197 lines
20 KiB
XML
197 lines
20 KiB
XML
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<mods:title id="220707561DE456A843C62F0638CF956B">Taxonomy of the subgenus Quedius (Raphirus) Stephens (Coleoptera: Staphylinidae: Staphylinini: Quediina) with descriptions of four new species from China</mods:title>
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<mods:namePart id="38907562E329978D5045B65526082051">Cai, Yan-Peng</mods:namePart>
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<mods:namePart id="B27F503B0D78CC2884BD839B8C99493A">Zhou, Hong-Zhang</mods:namePart>
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<mods:title id="E6875DCCF20D64B50146589C3BFE78B2">Zootaxa</mods:title>
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<treatment id="03CC2578FFBA1458FF04F28BFEB7E783" ID-DOI="http://doi.org/10.5281/zenodo.6115645" ID-GBIF-Taxon="119638131" ID-Zenodo-Dep="6115645" LSID="urn:lsid:plazi:treatment:03CC2578FFBA1458FF04F28BFEB7E783" httpUri="http://treatment.plazi.org/id/03CC2578FFBA1458FF04F28BFEB7E783" lastPageId="18" lastPageNumber="169" pageId="15" pageNumber="166">
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<paragraph id="8BDA946EFFBA1445FF04F28BFDD6E3B1" blockId="15.[151,638,1412,1471]" box="[151,638,1412,1438]" pageId="15" pageNumber="166">
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<heading id="D0922302FFBA1445FF04F28BFDD6E3B1" bold="true" box="[151,638,1412,1438]" fontSize="11" level="1" pageId="15" pageNumber="166" reason="1">
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<taxonomicName id="4C65EFEDFFBA1445FF04F28BFDD6E3B1" ID-CoL="4R38T" authority="Smetana, 2011" authorityName="Smetana" authorityYear="2011" box="[151,638,1412,1438]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="oros" subGenus="Raphirus">
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<emphasis id="B911487CFFBA1445FF04F28BFDD6E3B1" bold="true" box="[151,638,1412,1438]" pageId="15" pageNumber="166">
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<emphasis id="B911487CFFBA1445FF04F28BFF54E3B1" bold="true" box="[151,252,1412,1438]" italics="true" pageId="15" pageNumber="166">Quedius</emphasis>
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(
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<emphasis id="B911487CFFBA1445FE9EF28BFED5E3B1" bold="true" box="[269,381,1412,1438]" italics="true" pageId="15" pageNumber="166">Raphirus</emphasis>
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)
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<emphasis id="B911487CFFBA1445FE1DF28BFE6AE3B1" bold="true" box="[398,450,1412,1438]" italics="true" pageId="15" pageNumber="166">oros</emphasis>
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<bibRefCitation id="EFF4E99FFFBA1445FE59F28BFDD6E3B1" author="Smetana" box="[458,638,1412,1438]" pageId="15" pageNumber="166" refString="Smetana, A. (2011) Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae, 51 (1), 145 - 156." type="journal article" year="2011">Smetana, 2011</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BDA946EFFBA1445FF04F2A9FE81E390" blockId="15.[151,638,1412,1471]" box="[151,297,1446,1471]" pageId="15" pageNumber="166">
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(
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<figureCitation id="135E88EBFFBA1445FF0CF2A9FEB7E390" box="[159,287,1446,1471]" captionStart="FIGURE 6" captionStartId="16.[151,250,1877,1899]" captionTargetBox="[180,1433,194,1855]" captionTargetId="figure@16.[151,1436,193,1856]" captionTargetPageId="16" captionText="FIGURE 6. Quedius (Raphirus) oros Smetana, 2011: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245451/files/figure.png" pageId="15" pageNumber="166">Figs. 6, 6-1</figureCitation>
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)
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</paragraph>
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<paragraph id="8BDA946EFFBA1445FF04F2E1FE00E00C" blockId="15.[151,1436,1518,1571]" pageId="15" pageNumber="166">
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<bibRefCitation id="EFF4E99FFFBA1445FF04F2E1FE9BE02B" author="Smetana" box="[151,307,1518,1540]" pageId="15" pageNumber="166" refString="Smetana, A. (2011) Contributions to the knowledge of the Quediina of China (Coleoptera, Staphylinidae, Staphylinini). Part 41. Genus Quedius Stephens, 1829. Subgenus Raphirus Stephens, 1829. Section 9. Acta Entomologica Musei Nationalis Pragae, 51 (1), 145 - 156." type="journal article" year="2011">Smetana, 2011</bibRefCitation>
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: 150 (
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<taxonomicName id="4C65EFEDFFBA1445FEEAF2E1FE65E02B" box="[377,461,1518,1540]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
