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<document id="C9095782B6D6E77E0880FDC47FA4326A" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66A64FEC3AF89FF64FD103835" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera shangraoensis Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="263" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="262" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="64DD423BE309AF11EE1F92F7B16606B3">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="C4C856D3C024E8EA278EEC0C45D52A4E">Fabrizi, Silvia</mods:namePart>
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<mods:nameIdentifier id="41FED1C54BD9C9D51E73E2D11AD7F1CC" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:namePart id="8EB9CE6670EF883C91D8E9E7527B279C">Liu, Wan-Gang</mods:namePart>
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<mods:nameIdentifier id="43DD7991BA5AAD11D73F0EAEC19F91B9" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="43C03C68B88BEA832908E18B58A9D3F0">Bai, Ming</mods:namePart>
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<mods:nameIdentifier id="2B6BA552A7833FDE0EE4D1C853FD60A0" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
</mods:name>
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<mods:namePart id="73DA0D70512BD0FA5AEC657DF7AE6B2C">Yang, Xing-Ke</mods:namePart>
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<mods:nameIdentifier id="7F7684817F1D6AD169FCF45BED129B5F" type="email">liuwangang@ieecas.cn</mods:nameIdentifier>
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<mods:namePart id="5C6E13A149F215DB951DD9497FEEA480">Ahrens, Dirk</mods:namePart>
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<mods:nameIdentifier id="D32373A274C6BAC5E5CAFBA73CEF8852" type="email">ahrens.dirk_col@gmx.de</mods:nameIdentifier>
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<mods:title id="747BEF4AA46B7A41D8A8BE64F2EBBDC9">Zootaxa</mods:title>
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<mods:date id="8B85790340302CCD5EF521B75715D581">2021</mods:date>
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<mods:number id="9543B5CEF3D12663FCAEF14C84778A5B">2021-02-03</mods:number>
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<mods:number id="91E99CF5833FFF31E4057AC0F3C12051">4922</mods:number>
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<mods:classification id="AEE32413C91FBCC16A664439770FC1F8">journal article</mods:classification>
<mods:identifier id="217C43ACD5EE3C067C35CB5320AD05B6" type="CLB-Dataset">8257</mods:identifier>
<mods:identifier id="6B3FBDECBF3F5BFA3FDE974045BA7FAF" type="DOI">10.11646/zootaxa.4922.1.1</mods:identifier>
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<treatment id="03BD87E66A64FEC3AF89FF64FD103835" ID-DOI="http://doi.org/10.5281/zenodo.4534815" ID-GBIF-Taxon="177323771" ID-Zenodo-Dep="4534815" LSID="urn:lsid:plazi:treatment:03BD87E66A64FEC3AF89FF64FD103835" httpUri="http://treatment.plazi.org/id/03BD87E66A64FEC3AF89FF64FD103835" lastPageId="262" lastPageNumber="263" pageId="261" pageNumber="262">
<subSubSection id="C30E657B6A64FEC0AF89FF64FCEC3A65" box="[151,818,1806,1833]" pageId="261" pageNumber="262" type="nomenclature">
<paragraph id="8BAB36F06A64FEC0AF89FF64FCEC3A65" blockId="261.[151,818,1806,1869]" box="[151,818,1806,1833]" pageId="261" pageNumber="262">
<heading id="D0E3819C6A64FEC0AF89FF64FCEC3A65" bold="true" box="[151,818,1806,1833]" fontSize="11" level="2" pageId="261" pageNumber="262" reason="4">
<emphasis id="B960EAE26A64FEC0AF89FF64FCEC3A65" bold="true" box="[151,818,1806,1833]" pageId="261" pageNumber="262">
<taxonomicName id="4C144D736A64FEC0AF89FF64FD3D3A64" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,739,1806,1833]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="261" pageNumber="262" phylum="Arthropoda" rank="species" species="shangraoensis" status="sp. nov.">
<emphasis id="B960EAE26A64FEC0AF89FF64FE1C3A65" bold="true" box="[151,450,1806,1833]" italics="true" pageId="261" pageNumber="262">Maladera shangraoensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A64FEC0ADEEFF65FCEC3A65" box="[752,818,1807,1833]" pageId="261" pageNumber="262" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A64FEC0AF89FF59FEA43A01" box="[151,378,1843,1869]" pageId="261" pageNumber="262" type="description">
<paragraph id="8BAB36F06A64FEC0AF89FF59FEA43A01" blockId="261.[151,818,1806,1869]" box="[151,378,1843,1869]" pageId="261" pageNumber="262">
