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<document id="1E3836610387F2A841744B8E6C9586F0" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66BC8FF6FAF89FB28FD103E75" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera rufonitida Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="171" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="170" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="88095D685311DE8A3C1FEAB36F1BC88C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="F703804491347D10DD0E4572935241A2">
<mods:title id="ED297DC1341C7FA389AA414FB753EF4D">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
</mods:titleInfo>
<mods:name id="07ED4B9DE14E5707AEFA08DBCE39C5AE" type="personal">
<mods:role id="6AC812A4FDA0456E7E65BE5BDF03A37C">
<mods:roleTerm id="6A44289B2DF047FB80CBDBAC40AAB41E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A8DC4180ED1F146130131E5901DF2CF1">Fabrizi, Silvia</mods:namePart>
<mods:nameIdentifier id="5BC05668455D87B063D1E038122141E1" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="64C08674A423B3B388FEA747FF23C1BE" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:name id="8DEE84A391795059EFD9A99ABEC4A716" type="personal">
<mods:role id="7AC2FB82CC60B70EF664FC790B100E73">
<mods:roleTerm id="D1BEADFDEF75B60D793128284B036F34">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B8F002D64A2F7A00BF4EE3D7E004A875">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="FEC4167B0C4D847D099CCFC4F55D6FBC" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="300C65433A53263A08DE1D879C1EF34F" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="6B2B9E5B23213BF7C13C3913132D0E73" type="personal">
<mods:role id="F7C035D5381CFD8FCD18AEF1987C4206">
<mods:roleTerm id="895CDAA43A5A86A0846EBE9C0596C016">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="9AFB0B8EA8D860B649154230C70C3D7B">Bai, Ming</mods:namePart>
<mods:nameIdentifier id="805FA44A3CAABEFC1B9ED6BE7374E899" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="A18A624B099C4BCDA7AF7AE463EDD8C4" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="D3E60F0A8AD860B42A092B9E357185EC" type="personal">
<mods:role id="EBE333BF7C5C41AAEDA1616007665C96">
<mods:roleTerm id="7115F8D9E36D7CA7A079B1E6A1254E3D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="D62119A5115ADEE26C1B454D657952A6">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="B4F8A302BD24893790EF3F73204B2AC1" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
<mods:nameIdentifier id="57398A0589EC8DA43EC5B89E47F85A1A" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
<mods:name id="41B205E4713D27DC884962E641093805" type="personal">
<mods:role id="915EFEA5918F13479A4283F890422DF6">
<mods:roleTerm id="B759D050D29AB902D2D37D14AC52B4BD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="889C18A2AA55F0ACF018B5714D866690">Ahrens, Dirk</mods:namePart>
<mods:nameIdentifier id="A58E43940EB7D72050D45D5CAF55881C" type="ORCID">0000-0003-3524-7153</mods:nameIdentifier>
<mods:nameIdentifier id="F4A81121499FCE582D182ABE34A1BF08" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="4AC65D22F56E121EC008E1E7450448A8">text</mods:typeOfResource>
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<mods:title id="1D8102C8D8B1D5E50FC14A6DB25F2EE5">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="21C2071BD48717970FE4BEE8DAA58CD3">
<mods:date id="8D14A8E69F27BCD6BF50AD343FEE87E6">2021</mods:date>
<mods:detail id="97EC8916C22D9B225E15CA1DD014E313" type="pubDate">
<mods:number id="090B14E2754B58C66FC4C8A7CFF1BB05">2021-02-03</mods:number>
</mods:detail>
<mods:detail id="3C5E974EF458F6490E30CAC726BE6D12" type="volume">
<mods:number id="4C9244CF89C4200A1A0885A03EC1C5DE">4922</mods:number>
</mods:detail>
<mods:detail id="4A7F505018881E0E9AF69A919DF45CD5" type="issue">