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<emphasis id="B911487CFFBA1445FEEAF2E1FE65E02B" box="[377,461,1518,1540]" italics="true" pageId="15" pageNumber="166">Quedius</emphasis>
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</taxonomicName>
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; subgenus
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<taxonomicName id="4C65EFEDFFBA1445FDD5F2E1FD0BE02B" box="[582,675,1518,1540]" class="Insecta" family="Staphylinidae" genus="Raphirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
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<emphasis id="B911487CFFBA1445FDD5F2E1FD0BE02B" box="[582,675,1518,1540]" italics="true" pageId="15" pageNumber="166">Raphirus</emphasis>
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</taxonomicName>
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||
;
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<typeStatus id="54DE2ACCFFBA1445FD27F2E0FD4EE02B" box="[692,742,1519,1540]" pageId="15" pageNumber="166">Type</typeStatus>
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locality:
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<collectingCountry id="F372D4FEFFBA1445FCDEF2E1FC24E02B" box="[845,908,1518,1540]" name="China" pageId="15" pageNumber="166">China</collectingCountry>
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, W Sichuan, pass SE Barkam, between Zhuokeji- Lianghekou,
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<quantity id="4C9D398BFFBA1445FEDFF102FE34E00C" box="[332,412,1549,1571]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="15" pageNumber="166" unit="m" value="4100.0">4100 m</quantity>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C37FC7E5FFBA1445FF04F140FB7CE1A8" pageId="15" pageNumber="166" type="materials_examined">
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<paragraph id="8BDA946EFFBA1445FF04F140FB7CE1A8" blockId="15.[151,1437,1615,2001]" pageId="15" pageNumber="166">
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<emphasis id="B911487CFFBA1445FF04F140FDC2E047" bold="true" box="[151,618,1615,1640]" pageId="15" pageNumber="166">
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Material examined.
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<collectingCountry id="F372D4FEFFBA1445FE06F140FE5FE047" box="[405,503,1615,1640]" name="China" pageId="15" pageNumber="166">CHINA</collectingCountry>
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: Sichuan
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||
</emphasis>
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Province:
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||
<specimenCount id="9D635FE7FFBA1445FD60F15FFC8EE048" box="[755,806,1616,1640]" pageId="15" pageNumber="166" type="generic">1 ♂</specimenCount>
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||
, 1 ♀, Wolong National Nature Reserve, Dengsheng Conservation Station,
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||
<quantity id="4C9D398BFFBA1445FE00F17BFE44E0A3" box="[403,492,1652,1677]" metricMagnitude="3" metricUnit="m" metricValue="2.735" pageId="15" pageNumber="166" unit="m" value="2735.0">2735 m</quantity>
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,
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<date id="FFDBB2AEFFBA1445FE6AF17BFD08E0A3" box="[505,672,1652,1676]" pageId="15" pageNumber="166" value="2003-08-24">24. VIII. 2003</date>
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||
, coll. Caiyun Zhao;
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||
<specimenCount id="9D635FE7FFBA1445FC1AF17BFC1FE0A3" box="[905,951,1652,1676]" pageId="15" pageNumber="166" type="generic">1 ♂</specimenCount>
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||
, Wolong National Nature Reserve, Balang Mountain,
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||
<quantity id="4C9D398BFFBA1445FE87F198FEC5E080" box="[276,365,1687,1712]" metricMagnitude="3" metricUnit="m" metricValue="3.4549999999999996" pageId="15" pageNumber="166" unit="m" value="3455.0">3455 m</quantity>
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||
,
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||
<date id="FFDBB2AEFFBA1445FEE9F197FDB5E09F" box="[378,541,1688,1712]" pageId="15" pageNumber="166" value="2004-05-07" valueMax="2004-05-10" valueMin="2004-05-07">7–10. V. 2004</date>
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||
, coll. Xiaodong Yu;
|
||
<specimenCount id="9D635FE7FFBA1445FC98F197FC91E080" box="[779,825,1688,1712]" pageId="15" pageNumber="166" type="generic">1 ♂</specimenCount>
|
||
, Wolong National Nature Reserve, Balang Mountain,
|
||