<figureCitation id="132F2A756A64FEC0AF89FF59FED33A01" box="[151,269,1843,1869]" captionStart="FIGURE 61" captionStartId="260.[151,250,1909,1933]" captionTargetBox="[203,1383,191,1871]" captionTargetId="figure-19@260.[192,1396,181,1884]" captionTargetPageId="260" captionText="FIGURE 61. AD: Maladera dayaoshanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); EH: M. shangraoensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); IL: M. hunanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496474" httpUri="https://zenodo.org/record/4496474/files/figure.png" pageId="261" pageNumber="262">Figures 61</figureCitation>
EH, 123
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A64FEC0AF89FF11FEA73AF5" pageId="261" pageNumber="262" type="materials_examined">
<paragraph id="8BAB36F06A64FEC0AF89FF11FEA73AF5" blockId="261.[151,1436,1914,2013]" pageId="261" pageNumber="262">
<emphasis id="B960EAE26A64FEC0AF89FF11FE613AD8" bold="true" box="[151,447,1914,1941]" pageId="261" pageNumber="262">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6A64FEC0AED8FF11FEA73AF5" ID-GBIF-Occurrence="3034556003" collectingDate="2007-04-15" collectingDateMax="2007-04-20" collectingDateMin="2007-04-15" collectionCode="LSSYU" collectorName="Shi Lei" country="China" location="Shangrao" municipality="Mt. Sanqingshan" pageId="261" pageNumber="262" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="holotype">
<typeStatus id="54AF88526A64FEC0AED8FF11FDED3AD9" box="[454,563,1915,1941]" pageId="261" pageNumber="262" type="holotype">Holotype</typeStatus>
:
<specimenCount id="9D12FD796A64FEC0AD23FF11FD8C3AD9" box="[573,594,1915,1941]" pageId="261" pageNumber="262" type="male"></specimenCount>
“[
<collectingCountry id="F30376606A64FEC0AD77FF10FD6D3AD9" box="[617,691,1914,1941]" name="China" pageId="261" pageNumber="262">China</collectingCountry>
]
<collectingMunicipality id="6BCFAC8A6A64FEC0ADDFFF11FC5C3AD9" box="[705,898,1915,1941]" pageId="261" pageNumber="262">Mt. Sanqingshan</collectingMunicipality>
,
<location id="8ECB602B6A64FEC0AC93FF11FC243AD9" LSID="urn:lsid:plazi:treatment:03BD87E66A64FEC3AF89FF64FD103835:8ECB602B6A64FEC0AC93FF11FC243AD9" box="[909,1018,1915,1941]" country="China" municipality="Mt. Sanqingshan" name="Shangrao" pageId="261" pageNumber="262" stateProvince="Jiangxi">Shangrao</location>
,
<collectingRegion id="49D0F8126A64FEC0AB1BFF11FB873AD9" box="[1029,1113,1915,1941]" country="China" name="Jiangxi" pageId="261" pageNumber="262">Jiangxi</collectingRegion>
,
<date id="FFAA10306A64FEC0AB7AFF10FAD93AD9" box="[1124,1287,1914,1941]" pageId="261" pageNumber="262" value="2007-04-15" valueMax="2007-04-20" valueMin="2007-04-15">
<collectingDate id="EFEEE9D86A64FEC0AB7AFF10FAD93AD9" box="[1124,1287,1914,1941]" pageId="261" pageNumber="262" value="2007-04-15" valueMax="2007-04-20" valueMin="2007-04-15">15-20.IV.2007</collectingDate>
</date>
, leg.
<collectorName id="26E153266A64FEC0AA5DFF11FA4A3AD9" box="[1347,1428,1915,1941]" pageId="261" pageNumber="262">Shi Lei</collectorName>
/ LW-825” (
<collectionCode id="ED05AE356A64FEC0AE10FFF5FEB33AF5" box="[270,365,1951,1977]" pageId="261" pageNumber="262">LSSYU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A64FEC3AFD9FFA8FCA83989" lastPageId="262" lastPageNumber="263" pageId="261" pageNumber="262" type="description">
<paragraph id="8BAB36F06A64FEC3AFD9FFA8FEAB3D99" blockId="261.[151,1436,1914,2013]" lastBlockId="262.[151,1437,151,1401]" lastPageId="262" lastPageNumber="263" pageId="261" pageNumber="262">
<emphasis id="B960EAE26A64FEC0AFD9FFA8FE873A90" bold="true" box="[199,345,1986,2012]" pageId="261" pageNumber="262">Description.</emphasis>
Length:
<quantity id="4CEC9B156A64FEC0AEDFFFA9FDF63A91" box="[449,552,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.06" pageId="261" pageNumber="262" unit="mm" value="10.6">10.6 mm</quantity>
, length of elytra:
<quantity id="4CEC9B156A64FEC0ADEAFFA9FC933A91" box="[756,845,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="7.7" pageId="261" pageNumber="262" unit="mm" value="7.7">7.7 mm</quantity>
, width:
<quantity id="4CEC9B156A64FEC0ACB4FFA9FBDB3A91" box="[938,1029,1987,2013]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="261" pageNumber="262" unit="mm" value="6.9">6.9 mm</quantity>
. Body oval, light brown, frons with some greenish tomentum, labroclypeus shiny, remainder of dorsal surface dull, except a few small setae on head and elytra glabrous.
