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<mods:title id="7E74FD73AC8BFB17BE2386A19E61C1F0">Taxonomy of Serpulidae (Annelida, Polychaeta): The state of affairs</mods:title>
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<paragraph id="8BAB36EEC949FFFE7E93F8941C491714" blockId="54.[151,512,1868,1929]" box="[151,512,1868,1894]" pageId="54" pageNumber="55">
<heading id="D0E38182C949FFFE7E93F8941C491714" bold="true" box="[151,512,1868,1894]" fontSize="11" level="1" pageId="54" pageNumber="55" reason="1">
<emphasis id="B960EAFCC949FFFE7E93F8941C491714" bold="true" box="[151,512,1868,1894]" pageId="54" pageNumber="55">
17.
<taxonomicName id="4C144D6DC949FFFE7EC7F8941C491714" ID-CoL="84QSJ" authority="Saint-Joseph, 1894" authorityName="Saint-Joseph" authorityYear="1894" box="[195,512,1868,1894]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="54" pageNumber="55" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC949FFFE7EC7F8941F461714" bold="true" box="[195,271,1868,1894]" italics="true" pageId="54" pageNumber="55">Janita</emphasis>
<bibRefCitation id="EF854B1FC949FFFE7F12F8941C491714" author="Saint-Joseph, A. de" box="[278,512,1868,1894]" pageId="54" pageNumber="55" pagination="1 - 395" refId="ref67052" refString="Saint-Joseph, A. de (1894) Les annelides polychetes des Cotes de Dinard. Pt. 3. Annales des Sciences Naturelles (Zoologie et Paleontologie, serie 7), 17, 1 - 395." type="journal article" year="1894">Saint-Joseph, 1894</bibRefCitation>
</taxonomicName>
</emphasis>
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</paragraph>
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<paragraph id="8BAB36EEC949FFFE7E93F8B71EB417FB" blockId="54.[151,512,1868,1929]" box="[151,253,1903,1929]" pageId="54" pageNumber="55">
(
<figureCitation id="132F2A6BC949FFFE7E9BF8B71EBC17FB" box="[159,245,1903,1929]" captionStart="FIGURE 24" captionStartId="55.[151,255,1669,1693]" captionTargetBox="[163,1427,216,1621]" captionTargetId="figure-147@55.[151,1436,197,1645]" captionTargetPageId="55" captionText="FIGURE 24. SEM micrographs of chaetae in SEM micrographs of Janita fimbriata. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—Spirobranchus type collar chaeta, C—Apomatus chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini." pageId="54" pageNumber="55">Fig. 24</figureCitation>
)
</paragraph>
<paragraph id="8BAB36EEC949FFFE7E93F8641F331781" blockId="54.[151,1381,1980,2035]" pageId="54" pageNumber="55">
<emphasis id="B960EAFCC949FFFE7E93F8641F6F17A6" bold="true" box="[151,294,1980,2004]" pageId="54" pageNumber="55">Type-species</emphasis>
:
<taxonomicName id="4C144D6DC949FFFE7F30F8641CBB17A1" authority="Langerhans, 1884" authorityName="Langerhans" authorityYear="1884" box="[308,754,1980,2003]" class="Polychaeta" family="Serpulidae" genus="Omphalopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="54" pageNumber="55" phylum="Annelida" rank="species" species="spinosa">
<emphasis id="B960EAFCC949FFFE7F30F8641C6317A1" box="[308,554,1980,2003]" italics="true" pageId="54" pageNumber="55">Omphalopoma spinosa</emphasis>
<bibRefCitation id="EF854B1FC949FFFE7C35F8641CBB17A1" author="Langerhans, P." box="[561,754,1980,2003]" pageId="54" pageNumber="55" pagination="247 - 285" refId="ref63321" refString="Langerhans, P. (1884) Die Wurmfauna von Madeira. 4. Zeitschrift fur Wissenschaftliche Zoologie, 40, 247 - 285." type="journal article" year="1884">Langerhans, 1884</bibRefCitation>
</taxonomicName>
, = junior synonym of
<taxonomicName id="4C144D6DC949FFFE7DDBF8641AD217A1" authorityName="Delle Chiaje" authorityYear="1822" box="[991,1179,1980,2003]" class="Agaricomycetes" family="Serpulaceae" genus="Serpula" kingdom="Fungi" order="Boletales" pageId="54" pageNumber="55" phylum="Basidiomycota" rank="species" species="fimbriata">
<emphasis id="B960EAFCC949FFFE7DDBF8641AD217A1" box="[991,1179,1980,2003]" italics="true" pageId="54" pageNumber="55">Serpula fimbriata</emphasis>
</taxonomicName>
delle Chiaje, 1822 Number of species: 1
</paragraph>