<quantity id="4C9D398BFFBA1445FF04F1B3FF46E0FC" box="[151,238,1724,1748]" metricMagnitude="3" metricUnit="m" metricValue="3.5799999999999996" pageId="15" pageNumber="166" unit="m" value="3580.0">3580 m</quantity>
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||
,
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||
<date id="FFDBB2AEFFBA1445FF69F1B3FE32E0FB" box="[250,410,1724,1748]" pageId="15" pageNumber="166" value="2004-05-07" valueMax="2004-05-10" valueMin="2004-05-07">7–10. V. 2004</date>
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||
, coll. Xiaodong Yu; 1 ♀, Wolong National Nature Reserve, Balang Mountain,
|
||
<quantity id="4C9D398BFFBA1445FA9CF1B3FACEE0FC" box="[1295,1382,1724,1748]" metricMagnitude="3" metricUnit="m" metricValue="3.35" pageId="15" pageNumber="166" unit="m" value="3350.0">3350 m</quantity>
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,
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<date id="FFDBB2AEFFBA1445FAE1F1B3FE8FE0D7" pageId="15" pageNumber="166" value="2004-06-20" valueMax="2004-06-23" valueMin="2004-06-20">20– 23. VI. 2004</date>
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, coll. Xiaodong Yu; 1 ♀, Wolong National Nature Reserve, Balang Mountain,
|
||
<quantity id="4C9D398BFFBA1445FB04F1EFFB46E0D8" box="[1175,1262,1760,1784]" metricMagnitude="3" metricUnit="m" metricValue="3.5799999999999996" pageId="15" pageNumber="166" unit="m" value="3580.0">3580 m</quantity>
|
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,
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<date id="FFDBB2AEFFBA1445FB69F1EFFA30E0D7" box="[1274,1432,1760,1784]" pageId="15" pageNumber="166" value="2004-06-05" valueMax="2004-06-08" valueMin="2004-06-05">5–8. VI. 2004</date>
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, coll. Xiaodong Yu;
|
||
<specimenCount id="9D635FE7FFBA1445FE16F00BFE1EE133" box="[389,438,1796,1820]" pageId="15" pageNumber="166" type="generic">1 ♂</specimenCount>
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, Wolong National Nature Reserve, Balang Mountain,
|
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<quantity id="4C9D398BFFBA1445FBD3F00BFB35E133" box="[1088,1181,1796,1820]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="166" unit="m" value="3800.0">3800 m</quantity>
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,
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<date id="FFDBB2AEFFBA1445FB3CF00BFAF2E133" box="[1199,1370,1796,1821]" pageId="15" pageNumber="166" value="2004-06-05" valueMax="2004-06-08" valueMin="2004-06-05">5–8. VI. 2004</date>
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, coll. Xiaodong Yu; 1 ♀, Wolong National Nature Reserve, Balang Mountain,
|
||
<quantity id="4C9D398BFFBA1445FC58F027FB8AE110" box="[971,1058,1832,1856]" metricMagnitude="3" metricUnit="m" metricValue="3.17" pageId="15" pageNumber="166" unit="m" value="3170.0">3170 m</quantity>
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,
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<date id="FFDBB2AEFFBA1445FBBDF027FB42E16F" box="[1070,1258,1832,1856]" pageId="15" pageNumber="166" value="2004-06-20" valueMax="2004-06-23" valueMin="2004-06-20">20–23. VI. 2004</date>
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, coll. Xiaodong Yu;
|
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<specimenCount id="9D635FE7FFBA1445FF55F043FF46E14B" box="[198,238,1868,1892]" pageId="15" pageNumber="166" type="generic">1 ♂</specimenCount>
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, Wolong National Nature Reserve, Balang Mountain,
|
||
<quantity id="4C9D398BFFBA1445FCCFF043FC1CE14C" box="[860,948,1868,1892]" metricMagnitude="3" metricUnit="m" metricValue="3.4549999999999996" pageId="15" pageNumber="166" unit="m" value="3455.0">3455 m</quantity>
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,
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<date id="FFDBB2AEFFBA1445FC52F043FBD6E14B" box="[961,1150,1868,1892]" pageId="15" pageNumber="166" value="2004-06-20" valueMax="2004-06-23" valueMin="2004-06-20">20–23. VI. 2004</date>
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, coll. Xiaodong Yu; 1 ♀, Wolong National Nature Reserve, Balang Mountain,
|
||
<quantity id="4C9D398BFFBA1445FD79F07FFCE9E1A8" box="[746,833,1904,1928]" metricMagnitude="3" metricUnit="m" metricValue="3.8" pageId="15" pageNumber="166" unit="m" value="3800.0">3800 m</quantity>
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,
|
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<date id="FFDBB2AEFFBA1445FCDEF060FC5DE1A7" box="[845,1013,1903,1928]" pageId="15" pageNumber="166" value="2004-07-05" valueMax="2004-07-08" valueMin="2004-07-05">5–8. VII. 2004</date>
|
||
, coll. Xiaodong Yu.