</paragraph>
<paragraph id="8BAB36F06A67FEC3AFD9F8B5FB8D3CB9" blockId="262.[151,1437,151,1401]" pageId="262" pageNumber="263">
Labroclypeus subtrapezoidal, wider than long, widest at base, lateral margins weakly convex and convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a distinct blunt angle, margins strongly reflexed, anteriorly distinctly emarginate medially; surface moderately convex medially, shiny, coarsely and moderately punctate, distance between punctures larger than their diameter, with few long, erect setae anteriorly; frontoclypeal suture feebly impressed and weakly angled medially; smooth area in front of eye approximately twice as wide as long; ocular canthus short and narrow, finely punctate, without terminal seta. Frons dull, with fine, moderately dense punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.62. Antenna of
<typeStatus id="54AF88526A67FEC3AEB9F9B1FDD33CB9" box="[423,525,475,501]" pageId="262" pageNumber="263" type="holotype">holotype</typeStatus>
: missing.
<collectingRegion id="49D0F8126A67FEC3AD63F9B1FD023CB9" box="[637,732,475,501]" country="Myanmar" name="Chin" pageId="262" pageNumber="263">Mentum</collectingRegion>
anteriorly elevated and flattened.
</paragraph>
<paragraph id="8BAB36F06A67FEC3AFD9F995FCFE3FE5" blockId="262.[151,1437,151,1401]" pageId="262" pageNumber="263">Pronotum subtrapezoidal, widest at base, lateral margins subparallel in basal half, in anterior half weakly convey and convergent anteriorly, anterior angles sharp and produced, anterior margin weakly convex medially, anterior marginal line widely interrupted; surface finely and moderately densely punctate, with very minute setae in punctures only; anterior and lateral margins sparsely setose. Scutellum wide, triangular, with fine and sparse punctures each bearing a single minute seta, smooth at middle.</paragraph>
<paragraph id="8BAB36F06A67FEC3AFD9FAD9FB573E75" blockId="262.[151,1437,151,1401]" pageId="262" pageNumber="263">Elytra oblong, widest at posterior quarter (left elytra broken at base), striae weakly impressed, finely and densely punctate, intervals convex, with fine, dense punctures along striae, with a few fine, erect, short setae, punctures with very minute setae only; epipleural margin robust, ending at the strongly rounded external apical angle of elytra, epipleura densely setose, apical border membraneous, with a rim of short microtrichomes.</paragraph>
<paragraph id="8BAB36F06A67FEC3AFD9FB29FC26395D" blockId="262.[151,1437,151,1401]" pageId="262" pageNumber="263">
Ventral surface dull, meso-and metathorax as well as metacoxa with large and moderately dense punctures, sparsely setose, metacoxa glabrous except for a several long setae laterally; each abdominal sternite in addition to generally distributed fine and moderately dense punctures with a distinct transverse row of coarse punctures each bearing a short and robust seta, punctures with microscopic setae; penultimate sternite apically with a shiny, smooth, chitinous border being one third as long as sternite. Mesosternum between mesocoxae as wide as mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.45. Pygidium of
<typeStatus id="54AF88526A67FEC3AC2EFB9DFC4C395D" box="[816,914,1015,1041]" pageId="262" pageNumber="263" type="holotype">holotype</typeStatus>
missing.
</paragraph>
<paragraph id="8BAB36F06A67FEC3AFD9FC71FE1939ED" blockId="262.[151,1437,151,1401]" pageId="262" pageNumber="263">
Legs moderately wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and not serrate, dorsally not serrated, finely setose. Metatibia and subsequent tarsomeres of
<typeStatus id="54AF88526A67FEC3AFE1FCEDFEBF39ED" box="[255,353,1159,1185]" pageId="262" pageNumber="263" type="holotype">holotype</typeStatus>
missing.