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<mods:start id="2C053B176CD021944CEB805ABEF6865B">1</mods:start>
<mods:end id="E80B61012AFAD50D9DBD270D240A848B">400</mods:end>
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<mods:classification id="2B2E5F1F46A02368A1B79DAD941CC921">journal article</mods:classification>
<mods:identifier id="BC31DEF4CBC92FDAA6EB422D34974B39" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="EF73E448B14FC8ECDC2E52A2D71C2619" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
<mods:identifier id="1004D073946012B1BF4D7CE73A672499" type="GBIF-Dataset">188411de-569c-4408-a016-1a04b858c142</mods:identifier>
<mods:identifier id="701731A7F7A7511A1402BCD3E001F761" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="7C6653DDF5BA32F4ADD9D8CD85F7ADC4" type="Zenodo-Dep">4496316</mods:identifier>
<mods:identifier id="5549BF8CEF9DBE6CB25C270341198D0E" type="ZooBank">D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C</mods:identifier>
</mods:mods>
<treatment id="03BD87E66BC8FF6FAF89FB28FD103E75" ID-DOI="http://doi.org/10.5281/zenodo.4534625" ID-GBIF-Taxon="177323575" ID-Zenodo-Dep="4534625" LSID="urn:lsid:plazi:treatment:03BD87E66BC8FF6FAF89FB28FD103E75" httpUri="http://treatment.plazi.org/id/03BD87E66BC8FF6FAF89FB28FD103E75" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170">
<subSubSection id="C30E657B6BC8FF6CAF89FB28FD203E11" box="[151,766,834,861]" pageId="169" pageNumber="170" type="nomenclature">
<paragraph id="8BAB36F06BC8FF6CAF89FB28FD203E11" blockId="169.[151,766,834,897]" box="[151,766,834,861]" pageId="169" pageNumber="170">
<heading id="D0E3819C6BC8FF6CAF89FB28FD203E11" bold="true" box="[151,766,834,861]" fontSize="11" level="2" pageId="169" pageNumber="170" reason="4">
<emphasis id="B960EAE26BC8FF6CAF89FB28FD203E11" bold="true" box="[151,766,834,861]" pageId="169" pageNumber="170">
<taxonomicName id="4C144D736BC8FF6CAF89FB28FD713E10" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,687,834,861]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="169" pageNumber="170" phylum="Arthropoda" rank="species" species="rufonitida" status="sp. nov.">
<emphasis id="B960EAE26BC8FF6CAF89FB28FE503E11" bold="true" box="[151,398,834,861]" italics="true" pageId="169" pageNumber="170">Maladera rufonitida</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BC8FF6CADA2FB29FD203E11" box="[700,766,835,861]" pageId="169" pageNumber="170" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BC8FF6CAF89FB0DFEA33ECD" box="[151,381,871,897]" pageId="169" pageNumber="170" type="description">
<paragraph id="8BAB36F06BC8FF6CAF89FB0DFEA33ECD" blockId="169.[151,766,834,897]" box="[151,381,871,897]" pageId="169" pageNumber="170">
<figureCitation id="132F2A756BC8FF6CAF89FB0DFED03ECD" box="[151,270,871,897]" captionStart="FIGURE 38" captionStartId="171.[151,250,1943,1967]" captionTargetBox="[222,1371,191,1903]" captionTargetId="figure-20@171.[205,1382,181,1920]" captionTargetPageId="171" captionText="FIGURE 38. A–D: Maladera rufonitida Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. songi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. detersa (Erichson, 1834) (Vietnam: DaNang). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496408" httpUri="https://zenodo.org/record/4496408/files/figure.png" pageId="169" pageNumber="170">Figures 38</figureCitation>
A–D, 106
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BC8FF6CAF89FBC5FF32395D" pageId="169" pageNumber="170" type="materials_examined">
<paragraph id="8BAB36F06BC8FF6CAF89FBC5FF32395D" blockId="169.[151,1437,942,2013]" pageId="169" pageNumber="170">
<emphasis id="B960EAE26BC8FF6CAF89FBC5FE183E84" bold="true" box="[151,454,942,969]" pageId="169" pageNumber="170">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6BC8FF6CAECFFBC5FDC63EA1" ID-GBIF-Occurrence="3034555372" collectingDate="2003-08" collectionCode="ZFMK" collectorName="Li" country="China" elevation="1600" location="Mupu Mt." pageId="169" pageNumber="170" specimenCount="1" specimenCount-male="1" stateProvince="Hunan" typeStatus="holotype">