|
||
</paragraph>
|
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</subSubSection>
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<subSubSection id="C37FC7E5FFBA1458FF54F09BFCBDE733" lastPageId="18" lastPageNumber="169" pageId="15" pageNumber="166" type="description">
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<paragraph id="8BDA946EFFBA1445FF54F09BFB69E1FF" blockId="15.[151,1437,1615,2001]" pageId="15" pageNumber="166">
|
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<emphasis id="B911487CFFBA1445FF54F09BFED5E182" bold="true" box="[199,381,1940,1965]" pageId="15" pageNumber="166">Measurements.</emphasis>
|
||
BL = 6.0 mm, BW =
|
||
<quantity id="4C9D398BFFBA1445FDE8F09BFD7CE183" box="[635,724,1940,1964]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="15" pageNumber="166" unit="mm" value="1.1">1.1 mm</quantity>
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||
, HL/PL/EL = 1.00: 1.22: 1.14, HW/PW/EW/AW = 1.00: 1.12: 1.20: 1.29, HW/HL = 1.12, HEL/HTL = 10.36, PW/PL = 1.03, EW/EL = 1.18, ESL/EL = 0.48.
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||
</paragraph>
|
||
<caption id="DF1AC4E6FFA5145AFF04F05AFA99E1A5" httpUri="https://zenodo.org/record/245451/files/figure.png" pageId="16" pageNumber="167" targetBox="[180,1433,194,1855]" targetPageId="16">
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<paragraph id="8BDA946EFFA5145AFF04F05AFA99E1A5" blockId="16.[151,1436,1877,1930]" pageId="16" pageNumber="167">
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||
<emphasis id="B911487CFFA5145AFF04F05AFEBEE145" bold="true" box="[151,278,1877,1899]" pageId="16" pageNumber="167">FIGURE 6.</emphasis>
|
||
<taxonomicName id="4C65EFEDFFA5145AFEB2F05AFD62E144" authority="Smetana, 2011" authorityName="Smetana" authorityYear="2011" box="[289,714,1877,1899]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="167" phylum="Arthropoda" rank="species" species="oros" subGenus="Raphirus">
|
||
<emphasis id="B911487CFFA5145AFEB2F05AFEDDE144" box="[289,373,1877,1899]" italics="true" pageId="16" pageNumber="167">Quedius</emphasis>
|
||
(
|
||
<emphasis id="B911487CFFA5145AFE1BF05AFE4DE144" box="[392,485,1877,1899]" italics="true" pageId="16" pageNumber="167">Raphirus</emphasis>
|
||
)
|
||
<emphasis id="B911487CFFA5145AFE64F059FD8BE144" box="[503,547,1878,1899]" italics="true" pageId="16" pageNumber="167">oros</emphasis>
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||
Smetana, 2011
|
||
</taxonomicName>
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||
: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses).
