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<document id="31FF0361F2B186100649CBEDB51B4697" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66B01FFA7AF89FF58FD1E3F2D" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera fereobscurata Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="9" lastPageNumber="99" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="97" updateTime="1698926631736" updateUser="ExternalLinkService">
<mods:mods id="E92DCCD41773998E8D57BA3E0C6A2F4E" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:title id="C5A86E07470D4FEB0C1A521E897CB64D">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="72F324EF5974CEB46D7F188538BB3CCD">Fabrizi, Silvia</mods:namePart>
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<mods:nameIdentifier id="7ACADAC9273C4BE4EA224666D32610B7" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:namePart id="2F6872634F5F2A34F1981BC87EAD70CD">Liu, Wan-Gang</mods:namePart>
<mods:nameIdentifier id="FAF6123BF58EDA2A6677565655BB6BD8" type="ORCID">0000-0003-4788-7967</mods:nameIdentifier>
<mods:nameIdentifier id="1DE3BF3D3C87330C44F50B95E7901180" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:roleTerm id="5380FFF95B12896763A46A277CFCEDFB">Author</mods:roleTerm>
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<mods:namePart id="BBDE9914A310DE0F1B4C9F1C65101B02">Bai, Ming</mods:namePart>
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<mods:namePart id="B1148BEA363903D877E81A7303F7F4F6">Yang, Xing-Ke</mods:namePart>
<mods:nameIdentifier id="6587365F2E691718862196E56FD7E660" type="ORCID">0000-0003-3676-6828</mods:nameIdentifier>
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<mods:namePart id="777EB8F2F8FB03C025AAAAAAFEA8CBA3">Ahrens, Dirk</mods:namePart>
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<mods:title id="F4D459BC185EBDA57844F988CC7E7939">Zootaxa</mods:title>
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<mods:date id="DEE007DF36D1A6227E680C5ECA0C6CCB">2021</mods:date>
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<treatment id="03BD87E66B01FFA7AF89FF58FD1E3F2D" ID-DOI="http://doi.org/10.5281/zenodo.4534532" ID-GBIF-Taxon="177323708" ID-Zenodo-Dep="4534532" LSID="urn:lsid:plazi:treatment:03BD87E66B01FFA7AF89FF58FD1E3F2D" httpUri="http://treatment.plazi.org/id/03BD87E66B01FFA7AF89FF58FD1E3F2D" lastPageId="98" lastPageNumber="99" pageId="96" pageNumber="97">
<subSubSection id="C30E657B6B01FFA5AF89FF58FCF43A01" box="[151,810,1842,1869]" pageId="96" pageNumber="97" type="nomenclature">
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<heading id="D0E3819C6B01FFA5AF89FF58FCF43A01" bold="true" box="[151,810,1842,1869]" fontSize="11" level="2" pageId="96" pageNumber="97" reason="4">
<emphasis id="B960EAE26B01FFA5AF89FF58FCF43A01" bold="true" box="[151,810,1842,1869]" pageId="96" pageNumber="97">
<taxonomicName id="4C144D736B01FFA5AF89FF58FD053A00" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,731,1842,1869]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="fereobscurata" status="sp. nov.">
<emphasis id="B960EAE26B01FFA5AF89FF58FE643A01" bold="true" box="[151,442,1842,1869]" italics="true" pageId="96" pageNumber="97">Maladera fereobscurata</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B01FFA5ADF6FF59FCF43A01" box="[744,810,1843,1869]" pageId="96" pageNumber="97" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B01FFA5AF89FF3DFEB23A3D" box="[151,364,1878,1905]" pageId="96" pageNumber="97" type="description">
<paragraph id="8BAB36F06B01FFA5AF89FF3DFEB23A3D" blockId="96.[151,810,1842,1905]" box="[151,364,1878,1905]" pageId="96" pageNumber="97">
<figureCitation id="132F2A756B01FFA5AF89FF3DFED03A3D" box="[151,270,1879,1905]" captionStart="FIGURE 20" captionStartId="95.[151,250,1890,1914]" captionTargetBox="[222,1366,195,1857]" captionTargetId="figure-20@95.[211,1377,181,1865]" captionTargetPageId="95" captionText="FIGURE 20. A–D: Maladera shengqiaoae Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. fereobscurata Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. saitoi (Niijima &amp; Kinoshita, 1927) (Laos: Borikhamai). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496358" httpUri="https://zenodo.org/record/4496358/files/figure.png" pageId="96" pageNumber="97">Figures 20</figureCitation>
E–H, 95
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B01FFA4AF89FFF5FAA33FBD" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="materials_examined">
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<emphasis id="B960EAE26B01FFA5AF89FFF5FE1F3AF4" bold="true" box="[151,449,1950,1977]" pageId="96" pageNumber="97">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6B01FFA5AED6FFF5FC9E3A91" ID-GBIF-Occurrence="3034555564" lastPageId="96" lastPageNumber="98" pageId="96" pageNumber="97" typeStatus="holotype">
<typeStatus id="54AF88526B01FFA5AED6FFF5FDEB3AF5" box="[456,565,1951,1977]" pageId="96" pageNumber="97" type="holotype">Holotype</typeStatus>
:
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“China, W
<collectingRegion id="49D0F8126B01FFA5ADC5FFF5FCAA3AF5" box="[731,884,1951,1977]" country="China" name="Yunnan" pageId="96" pageNumber="97">Yunnan prov.</collectingRegion>
, mts.