<typeStatus id="54AF88526BC8FF6CAECFFBC5FDE03E85" box="[465,574,943,969]" pageId="169" pageNumber="170" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796BC8FF6CAD55FBC5FDBE3E85" box="[587,608,943,969]" pageId="169" pageNumber="170" type="male">♂</specimenCount>
“
<collectingCountry id="F30376606BC8FF6CAD6BFBC5FD603E85" box="[629,702,943,969]" name="China" pageId="169" pageNumber="170">China</collectingCountry>
:
<collectingRegion id="49D0F8126BC8FF6CADD3FBC5FCC53E85" box="[717,795,943,969]" country="China" name="Hunan" pageId="169" pageNumber="170">Hunan</collectingRegion>
;
<location id="8ECB602B6BC8FF6CAC34FBC5FC413E85" LSID="urn:lsid:plazi:treatment:03BD87E66BC8FF6FAF89FB28FD103E75:8ECB602B6BC8FF6CAC34FBC5FC413E85" box="[810,927,943,969]" country="China" name="Mupu Mt." pageId="169" pageNumber="170" stateProvince="Hunan">Mupu Mt.</location>
<quantity id="4CEC9B156BC8FF6CACB4FBC5FC243E85" box="[938,1018,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="169" pageNumber="170" unit="m" value="1600.0">
<elevation id="0039D1C36BC8FF6CACB4FBC5FC243E85" box="[938,1018,943,969]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="169" pageNumber="170" unit="m" value="1600.0">1600m</elevation>
</quantity>
, Pingjiang
<date id="FFAA10306BC8FF6CAB60FBC5FB2C3E85" box="[1150,1266,943,969]" pageId="169" pageNumber="170" value="2003-08">
<collectingDate id="EFEEE9D86BC8FF6CAB60FBC5FB2C3E85" box="[1150,1266,943,969]" pageId="169" pageNumber="170" value="2003-08">VIII-2003</collectingDate>
</date>
, leg.
<collectorName id="26E153266BC8FF6CAA2BFBC5FA903E85" box="[1333,1358,943,969]" pageId="169" pageNumber="170">Li</collectorName>
<emphasis id="B960EAE26BC8FF6CAA47FBC5FA4B3E85" box="[1369,1429,943,969]" italics="true" pageId="169" pageNumber="170">et al.</emphasis>
/ 719
<taxonomicName id="4C144D736BC8FF6CAFDCFBB9FEC33EA1" box="[194,285,979,1005]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="169" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
: Asia spec.” (
<collectionCode id="ED05AE356BC8FF6CAEA5FBB9FDD13EA1" box="[443,527,979,1005]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="169" pageNumber="170" type="Museum">ZFMK</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B7C3CAD6BC8FF6CAD3AFBB9FF36395D" ID-GBIF-Occurrence="3034556528" collectingDate="1982-06-11" collectionCode="CAU" collectorName="Yang Jikun" country="China" location="Jinxiu" pageId="169" pageNumber="170" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<typeStatus id="54AF88526BC8FF6CAD3AFBB9FD543EA1" box="[548,650,979,1005]" pageId="169" pageNumber="170" type="paratype">Paratype</typeStatus>
:
<specimenCount id="9D12FD796BC8FF6CAD86FBB9FD1A3EA1" box="[664,708,979,1005]" pageId="169" pageNumber="170" type="male">1 ♂</specimenCount>
“[
<collectingCountry id="F30376606BC8FF6CADC1FBB8FCF73EA1" box="[735,809,978,1005]" name="China" pageId="169" pageNumber="170">China</collectingCountry>
]
<location id="8ECB602B6BC8FF6CAC27FBB9FC5E3EA1" LSID="urn:lsid:plazi:treatment:03BD87E66BC8FF6FAF89FB28FD103E75:8ECB602B6BC8FF6CAC27FBB9FC5E3EA1" box="[825,896,979,1005]" country="China" name="Jinxiu" pageId="169" pageNumber="170" stateProvince="Guangxi">Jinxiu</location>
,
<collectingRegion id="49D0F8126BC8FF6CAC90FBB9FC2F3EA1" box="[910,1009,979,1005]" country="China" name="Guangxi" pageId="169" pageNumber="170">Guangxi</collectingRegion>
,
<date id="FFAA10306BC8FF6CACE1FBB9FB5E3EA1" box="[1023,1152,979,1005]" pageId="169" pageNumber="170" value="1982-06-11">
<collectingDate id="EFEEE9D86BC8FF6CACE1FBB9FB5E3EA1" box="[1023,1152,979,1005]" pageId="169" pageNumber="170" value="1982-06-11">
11.VI.