<caption id="DF6B6666C948FFFF7E93F95D1A291690" pageId="55" pageNumber="56" startId="55.[151,255,1669,1693]" targetBox="[163,1427,216,1621]" targetPageId="55">
<paragraph id="8BAB36EEC948FFFF7E93F95D1A291690" blockId="55.[151,1436,1669,1762]" pageId="55" pageNumber="56">
<emphasis id="B960EAFCC948FFFF7E93F95D1F6E16EF" bold="true" box="[151,295,1669,1693]" pageId="55" pageNumber="56">FIGURE 24.</emphasis>
SEM micrographs of chaetae in SEM micrographs of
<taxonomicName id="4C144D6DC948FFFF7D6AF95D1A5216EE" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1822" box="[878,1051,1669,1692]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="55" pageNumber="56" phylum="Annelida" rank="species" species="fimbriata">
<emphasis id="B960EAFCC948FFFF7D6AF95D1A5216EE" box="[878,1051,1669,1692]" italics="true" pageId="55" pageNumber="56">Janita fimbriata</emphasis>
</taxonomicName>
. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—
<taxonomicName id="4C144D6DC948FFFF7D89F9701A6016CD" authorityName="de Blainville" authorityYear="1818" box="[909,1065,1704,1727]" class="Polychaeta" family="Serpulidae" genus="Spirobranchus" kingdom="Animalia" order="Sabellida" pageId="55" pageNumber="56" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC948FFFF7D89F9701A6016CD" box="[909,1065,1704,1727]" italics="true" pageId="55" pageNumber="56">Spirobranchus</emphasis>
</taxonomicName>
type collar chaeta, C—
<taxonomicName id="4C144D6DC948FFFF7B35F9701BD516CD" authorityName="Philippi" authorityYear="1844" box="[1329,1436,1704,1727]" class="Polychaeta" family="Serpulidae" genus="Apomatus" kingdom="Animalia" order="Sabellida" pageId="55" pageNumber="56" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC948FFFF7B35F9701BD516CD" box="[1329,1436,1704,1727]" italics="true" pageId="55" pageNumber="56">Apomatus</emphasis>
</taxonomicName>
chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini.
</paragraph>
</caption>
<paragraph id="8BAB36EEC948FFF07E93F8CC1FD2127C" blockId="55.[151,1436,1812,2031]" lastBlockId="56.[151,1436,152,1222]" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
Tube white, sub-circular in cross-section, with 5 longitudinal winding ridges. Granular overlay absent. Operculum bell-shaped, ending in simple thick brown concave endplate; opercular base surrounded by three fleshy processes, one triangular and two rounded ones, not unlike those figured for
<taxonomicName id="4C144D6DC948FFFF7A7DF8B91B3F1708" baseAuthorityName="Johnson" baseAuthorityYear="1901" box="[1145,1398,1889,1914]" class="Polychaeta" family="Serpulidae" genus="Crucigera" kingdom="Animalia" order="Sabellida" pageId="55" pageNumber="56" phylum="Annelida" rank="species" species="zygophora">
<emphasis id="B960EAFCC948FFFF7A7DF8B91B3F1708" box="[1145,1398,1889,1914]" italics="true" pageId="55" pageNumber="56">Crucigera zygophora</emphasis>
</taxonomicName>
by ten
<bibRefCitation id="EF854B1FC948FFFF7EC3F8501C6417D0" author="Hove, H. A. ten &amp; Jansen-Jacobs, M. J." box="[199,557,1928,1954]" pageId="55" pageNumber="56" pagination="143 - 180" refId="ref61117" refString="Hove, H. A. ten &amp; Jansen-Jacobs, M. J. (1984) A revision of the genus Crucigera (Polychaeta; Serpulidae); a proposed methodical approach to serpulids, with special reference to variation in Serpula and Hydroides. In: Hutchings, P. A. (Ed.) Proceedings of the First International Polychaete Conference, Sydney, Australia, July 1983. Linnean Society of New South Wales, Sydney, pp. 143 - 180." type="book chapter" year="1984">Hove &amp; Jansen-Jacobs (1984</bibRefCitation>
fig. 9C). Peduncle cylindrical, slightly compressed dorso-ventrally and wrinkled; inserted below and between first and second normal radiole (below second in larger specimens). Pseudoperculum absent. Arrangement of radioles short pectinate, up to 12 radioles per lobe. Inter-radiolar membrane and stylodes absent. Branchial eyes present, reported as stalked eyes at base of pinnules by