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SW
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,
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</quantity>
,
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</date>
,
<collectorName id="26E153266B01FFA5AF89FFA9FED63A91" box="[151,264,1987,2013]" pageId="96" pageNumber="97">S. Murzin</collectorName>
&amp;
<collectorName id="26E153266B01FFA5AE2EFFA9FE7B3A91" box="[304,421,1987,2013]" pageId="96" pageNumber="97">I. Shokhin</collectorName>
/ 860
<taxonomicName id="4C144D736B01FFA5AEF6FFA9FD9A3A91" box="[488,580,1987,2013]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="380" phylum="Arthropoda" rank="tribe" tribe="Sericini">Sericini</taxonomicName>
: Asia spec.” (
<collectionCode id="ED05AE356B01FFA5ADFEFFA9FCEA3A91" box="[736,820,1987,2013]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="96" pageNumber="97" type="Museum">ZFMK</collectionCode>
).
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<materialsCitation id="3B7C3CAD6B01FFA4AC56FFA9FB133DFD" ID-GBIF-Occurrence="3034555619" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" typeStatus="paratype">
<typeStatus id="54AF88526B01FFA5AC56FFA9FC673A91" box="[840,953,1987,2013]" pageId="96" pageNumber="97" type="paratype">Paratypes</typeStatus>
:
<emphasis id="B960EAE26B01FFA5ACDBFFA8FBC93A90" bold="true" box="[965,1047,1986,2012]" pageId="96" pageNumber="97">
<collectingCountry id="F30376606B01FFA5ACDBFFA8FBCC3A90" box="[965,1042,1986,2012]" name="China" pageId="96" pageNumber="97">China</collectingCountry>
:
</emphasis>
<specimenCount id="9D12FD796B01FFA5AB3EFFA9FBAE3A91" box="[1056,1136,1987,2013]" pageId="96" pageNumber="97" type="male">17 ♂♂</specimenCount>
,
<specimenCount id="9D12FD796B01FFA5AB60FFA9FB603A91" box="[1150,1214,1987,2013]" pageId="96" pageNumber="97" type="female">4 ♀♀</specimenCount>
“
<collectorName id="26E153266B01FFA5ABCFFFA9FAE13A91" box="[1233,1343,1987,2013]" pageId="96" pageNumber="97">China, W</collectorName>
<collectingRegion id="49D0F8126B01FFA5AA59FFA9FA433A91" box="[1351,1437,1987,2013]" country="China" name="Yunnan" pageId="96" pageNumber="97">Yunnan</collectingRegion>
prov., mts.
<quantity id="4CEC9B156B00FFA4AE07F8FDFE873DFD" box="[281,345,151,177]" metricMagnitude="4" metricUnit="m" metricValue="2.0" pageId="97" pageNumber="98" unit="km" value="20.0">20km</quantity>
SW Baoshan,
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,
<date id="FFAA10306B00FFA4AD7AF8FCFCDF3DFD" box="[612,769,150,177]" pageId="97" pageNumber="98" value="2006-05-23" valueMax="2006-05-25" valueMin="2006-05-23">
<collectingDate id="EFEEE9D86B00FFA4AD7AF8FCFCDF3DFD" box="[612,769,150,177]" pageId="97" pageNumber="98" value="2006-05-23" valueMax="2006-05-25" valueMin="2006-05-23">23.-25.v.2006</collectingDate>
</date>
,
<collectorName id="26E153266B00FFA4AC11F8FDFC5F3DFD" box="[783,897,151,177]" pageId="97" pageNumber="98">S. Murzin</collectorName>
&amp;
<collectorName id="26E153266B00FFA4ACB4F8FDFBFD3DFD" box="[938,1059,151,177]" pageId="97" pageNumber="98">I. Shokhin</collectorName>
” (
<collectionCode id="ED05AE356B00FFA4AB20F8FDFBBC3DFD" box="[1086,1122,151,177]" pageId="97" pageNumber="98">CP</collectionCode>
,
