520 lines
51 KiB
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520 lines
51 KiB
XML
<document ID-DOI="10.11646/zootaxa.5277.1.4" ID-GBIF-Dataset="ad5ba4d1-c2ec-42e3-a76d-a5c9b395a6da" ID-ISSN="1175-5326" ID-Zenodo-Dep="7892052" ID-ZooBank="272D07DF-8471-4ADE-9689-DB8F588ACF35" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1682930765663" checkinUser="plazi" docAuthor="Short, Andrew E. Z., Torres-Gavosto, Juan Martin & Hettinger, Ty" docDate="2023" docId="03BD87B6FFB4F438CFDB7787FA7DFA2C" docLanguage="en" docName="zootaxa.5277.1.4.pdf" docOrigin="Zootaxa 5277 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5277.1.4" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oocyclus" docType="key" docVersion="2" lastPageNumber="93" masterDocId="FF84FFCEFFB5F43ACF4C7204FFE1FFDE" masterDocTitle="A review of the Oocyclus Sharp of Ecuador with description of 12 new species (Coleoptera: Hydrophilidae: Laccobiini)" masterLastPageNumber="112" masterPageNumber="91" pageNumber="92" updateTime="1683122599468" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:titleInfo>
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<mods:title>A review of the Oocyclus Sharp of Ecuador with description of 12 new species (Coleoptera: Hydrophilidae: Laccobiini)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Short, Andrew E. Z.</mods:namePart>
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<mods:affiliation>Division of Entomology, Biodiversity Institute & Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A. & aezshort @ gmail. com; https: // orcid. org / 0000 - 0002 - 7467 - 7116</mods:affiliation>
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<mods:nameIdentifier type="email">aezshort@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Torres-Gavosto, Juan Martin</mods:namePart>
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<mods:nameIdentifier type="ORCID">0009-0007-9274-2923</mods:nameIdentifier>
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<mods:affiliation>Division of Entomology, Biodiversity Institute & Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A. & https: // orcid. org / 0009 - 0007 - 9274 - 2923</mods:affiliation>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Hettinger, Ty</mods:namePart>
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<mods:affiliation>Division of Entomology, Biodiversity Institute & Department of Ecology and Evolutionary Biology, University of Kansas, 1501 Crestline Drive, Suite 140, Lawrence, KS 66045, U. S. A.</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Zootaxa</mods:title>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-05-01</mods:number>
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<mods:detail type="volume">
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<mods:number>5277</mods:number>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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<mods:start>91</mods:start>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5277.1.4</mods:url>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5277.1.4</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7892056" ID-Zenodo-Dep="7892056" LSID="urn:lsid:plazi:treatment:03BD87B6FFB4F438CFDB7787FA7DFA2C" httpUri="http://treatment.plazi.org/id/03BD87B6FFB4F438CFDB7787FA7DFA2C" lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
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<subSubSection box="[151,842,1411,1438]" pageId="1" pageNumber="92" type="nomenclature">
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<paragraph blockId="1.[151,842,1411,1438]" box="[151,842,1411,1438]" pageId="1" pageNumber="92">
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<heading bold="true" box="[151,842,1411,1438]" fontSize="11" level="1" pageId="1" pageNumber="92" reason="1">
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<emphasis bold="true" box="[151,842,1411,1438]" pageId="1" pageNumber="92">
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Key to the species of
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<taxonomicName ID-CoL="8VZW4" authority="Sharp, 1882" box="[413,523,1411,1438]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
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<emphasis bold="true" box="[413,512,1411,1438]" italics="true" pageId="1" pageNumber="92">Oocyclus</emphasis>
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</taxonomicName>
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from the northern Andes
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92" type="key">
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<key pageId="1" pageNumber="92">
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<keyStep pageId="1" pageNumber="92">
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1480,1502]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1480,1502]" pageId="1" pageNumber="92">
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1 Elytra with distinct pale spot in posterior third (e.g.
