, Valle La Estrella, Prov. Limon,
<collectingCountry id="F36C7607FFFAFFDF5E5A6453FABBF921" box="[1131,1299,1662,1686]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
.
<quantity id="4C839B72FFFAFFDF5F2C6453FA30F921" box="[1309,1432,1662,1686]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="268" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
100–
<specimenCount id="9D7DFD1EFFFAFFDF5F676453FA30F921" box="[1366,1432,1662,1686]" pageId="13" pageNumber="268" type="generic">200m</specimenCount>
</quantity>
.
<date id="FFC51057FFFAFFDF5AA6648FFEE3F90D" box="[151,331,1698,1723]" pageId="13" pageNumber="268" value="1994-05-09" valueMax="1994-05-14" valueMin="1994-05-09">9–14 May 1994</date>
, G. Carballo, L N 643400_184600 # 2856 /
<collectingCountry id="F36C7607FFFAFFDF5907648FFC72F90D" box="[822,986,1698,1722]" name="Costa Rica" pageId="13" pageNumber="268">COSTA RICA</collectingCountry>
INBIOCRI001799052&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFFAFFDC5AF664EBFDF2FACF" lastPageId="14" lastPageNumber="269" pageId="13" pageNumber="268" type="description">
<paragraph id="8BC43697FFFAFFDF5AF664EBFBB8F891" blockId="13.[151,1437,690,1975]" pageId="13" pageNumber="268">
<emphasis id="B90FEA85FFFAFFDF5AF664EBFEF1F968" bold="true" box="[199,345,1734,1759]" pageId="13" pageNumber="268">Description.</emphasis>
Male. Body shape oval. Length
<quantity id="4C839B72FFFAFFDF58EA64EBFCFBF969" box="[731,851,1734,1758]" metricMagnitude="-2" metricUnit="m" metricValue="1.216" pageId="13" pageNumber="268" unit="mm" value="12.16">12.16 mm</quantity>
. Width
<quantity id="4C839B72FFFAFFDF598064EBFBB3F969" box="[945,1051,1734,1758]" metricMagnitude="-3" metricUnit="m" metricValue="6.57" pageId="13" pageNumber="268" unit="mm" value="6.57">6.57 mm</quantity>
. Head, pronotum, scutellum, and pygidium dark reddish brown; legs and underside reddish brown; elytra dark brown with a light horizontal band along the anterior margin. The entire surface is covered with fine, blonde setae.
</paragraph>
<paragraph id="8BC43697FFFAFFDF5AF6651FFDB6F801" blockId="13.[151,1437,690,1975]" pageId="13" pageNumber="268">Clypeus trapezoidal, with shallow and large reticulate punctures; ratio width/length 2.04. Anterior angles rounded. Frons densely punctate, flat, gradually curving towards vertex (no boundary clearly visible between the two parts). Ocular canthum long, thin, and rounded at apex. Interocular ratio (interocular width/width of eye): 3.03. Antenna: ratio funiculus/club 0.69.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF662B5FBADFF40" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">Pronotum trapezoidal, width 1.58 times length. Lateral margins forming an obtuse angle at 1/3 of pronotum length. Anterior angles right and sharp, posterior angles obtuse and rounded. Basal margin thin and complete but partially obliterated in the middle, sinuate. Entire surface with deep punctures.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF66329FEB4FE88" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">Scutellum subpentagonal to subtriangular in shape, with rounded sides, surface finely punctate. Ratio width/ length 1.47.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF66361FC95FE1B" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">Elytra with striae defined by shallow grooves, surface with punctures coalescent. On light bands, dark spots show the striae-defining punctures and sparse punctures in the interstices. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF6639AFE00FE43" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">Pygidium finely granulate, with all setae equal in size. Triangular in shape. Slightly convex in lateral view. Ratio width/length 1.94.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF663D2FC41FD8B" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
Metasternal disc slightly sulcated, densely setose. Space between the mesocoxae narrow, slightly convex, width
<quantity id="4C839B72FFF9FFDC5AEC6009FEECFD8B" box="[221,324,548,572]" metricMagnitude="-4" metricUnit="m" metricValue="1.7" pageId="14" pageNumber="269" unit="mm" value="0.17">0.17 mm</quantity>
. Mesometasternal suture well defined at base of mesocoxae.