<bibRefCitation id="EF854B1FC947FFF07E93FF661F3810AA" author="Langerhans, P." box="[151,369,190,216]" pageId="56" pageNumber="57" pagination="247 - 285" refId="ref63321" refString="Langerhans, P. (1884) Die Wurmfauna von Madeira. 4. Zeitschrift fur Wissenschaftliche Zoologie, 40, 247 - 285." type="journal article" year="1884">Langerhans (1884</bibRefCitation>
fig. 45a). Mouth palps present. 7 thoracic chaetigerous segments. Collar pentalobate, medioventral lobe divided by deep median and two shallow incisions. Tonguelets absent. Thoracic membranes short, ending at second thoracic chaetiger. Collar chaetae of
<taxonomicName id="4C144D6DC947FFF07A1AFED41A801157" authorityName="de Blainville" authorityYear="1818" box="[1054,1225,268,293]" class="Polychaeta" family="Serpulidae" genus="Spirobranchus" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07A1AFED41A801157" box="[1054,1225,268,293]" italics="true" pageId="56" pageNumber="57">Spirobranchus</emphasis>
</taxonomicName>
<typeStatus id="54AF884CC947FFF07AD2FED41B411154" box="[1238,1288,268,294]" pageId="56" pageNumber="57">type</typeStatus>
(
<figureCitation id="132F2A6BC947FFF07B1AFED41BC71154" box="[1310,1422,268,294]" captionStart="FIGURE 24" captionStartId="55.[151,255,1669,1693]" captionTargetBox="[163,1427,216,1621]" captionTargetId="figure-147@55.[151,1436,197,1645]" captionTargetPageId="55" captionText="FIGURE 24. SEM micrographs of chaetae in SEM micrographs of Janita fimbriata. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—Spirobranchus type collar chaeta, C—Apomatus chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini." pageId="56" pageNumber="57">Fig. 24B</figureCitation>
), acicular and limbate.
<taxonomicName id="4C144D6DC947FFF07F93FEEA1C451139" authorityName="Philippi" authorityYear="1844" box="[407,524,306,331]" class="Polychaeta" family="Serpulidae" genus="Apomatus" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07F93FEEA1C451139" box="[407,524,306,331]" italics="true" pageId="56" pageNumber="57">Apomatus</emphasis>
</taxonomicName>
chaetae present (
<figureCitation id="132F2A6BC947FFF07CD8FEEA1D03113E" box="[732,842,306,332]" captionStart="FIGURE 24" captionStartId="55.[151,255,1669,1693]" captionTargetBox="[163,1427,216,1621]" captionTargetId="figure-147@55.[151,1436,197,1645]" captionTargetPageId="55" captionText="FIGURE 24. SEM micrographs of chaetae in SEM micrographs of Janita fimbriata. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—Spirobranchus type collar chaeta, C—Apomatus chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini." pageId="56" pageNumber="57">Fig. 24C</figureCitation>
). Thoracic uncini saw-shaped with up to 16 teeth, anterior peg blunt, questionably gouged (
<figureCitation id="132F2A6BC947FFF07C7BFE811CA21101" box="[639,747,345,371]" captionStart="FIGURE 24" captionStartId="55.[151,255,1669,1693]" captionTargetBox="[163,1427,216,1621]" captionTargetId="figure-147@55.[151,1436,197,1645]" captionTargetPageId="55" captionText="FIGURE 24. SEM micrographs of chaetae in SEM micrographs of Janita fimbriata. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—Spirobranchus type collar chaeta, C—Apomatus chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini." pageId="56" pageNumber="57">Fig. 24E</figureCitation>
). Triangular depression absent. Anterior abdominal uncini saw-shaped, posterior rasp-shaped (
<figureCitation id="132F2A6BC947FFF07C4DFE581CFF11E8" box="[585,694,384,410]" captionStart="FIGURE 24" captionStartId="55.[151,255,1669,1693]" captionTargetBox="[163,1427,216,1621]" captionTargetId="figure-147@55.[151,1436,197,1645]" captionTargetPageId="55" captionText="FIGURE 24. SEM micrographs of chaetae in SEM micrographs of Janita fimbriata. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—Spirobranchus type collar chaeta, C—Apomatus chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini." pageId="56" pageNumber="57">Fig. 24F</figureCitation>
), with approximately 13 teeth in profile, 35 teeth per row. Abdominal chaetae flat narrow geniculate, with a more or less crenulated edge to the blade (