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<figureCitation box="[724,801,1480,1502]" captionStart="FIGURE 4" captionStartId="13.[151,250,1523,1547]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-20@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) lepidus sp. n. (B) morgani/trujillo group" figureDoi="http://doi.org/10.5281/zenodo.7892068" httpUri="https://zenodo.org/record/7892068/files/figure.png" pageId="1" pageNumber="92">Fig. 4A</figureCitation>
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); size smaller (not more than 4.0 mm)..................... 2
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</keyLead>
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</paragraph>
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1509,1531]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1509,1531]" pageId="1" pageNumber="92">- Elytra without distinct pale spot in posterior third; size variable................................................. 4</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="1" pageNumber="92">
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1538,1561]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1538,1561]" pageId="1" pageNumber="92">
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2(1) Elytra without rows of setiferous punctures; posterolateral corners of pronotum broadly rounded.............
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<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1310,1436,1538,1560]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="sapphirus">
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<emphasis bold="true" box="[1310,1436,1538,1560]" italics="true" pageId="1" pageNumber="92">O. sapphirus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="1.[151,1437,1480,2030]" pageId="1" pageNumber="92">
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<keyLead pageId="1" pageNumber="92">
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<emphasis bold="true" box="[151,159,1568,1589]" italics="true" pageId="1" pageNumber="92">-</emphasis>
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Elytra with distinct rows of setiferous punctures (e.g.
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<figureCitation box="[750,828,1568,1590]" captionStart="FIGURE 4" captionStartId="13.[151,250,1523,1547]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-20@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) lepidus sp. n. (B) morgani/trujillo group" figureDoi="http://doi.org/10.5281/zenodo.7892068" httpUri="https://zenodo.org/record/7892068/files/figure.png" pageId="1" pageNumber="92">Fig. 4A</figureCitation>
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); posterolateral corners of pronotum usually appearing weakly spinose............................................................................................. 3
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="1" pageNumber="92">
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<paragraph blockId="1.[151,1437,1480,2030]" pageId="1" pageNumber="92">
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<keyLead pageId="1" pageNumber="92">
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3(2’) Elytra with first line of erect setae distinct and continuous (
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<figureCitation box="[802,880,1627,1649]" captionStart="FIGURE 4" captionStartId="13.[151,250,1523,1547]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-20@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) lepidus sp. n. (B) morgani/trujillo group" figureDoi="http://doi.org/10.5281/zenodo.7892068" httpUri="https://zenodo.org/record/7892068/files/figure.png" pageId="1" pageNumber="92">Fig. 4A</figureCitation>
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); aedeagus with parameres tapering to a nearly flat, blunt tip at apex (
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<figureCitation box="[339,411,1656,1678]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="1" pageNumber="92">Fig. 6F</figureCitation>
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).......................................................................
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<emphasis bold="true" box="[1276,1436,1656,1678]" pageId="1" pageNumber="92">
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||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1276,1377,1656,1678]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="lepidus" status="sp. n.">
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<emphasis bold="true" box="[1276,1377,1656,1678]" italics="true" pageId="1" pageNumber="92">O. lepidus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1382,1436,1656,1677]" pageId="1" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</keyLead>
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</paragraph>
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||
<paragraph blockId="1.[151,1437,1480,2030]" pageId="1" pageNumber="92">
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||
<keyLead pageId="1" pageNumber="92">
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||
- Elytra with first line of erect setae discontinuous, with distinct gaps in the line; aedeagus with parameres broadly rounded and parallel sided at apex, not tapering (
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||
<figureCitation box="[570,644,1715,1737]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="1" pageNumber="92">Fig. 6G</figureCitation>
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)....................................................
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<emphasis bold="true" box="[1288,1436,1714,1736]" pageId="1" pageNumber="92">
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<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1288,1377,1714,1736]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="bellus" status="sp. n.">
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<emphasis bold="true" box="[1288,1377,1714,1736]" italics="true" pageId="1" pageNumber="92">O. bellus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1382,1436,1715,1736]" pageId="1" pageNumber="92" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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||
</keyLead>
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||
</paragraph>
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||
</keyStep>
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||
<keyStep pageId="1" pageNumber="92">
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||
<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1744,1766]" pageId="1" pageNumber="92">
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||
<keyLead box="[151,1436,1744,1766]" pageId="1" pageNumber="92">
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||
4(1’) Posterolateral corners of pronotum broadly rounded (e.g.