<quantity id="4CEC9B156BC8FF6CAB5CFBB9FB5E3EA1" box="[1090,1152,979,1005]" metricMagnitude="6" metricUnit="m" metricValue="1.9827199999999998" pageId="169" pageNumber="170" unit="m" value="1982720.0">1982</quantity>
</collectingDate>
</date>
, 720m, leg.
<collectorName id="26E153266BC8FF6CAA0EFBB9FA4D3EA1" box="[1296,1427,979,1005]" pageId="169" pageNumber="170">Yang Jikun</collectorName>
” (
<collectionCode id="ED05AE356BC8FF6CAF81FB9DFF3E395D" box="[159,224,1015,1041]" country="China" lsid="urn:lsid:biocol.org:col:33074" name="China Agricultural University" pageId="169" pageNumber="170">CAU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BC8FF6FAFD9FC70FCA93F2D" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170" type="description">
<paragraph id="8BAB36F06BC8FF6CAFD9FC70FD133915" blockId="169.[151,1437,942,2013]" pageId="169" pageNumber="170">
<emphasis id="B960EAE26BC8FF6CAFD9FC70FE873978" bold="true" box="[199,345,1050,1076]" pageId="169" pageNumber="170">Description.</emphasis>
Length:
<quantity id="4CEC9B156BC8FF6CAEDDFC71FDC03979" box="[451,542,1051,1077]" metricMagnitude="-3" metricUnit="m" metricValue="8.9" pageId="169" pageNumber="170" unit="mm" value="8.9">8.9 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BC8FF6CADF1FC70FC943979" box="[751,842,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" pageId="169" pageNumber="170" unit="mm" value="6.5">6.5 mm</quantity>
, width:
<quantity id="4CEC9B156BC8FF6CACB4FC70FBDB3979" box="[938,1029,1050,1077]" metricMagnitude="-3" metricUnit="m" metricValue="5.1" pageId="169" pageNumber="170" unit="mm" value="5.1">5.1 mm</quantity>
. Body oblong-oval, reddish brown, antenna yellow, shiny, dorsal face nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06BC8FF6CAFD9FC09FC9A38D1" blockId="169.[151,1437,942,2013]" pageId="169" pageNumber="170">
Labroclypeus moderately wide and subtrapezoidal, widest at base, lateral margins strongly convex and convergent anteriorly, anterior angles strongly rounded, anterior margin distinctly emarginate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface flat, coarsely, very densely punctate, with a few long setae anteriorly; frontoclypeal suture distinctly incised, weakly curved; smooth area anterior to eye convex, 2.5 times as wide as long; ocular canthus moderately long and wide (1/3 of ocular diameter), finely densely punctate, with two terminal setae. Frons shiny, with irregularly dense and coarse punctures, partly completely impunctate, with a few single setae beside eyes. Eyes large, ratio diameter/ interocular width: 0.83. Antenna with nine antennomeres; club with three antennomeres and strongly reflexed, twice as long as remaining antennomeres combined.