<figureCitation id="132F2A6BC947FFF07B18FE7E1BC611B2" box="[1308,1423,422,448]" captionStart="FIGURE 24" captionStartId="55.[151,255,1669,1693]" captionTargetBox="[163,1427,216,1621]" captionTargetId="figure-147@55.[151,1436,197,1645]" captionTargetPageId="55" captionText="FIGURE 24. SEM micrographs of chaetae in SEM micrographs of Janita fimbriata. France, Marseille, det. and don. H. Zibrowius, ZMA V.Pol. 3033. A—lateral view of entire animal, B—Spirobranchus type collar chaeta, C—Apomatus chaeta, D—anterior abdominal chaeta, E—thoracic uncini, F—posterior abdominal uncini." pageId="56" pageNumber="57">Fig. 24D</figureCitation>
). Achaetous anterior abdominal zone very short or absent. Long posterior capillary chaetae absent. Posterior glandular pad present.
</paragraph>
</subSubSection>
<subSubSection id="C30E6565C947FFF07EC2FDC21AB81561" pageId="56" pageNumber="57" type="discussion">
<paragraph id="8BAB36EEC947FFF07EC2FDC21F141331" blockId="56.[151,1436,152,1222]" pageId="56" pageNumber="57">
<emphasis id="B960EAFCC947FFF07EC2FDC21F711246" bold="true" box="[198,312,538,564]" pageId="56" pageNumber="57">Remarks</emphasis>
. The genus
<taxonomicName id="4C144D6DC947FFF07FC9FDC21C5F1241" authorityName="Saint-Joseph" authorityYear="1894" box="[461,534,538,563]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07FC9FDC21C5F1241" box="[461,534,538,563]" italics="true" pageId="56" pageNumber="57">Janita</emphasis>
</taxonomicName>
was erected by Saint-Joseph for
<taxonomicName id="4C144D6DC947FFF07DA2FDC21BDE1246" authority="Langerhans, 1884" authorityName="Langerhans" authorityYear="1884" box="[934,1431,538,564]" class="Polychaeta" family="Serpulidae" genus="Omphalopoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="species" species="spinosa">
<emphasis id="B960EAFCC947FFF07DA2FDC21AFE1241" box="[934,1207,538,563]" italics="true" pageId="56" pageNumber="57">Omphalopoma spinosa</emphasis>
<bibRefCitation id="EF854B1FC947FFF07AC4FDC21BDE1246" author="Langerhans, P." box="[1216,1431,538,564]" pageId="56" pageNumber="57" pagination="247 - 285" refId="ref63321" refString="Langerhans, P. (1884) Die Wurmfauna von Madeira. 4. Zeitschrift fur Wissenschaftliche Zoologie, 40, 247 - 285." type="journal article" year="1884">Langerhans, 1884</bibRefCitation>
</taxonomicName>
, which is a junior synonym of
<taxonomicName id="4C144D6DC947FFF07C29FD991D4F1228" authorityName="Delle Chiaje" authorityYear="1822" box="[557,774,577,602]" class="Agaricomycetes" family="Serpulaceae" genus="Serpula" kingdom="Fungi" order="Boletales" pageId="56" pageNumber="57" phylum="Basidiomycota" rank="species" species="fimbriata">
<emphasis id="B960EAFCC947FFF07C29FD991D4F1228" box="[557,774,577,602]" italics="true" pageId="56" pageNumber="57">Serpula fimbriata</emphasis>
</taxonomicName>
delle Chiaje, 1822 (see e.g.,
<bibRefCitation id="EF854B1FC947FFF07A96FD991BDF1229" author="Fauchald, K." box="[1170,1430,577,603]" pageId="56" pageNumber="57" pagination="1 - 188" refId="ref58220" refString="Fauchald, K. (1977) The polychaete worms. Definitions and keys to the orders, families and genera. Natural History Museum of Los Angeles County Science Series, 28, 1 - 188." type="journal article" year="1977">Fauchald, 1977: 144</bibRefCitation>
,
<bibRefCitation id="EF854B1FC947FFF07E93FDB01F8612F0" author="Lommerzheim, A." box="[151,463,616,642]" pageId="56" pageNumber="57" pagination="110 - 195" refId="ref63571" refString="Lommerzheim, A. (1979) Monographische Bearbeitung der Serpulidae (Polychaeta sedentaria) aus dem Cenoman (Oberkreide) am Sudwestrand des Munsterlander Beckens. Decheniana, 132, 110 - 195." type="journal article" year="1979">Lommerzheim 1979: 157</bibRefCitation>
). There was a considerable confusion about generic attribution of this species, generally it has been attributed to
<emphasis id="B960EAFCC947FFF07C48FD571D6612DA" box="[588,815,655,680]" italics="true" pageId="56" pageNumber="57">
<taxonomicName id="4C144D6DC947FFF07C48FD571D6512DA" authorityName="Saint-Joseph" authorityYear="1894" box="[588,812,655,680]" class="Polychaeta" family="Serpulidae" genus="Omphalopomopsis" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">Omphalopomopsis</taxonomicName>
.