<collectionCode id="ED05AE356B00FFA4AB71F8FDFB1F3DFD" box="[1135,1217,151,177]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34613" lsid="urn:lsid:biocol.org:col:34613" name="Zoologisches Forschungsmuseum Alexander Koenig" pageId="97" pageNumber="98" type="Museum">ZFMK</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B7C3CAD6B00FFA4ABC9F8FDFE5E3D99" ID-GBIF-Occurrence="3034556688" lastPageId="97" lastPageNumber="98" pageId="97" pageNumber="97">
<specimenCount id="9D12FD796B00FFA4ABC9F8FDFADD3DFD" box="[1239,1283,151,177]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6B00FFA4AA08F8FDFA463DFD" LSID="urn:lsid:plazi:treatment:03BD87E66B01FFA7AF89FF58FD1E3F2D:8ECB602B6B00FFA4AA08F8FDFA463DFD" box="[1302,1432,151,177]" name="Tengchong" pageId="97" pageNumber="98">Tengchong</location>
,
<date id="FFAA10306B00FFA4AF89F8D1FEC93D99" box="[151,279,185,213]" pageId="97" pageNumber="98" value="1983-05-24">
<collectingDate id="EFEEE9D86B00FFA4AF89F8D1FEC93D99" box="[151,279,185,213]" pageId="97" pageNumber="98" value="1983-05-24">24.Ⅴ.1983</collectingDate>
</date>
” (
<collectionCode id="ED05AE356B00FFA4AE2EF8D1FEAD3D99" box="[304,371,187,213]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="97" pageNumber="98">IZAS</collectionCode>
),
</materialsCitation>
<materialsCitation id="3B7C3CAD6B00FFA4AE99F8D1FEDE3DB5" ID-GBIF-Occurrence="3034556724" lastPageId="97" lastPageNumber="98" pageId="97" pageNumber="97">
<specimenCount id="9D12FD796B00FFA4AE99F8D1FE6E3D99" box="[391,432,187,213]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6B00FFA4AEDEF8D1FDC73D99" LSID="urn:lsid:plazi:treatment:03BD87E66B01FFA7AF89FF58FD1E3F2D:8ECB602B6B00FFA4AEDEF8D1FDC73D99" box="[448,537,187,213]" name="Yunnan" pageId="97" pageNumber="98">Yunnan</location>
<quantity id="4CEC9B156B00FFA4AD3EF8D0FD713D99" box="[544,687,186,213]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="97" pageNumber="98" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">
<elevation id="0039D1C36B00FFA4AD3EF8D0FD713D99" box="[544,687,186,213]" metricMagnitude="3" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="97" pageNumber="98" unit="m" value="2250.0" valueMax="2500.0" valueMin="2000.0">2000-2500m</elevation>
</quantity>
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“China: N-Yunnan, Baiyungshan (Rai Railing Mts.)
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, Yong Ren,
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<collectingDate id="EFEEE9D86B00FFA4ABB2F8B5FACA3DB5" box="[1196,1300,223,249]" pageId="97" pageNumber="98" value="2003-07">VII-2003</collectingDate>
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leg.
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<emphasis id="B960EAE26B00FFA4AA96F8B5FF6A3C51" italics="true" pageId="97" pageNumber="98">et al.</emphasis>
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Asia spec.” (
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<collectingDate id="EFEEE9D86B00FFA4AC87F969FC373C51" box="[921,1001,259,285]" pageId="97" pageNumber="98" value="1940-05">V.1940</collectingDate>
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<collectorName id="26E153266B00FFA4ACF0F969FB693C51" box="[1006,1207,259,285]" pageId="97" pageNumber="98">M.S. King T. Maa</collectorName>
” (
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spTHAI_DE09_4teinzoanaGr
<collectingMunicipality id="6BCFAC8A6B00FFA4ADEFF94DFC8B3C0D" box="[753,853,295,321]" pageId="97" pageNumber="98">Thailand</collectingMunicipality>
L.
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2.-
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<collectingDate id="EFEEE9D86B00FFA4AB15F94CFBAD3C0D" box="[1035,1139,294,321]" pageId="97" pageNumber="98" value="2009-05-07">7.5.2009</collectingDate>
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<collectorName id="26E153266B00FFA4AB69F94DFB263C0D" box="[1143,1272,295,321]" pageId="97" pageNumber="98">Ang Khang</collectorName>
<collectingRegion id="49D0F8126B00FFA4ABE0F94DFED63C29" country="Thailand" name="Chiang Mai" pageId="97" pageNumber="98">region Chiang Mai Prov.</collectingRegion>
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” (
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<specimenCount id="9D12FD796B00FFA4AC14F921FCE83C29" box="[778,822,331,357]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
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,
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<quantity id="4CEC9B156B00FFA4AB66F921FB663C29" box="[1144,1208,331,357]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="97" pageNumber="98" unit="km" value="80.0">80km</quantity>
NE of
<collectingRegion id="49D0F8126B00FFA4AA10F921FA463C29" box="[1294,1432,331,357]" country="Thailand" name="Chiang Mai" pageId="97" pageNumber="98">Chiang Mai</collectingRegion>
</locationDeviation>
; 1995” (
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),
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<specimenCount id="9D12FD796B00FFA4AE4AF905FE4B3CC5" box="[340,405,367,393]" pageId="97" pageNumber="98" type="male">2 ♂♂</specimenCount>
“N-Thailand 29.5.-
<date id="FFAA10306B00FFA4AD6EF904FD093CC5" box="[624,727,366,393]" pageId="97" pageNumber="98" value="1989-06-05">
<collectingDate id="EFEEE9D86B00FFA4AD6EF904FD093CC5" box="[624,727,366,393]" pageId="97" pageNumber="98" value="1989-06-05">5.6.1989</collectingDate>
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<collectorName id="26E153266B00FFA4ADFEF905FCA73CC5" box="[736,889,367,393]" pageId="97" pageNumber="98">Doi Inthanon</collectorName>
,
<collectorName id="26E153266B00FFA4AC98F905FC283CC5" box="[902,1014,367,393]" pageId="97" pageNumber="98">Lichtfalle</collectorName>
,
<location id="8ECB602B6B00FFA4AB1CF905FB0F3CC5" LSID="urn:lsid:plazi:treatment:03BD87E66B01FFA7AF89FF58FD1E3F2D:8ECB602B6B00FFA4AB1CF905FB0F3CC5" box="[1026,1233,367,393]" country="Thailand" name="Bang Khun Klang" pageId="97" pageNumber="98" stateProvince="Chiang Mai">Bang Khun Klang</location>
<quantity id="4CEC9B156B00FFA4ABC4F905FAF43CC5" box="[1242,1322,367,393]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="97" pageNumber="98" unit="m" value="1200.0">
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,
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<collectorName id="26E153266B00FFA4AE1CF9F9FE793CE1" box="[258,423,403,429]" pageId="97" pageNumber="98">Chantamonkol</collectorName>
&amp;
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leg.” (
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),
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<collectingDate id="EFEEE9D86B00FFA4ACA8F9F8FB8B3CE1" box="[950,1109,402,429]" pageId="97" pageNumber="98" value="1990-05-25" valueMax="1990-05-29" valueMin="1990-05-25">25.-29.V.1990</collectingDate>
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<collectorName id="26E153266B00FFA4AB41F9F9FA833CE1" box="[1119,1373,403,429]" pageId="97" pageNumber="98">Doi Inthanon Malicky</collectorName>
leg.” (
<collectionCode id="ED05AE356B00FFA4AF81F9DDFF003C9D" box="[159,222,439,465]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="97" pageNumber="98" type="Museum">ZSM</collectionCode>
),
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“N-Thailand
<date id="FFAA10306B00FFA4AED3F9DDFDC33C9D" box="[461,541,439,465]" pageId="97" pageNumber="98" value="1990-05">
<collectingDate id="EFEEE9D86B00FFA4AED3F9DDFDC33C9D" box="[461,541,439,465]" pageId="97" pageNumber="98" value="1990-05">V.1990</collectingDate>
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<collectorName id="26E153266B00FFA4AD3AF9DDFCC33C9D" box="[548,797,439,465]" pageId="97" pageNumber="98">Doi Inthanon Malicky</collectorName>
leg.” (
<collectionCode id="ED05AE356B00FFA4AC74F9DDFC773C9D" box="[874,937,439,465]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="97" pageNumber="98" type="Museum">ZSM</collectionCode>
),
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<specimenCount id="9D12FD796B00FFA4ACA2F9DDFC393C9D" box="[956,999,439,465]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
“
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<date id="FFAA10306B00FFA4AB2BF9DDFB6F3C9D" box="[1077,1201,439,465]" pageId="97" pageNumber="98" value="1993-05-01" valueMax="1993-05-08" valueMin="1993-05-01">
<collectingDate id="EFEEE9D86B00FFA4AB2BF9DDFB6F3C9D" box="[1077,1201,439,465]" pageId="97" pageNumber="98" value="1993-05-01" valueMax="1993-05-08" valueMin="1993-05-01">1-8.V.1993</collectingDate>
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Soppong Pai
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<quantity id="4CEC9B156B00FFA4AA50F9DDFA433C9D" box="[1358,1437,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.8" pageId="97" pageNumber="98" unit="m" value="1800.0">
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<locationDeviation id="9A192FD26B00FFA4AF89F9B1FECD3CB9" box="[151,275,475,501]" pageId="97" pageNumber="98">Pacholatko</locationDeviation>
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&amp;
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/ TS3/ 2
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Asia spec.” (
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).