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<figureCitation box="[778,890,1744,1766]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1994]" captionTargetBox="[180,1407,181,1952]" captionTargetId="figure-20@3.[180,1407,181,1952]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ancho sp. n. (B) zamora sp. n. (C) yantzaza sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892054" httpUri="https://zenodo.org/record/7892054/files/figure.png" pageId="1" pageNumber="92">Figs. 1A–C</figureCitation>
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).......................................... 19
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</keyLead>
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</paragraph>
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1773,1795]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1773,1795]" pageId="1" pageNumber="92">
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- Posterolateral corners of pronotum angulate or spinose (e.g.
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<figureCitation box="[806,918,1773,1795]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="1" pageNumber="92">Figs. 2A–B</figureCitation>
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,
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<figureCitation box="[928,987,1774,1795]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="1" pageNumber="92">3A–B</figureCitation>
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)................................... 5
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="1" pageNumber="92">
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1803,1825]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1803,1825]" pageId="1" pageNumber="92">
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5(4’) Posterolateral corners of pronotum angulate but not prolonged into an acute spine (e.g.
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<figureCitation box="[1064,1138,1803,1825]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="1" pageNumber="92">Fig. 3A</figureCitation>
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)....................... 6
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</keyLead>
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</paragraph>
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1832,1854]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1832,1854]" pageId="1" pageNumber="92">
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- Posterolateral corners of pronotum spinose (
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<figureCitation box="[651,735,1832,1854]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="1" pageNumber="92">Figs. 3B</figureCitation>
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,
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<figureCitation box="[745,803,1832,1853]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="1" pageNumber="92">2A–B</figureCitation>
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||
).................................................. 10
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="1" pageNumber="92">
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1861,1883]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1861,1883]" pageId="1" pageNumber="92">6(5) Abdominal ventrites uniformly pale yellow................................................................. 7</keyLead>
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</paragraph>
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1891,1913]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1891,1913]" pageId="1" pageNumber="92">- Abdominal ventrites darkly colored, at least in part........................................................... 8</keyLead>
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</paragraph>
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</keyStep>
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<keyStep pageId="1" pageNumber="92">
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1920,1942]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1920,1942]" pageId="1" pageNumber="92">
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7(6) First row of elytra punctures formed by an irregular line of very coarse punctures, not bearing erect setae........
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<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1342,1436,1920,1942]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="galbus">
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<emphasis bold="true" box="[1342,1436,1920,1942]" italics="true" pageId="1" pageNumber="92">O. galbus</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1949,1971]" pageId="1" pageNumber="92">
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<keyLead box="[151,1436,1949,1971]" pageId="1" pageNumber="92">
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- First row of elytra punctures forming a dense, continuous line bearing erect setae...........................
|
||
<taxonomicName authorityName="Garcia-Hernandez" authorityYear="2009" box="[1347,1436,1949,1971]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="luteus">
|
||
<emphasis bold="true" box="[1347,1436,1949,1971]" italics="true" pageId="1" pageNumber="92">O. luteus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
|
||
<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,1979,2001]" pageId="1" pageNumber="92">
|
||
<keyLead box="[151,1436,1979,2001]" pageId="1" pageNumber="92">8(6’) First row of elytra punctures with row of erect setae along at least half of its length................................. 9</keyLead>
|
||
</paragraph>
|
||
<keyLead lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
|
||
<paragraph blockId="1.[151,1437,1480,2030]" box="[151,1436,2008,2030]" pageId="1" pageNumber="92">
|
||
- First row of elytra punctures not bearing long setae along its length.......................................
|
||
<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1342,1436,2008,2029]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="pittieri">
|
||
<emphasis bold="true" box="[1342,1436,2008,2029]" italics="true" pageId="1" pageNumber="92">O. pittieri</emphasis>
|
||
</taxonomicName>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,151,173]" pageId="2" pageNumber="93">
|
||
9(8) First row of elytra punctures bearing dense and continuous row of erect setae.........................