<collectingRegion id="49D0F8126BC8FF6CAE0AFDE9FEAD38D1" box="[276,371,1411,1437]" country="Myanmar" name="Chin" pageId="169" pageNumber="170">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06BC8FF6CAFD9FDCDFE523B39" blockId="169.[151,1437,942,2013]" pageId="169" pageNumber="170">Pronotum moderately transverse, widest at base, lateral margins in basal half straight and weakly convergent anteriorly, in anterior half convex and more convergent, anterior angles distinctly produced and sharp, posterior angles blunt; anterior margin weakly convex, with fine marginal line, base without marginal line; surface densely and coarsely punctate, with minute setae in punctures being as long as puncture diameter, otherwise glabrous; anterior and lateral margin densely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, with fine, dense punctures.</paragraph>
<paragraph id="8BAB36F06BC8FF6CAFD9FE15FAA33A4A" blockId="169.[151,1437,942,2013]" pageId="169" pageNumber="170">Elytra widest in posterior third, striae finely impressed, finely and densely punctate, intervals weakly convex, with fine, sparse punctures concentrated along striae, with minute setae in punctures, penultimate lateral interval with a few single setae; epipleural margin robust, ending at apical external angle of elytra, epipleura densely setose; apical border of elytra membraneous, with a very fine rim of microtrichomes (visible at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06BC8FF6CAFD9FF65FA983AF5" blockId="169.[151,1437,942,2013]" pageId="169" pageNumber="170">Ventral surface shiny, coarsely and densely punctate, metasternum including disc densely setose; metacoxa glabrous, with a few longer setae laterally. Abdominal sternites finely and densely punctate, punctures with minute setae, each sternite with a transverse row of punctures each bearing a fine seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.7. Pygidium moderately convex, shiny, coarsely and densely punctate, without smooth midline, with a few long but fine setae along apical margin.</paragraph>
<paragraph id="8BAB36F06BC8FF6FAFD9FFA9FCF13F71" blockId="169.[151,1437,942,2013]" lastBlockId="170.[151,1437,151,826]" lastPageId="170" lastPageNumber="171" pageId="169" pageNumber="170">Legs long and moderately wide, shiny; femora with two longitudinal rows of setae, coarsely and densely punctate in basal half, otherwise punctures sparse. Anterior margin of metafemur acute, with adjacent continuous serrated line, anterior row of setae complete; posterior ventral margin smooth, weakly widened at ventral apex, dorsal posterior margin smooth, neither serrate, glabrous. Metatibia moderately long and moderately wide, widest at apex, ratio of width/length: 1/2.5, sharply carinate dorsally, with two groups of spines, basal group at middle, apical group at three quarters of metatibial length, in basal half with a few short and robust single setae and a continuous serrated line beside basal half of dorsal margin; lateral face longitudinally convex, except a few punctures on base smooth and glabrous; ventral margin finely serrate, with four equidistant robust setae; medial face smooth and glabrous; apex finely serrate, distinctly concavely emarginate interiorly near tarsal articulation. Tarsomeres dorsally impunctate, glabrous, neither laterally nor dorsally carinate, moderately setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina, glabrous; first metatarsomere distinctly longer than following tarsomere and longer than dorsal tibial spur. Protibia moderately long, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06BCBFF6FAFD9FA2DFCA93F2D" blockId="170.[151,1437,151,826]" box="[199,887,583,609]" pageId="170" pageNumber="171">
Aedeagus:
<figureCitation id="132F2A756BCBFF6FAE5AFA2DFE4A3F2D" box="[324,404,583,609]" captionStart="FIGURE 38" captionStartId="171.[151,250,1943,1967]" captionTargetBox="[222,1371,191,1903]" captionTargetId="figure-20@171.[205,1382,181,1920]" captionTargetPageId="171" captionText="FIGURE 38. A–D: Maladera rufonitida Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. songi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. detersa (Erichson, 1834) (Vietnam: DaNang). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496408" httpUri="https://zenodo.org/record/4496408/files/figure.png" pageId="170" pageNumber="171">Fig. 38</figureCitation>
A–C. Habitus:
<figureCitation id="132F2A756BCBFF6FAD23FA2DFD7D3F2D" box="[573,675,583,609]" captionStart="FIGURE 38" captionStartId="171.[151,250,1943,1967]" captionTargetBox="[222,1371,191,1903]" captionTargetId="figure-20@171.[205,1382,181,1920]" captionTargetPageId="171" captionText="FIGURE 38. A–D: Maladera rufonitida Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. songi Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. detersa (Erichson, 1834) (Vietnam: DaNang). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496408" httpUri="https://zenodo.org/record/4496408/files/figure.png" pageId="170" pageNumber="171">Fig. 38D</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCBFF6FAFD9FA00FCAC3FE5" pageId="170" pageNumber="171" type="diagnosis">