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<collectingCountry id="F30376606B00FFA4AC21F9B1FCA33CB9" box="[831,893,475,501]" name="Laos" pageId="97" pageNumber="98">Laos</collectingCountry>
:
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<specimenCount id="9D12FD796B00FFA4AC94F9B1FC683CB9" box="[906,950,475,501]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
“
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,
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,
<collectingRegion id="49D0F8126B00FFA4ABEAF9B1FA423CB9" box="[1268,1436,475,501]" country="Laos" name="Louangnamtha" pageId="97" pageNumber="98">Louangnamtha</collectingRegion>
pr. Namtha-MuangSing,
<date id="FFAA10306B00FFA4AEADF994FDE53F55" box="[435,571,510,537]" pageId="97" pageNumber="98" value="1997-05-05" valueMax="1997-05-31" valueMin="1997-05-05">5-31.v.1997</date>
, 900-
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,
<collectorName id="26E153266B00FFA4ADFBF995FC853F55" box="[741,859,511,537]" pageId="97" pageNumber="98">Vit Kubán</collectorName>
leg.” (
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),
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<specimenCount id="9D12FD796B00FFA4ACF4F995FBC93F55" box="[1002,1047,511,537]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
“
<location id="8ECB602B6B00FFA4AB34F995FBBD3F55" LSID="urn:lsid:plazi:treatment:03BD87E66B01FFA7AF89FF58FD1E3F2D:8ECB602B6B00FFA4AB34F995FBBD3F55" box="[1066,1123,511,537]" country="Laos" name="Laos" pageId="97" pageNumber="98" stateProvince="Louangnamtha">Laos</location>
north,
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, 20 km NW
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N 21°09,2‘, E 101°18,7‘, alt. 900+
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&amp;
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:
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“N-Vietnam,
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<collectingRegion id="49D0F8126B00FFA4AEA8FA2DFDCD3F2D" box="[438,531,583,609]" country="Vietnam" name="Lao Cai" pageId="97" pageNumber="98">Lao Cai</collectingRegion>
,
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, N22°19‘52‘‘; E103°50‘;
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leg.
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,
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,
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” (
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<collectorName id="26E153266B00FFA4AEBEFA01FDF93FC9" box="[416,551,619,645]" pageId="97" pageNumber="98">N Viet Nam</collectorName>
(
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) pr.
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<date id="FFAA10306B00FFA4ACA7FA00FB8C3FC9" box="[953,1106,618,645]" pageId="97" pageNumber="98" value="1990-05-11" valueMax="1990-05-15" valueMin="1990-05-11">
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<collectorName id="26E153266B00FFA4AB47FA01FB303FC9" box="[1113,1262,619,645]" pageId="97" pageNumber="98">P. Pacholátko</collectorName>
leg.” (
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),
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“S
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,
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NW of An Khe Buon Luoi
</location>
,
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28.3.-
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<collectingDate id="EFEEE9D86B00FFA4AB11FAE4FB5D3FE5" box="[1039,1155,654,681]" pageId="97" pageNumber="98" value="1995-12-04">12.4.1995</collectingDate>
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&amp;
<collectorName id="26E153266B00FFA4AA36FAE5FA433FE5" box="[1320,1437,655,681]" pageId="97" pageNumber="98">Dembicky</collectorName>
leg.” (
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),
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<specimenCount id="9D12FD796B00FFA4AE09FAD9FE9F3F81" box="[279,321,691,717]" pageId="97" pageNumber="98" type="male">1 ♂</specimenCount>
“
<collectorName id="26E153266B00FFA4AE4AFAD9FE123F81" box="[340,460,691,717]" pageId="97" pageNumber="98">S Vietnam</collectorName>
,
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N of Dalat-Lang Bian
</locationDeviation>
,
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<date id="FFAA10306B00FFA4AB68FAD8FAC63F81" box="[1142,1304,690,717]" pageId="97" pageNumber="98" value="1995-04-17" valueMax="1995-04-21" valueMin="1995-04-17">
<collectingDate id="EFEEE9D86B00FFA4AB68FAD8FAC63F81" box="[1142,1304,690,717]" pageId="97" pageNumber="98" value="1995-04-17" valueMax="1995-04-21" valueMin="1995-04-17">17.-21.iv.1995</collectingDate>
</date>
<collectingCounty id="62CA4E7C6B00FFA4AA3EFAD9FA423F81" box="[1312,1436,691,717]" pageId="97" pageNumber="98">Pacholátko</collectingCounty>
&amp;
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leg.” (
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).