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||
</paragraph>
|
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</caption>
|
||
<caption id="DF1AC4E6FFA4145BFF04F0AEFD31E1DB" httpUri="https://zenodo.org/record/245452/files/figure.png" pageId="17" pageNumber="168" targetBox="[181,1406,193,1931]" targetPageId="17">
|
||
<paragraph id="8BDA946EFFA4145BFF04F0AEFD31E1DB" blockId="17.[151,1436,1953,2036]" pageId="17" pageNumber="168">
|
||
<emphasis id="B911487CFFA4145BFF04F0AEFE83E199" bold="true" box="[151,299,1953,1975]" pageId="17" pageNumber="168">FIGURE 6-1.</emphasis>
|
||
<taxonomicName id="4C65EFEDFFA4145BFEA6F0AEFD77E198" authority="Smetana, 2011" authorityName="Smetana" authorityYear="2011" box="[309,735,1953,1975]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="oros" subGenus="Raphirus">
|
||
<emphasis id="B911487CFFA4145BFEA6F0AEFE21E198" box="[309,393,1953,1975]" italics="true" pageId="17" pageNumber="168">Quedius</emphasis>
|
||
(
|
||
<emphasis id="B911487CFFA4145BFE0FF0AEFE51E198" box="[412,505,1953,1975]" italics="true" pageId="17" pageNumber="168">Raphirus</emphasis>
|
||
)
|
||
<emphasis id="B911487CFFA4145BFD9FF0ADFD90E198" box="[524,568,1954,1975]" italics="true" pageId="17" pageNumber="168">oros</emphasis>
|
||
Smetana, 2011
|
||
</taxonomicName>
|
||
: A. male sternite VIII; B. male sternite IX; C. male tergite X; D. underside of paramere; E. apical portion of median lobe, parameral view; F. aedeagus, lateral view; G. aedeagus, parameral view; H. female tergite X. (Scale bars = 0.20 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA946EFFA71458FF54F798FCBDE733" blockId="18.[151,1436,151,429]" pageId="18" pageNumber="169">
|
||
<emphasis id="B911487CFFA71458FF54F798FE7CE69F" bold="true" box="[199,468,151,176]" pageId="18" pageNumber="169">Description of female.</emphasis>
|
||
Female with first four segments of foretarsus simple, not dilated; tergite VIII with basal ridge complete, slightly sinuate, with three long setae on each side; sternite VIII with basal ridge complete, nearly straight, with two long setae on each side; tergite X (
|
||
<figureCitation id="135E88EBFFA71458FCCEF7D0FC1FE6D7" box="[861,951,223,248]" captionStart="FIGURE 6" captionStartId="16.[151,250,1877,1899]" captionTargetBox="[180,1433,194,1855]" captionTargetId="figure@16.[151,1436,193,1856]" captionTargetPageId="16" captionText="FIGURE 6. Quedius (Raphirus) oros Smetana, 2011: A. head (1.0 mm); B. pronotum (1.0 mm); C. elytra (1.0 mm); D. underside of paramere (0.2 mm); E. apical portion of median lobe, parameral view (0.2 mm). (Scale in parentheses)." httpUri="https://zenodo.org/record/245451/files/figure.png" pageId="18" pageNumber="169">Fig. 6-1</figureCitation>
|
||
H) with basal side broadly and shallowly concave, apical portion nearly evenly converged to apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37FC7E5FFA71458FF54F628FEB7E783" pageId="18" pageNumber="169" type="distribution">
|
||
<paragraph id="8BDA946EFFA71458FF54F628FEB7E783" blockId="18.[151,1436,151,429]" pageId="18" pageNumber="169">
|
||
<emphasis id="B911487CFFA71458FF54F628FEC8E76F" bold="true" box="[199,352,295,320]" pageId="18" pageNumber="169">Distribution.</emphasis>
|
||
<taxonomicName id="4C65EFEDFFA71458FEFFF627FDDBE76F" authority="Smetana" authorityName="Smetana" box="[364,627,296,320]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="oros">
|
||
<emphasis id="B911487CFFA71458FEFFF627FDAFE76F" box="[364,519,296,320]" italics="true" pageId="18" pageNumber="169">Quedius oros</emphasis>
|
||
Smetana
|
||
</taxonomicName>
|
||
is known only from high mountains in central Sichuan Province at altitudes between
|
||
<quantity id="4C9D398BFFA71458FEE2F643FE67E74B" box="[369,463,332,357]" metricMagnitude="3" metricUnit="m" metricValue="2.735" pageId="18" pageNumber="169" unit="m" value="2735.0">2735 m</quantity>
|
||
and
|
||
<quantity id="4C9D398BFFA71458FD86F643FDDEE74B" box="[533,630,332,356]" metricMagnitude="3" metricUnit="m" metricValue="4.1" pageId="18" pageNumber="169" unit="m" value="4100.0">4100 m</quantity>
|
||
. The examined specimens from Wolong National Nature Reserve, Dengsheng Conservation Station were collected from mushrooms by hand near a creek, others were collected using pitfall traps.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |