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<document id="E8E524981EC8E64C494DCED9851E66F0" ID-CLB-Dataset="8257" ID-DOI="10.11646/zootaxa.4922.1.1" ID-GBIF-Dataset="188411de-569c-4408-a016-1a04b858c142" ID-ISSN="1175-5326" ID-Zenodo-Dep="4496316" ID-ZooBank="D7F9C6A3-9C28-4F4C-8E81-BF24849FDD8C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612331093960" checkinUser="plazi" docAuthor="Fabrizi, Silvia, Liu, Wan-Gang, Bai, Ming, Yang, Xing-Ke &amp; Ahrens, Dirk" docDate="2021" docId="03BD87E66A63FEC6AF89FEACFD103B1D" docLanguage="en" docName="zootaxa.4922.1.1.pdf" docOrigin="Zootaxa 4922 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Maladera excisilabrata Ahrens, Fabrizi &amp; Liu 2021, sp. n." docType="treatment" docVersion="8" lastPageNumber="260" masterDocId="FF84FF9E6B61FFC5AF1EF86AFFDE3D4C" masterDocTitle="A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)" masterLastPageNumber="400" masterPageNumber="1" pageNumber="259" updateTime="1698926631736" updateUser="ExternalLinkService">
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<mods:title id="28F9630984A14C36E23B1D841F92E6F1">A monograph of the genus Maladera Mulsant &amp; Rey, 1871 of China (Coleoptera: Scarabaeidae: Melolonthinae: Sericini)</mods:title>
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<mods:namePart id="4FE27579723C510D298622079445691F">Fabrizi, Silvia</mods:namePart>
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<mods:namePart id="646BB5D33065C45248D4C173CA85CBA8">Liu, Wan-Gang</mods:namePart>
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<mods:namePart id="BBEB50B0EDB35E91D3B368419358150C">Bai, Ming</mods:namePart>
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<mods:namePart id="2E994AF4CB8EE29018FC51DBCF9443AC">Yang, Xing-Ke</mods:namePart>
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<mods:namePart id="A78C2CBD841B053C4D3BF80F62E5398C">Ahrens, Dirk</mods:namePart>
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<mods:date id="34A249C2765AC468278FC972E9CDB95B">2021</mods:date>
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<mods:number id="BCC20E083B322CDBB90655DE23BD8454">4922</mods:number>
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<treatment id="03BD87E66A63FEC6AF89FEACFD103B1D" ID-DOI="http://doi.org/10.5281/zenodo.4534808" ID-GBIF-Taxon="177323772" ID-Zenodo-Dep="4534808" LSID="urn:lsid:plazi:treatment:03BD87E66A63FEC6AF89FEACFD103B1D" httpUri="http://treatment.plazi.org/id/03BD87E66A63FEC6AF89FEACFD103B1D" lastPageId="259" lastPageNumber="260" pageId="258" pageNumber="259">
<subSubSection id="C30E657B6A63FEC7AF89FEACFCC13BAD" box="[151,799,1734,1761]" pageId="258" pageNumber="259" type="nomenclature">
<paragraph id="8BAB36F06A63FEC7AF89FEACFCC13BAD" blockId="258.[151,799,1734,1797]" box="[151,799,1734,1761]" pageId="258" pageNumber="259">
<heading id="D0E3819C6A63FEC7AF89FEACFCC13BAD" bold="true" box="[151,799,1734,1761]" fontSize="11" level="2" pageId="258" pageNumber="259" reason="4">
<emphasis id="B960EAE26A63FEC7AF89FEACFCC13BAD" bold="true" box="[151,799,1734,1761]" pageId="258" pageNumber="259">
<taxonomicName id="4C144D736A63FEC7AF89FEACFD0E3BAC" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[151,720,1734,1761]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="258" pageNumber="259" phylum="Arthropoda" rank="species" species="excisilabrata" status="sp. nov.">