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<collectingCountry id="F30376606B00FFA4AEB1FABDFDF53FBD" box="[431,555,727,753]" name="Myanmar" pageId="97" pageNumber="98">Myanmar</collectingCountry>
:
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“
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: SW
<collectingRegion id="49D0F8126B00FFA4AC1EFABDFCA43FBD" box="[768,890,727,753]" country="Myanmar" name="Shan" pageId="97" pageNumber="98">Shan State</collectingRegion>
,
<location id="8ECB602B6B00FFA4AC9BFABDFC2C3FBD" LSID="urn:lsid:plazi:treatment:03BD87E66B01FFA7AF89FF58FD1E3F2D:8ECB602B6B00FFA4AC9BFABDFC2C3FBD" box="[901,1010,727,753]" country="Myanmar" county="Pacholatko" municipality="Burma" name="Taunggyi" pageId="97" pageNumber="98" stateProvince="Shan">Taunggyi</location>
;
<date id="FFAA10306B00FFA4ACE3FABDFB4B3FBD" box="[1021,1173,727,753]" pageId="97" pageNumber="98" value="1997-06-01" valueMax="1997-06-18" valueMin="1997-06-01">
<collectingDate id="EFEEE9D86B00FFA4ACE3FABDFB4B3FBD" box="[1021,1173,727,753]" pageId="97" pageNumber="98" value="1997-06-01" valueMax="1997-06-18" valueMin="1997-06-01">1.-18.vi.1997</collectingDate>
</date>
;
<collectorName id="26E153266B00FFA4ABBCFABDFB203FBD" box="[1186,1278,727,753]" pageId="97" pageNumber="98">J. Kalab</collectorName>
leg.” (
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)
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.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B00FFA7AFD9FA90FD783C51" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98" type="description">
<paragraph id="8BAB36F06B00FFA4AFD9FA90FBA23E75" blockId="97.[151,1437,150,2013]" pageId="97" pageNumber="98">
<emphasis id="B960EAE26B00FFA4AFD9FA90FE873E58" bold="true" box="[199,345,762,788]" pageId="97" pageNumber="98">Description.</emphasis>
Length:
<quantity id="4CEC9B156B00FFA4AEA2FA91FDCD3E59" box="[444,531,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="6.4" pageId="97" pageNumber="98" unit="mm" value="6.4">6.4 mm</quantity>
, length of elytra: 4.0 mm, width:
<quantity id="4CEC9B156B00FFA4AC9AFA91FC023E59" box="[900,988,763,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="97" pageNumber="98" unit="mm" value="3.6">3.6 mm</quantity>
. Body oval, reddish brown, antenna yellow, dorsal surface dull, except some single setae on head dorsal surface nearly glabrous.
</paragraph>
<paragraph id="8BAB36F06B00FFA4AFD9FB29FB7A3931" blockId="97.[151,1437,150,2013]" pageId="97" pageNumber="98">
Labroclypeus moderately wide and trapezoidal, shiny, widest at base, lateral margins weakly convex and convergent anteriorly, anterior angles blunt and slightly reflexed, anterior margin moderately emarginate, margins moderately reflexed; lateral margin and ocular canthus produce a blunt angle; surface flat, finely and densely punctate, with numerous coarse punctures each bearing an erect seta; frontoclypeal suture finely incised, nearly straight; smooth area anterior to eye convex, as twice as wide as long; ocular canthus long and moderately wide (1/3 of ocular diameter), finely and sparsely punctate, with a terminal seta. Frons dull, with fine, sparse punctures and a few single erect setae beside eyes. Eyes very large, ratio diameter/ interocular width: 0.93. Antenna with ten antennomeres; club with three antennomeres and weakly reflexed, 1.7 times as long as remaining antennomeres combined, antennomeres 3-6 nearly twice as long as wide.
<collectingRegion id="49D0F8126B00FFA4AD6AFC09FD0D3931" box="[628,723,1123,1149]" country="Myanmar" name="Chin" pageId="97" pageNumber="98">Mentum</collectingRegion>
elevated and slightly flattened anteriorly.
</paragraph>
<paragraph id="8BAB36F06B00FFA4AFD9FCEDFA5B387D" blockId="97.[151,1437,150,2013]" pageId="97" pageNumber="98">Pronotum moderately transverse, widest at base, lateral margins evenly convex, moderately convergent anteriorly, anterior angles moderately produced and right-angled, posterior angles blunt, weakly rounded at tip; anterior margin convex, with fine and complete marginal line being interrupted medially, base without marginal line; surface finely and densely punctate, with minute setae in punctures, otherwise glabrous; lateral and anterior margin finely setose; hypomeron carinate, not produced ventrally. Scutellum wide, triangular, dull, with fine, sparse punctures.</paragraph>
<paragraph id="8BAB36F06B00FFA4AFD9FD51FA57388E" blockId="97.[151,1437,150,2013]" pageId="97" pageNumber="98">Elytra widest shortly behind middle, striae distinctly impressed, finely and sparsely punctate, intervals weakly convex, with fine and sparse punctures concentrated along striae, with only minute setae in punctures, otherwise glabrous; epipleural margin robust, ending at strongly curved external apical angle of elytra, epipleura densely setose; apical border of elytra not membraneous, without visible rim of microtrichomes (at ca 100x magnification).</paragraph>