|
||
<emphasis bold="true" box="[1275,1436,151,173]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1275,1376,151,173]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="pakcha" status="sp. n.">
|
||
<emphasis bold="true" box="[1275,1376,151,173]" italics="true" pageId="2" pageNumber="93">O. pakcha</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,152,173]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</keyLead>
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,180,203]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,180,203]" pageId="2" pageNumber="93">
|
||
- First row of elytra punctures bearing an irregular, broken row of erect setae (
|
||
<figureCitation box="[961,1036,181,203]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="2" pageNumber="93">Fig. 3A</figureCitation>
|
||
).................
|
||
<emphasis bold="true" box="[1266,1436,180,202]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1266,1376,180,202]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="radiatus" status="sp. n.">
|
||
<emphasis bold="true" box="[1266,1376,180,202]" italics="true" pageId="2" pageNumber="93">O. radiatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,181,202]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,210,232]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,210,232]" pageId="2" pageNumber="93">
|
||
10(4’) Abdominal ventrites uniformly pale (
|
||
<figureCitation box="[584,659,210,232]" captionStart="FIGURE 3" captionStartId="11.[151,250,1726,1750]" captionTargetBox="[151,1436,453,1709]" captionTargetId="figure-137@11.[151,1436,453,1709]" captionTargetPageId="11" captionText="FIGURE 3. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) radiatus sp. n. (B) ustulatus sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892066" httpUri="https://zenodo.org/record/7892066/files/figure.png" pageId="2" pageNumber="93">Fig. 3B</figureCitation>
|
||
); size very large (>
|
||
<quantity box="[843,919,210,232]" metricMagnitude="-3" metricUnit="m" metricValue="5.2" pageId="2" pageNumber="93" unit="mm" value="5.2">5.2 mm</quantity>
|
||
)...........................
|
||
<emphasis bold="true" box="[1256,1436,210,232]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1256,1376,210,232]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="ustulatus" status="sp. n.">
|
||
<emphasis bold="true" box="[1256,1376,210,232]" italics="true" pageId="2" pageNumber="93">O. ustulatus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,210,231]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- Abdominal ventrites dark brown, at most with paler or orange spots laterally (e.g.
|
||
<figureCitation box="[1015,1130,239,261]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="2" pageNumber="93">Figs. 2A–B</figureCitation>
|
||
); size variable (
|
||
<quantity box="[1280,1398,239,261]" metricMagnitude="-3" metricUnit="m" metricValue="4.35" metricValueMax="5.5" metricValueMin="3.2" pageId="2" pageNumber="93" unit="mm" value="4.35" valueMax="5.5" valueMin="3.2">3.2–5.5 mm</quantity>
|
||
).. .................................................................................................. 11
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
11(10’) First row of systematic punctures bearing a more or less continuous like of erect setae (the line may have small breaks but these are short); size large (>
|
||
<quantity box="[447,523,328,349]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="2" pageNumber="93" unit="mm" value="4.2">4.2 mm</quantity>
|
||
)......................................................................... 16
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- First row of systematic punctures at most bearing a few small clumps or erect setae along its length or occasionally without any distinct line or clups of erect setae; size usually smaller, rarely greater than 4.0 mm (except
|
||
<taxonomicName authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1105,1206,386,408]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="zamora">
|
||
<emphasis box="[1105,1206,386,408]" italics="true" pageId="2" pageNumber="93">O. zamora</emphasis>
|
||
</taxonomicName>
|
||
:
|
||
<quantity box="[1219,1336,386,407]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="2" pageNumber="93" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
|
||
)..... 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,415,437]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,415,437]" pageId="2" pageNumber="93">
|
||
12(11’) Elytra without clumps of erect setae.................................................................
|
||
<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1366,1436,415,436]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="pico">
|
||
<emphasis bold="true" box="[1366,1436,415,436]" italics="true" pageId="2" pageNumber="93">O. pico</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,445,467]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,445,467]" pageId="2" pageNumber="93">
|
||
- Elytra bearing clumps of erect setae, at least mesally near the elytra suture (e.g.