<emphasis id="B960EAE26A63FEC7AF89FEACFE713BAD" bold="true" box="[151,431,1734,1761]" italics="true" pageId="258" pageNumber="259">Maladera excisilabrata</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A63FEC7ADC2FEADFCC13BAD" box="[732,799,1735,1761]" pageId="258" pageNumber="259" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
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<subSubSection id="C30E657B6A63FEC7AF89FE81FEB13A49" box="[151,367,1771,1797]" pageId="258" pageNumber="259" type="description">
<paragraph id="8BAB36F06A63FEC7AF89FE81FEB13A49" blockId="258.[151,799,1734,1797]" box="[151,367,1771,1797]" pageId="258" pageNumber="259">
<figureCitation id="132F2A756A63FEC7AF89FE81FED03A49" box="[151,270,1771,1797]" captionStart="FIGURE 60" captionStartId="256.[151,250,1908,1932]" captionTargetBox="[210,1387,181,1874]" captionTargetId="figure-19@256.[199,1388,181,1884]" captionTargetPageId="256" captionText="FIGURE 60. AD: Maladera tiammushanicaAhrens, Fabrizi &amp; Liu, sp. n. (holotype); EH: M. putaodiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); IL: M. excisilabrata Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496472" httpUri="https://zenodo.org/record/4496472/files/figure.png" pageId="258" pageNumber="259">Figures 60</figureCitation>
IL, 121
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A63FEC7AF89FF59FAAE3A01" box="[151,1392,1842,1869]" pageId="258" pageNumber="259" type="materials_examined">
<paragraph id="8BAB36F06A63FEC7AF89FF59FAAE3A01" blockId="258.[151,1436,1842,2013]" box="[151,1392,1842,1869]" pageId="258" pageNumber="259">
<emphasis id="B960EAE26A63FEC7AF89FF59FE613A00" bold="true" box="[151,447,1842,1869]" pageId="258" pageNumber="259">Type material examined.</emphasis>
<materialsCitation id="3B7C3CAD6A63FEC7AED8FF59FAAE3A01" ID-GBIF-Occurrence="3034556009" box="[454,1392,1842,1869]" collectingDate="1977-05-10" collectionCode="IZAS" collectorName="Zhang Youwei" country="China" location="Mt. Lushan" pageId="258" pageNumber="259" specimenCount="1" specimenCount-male="1" stateProvince="Jiangxi" typeStatus="holotype">
<typeStatus id="54AF88526A63FEC7AED8FF59FDED3A01" box="[454,563,1843,1869]" pageId="258" pageNumber="259" type="holotype">Holotype</typeStatus>
:
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“[
<collectingCountry id="F30376606A63FEC7AD74FF58FD6A3A01" box="[618,692,1842,1869]" name="China" pageId="258" pageNumber="259">China</collectingCountry>
]
<location id="8ECB602B6A63FEC7ADDCFF59FC983A01" LSID="urn:lsid:plazi:treatment:03BD87E66A63FEC6AF89FEACFD103B1D:8ECB602B6A63FEC7ADDCFF59FC983A01" box="[706,838,1843,1869]" country="China" name="Mt. Lushan" pageId="258" pageNumber="259" stateProvince="Jiangxi">Mt. Lushan</location>
,
<collectingRegion id="49D0F8126A63FEC7AC4FFF59FC7B3A01" box="[849,933,1843,1869]" country="China" name="Jiangxi" pageId="258" pageNumber="259">Jiangxi</collectingRegion>
,
<date id="FFAA10306A63FEC7ACAEFF59FBFB3A01" box="[944,1061,1843,1869]" pageId="258" pageNumber="259" value="1977-05-10">
<collectingDate id="EFEEE9D86A63FEC7ACAEFF59FBFB3A01" box="[944,1061,1843,1869]" pageId="258" pageNumber="259" value="1977-05-10">10.V.1977</collectingDate>
</date>
, leg.