</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF6606AFB9DFD33" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">Abdominal sternites with numerous (5–10) irregular transverse rows of setae, and elongate punctures that fuse at sides. Last sternite strigate, with apical margin well defined and slightly sinuate.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF660A2FABDFD58" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
Protibia with 2 teeth (
<figureCitation id="13402A12FFF9FFDC5B8C60A2FDA1FD10" box="[445,521,655,679]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,269,1156,1180]" captionTargetBox="[174,1395,662,1128]" captionTargetId="figure@5.[151,1436,658,1144]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Shape of protibia in 7: Anomala eucoma (paralectotype, Las Mercedes, Guatemala); 8: A. flavacoma (3 holotype, Ƥ Hitoy Cerere, Costa Rica); 9: A. megaparamera (3 holotype, Ƥ paratype, Cuatro Esquinas, Costa Rica); 10: A. pseudoeucoma (3 holotype, Ƥ paratype, Reserva de San Ramón, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223580/files/figure.png" pageId="14" pageNumber="269">Fig.10</figureCitation>
), apical tooth long and slightly curved; second tooth at the same level of internal apex of protibia, triangular in shape, obtuse. Metatibia stout, slightly narrower subapically. Ratio length/width 3.97. First carina poorly defined by isolated spines. Surface punctate above second carina and rugose below.
</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF660D1FB19FC80" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">Protarsal claws: external claw strongly curved; ratio length/height: 1.82; internal claw bifurcate, with upper branch slightly shorter and more than half the width of the lower one. Inferior margin sinuate.</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF66169FEB5FC70" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
Aedeagus (
<figureCitation id="13402A12FFF9FFDC5B786169FE70FCEB" box="[329,472,836,860]" captionStart="FIGURES 11 – 25. 11 – 16" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[286,1306,190,1615]" captionTargetId="figure@6.[276,1317,176,1615]" captionTargetPageId="6" captionText="FIGURES 11 – 25. 11 – 16. Aedeagus of A. eucoma (paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado. 17 – 19. Aedeagus of Anomala flavacoma (paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect. 20 – 22. Aedeagus of Anomala megaparamera (holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect. 23 – 25. Aedeagus of Anomala pseudoeucoma (paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm." httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="14" pageNumber="269">Figs. 23–25</figureCitation>
): parameres with blunt apex, protruding from the strongly sinuate ventral margin. Endophallus (
<figureCitation id="13402A12FFF9FFDC5B05614AFE20FCC8" box="[308,392,871,895]" captionStart="FIGURES 26 – 32. 26 – 29" captionStartId="7.[151,269,1798,1822]" captionTargetBox="[279,1291,205,1753]" captionTargetId="figure@7.[254,1296,194,1767]" captionTargetPageId="7" captionText="FIGURES 26 – 32. 26 – 29. Endophallus of Anomala eucoma showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica. 30. Endophallus of Anomala flavacoma (holotype). 31. Endophallus of Anomala megaparamera (holotype). 32. Endophallus of Anomala pseudoeucoma (holotype). Scale = 1 mm. The arrows show the position of the gonopore." httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="14" pageNumber="269">Fig. 32</figureCitation>
) composed by a long ventral sacculus with a ridge of thick spines on dorsal margin, and a swollen shorter dorsal sacculus with a wide patch of sclerotized setae on left side. Gonopore ventral, between the two sacculi.
</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF661F9FAFAFB83" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
Female: similar to male, but the antennal club is slightly shorter than in males. Apical tooth of protibia (
<figureCitation id="13402A12FFF9FFDC5F5F61F9FF1CFBB8" captionStart="FIGURES 7 – 10" captionStartId="5.[151,269,1156,1180]" captionTargetBox="[174,1395,662,1128]" captionTargetId="figure@5.[151,1436,658,1144]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Shape of protibia in 7: Anomala eucoma (paralectotype, Las Mercedes, Guatemala); 8: A. flavacoma (3 holotype, Ƥ Hitoy Cerere, Costa Rica); 9: A. megaparamera (3 holotype, Ƥ paratype, Cuatro Esquinas, Costa Rica); 10: A. pseudoeucoma (3 holotype, Ƥ paratype, Reserva de San Ramón, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223580/files/figure.png" pageId="14" pageNumber="269">Fig. 10</figureCitation>
) wider and longer than in males; second tooth above internal apex. Internal protarsal claw narrower than in males, inferior margin straight. Metatibia stouter than in males. Last abdominal sternite with curved margin.