</emphasis>
<bibRefCitation id="EF854B1FC947FFF07D38FD561A5612DA" author="Zibrowius, H." box="[828,1055,654,680]" pageId="56" pageNumber="57" pagination="113 - 126" refId="ref69643" refString="Zibrowius, H. (1972 b) Mise au point sur les especes Mediterraneennes de Serpulidae (Annelida Polychaeta) decrites par Stefano delle Chiaje (1822 - 1829, 1841 - 1844) et Orionzo Costa (1861). Tethys, 4, 113 - 126." type="journal article" year="1972">Zibrowius (1972b)</bibRefCitation>
points out that the distinction between
<taxonomicName id="4C144D6DC947FFF07F04FD6D1F0012BC" authorityName="Saint-Joseph" authorityYear="1894" box="[256,329,693,718]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07F04FD6D1F0012BC" box="[256,329,693,718]" italics="true" pageId="56" pageNumber="57">Janita</emphasis>
</taxonomicName>
and
<taxonomicName id="4C144D6DC947FFF07F86FD6D1C1712BC" authorityName="Saint-Joseph" authorityYear="1894" box="[386,606,693,718]" class="Polychaeta" family="Serpulidae" genus="Omphalopomopsis" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07F86FD6D1C1712BC" box="[386,606,693,718]" italics="true" pageId="56" pageNumber="57">Omphalopomopsis</emphasis>
</taxonomicName>
is justified due to having very different opercula:
<taxonomicName id="4C144D6DC947FFF07AAEFD6D1B1012BC" authorityName="Ehlers" authorityYear="1887" box="[1194,1369,693,718]" class="Polychaeta" family="Serpulidae" genus="Hyalopomatus" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="species" species="langerhansi">
<emphasis id="B960EAFCC947FFF07AAEFD6D1B1012BC" box="[1194,1369,693,718]" italics="true" pageId="56" pageNumber="57">O. langerhansi</emphasis>
</taxonomicName>
has a simple globular operculum with a shallow concave calcareous endplate,
<taxonomicName id="4C144D6DC947FFF07A08FD041AD01287" authorityName="Delle Chiaje" authorityYear="1822" box="[1036,1177,732,757]" class="Agaricomycetes" family="Serpulaceae" genus="Serpula" kingdom="Fungi" order="Boletales" pageId="56" pageNumber="57" phylum="Basidiomycota" rank="species" species="fimbriata">
<emphasis id="B960EAFCC947FFF07A08FD041AD01287" box="[1036,1177,732,757]" italics="true" pageId="56" pageNumber="57">J. fimbriata</emphasis>
</taxonomicName>
has a more complex operculum with a deeply cupped chitinous endplate, which has a horny talon into the fleshy opercular ampulla (
<bibRefCitation id="EF854B1FC947FFF07E9BFCF11F191331" author="Imajima, M." box="[159,336,809,835]" pageId="56" pageNumber="57" pagination="159 - 183" refId="ref61932" refString="Imajima, M. (1979) Serpulidae (Annelida, Polychaeta) collected around Cape Shionomisaki, Kii Peninsula. Memoirs of the National Science Museum, Tokyo, 12, 159 - 183." type="journal article" year="1979">Imajima, 1979</bibRefCitation>
).