<collectorName id="26E153266A63FEC7AB7FFF59FAD93A01" box="[1121,1287,1843,1869]" pageId="258" pageNumber="259">Zhang Youwei</collectorName>
” (
<collectionCode id="ED05AE356A63FEC7AA3EFF59FABA3A01" box="[1312,1380,1843,1869]" country="0" httpUri="http://grbio.org/cool/043p-ts4f" name="Institut Zoologii Akademii Nauk Ukraini - Institute of Zoology of the Academy of Sciences of Ukraine" pageId="258" pageNumber="259">IZAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A63FEC6AFD9FF3CFCB53819" lastPageId="259" lastPageNumber="260" pageId="258" pageNumber="259" type="description">
<paragraph id="8BAB36F06A63FEC7AFD9FF3CFB963AD9" blockId="258.[151,1436,1842,2013]" pageId="258" pageNumber="259">
<emphasis id="B960EAE26A63FEC7AFD9FF3CFE873A3C" bold="true" box="[199,345,1878,1904]" pageId="258" pageNumber="259">Description.</emphasis>
Length: 9.0 mm, length of elytra:
<quantity id="4CEC9B156A63FEC7ADF5FF3DFC9B3A3D" box="[747,837,1879,1905]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="258" pageNumber="259" unit="mm" value="6.6">6.6 mm</quantity>
, width:
<quantity id="4CEC9B156A63FEC7ACBDFF3CFC213A3D" box="[931,1023,1878,1905]" metricMagnitude="-3" metricUnit="m" metricValue="5.4" pageId="258" pageNumber="259" unit="mm" value="5.4">5.4 mm</quantity>
. Body oval, reddish brown, antenna brown, head and pronotum weakly shiny, remainder of dorsal surface dull, glabrous.
</paragraph>
<paragraph id="8BAB36F06A63FEC6AFD9FFF5FD373CC5" blockId="258.[151,1436,1842,2013]" lastBlockId="259.[151,1437,151,1617]" lastPageId="259" lastPageNumber="260" pageId="258" pageNumber="259">
Labroclypeus subtrapezoidal, little wider than long, widest at base, lateral margins weakly convex and strongly convergent to moderately rounded anterior angles, lateral margin and ocular canthus producing a blunt angle, margins moderately reflexed, anterior margin weakly emarginate medially; surface transversely convex medially, shiny, coarsely and moderately densely punctate, distance between punctures equal their diameter, with few long, erect setae anteriorly; frontoclypeal suture weakly impressed and angled medially; smooth area in front of eye approximately three times as wide as long; ocular canthus short and wide, finely punctate, without terminal seta. Frons with fine, dense punctures, glabrous except for a few setae beside eyes. Eyes small, ratio of diameter/interocular width: 0.53. Antenna with 10 antennomeres; club with three antennomeres, slightly longer than remaining antennomeres combined.
<collectingRegion id="49D0F8126A62FEC6AE0AF905FEAD3CC5" box="[276,371,367,393]" country="Myanmar" name="Chin" pageId="259" pageNumber="260">Mentum</collectingRegion>
anteriorly elevated and flattened.
</paragraph>
<paragraph id="8BAB36F06A62FEC6AFD9F9F9FC383F55" blockId="259.[151,1437,151,1617]" pageId="259" pageNumber="260">Pronotum subtrapezoidal, widest at base, lateral margins weakly convergent anteriorly, anterior angles moderately produced but sharp, anterior margin weakly convex, marginal line widely interrupted medially; surface finely and densely punctate, glabrous; anterior and lateral margins sparsely setose; hypomeron carinate. Scutellum wide, triangular, with fine and dense punctures, each bearing a single minute seta.</paragraph>
<paragraph id="8BAB36F06A62FEC6AFD9FA49FAA63FE5" blockId="259.[151,1437,151,1617]" pageId="259" pageNumber="260">Elytra oblong, widest at middle, striae weakly impressed, finely and densely punctate, intervals weakly elevated, with fine, moderately dense punctures concentrated along striae, odd intervals with a few fine, erect, short setae, otherwise punctures with a very minute seta; epipleural margin robust, ending at strongly rounded external apical angle of elytra, epipleura sparsely setose, apical border membraneous, with a rim of very short microtrichomes.</paragraph>
<paragraph id="8BAB36F06A62FEC6AFD9FAD9FDAA3ECD" blockId="259.[151,1437,151,1617]" pageId="259" pageNumber="260">
Ventral surface dull, metasternum and metacoxa with large and dense punctures, sparsely setose; metacoxa glabrous except for a several long setae laterally. Abdominal sternites, in addition to generally distributed fine and moderately dense punctures, each with a distinct transverse row of coarse punctures each bearing a short and robust seta, punctures with microscopic setae, penultimate sternite apically with a shiny smooth short chitinous border. Mesosternum between mesocoxae narrower than mesofemur. Ratio of length of metepisternum/metacoxa: 1/1.43. Pygidium of
<typeStatus id="54AF88526A62FEC6AE37FB0DFE553ECD" box="[297,395,871,897]" pageId="259" pageNumber="260" type="holotype">holotype</typeStatus>
lost after dissection.