</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF66612FB15FB5F" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
<emphasis id="B90FEA85FFF9FFDC5AF66612FE96FBEF" bold="true" box="[199,318,1087,1112]" pageId="14" pageNumber="269">Variation:</emphasis>
Underside from reddish brown to dark brown; elytra dark brown to bluish black; in some specimens with homogeneous colour (without the basal lighter band), and rarely, with a barely distinguishable median band. In some specimens, particularly the darker ones, the denser and thicker setae give the pronotum a velvety blond appearance. Second tooth of protibia at the same level or slightly above internal apex of protibia. On endophallus, the patch of setae on dorsal sacculus extends further frontally in some specimens.
</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF666D9FDF2FACF" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
Body length
<quantity id="4C839B72FFF9FFDC5B5D66D9FD9FFABB" box="[364,567,1268,1292]" metricMagnitude="-2" metricUnit="m" metricValue="1.1985" metricValueMax="1.263" metricValueMin="1.134" pageId="14" pageNumber="269" unit="mm" value="11.985" valueMax="12.63" valueMin="11.34">11.34–12.63 mm</quantity>
, body width
<quantity id="4C839B72FFF9FFDC58D766D9FC3EFABB" box="[742,918,1268,1292]" metricMagnitude="-3" metricUnit="m" metricValue="6.885" metricValueMax="7.2" metricValueMin="6.57" pageId="14" pageNumber="269" unit="mm" value="6.885" valueMax="7.2" valueMin="6.57">6.57–7.20 mm</quantity>
. Clypeus w/l: 1.81–2.10. Interocular ratio (interocular width/width of eye): 2.94–3.45. Antenna: ratio funiculus/club 0.69–0.71. Pronotum w/l: 1.58–1.65. Scutellum w/l: 1.26–1.47. Pygidium w/l: 1.65–1.94. Width between mesocoxae:
<quantity id="4C839B72FFF9FFDC5E736711FB44FAE3" box="[1090,1260,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" metricValueMax="2.4" metricValueMin="1.6" pageId="14" pageNumber="269" unit="mm" value="0.2" valueMax="0.24" valueMin="0.16">0.16–0.24 mm</quantity>
. Metatibia w/l: 2.79–3.97. External claw l/h: 1.37–1.82.
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFF9FFDC5AF667AEFBC0F8B3" pageId="14" pageNumber="269" type="diagnosis">
<paragraph id="8BC43697FFF9FFDC5AF667AEFAA5F9BF" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
<emphasis id="B90FEA85FFF9FFDC5AF667AEFEE9FA2B" bold="true" box="[199,321,1411,1436]" pageId="14" pageNumber="269">Diagnosis.</emphasis>
<taxonomicName id="4C7B4D14FFF9FFDC5B7967A9FDF2FA2B" box="[328,602,1412,1436]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFF9FFDC5B7967A9FDF2FA2B" box="[328,602,1412,1436]" italics="true" pageId="14" pageNumber="269">Anomala pseudoeucoma</emphasis>
</taxonomicName>
can be distinguished from the other species described here by its dark and uniform brown colour, median size, frons nearly continuous with apex, first carina of metatibia obliterated, surface of metatibia punctate above second carina, parameres with strongly sinuate inferior margin, and endophallus with two patches of sclerotized spines and setae that can be distinguished even in the inverted endophallus.
</paragraph>
<paragraph id="8BC43697FFF9FFDC5AF66439FBC0F8B3" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
This species is very similar to
<taxonomicName id="4C7B4D14FFF9FFDC580D6439FD1FF99B" box="[572,695,1556,1580]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF9FFDC580D6439FD1FF99B" box="[572,695,1556,1580]" italics="true" pageId="14" pageNumber="269">A. eucoma</emphasis>
</taxonomicName>
, especially in the lighter specimens;
<taxonomicName id="4C7B4D14FFF9FFDC5E466439FAEAF99B" box="[1143,1346,1556,1580]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFF9FFDC5E466439FAEAF99B" box="[1143,1346,1556,1580]" italics="true" pageId="14" pageNumber="269">A. pseudoeucoma</emphasis>
</taxonomicName>
can be distinguished by the following characters: shorter and less curved apical tooth in male protibia; large internal claw in males, with inferior border more strongly sinuate; narrower intermesocoxal space, and less convex than in
<taxonomicName id="4C7B4D14FFF9FFDC5FB56471FF46F92F" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF9FFDC5FB56471FF46F92F" italics="true" pageId="14" pageNumber="269">A. eucoma</emphasis>
</taxonomicName>
. As in the case of
<taxonomicName id="4C7B4D14FFF9FFDC5B8D64ADFD9BF92F" box="[444,563,1664,1688]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF9FFDC5B8D64ADFD9BF92F" box="[444,563,1664,1688]" italics="true" pageId="14" pageNumber="269">A. eucoma</emphasis>
</taxonomicName>
, the brown and trapezoidal pronotum easily distinguishes this species from
<taxonomicName id="4C7B4D14FFF9FFDC5FB564ADFEB2F90B" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="amphicoma">
<emphasis id="B90FEA85FFF9FFDC5FB564ADFEB2F90B" italics="true" pageId="14" pageNumber="269">A. amphicoma</emphasis>
</taxonomicName>
.