<paragraph id="8BAB36F06B00FFA4AFD9FDA1FA493B39" blockId="97.[151,1437,150,2013]" pageId="97" pageNumber="98">Ventral surface dull, coarsely and moderately densely punctate, metasternum sparsely covered with sparse fine setae; metacoxa with a few longer setae laterally. Abdominal sternites finely and irregularly densely punctate, glabrous, each sternite with a transverse row of coarse punctures each bearing a robust long seta. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.5. Pygidium moderately convex and dull, coarsely and densely punctate, without smooth midline, glabrous except a few setae along apical margin.</paragraph>
<paragraph id="8BAB36F06B00FFA7AFD9FE15FE333DB5" blockId="97.[151,1437,150,2013]" lastBlockId="98.[151,1437,151,609]" lastPageId="98" lastPageNumber="99" pageId="97" pageNumber="98">Legs moderately wide and short, weakly shiny; femora with two longitudinal rows of setae, finely and sparsely punctate. Metafemur dull, with anterior margin acute, with continuous serrated line behind anterior margin, setae of anterior row very short; posterior margin smooth and moderately widened at apex ventrally, not serrate dorsally, with just a few short setae basally. Metatibia short, widest at apex, ratio of width/length: 1/3.16, sharply carinate dorsally, with two groups of spines, basal group at two thirds, apical group at five sixths of metatibial length, with a few short robust setae basally and a continuously serrated line beside dorsal margin from base to basal group of spines; lateral face weakly longitudinally convex, sparsely punctate, with minute setae in punctures; ventral margin finely serrate, with four robust setae of which the apical one is more distant; medial face smooth and glabrous; apex finely serrate, moderately truncate interiorly near tarsal articulation. Tarsomeres dorsally smooth and glabrous, neither laterally nor dorsally carinate, sparsely setose ventrally; metatarsomeres with a strongly serrated ridge ventrally and a smooth subventral longitudinal carina; first metatarsomere slightly shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate; anterior claws symmetrical, basal tooth of both claws bluntly truncate at apex.</paragraph>
<paragraph id="8BAB36F06B03FFA7AFD9F969FD783C51" blockId="98.[151,1437,151,609]" box="[199,678,259,285]" pageId="98" pageNumber="99">
Aedeagus:
<figureCitation id="132F2A756B03FFA7AE5AF969FE4B3C51" box="[324,405,259,285]" captionStart="FIGURE 20" captionStartId="95.[151,250,1890,1914]" captionTargetBox="[222,1366,195,1857]" captionTargetId="figure-20@95.[211,1377,181,1865]" captionTargetPageId="95" captionText="FIGURE 20. A–D: Maladera shengqiaoae Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. fereobscurata Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. saitoi (Niijima &amp; Kinoshita, 1927) (Laos: Borikhamai). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496358" httpUri="https://zenodo.org/record/4496358/files/figure.png" pageId="98" pageNumber="99">Fig. 20</figureCitation>
E–G. Habitus:
<figureCitation id="132F2A756B03FFA7AD25F969FD7C3C51" box="[571,674,259,285]" captionStart="FIGURE 20" captionStartId="95.[151,250,1890,1914]" captionTargetBox="[222,1366,195,1857]" captionTargetId="figure-20@95.[211,1377,181,1865]" captionTargetPageId="95" captionText="FIGURE 20. A–D: Maladera shengqiaoae Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); E–H: M. fereobscurata Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); I–L: M. saitoi (Niijima &amp; Kinoshita, 1927) (Laos: Borikhamai). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496358" httpUri="https://zenodo.org/record/4496358/files/figure.png" pageId="98" pageNumber="99">Fig. 20H</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B03FFA7AFD9F94CFA8C3C29" pageId="98" pageNumber="99" type="diagnosis">
<paragraph id="8BAB36F06B03FFA7AFD9F94CFA8C3C29" blockId="98.[151,1437,151,609]" pageId="98" pageNumber="99">
<emphasis id="B960EAE26B03FFA7AFD9F94CFE9C3C0C" bold="true" box="[199,322,294,320]" pageId="98" pageNumber="99">Diagnosis.</emphasis>
<taxonomicName id="4C144D736B03FFA7AE58F94DFC923C0D" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[326,844,295,321]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="fereobscurata" status="sp. nov.">
<emphasis id="B960EAE26B03FFA7AE58F94DFD8A3C0D" box="[326,596,295,321]" italics="true" pageId="98" pageNumber="99">Maladera fereobscurata</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996B03FFA7AC4BF94DFC513C0D" box="[853,911,295,321]" pageId="98" pageNumber="99" rank="species">sp. n.</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C144D736B03FFA7AB50F94DFA493C0D" authority="(Brenske, 1898)" baseAuthorityName="Brenske" baseAuthorityYear="1898" box="[1102,1431,295,321]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="teinzoana">