</paragraph>
<paragraph id="8BAB36F06A62FEC6AFD9FBE1FE4D387D" blockId="259.[151,1437,151,1617]" pageId="259" pageNumber="260">Legs moderately long and wide; femora with two longitudinal rows of setae, finely and moderately densely punctate; metafemur dull, anterior margin acute, lacking an adjacent serrated line, posterior ventral margin medially straight, weakly widened in apical half and entirely strongly serrate, dorsally also serrate, finely setose. Metatibia moderately wide and long, widest at apex, ratio width/length: 1/3.3, dorsally sharply carinate, with two groups of spines, basal one at middle, apical one at four fifths of metatibial length, in basal half with a blunt subdorsal margin, beside it with a few punctures, and short spines; lateral face longitudinally convex, with moderately dense and fine punctures, punctures with minute setae; ventral margin serrate, with two widely distant spines; medial face finely sparsely punctate, apex interiorly near tarsal articulation shallowly concave. Tarsomeres impunctate dorsally, ventrally with sparse, short setae; metatarsomeres ventrally glabrous, with a strongly serrated ridge, and a subventral longitudinal carina beside it; first metatarsomere shorter than following two tarsomeres combined and slightly longer than dorsal tibial spur. Protibia short, bidentate. All claws symmetrical, feebly curved and long, with normally developed basal tooth.</paragraph>
<paragraph id="8BAB36F06A62FEC6AFD9FD51FCB53819" blockId="259.[151,1437,151,1617]" box="[199,875,1339,1365]" pageId="259" pageNumber="260">
Aedeagus:
<figureCitation id="132F2A756A62FEC6AE5AFD51FE4B3819" box="[324,405,1339,1365]" captionStart="FIGURE 60" captionStartId="256.[151,250,1908,1932]" captionTargetBox="[210,1387,181,1874]" captionTargetId="figure-19@256.[199,1388,181,1884]" captionTargetPageId="256" captionText="FIGURE 60. AD: Maladera tiammushanicaAhrens, Fabrizi &amp; Liu, sp. n. (holotype); EH: M. putaodiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); IL: M. excisilabrata Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496472" httpUri="https://zenodo.org/record/4496472/files/figure.png" pageId="259" pageNumber="260">Fig. 60</figureCitation>
IK. Habitus:
<figureCitation id="132F2A756A62FEC6AD2DFD51FD493819" box="[563,663,1339,1365]" captionStart="FIGURE 60" captionStartId="256.[151,250,1908,1932]" captionTargetBox="[210,1387,181,1874]" captionTargetId="figure-19@256.[199,1388,181,1884]" captionTargetPageId="256" captionText="FIGURE 60. AD: Maladera tiammushanicaAhrens, Fabrizi &amp; Liu, sp. n. (holotype); EH: M. putaodiensis Ahrens, Fabrizi &amp; Liu, sp. n. (holotype); IL: M. excisilabrata Ahrens, Fabrizi &amp; Liu, sp. n. (holotype). A, E, I: aedeagus, left side lateral view; C, G, K: aedeagus, right side lateral view; B, F, J: parameres, dorsal view; D, H, L: habitus. Scale: 0.5 mm. Habitus not to scale." figureDoi="http://doi.org/10.5281/zenodo.4496472" httpUri="https://zenodo.org/record/4496472/files/figure.png" pageId="259" pageNumber="260">
Fig.