</paragraph>
<paragraph id="8BAB36EEC947FFF07EC2FC881AAC1420" blockId="56.[151,1436,152,1222]" pageId="56" pageNumber="57">
It should be noted that
<bibRefCitation id="EF854B1FC947FFF07FD6FC881C2E1318" author="Rioja, E." box="[466,615,848,874]" pageId="56" pageNumber="57" pagination="1 - 144" refId="ref66509" refString="Rioja, E. (1923). Estudio sistematico de las especies Ibericas del suborden Sabelliformia. Trabajos del Museo Nacional de Ciencias Naturales Serie Zoologica, 48, 1 - 144." type="journal article" year="1923">
<collectingRegion id="49D0F80CC947FFF07FD6FC881C581318" box="[466,529,848,874]" country="Spain" name="La Rioja" pageId="56" pageNumber="57">Rioja</collectingRegion>
(1923)
</bibRefCitation>
and
<bibRefCitation id="EF854B1FC947FFF07C9AFC881D0D1318" author="Fauvel, P." box="[670,836,848,874]" pageId="56" pageNumber="57" pagination="1 - 494" refId="ref58461" refString="Fauvel, P. (1927) Polychetes sedentaires. Addenda aux errantes, Archiannelides, Myzostomaires. Faune de France, 16, 1 - 494." type="journal article" year="1927">Fauvel (1927)</bibRefCitation>
mentioned both “
<taxonomicName id="4C144D6DC947FFF07A11FC881A89131B" authorityName="de Blainville" authorityYear="1818" box="[1045,1216,848,873]" class="Polychaeta" family="Serpulidae" genus="Spirobranchus" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07A11FC881A89131B" box="[1045,1216,848,873]" italics="true" pageId="56" pageNumber="57">Spirobranchus</emphasis>
</taxonomicName>
<typeStatus id="54AF884CC947FFF07AD0FC881B4F1318" box="[1236,1286,848,874]" pageId="56" pageNumber="57">type</typeStatus>
and acicular collar chaetae for
<taxonomicName id="4C144D6DC947FFF07F6DFCAF1C6F13E2" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1822" box="[361,550,887,912]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="species" species="fimbriata">
<emphasis id="B960EAFCC947FFF07F6DFCAF1C6F13E2" box="[361,550,887,912]" italics="true" pageId="56" pageNumber="57">Janita fimbriata</emphasis>
</taxonomicName>
(as
<taxonomicName id="4C144D6DC947FFF07C53FCAF1D7A13E2" authorityName="Saint-Joseph" authorityYear="1894" box="[599,819,887,912]" class="Polychaeta" family="Serpulidae" genus="Omphalopomopsis" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07C53FCAF1D7A13E2" box="[599,819,887,912]" italics="true" pageId="56" pageNumber="57">Omphalopomopsis</emphasis>
</taxonomicName>
); (
<bibRefCitation id="EF854B1FC947FFF07D57FCAE1A6613E2" author="Zibrowius, H." box="[851,1071,886,912]" pageId="56" pageNumber="57" pagination="81 - 252" refId="ref69212" refString="Zibrowius, H. (1968 a) Etude morphologique, systematique et ecologique des Serpulidae (Annelida Polychaeta) de la region de Marseille. Recueil des Traveaux de la Station Marine d'Endoume, Bulletin 43, 81 - 252." type="journal article" year="1968">Zibrowius (1968a)</bibRefCitation>
on the other hand regarded the “acicular” chaetae as misinterpretation of “
<taxonomicName id="4C144D6DC947FFF07C94FC451D7213C4" authorityName="de Blainville" authorityYear="1818" box="[656,827,925,950]" class="Polychaeta" family="Serpulidae" genus="Spirobranchus" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07C94FC451D7213C4" box="[656,827,925,950]" italics="true" pageId="56" pageNumber="57">Spirobranchus</emphasis>
</taxonomicName>
<typeStatus id="54AF884CC947FFF07D54FC451DCC13C5" box="[848,901,925,951]" pageId="56" pageNumber="57">type</typeStatus>
, observed from the back (not in lateral view).