<taxonomicName id="4C7B4D14FFF9FFDC5B1B6489FDA6F90B" box="[298,526,1700,1724]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFF9FFDC5B1B6489FDA6F90B" box="[298,526,1700,1724]" italics="true" pageId="14" pageNumber="269">Anomala flavacoma</emphasis>
</taxonomicName>
is smaller and much lighter in colour, with marked sides of frons with vertex, metatibia with oblong punctures above second carina.
<taxonomicName id="4C7B4D14FFF9FFDC58C664E5FBA6F957" box="[759,1038,1736,1760]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFF9FFDC58C664E5FBA6F957" box="[759,1038,1736,1760]" italics="true" pageId="14" pageNumber="269">Anomala megaparamera</emphasis>
</taxonomicName>
is much smaller in size, with larger parameres and endophallus with diverticles and setae less sclerotized and more diffuse.
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFF9FFDC5AF66522FCE9F8C7" pageId="14" pageNumber="269" type="distribution">
<paragraph id="8BC43697FFF9FFDC5AF66522FCE9F8C7" blockId="14.[151,1437,151,1940]" pageId="14" pageNumber="269">
<emphasis id="B90FEA85FFF9FFDC5AF66522FEC8F89F" bold="true" box="[199,352,1807,1832]" pageId="14" pageNumber="269">Distribution.</emphasis>
Widely distributed, with localities recorded in Guanacaste and the Tilarán mountain ranges to the north, the Talamanca Mountain Range near the border with
<collectingCountry id="F36C7607FFF9FFDC59B06518FC72F8FB" box="[897,986,1845,1868]" name="Panama" pageId="14" pageNumber="269">Panama</collectingCountry>
to the south, and Caribbean lowlands. Altitudinal range varies from 0 m to above
<quantity id="4C839B72FFF9FFDC584D6575FD79F8C7" box="[636,721,1880,1904]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="269" unit="m" value="1500.0">1500 m</quantity>
(
<figureCitation id="13402A12FFF9FFDC58D16575FC9CF8C7" box="[736,820,1880,1904]" captionStart="FIGURE 36" captionStartId="15.[151,255,1195,1219]" captionTargetBox="[163,1411,216,1140]" captionTargetId="figure@15.[151,1436,194,1173]" captionTargetPageId="15" captionText="FIGURE 36. Distribution of Anomala pseudoeucoma." httpUri="https://zenodo.org/record/223586/files/figure.png" pageId="14" pageNumber="269">Fig. 36</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFF9FFDC5AF66551FB68F823" box="[199,1216,1916,1940]" pageId="14" pageNumber="269" type="etymology">
<paragraph id="8BC43697FFF9FFDC5AF66551FB68F823" blockId="14.[151,1437,151,1940]" box="[199,1216,1916,1940]" pageId="14" pageNumber="269">
<emphasis id="B90FEA85FFF9FFDC5AF66551FEE6F823" bold="true" box="[199,334,1916,1940]" pageId="14" pageNumber="269">Etymology.</emphasis>
From Greek &quot;pseudo-&quot;, false, and &quot;
<taxonomicName id="4C7B4D14FFF9FFDC58ED6551FC93F823" box="[732,827,1916,1940]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="eucoma">eucoma</taxonomicName>
&quot;, for its similarity with
<taxonomicName id="4C7B4D14FFF9FFDC5E726551FB11F823" box="[1091,1209,1916,1940]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="269" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF9FFDC5E726551FB11F823" box="[1091,1209,1916,1940]" italics="true" pageId="14" pageNumber="269">A. eucoma</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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