</paragraph>
<paragraph id="8BAB36F06A67FEC3AFD9FCC1FCA83989" blockId="262.[151,1437,151,1401]" box="[199,886,1195,1221]" pageId="262" pageNumber="263">
Aedeagus:
<figureCitation id="132F2A756A67FEC3AE5AFCC1FE4D3989" box="[324,403,1195,1221]" captionStart="FIGURE 61" captionStartId="260.[151,250,1909,1933]" captionTargetBox="[203,1383,191,1871]" captionTargetId="figure-19@260.[192,1396,181,1884]" captionTargetPageId="260" captionText="FIGURE 61. AD: Maladera dayaoshanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); EH: M. shangraoensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); IL: M. hunanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496474" httpUri="https://zenodo.org/record/4496474/files/figure.png" pageId="262" pageNumber="263">Fig. 61</figureCitation>
EG. Habitus:
<figureCitation id="132F2A756A67FEC3AD25FCC1FD7C3989" box="[571,674,1195,1221]" captionStart="FIGURE 61" captionStartId="260.[151,250,1909,1933]" captionTargetBox="[203,1383,191,1871]" captionTargetId="figure-19@260.[192,1396,181,1884]" captionTargetPageId="260" captionText="FIGURE 61. AD: Maladera dayaoshanica Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); EH: M. shangraoensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); IL: M. hunanensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496474" httpUri="https://zenodo.org/record/4496474/files/figure.png" pageId="262" pageNumber="263">Fig. 61H</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A67FEC3AFD9FCA4FF28387D" pageId="262" pageNumber="263" type="diagnosis">
<paragraph id="8BAB36F06A67FEC3AFD9FCA4FF28387D" blockId="262.[151,1437,151,1401]" pageId="262" pageNumber="263">
<emphasis id="B960EAE26A67FEC3AFD9FCA4FE9C39A4" bold="true" box="[199,322,1230,1256]" pageId="262" pageNumber="263">Diagnosis.</emphasis>
Having the right paramere straight separates
<taxonomicName id="4C144D736A67FEC3AC52FCA5FAAE39A5" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[844,1392,1231,1257]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="262" pageNumber="263" phylum="Arthropoda" rank="species" species="shangraoensis" status="sp. nov.">
<emphasis id="B960EAE26A67FEC3AC52FCA5FBBB39A5" box="[844,1125,1231,1257]" italics="true" pageId="262" pageNumber="263">Maladera shangraoensis</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A67FEC3AA62FCA5FF723841" pageId="262" pageNumber="263" rank="species">sp. n.</taxonomicNameLabel>
from all other species in
<taxonomicName id="4C144D736A67FEC3AED1FC99FDBE3840" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[463,608,1267,1292]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="262" pageNumber="263" phylum="Arthropoda" rank="species" species="uncipenis">
<emphasis id="B960EAE26A67FEC3AED1FC99FDBE3840" box="[463,608,1267,1292]" italics="true" pageId="262" pageNumber="263">M. uncipenis</emphasis>
</taxonomicName>
group that have the right paramere densely covered with fine short setae apically.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A67FEC3AFD9FD51FA51381A" box="[199,1423,1339,1366]" pageId="262" pageNumber="263" type="etymology">
<paragraph id="8BAB36F06A67FEC3AFD9FD51FA51381A" blockId="262.[151,1437,151,1401]" box="[199,1423,1339,1366]" pageId="262" pageNumber="263">
<emphasis id="B960EAE26A67FEC3AFD9FD51FE903819" bold="true" box="[199,334,1339,1365]" pageId="262" pageNumber="263">Etymology.</emphasis>
The new species is named after its
<typeStatus id="54AF88526A67FEC3ADC5FD51FCD53819" box="[731,779,1339,1365]" pageId="262" pageNumber="263">type</typeStatus>
locality, Shangrao (adjective in the nominative singular).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A67FEC3AFD9FD34FD103835" box="[199,718,1374,1401]" pageId="262" pageNumber="263" type="distribution">
<paragraph id="8BAB36F06A67FEC3AFD9FD34FD103835" blockId="262.[151,1437,151,1401]" box="[199,718,1374,1401]" pageId="262" pageNumber="263">
<emphasis id="B960EAE26A67FEC3AFD9FD34FEBF3834" bold="true" box="[199,353,1374,1400]" pageId="262" pageNumber="263">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756A67FEC3AEC8FD35FDE83835" box="[470,566,1375,1401]" captionStart="FIGURE 123" captionStartId="363.[151,250,1037,1061]" captionTargetBox="[208,1379,188,1006]" captionTargetId="figure-20@363.[202,1385,181,1014]" captionTargetPageId="363" captionText="FIGURE 123. Distribution of the Maladera uncipenis group: Maladera pieli sp. n., M. putaodiensis sp. n., M. qianqingtangensis sp. n., M. serripes (Moser, 1915), M. shangraoensis sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496652" httpUri="https://zenodo.org/record/4496652/files/figure.png" pageId="262" pageNumber="263">Fig. 123</figureCitation>
) and
<tableCitation id="C696034B6A67FEC3AD6AFD35FD163835" box="[628,712,1375,1401]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x records from material examined herein, o records from literature, with no material of this study)." pageId="262" pageNumber="263">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>