<bibRefCitation id="EF854B1FC947FFF07E93FC1C1F0913AC" author="Martin, D." box="[151,320,964,990]" pageId="56" pageNumber="57" pagination="61 - 75" refId="ref63982" refString="Martin, D. (1989) Anelidos poliquetos asociados a las concreciones de algas calcareas del litoral catalan. Miscellania Zoologica, Barcelona (1987), 11, 61 - 75." type="journal article" year="1989">Martín (1989)</bibRefCitation>
assumed that specimens with acicular collar chaetae and those with “
<taxonomicName id="4C144D6DC947FFF07AA1FC1C1B1913AF" authorityName="de Blainville" authorityYear="1818" box="[1189,1360,964,989]" class="Polychaeta" family="Serpulidae" genus="Spirobranchus" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07AA1FC1C1B1913AF" box="[1189,1360,964,989]" italics="true" pageId="56" pageNumber="57">Spirobranchus</emphasis>
</taxonomicName>
<typeStatus id="54AF884CC947FFF07B6DFC1C1BD213AC" box="[1385,1435,964,990]" pageId="56" pageNumber="57">type</typeStatus>
chaetae belong to different taxa.
<bibRefCitation id="EF854B1FC947FFF07C1EFC321D151476" author="Ben-Eliahu, M. N. &amp; Fiege, D." box="[538,860,1002,1028]" pageId="56" pageNumber="57" pagination="1 - 51" refId="ref56291" refString="Ben-Eliahu, M. N. &amp; Fiege, D. (1996) Serpulid tube-worms (Annelida, Polychaeta) of the central and eastern Mediterranean with particular attention to the Levant Basin. Senckenbergiana Maritima, 28, 1 - 51." type="journal article" year="1996">Ben-Eliahu &amp; Fiege (1996)</bibRefCitation>
mentioned specimens with one or the other
<typeStatus id="54AF884CC947FFF07B6DFC321BD21476" box="[1385,1435,1002,1028]" pageId="56" pageNumber="57">type</typeStatus>
of collar chaetae from a single population of what they regard to be
<taxonomicName id="4C144D6DC947FFF07DC4FBC91A031458" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1822" box="[960,1098,1041,1066]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="species" species="fimbriata">
<emphasis id="B960EAFCC947FFF07DC4FBC91A031458" box="[960,1098,1041,1066]" italics="true" pageId="56" pageNumber="57">J. fimbriata</emphasis>
</taxonomicName>
; ten Hove (in
<bibRefCitation id="EF854B1FC947FFF07AF3FBC91F521420" author="Ben-Eliahu, M. N. &amp; Fiege, D." pageId="56" pageNumber="57" pagination="1 - 51" refId="ref56291" refString="Ben-Eliahu, M. N. &amp; Fiege, D. (1996) Serpulid tube-worms (Annelida, Polychaeta) of the central and eastern Mediterranean with particular attention to the Levant Basin. Senckenbergiana Maritima, 28, 1 - 51." type="journal article" year="1996">Ben-Eliahu &amp; Fiege 1996</bibRefCitation>
) mentioned a specimen with both
<typeStatus id="54AF884CC947FFF07CABFBE01CB91420" box="[687,752,1080,1106]" pageId="56" pageNumber="57">types</typeStatus>
. The phenomenon merits further attention.