<paragraph id="8BAB36F06BCBFF6FAFD9FA00FCAC3FE5" blockId="170.[151,1437,151,826]" pageId="170" pageNumber="171">
<emphasis id="B960EAE26BCBFF6FAFD9FA00FE9C3FC8" bold="true" box="[199,322,618,644]" pageId="170" pageNumber="171">Diagnosis.</emphasis>
<taxonomicName id="4C144D736BCBFF6FAE56FA01FCF23FC9" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[328,812,619,645]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="rufonitida" status="sp. nov.">
<emphasis id="B960EAE26BCBFF6FAE56FA01FDF23FC9" box="[328,556,619,645]" italics="true" pageId="170" pageNumber="171">Maladera rufonitida</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996BCBFF6FAC29FA01FCAC3FC9" box="[823,882,619,645]" pageId="170" pageNumber="171" rank="species">sp. n.</taxonomicNameLabel>
differs from the similar
<taxonomicName id="4C144D736BCBFF6FAB9EFA01FA963FC9" authority="(Moser)" authorityName="Fabrizi &amp; Liu &amp; Bai &amp; Yang &amp; Ahrens" authorityYear="2021" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[1152,1352,619,645]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="170" pageNumber="171" phylum="Arthropoda" rank="species" species="nitens">
<emphasis id="B960EAE26BCBFF6FAB9EFA01FB363FC8" box="[1152,1256,619,644]" italics="true" pageId="170" pageNumber="171">M. nitens</emphasis>
(Moser)
</taxonomicName>
in having the right paramere abruptly and strongly bent after basal half.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCBFF6FAFD9FAD9FB393FBD" pageId="170" pageNumber="171" type="etymology">
<paragraph id="8BAB36F06BCBFF6FAFD9FAD9FB393FBD" blockId="170.[151,1437,151,826]" pageId="170" pageNumber="171">
<emphasis id="B960EAE26BCBFF6FAFD9FAD9FE903F81" bold="true" box="[199,334,691,717]" pageId="170" pageNumber="171">Etymology.</emphasis>
The name (adjective in nominative singular) of the new species is derrived from the combined Latin words rufus (red) and nitidus (shiny), with reference on the body integumaent of the species.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCBFF6FAFD9FA90FBBA3E59" box="[199,1124,762,789]" pageId="170" pageNumber="171" type="description">
<paragraph id="8BAB36F06BCBFF6FAFD9FA90FBBA3E59" blockId="170.[151,1437,151,826]" box="[199,1124,762,789]" pageId="170" pageNumber="171">
<emphasis id="B960EAE26BCBFF6FAFD9FA90FEE13E58" bold="true" box="[199,319,762,788]" pageId="170" pageNumber="171">Variation.</emphasis>
Length:
<quantity id="4CEC9B156BCBFF6FAEBAFA91FDF33E59" box="[420,557,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" metricValueMax="8.9" metricValueMin="8.1" pageId="170" pageNumber="171" unit="mm" value="8.5" valueMax="8.9" valueMin="8.1">8.1–8.9 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156BCBFF6FADECFA90FCA53E59" box="[754,891,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" metricValueMax="6.5" metricValueMin="5.9" pageId="170" pageNumber="171" unit="mm" value="6.2" valueMax="6.5" valueMin="5.9">5.9–6.5 mm</quantity>
, width:
<quantity id="4CEC9B156BCBFF6FACCBFA90FB813E59" box="[981,1119,762,789]" metricMagnitude="-3" metricUnit="m" metricValue="4.949999999999999" metricValueMax="5.1" metricValueMin="4.8" pageId="170" pageNumber="171" unit="mm" value="4.949999999999999" valueMax="5.1" valueMin="4.8">4.8–5.1 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6BCBFF6FAFD9FB74FD103E75" box="[199,718,798,826]" pageId="170" pageNumber="171" type="distribution">
<paragraph id="8BAB36F06BCBFF6FAFD9FB74FD103E75" blockId="170.[151,1437,151,826]" box="[199,718,798,826]" pageId="170" pageNumber="171">
<emphasis id="B960EAE26BCBFF6FAFD9FB74FEBF3E74" bold="true" box="[199,353,798,824]" pageId="170" pageNumber="171">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756BCBFF6FAEC8FB75FDE83E76" box="[470,566,799,826]" captionStart="FIGURE 106" captionStartId="354.[151,250,1982,2006]" captionTargetBox="[208,1378,1133,1950]" captionTargetId="figure-48@354.[202,1386,1127,1958]" captionTargetPageId="354" captionText="FIGURE 106. Distribution of the Maladera nitens group: Maladera paranitens sp. n., M. pseudonitens sp. n., M. rufonitida sp. n., M. songi sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496604" httpUri="https://zenodo.org/record/4496604/files/figure.png" pageId="170" pageNumber="171">Fig. 106</figureCitation>
) and
<tableCitation id="C696034B6BCBFF6FAD6AFB75FD163E75" box="[628,712,799,825]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="170" pageNumber="171">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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