<specimenCount id="9D12FD796A62FEC6AD79FD51FD493819" box="[615,663,1339,1365]" pageId="259" pageNumber="260" type="generic">60L</specimenCount>
</figureCitation>
. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A62FEC6AFD9FD34FC3F388D" pageId="259" pageNumber="260" type="diagnosis">
<paragraph id="8BAB36F06A62FEC6AFD9FD34FC3F388D" blockId="259.[151,1437,151,1617]" pageId="259" pageNumber="260">
<emphasis id="B960EAE26A62FEC6AFD9FD34FE9C3834" bold="true" box="[199,322,1374,1400]" pageId="259" pageNumber="260">Diagnosis.</emphasis>
<taxonomicName id="4C144D736A62FEC6AE54FD35FC803835" authority="Ahrens, Fabrizi &amp; Liu" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[330,862,1375,1401]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="species" species="excisilabrata" status="sp. nov.">
<emphasis id="B960EAE26A62FEC6AE54FD35FD8A3835" box="[330,596,1375,1401]" italics="true" pageId="259" pageNumber="260">Maladera excisilabrata</emphasis>
Ahrens, Fabrizi &amp; Liu
</taxonomicName>
,
<taxonomicNameLabel id="A25357996A62FEC6AC75FD35FC773835" box="[875,937,1375,1401]" pageId="259" pageNumber="260" rank="species">sp. n.</taxonomicNameLabel>
differs from all other Chinese
<taxonomicName id="4C144D736A62FEC6AA11FD35FAA33835" authorityName="Mulsant &amp; Rey" authorityYear="1871" box="[1295,1405,1375,1401]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="genus">
<emphasis id="B960EAE26A62FEC6AA11FD35FAA33835" box="[1295,1405,1375,1401]" italics="true" pageId="259" pageNumber="260">Maladera</emphasis>
</taxonomicName>
in having narrow lateral apophyses on the apex of the phallobase as well as having short, ventrally curved parameres. Additionally, it differs from
<taxonomicName id="4C144D736A62FEC6AECAFDCDFDBB388C" authorityName="Ahrens, Fabrizi &amp; Liu" authorityYear="2021" box="[468,613,1447,1472]" class="Insecta" family="Melolonthidae" genus="Maladera" kingdom="Animalia" order="Coleoptera" pageId="259" pageNumber="260" phylum="Arthropoda" rank="species" species="uncipenis">
<emphasis id="B960EAE26A62FEC6AECAFDCDFDBB388C" box="[468,613,1447,1472]" italics="true" pageId="259" pageNumber="260">M. uncipenis</emphasis>
</taxonomicName>
in having impunctate tarsomeres.
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A62FEC6AFD9FDA1FF063B61" pageId="259" pageNumber="260" type="etymology">
<paragraph id="8BAB36F06A62FEC6AFD9FDA1FF063B61" blockId="259.[151,1437,151,1617]" pageId="259" pageNumber="260">
<emphasis id="B960EAE26A62FEC6AFD9FDA1FE9038A9" bold="true" box="[199,334,1483,1509]" pageId="259" pageNumber="260">Etymology.</emphasis>
The name of the new species is derived from the combined Latin adjectives excisus- (emarginate) and labratus (with labrum), with reference to the deeply excavated labrum (adjective in nominative singular case).
</paragraph>
</subSubSection>
<subSubSection id="C30E657B6A62FEC6AFD9FE5CFD103B1D" box="[199,718,1590,1617]" pageId="259" pageNumber="260" type="distribution">
<paragraph id="8BAB36F06A62FEC6AFD9FE5CFD103B1D" blockId="259.[151,1437,151,1617]" box="[199,718,1590,1617]" pageId="259" pageNumber="260">
<emphasis id="B960EAE26A62FEC6AFD9FE5CFEBF3B1C" bold="true" box="[199,353,1590,1616]" pageId="259" pageNumber="260">Distribution.</emphasis>
See map (
<figureCitation id="132F2A756A62FEC6AEC8FE5DFDE83B1D" box="[470,566,1591,1617]" captionStart="FIGURE 121" captionStartId="362.[151,250,1008,1032]" captionTargetBox="[228,1362,187,978]" captionTargetId="figure-19@362.[220,1368,181,984]" captionTargetPageId="362" captionText="FIGURE 121. Distribution of the Maladera uncipenis group: Maladera baii sp. n., M. bawanglingana sp. n., M. beidouensis sp. n., M. chinensis (Moser, 1915), M. dadongshanica sp. n., M. dayaoshanica sp. n., M. emeifengensis sp. n., M. excisilabrata sp. n.." figureDoi="http://doi.org/10.5281/zenodo.4496646" httpUri="https://zenodo.org/record/4496646/files/figure.png" pageId="259" pageNumber="260">Fig. 121</figureCitation>
) and
<tableCitation id="C696034B6A62FEC6AD6AFE5DFD163B1D" box="[628,712,1591,1617]" captionStart="TABLE 1" captionStartId="368.[151,239,544,570]" captionText="TABLE 1. Overview on the distribuition of the Chinese mainland species of Maladera Mulsant &amp; Rey, 1871 (x records from material examined herein, o records from literature, with no material of this study)." pageId="259" pageNumber="260">Table 1</tableCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>