</paragraph>
<paragraph id="8BAB36EEC947FFF07EC2FB861D5A14B4" blockId="56.[151,1436,152,1222]" pageId="56" pageNumber="57">
The monotypic genus is distributed in the (sub) tropical Atlantic, Mediterranean (
<bibRefCitation id="EF854B1FC947FFF07AC7FB861BDE140A" author="Zibrowius, H." box="[1219,1431,1118,1144]" pageId="56" pageNumber="57" pagination="113 - 126" refId="ref69643" refString="Zibrowius, H. (1972 b) Mise au point sur les especes Mediterraneennes de Serpulidae (Annelida Polychaeta) decrites par Stefano delle Chiaje (1822 - 1829, 1841 - 1844) et Orionzo Costa (1861). Tethys, 4, 113 - 126." type="journal article" year="1972">Zibrowius 1972b</bibRefCitation>
,
<bibRefCitation id="EF854B1FC947FFF07E93FB5D1EAB14ED" author="Zibrowius, H." box="[151,226,1157,1183]" pageId="56" pageNumber="57" pagination="1 - 93" refId="ref69840" refString="Zibrowius, H. (1973 b) Serpulidae (Annelida Polychaeta) des cotes ouest de l'Afrique et des Archipels Voisins. Musee Royal de l'Afrique Centrale, Tervuren, Belgique Annales, Serie 8, Sciences Zoologiques, 207, 1 - 93." type="journal article" year="1973">1973b</bibRefCitation>
,
<bibRefCitation id="EF854B1FC947FFF07EEAFB5D1FC414ED" author="Bianchi, C. N." box="[238,397,1157,1183]" pageId="56" pageNumber="57" pagination="1 - 187" refId="ref56534" refString="Bianchi, C. N. (1981) Policheti Serpuloidei. Guide per il riconoscimento delle specie animali delle acqua laguna e costiere italiane AQ / 1 / 96, 5, 1 - 187." type="journal article" year="1981">Bianchi 1981</bibRefCitation>
,
<bibRefCitation id="EF854B1FC947FFF07F9DFB5D1C3314ED" author="Bianchi, C. N. &amp; Chessa, L. A. &amp; Morri, C." box="[409,634,1157,1183]" pageId="56" pageNumber="57" pagination="49 - 58" refId="ref56571" refString="Bianchi, C. N., Chessa, L. A. &amp; Morri, C. (1984) Serpuloidea (Annelida, Polychaeta) della Sardegna, con particolare riguardo alle lagune costiere. Ecologia della Sardegna e delle isole tirreniche supplemento, 54, 49 - 58." type="journal article" year="1984">
Bianchi
<emphasis id="B960EAFCC947FFF07FFFFB5D1C7C14EC" box="[507,565,1157,1182]" italics="true" pageId="56" pageNumber="57">et al.</emphasis>
1984
</bibRefCitation>
), and Indo-West Pacific (
<bibRefCitation id="EF854B1FC947FFF07D9BFB5D1A9414ED" author="Imajima, M. &amp; Hove, H. A. ten" box="[927,1245,1157,1183]" pageId="56" pageNumber="57" pagination="35 - 66" refId="ref62002" refString="Imajima, M. &amp; Hove, H. A. ten (1984) Serpulinae (Annelida, Polychaeta) from the Truk Islands, Ponape and Majuro Atoll, with some other new Indo-Pacific records. Proceedings of the Japanese Society of Systematic Zoology, 27, 35 - 66." type="journal article" year="1984">Imajima &amp; ten Hove, 1984</bibRefCitation>
,
<bibRefCitation id="EF854B1FC947FFF07AEDFB5D1B6F14ED" author="Imajima, M. &amp; Hove, H. A. ten" box="[1257,1318,1157,1183]" pageId="56" pageNumber="57" pagination="1 - 16" refId="ref62055" refString="Imajima, M. &amp; Hove, H. A. ten (1986) Serpulinae (Annelida, Polychaeta) from Nauru, the Gilbert Islands (Kiribati) and the Solomon Islands. Proceedings of the Japanese Society of Systematic Zoology, Tokyo, 32, 1 - 16." type="journal article" year="1986">1986</bibRefCitation>
, ten Hove 1994). See also remarks following
<taxonomicName id="4C144D6DC947FFF07C35FB741D4414B7" authorityName="Saint-Joseph" authorityYear="1894" box="[561,781,1196,1221]" class="Polychaeta" family="Serpulidae" genus="Omphalopomopsis" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="genus">
<emphasis id="B960EAFCC947FFF07C35FB741D4414B7" box="[561,781,1196,1221]" italics="true" pageId="56" pageNumber="57">Omphalopomopsis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BAB36EEC947FFF07E93FB211AB81561" blockId="56.[151,1265,1273,1299]" box="[151,1265,1273,1299]" pageId="56" pageNumber="57">
<taxonomicName id="4C144D6DC947FFF07E93FB211F1D1560" baseAuthorityName="Delle Chiaje" baseAuthorityYear="1822" box="[151,340,1273,1298]" class="Polychaeta" family="Serpulidae" genus="Janita" kingdom="Animalia" order="Sabellida" pageId="56" pageNumber="57" phylum="Annelida" rank="species" species="fimbriata">
<emphasis id="B960EAFCC947FFF07E93FB211F1D1560" box="[151,340,1273,1298]" italics="true" pageId="56" pageNumber="57">Janita fimbriata</emphasis>
</taxonomicName>
(delle Chiaje, 1822), (sub)tropical Atlantic, Mediterranean, Indo-West Pacific.
</paragraph>
</subSubSection>
</treatment>
</document>