<emphasis id="B960EAE26B03FFA7AB50F94DFB003C0C" box="[1102,1246,295,320]" italics="true" pageId="98" pageNumber="99">M. teinzoana</emphasis>
(
<bibRefCitation id="EF854B016B03FFA7ABF5F94DFA513C0D" author="Brenske, E." box="[1259,1423,295,321]" pageId="98" pageNumber="99" pagination="205 - 403" refId="ref331459" refString="Brenske, E. (1898) Die Serica - Arten der Erde. Berliner Entomologische Zeitschrift, 43, 205 - 403. https: // doi. org / 10.1002 / mmnd. 47918980303" type="journal article" year="1898">Brenske, 1898</bibRefCitation>
)
</taxonomicName>
, differing in having the metatibia longer and the more flattened and widened parameres that lack basal lobes.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B03FFA7AFD9F905FF273C9D" pageId="98" pageNumber="99" type="etymology">
<paragraph id="8BAB36F06B03FFA7AFD9F905FF273C9D" blockId="98.[151,1437,151,609]" pageId="98" pageNumber="99">
<emphasis id="B960EAE26B03FFA7AFD9F905FE903CC5" bold="true" box="[199,334,367,393]" pageId="98" pageNumber="99">Etymology.</emphasis>
The species name (adjective in the nominative singular) is derived from the combined Latin prefix ‘
<emphasis id="B960EAE26B03FFA7AFD6F9F9FF2C3CE1" box="[200,242,403,429]" italics="true" pageId="98" pageNumber="99">fere</emphasis>
-’ (nearly) and the species name
<taxonomicName id="4C144D736B03FFA7AD7AF9F9FD0A3CE1" baseAuthorityName="Moser" baseAuthorityYear="1915" box="[612,724,403,429]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="obscurata">
<emphasis id="B960EAE26B03FFA7AD7AF9F9FD0A3CE1" box="[612,724,403,429]" italics="true" pageId="98" pageNumber="99">obscurata</emphasis>
</taxonomicName>
, with reference to the strong similarity to
<taxonomicName id="4C144D736B03FFA7ABA8F9F9FF2B3C9D" authority="(Moser)" baseAuthorityName="Moser" baseAuthorityYear="1915" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="98" pageNumber="99" phylum="Arthropoda" rank="species" species="obscurata">
<emphasis id="B960EAE26B03FFA7ABA8F9F9FA423CE1" box="[1206,1436,403,429]" italics="true" pageId="98" pageNumber="99">Maladera obscurata</emphasis>
(Moser)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B03FFA7AFD9F9B0FE403F71" pageId="98" pageNumber="99" type="description">
<paragraph id="8BAB36F06B03FFA7AFD9F9B0FE403F71" blockId="98.[151,1437,151,609]" pageId="98" pageNumber="99">
<emphasis id="B960EAE26B03FFA7AFD9F9B0FEE13CB8" bold="true" box="[199,319,474,500]" pageId="98" pageNumber="99">Variation.</emphasis>
Length:
<quantity id="4CEC9B156B03FFA7AEB8F9B0FDEE3CB9" box="[422,560,474,501]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="6.5" metricValueMin="5.5" pageId="98" pageNumber="99" unit="mm" value="6.0" valueMax="6.5" valueMin="5.5">5.5–6.5 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156B03FFA7ADE7F9B1FC5D3CB9" box="[761,899,475,501]" metricMagnitude="-3" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="98" pageNumber="99" unit="mm" value="4.1" valueMax="4.4" valueMin="3.8">3.8–4.4 mm</quantity>
, width: 3.1–4.0 mm. Female: eyes smaller than in male, ratio of diameter/interocular width: 0.7; antennal club as long as remaining antennomeres combined; pygidium weakly convex.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6B03FFA7AFD9FA2CFD1E3F2D" box="[199,704,582,609]" pageId="98" pageNumber="99" type="distribution">
<paragraph id="8BAB36F06B03FFA7AFD9FA2CFD1E3F2D" blockId="98.[151,1437,151,609]" box="[199,704,582,609]" pageId="98" pageNumber="99">
<emphasis id="B960EAE26B03FFA7AFD9FA2CFEBF3F2C" bold="true" box="[199,353,582,608]" pageId="98" pageNumber="99">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756B03FFA7AEC8FA2DFDF63F2D" box="[470,552,582,609]" captionStart="FIGURE 95" captionStartId="349.[151,250,1031,1055]" captionTargetBox="[210,1378,187,1002]" captionTargetId="figure-20@349.[201,1386,181,1008]" captionTargetPageId="349" captionText="FIGURE 95. Distribution of the Maladera teizoana group: Maladera fereobscurata sp. n., M. futschauana (Brenske, 1897), M. invenusta (Moser, 1918), Maladera piceola (Moser, 1915) (not precisely localized collection site indicated by a “?”)." figureDoi="http://doi.org/10.5281/zenodo.4496572" httpUri="https://zenodo.org/record/4496572/files/figure.png" pageId="98" pageNumber="99">Fig. 95</figureCitation>
) and
<tableCitation id="C696034B6B03FFA7AD78FA2DFD643F2D" box="[614,698,583,609]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x – records from material examined herein, o – records from literature, with no material of this study)." pageId="98" pageNumber="99">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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