|
||
<figureCitation box="[993,1105,445,467]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="2" pageNumber="93">Figs. 2A–B</figureCitation>
|
||
,
|
||
<figureCitation box="[1116,1145,445,466]" captionStart="FIGURE 4" captionStartId="13.[151,250,1523,1547]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-20@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) lepidus sp. n. (B) morgani/trujillo group" figureDoi="http://doi.org/10.5281/zenodo.7892068" httpUri="https://zenodo.org/record/7892068/files/figure.png" pageId="2" pageNumber="93">4B</figureCitation>
|
||
)..................... 13
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
13(12’) Aedeaegus with parameres strongly narrowing in apical half (
|
||
<figureCitation box="[827,942,474,496]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="2" pageNumber="93">Figs. 6D–E</figureCitation>
|
||
); rows of elytra systematic punctures distinct from ground punctation; elytra with blue maculae (
|
||
<figureCitation box="[664,776,503,525]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="2" pageNumber="93">Figs. 2A–B</figureCitation>
|
||
)................................................... 14
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- Aedeaegus with parameres not narrowing in apical half, sometimes even appearing to swell towards apex (
|
||
<figureCitation box="[1262,1372,533,555]" captionStart="FIGURE 6" captionStartId="17.[151,250,1605,1629]" captionTargetBox="[157,1427,187,1586]" captionTargetId="figure-20@17.[151,1437,181,1587]" captionTargetPageId="17" captionText="FIGURE 6. Aedeagi of Oocyclus species in Ecuador with angulate or spinose posterolateral corners of the pronotum. (A) ustulatus sp. n. (B) radiatus sp. n. (C) pakcha sp. n. (D) sumak sp. n. (E) ankas sp. n. (F) lepidus sp. n. (G) bellus sp. n. (H) zamora sp. n. (I) specimen of the morgani/trujillo species group (J) specimen of the morgani/trujillo species group." figureDoi="http://doi.org/10.5281/zenodo.7892076" httpUri="https://zenodo.org/record/7892076/files/figure.png" pageId="2" pageNumber="93">Figs. 6H–J</figureCitation>
|
||
); rows of systematic punctures not distinct; elytra without blue maculae............................................... 15
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
14(13) First row of elytra systematic punctures represented by a short linear tuft of erect setae anteriorly, then the remainder of the row bearing sparse usually decumbant setae (
|
||
<figureCitation box="[618,693,621,643]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="2" pageNumber="93">Fig. 2B</figureCitation>
|
||
); anterolateral pale spot larger........................
|
||
<emphasis bold="true" box="[1282,1436,620,642]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1282,1376,620,642]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="sumak" status="sp. n.">
|
||
<emphasis bold="true" box="[1282,1376,620,642]" italics="true" pageId="2" pageNumber="93">O. sumak</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,621,642]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- First row of elytra systematic punctures represented by a line of tufts and erect setae along its entire length (
|
||
<figureCitation box="[1225,1301,650,672]" captionStart="FIGURE 2" captionStartId="5.[151,250,1603,1627]" captionTargetBox="[151,1436,181,1585]" captionTargetId="figure-20@5.[151,1436,181,1585]" captionTargetPageId="5" captionText="FIGURE 2. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ankas sp. n. (B) sumak sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892062" httpUri="https://zenodo.org/record/7892062/files/figure.png" pageId="2" pageNumber="93">Fig. 2A</figureCitation>
|
||
); anterolateral pale spot smaller..........................................................................
|
||
<emphasis bold="true" box="[1288,1436,679,701]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1288,1376,679,701]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="ankas" status="sp. n.">
|
||
<emphasis bold="true" box="[1288,1376,679,701]" italics="true" pageId="2" pageNumber="93">O. ankas</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,680,701]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
15(13’) Elytral ground punctation bearing short scattered setae; size larger (
|
||
<quantity box="[874,994,709,730]" metricMagnitude="-3" metricUnit="m" metricValue="4.25" metricValueMax="4.4" metricValueMin="4.1" pageId="2" pageNumber="93" unit="mm" value="4.25" valueMax="4.4" valueMin="4.1">4.1–4.4 mm</quantity>
|
||
), ground punctation coarser (
|
||
<figureCitation box="[1274,1347,709,731]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1994]" captionTargetBox="[180,1407,181,1952]" captionTargetId="figure-20@3.[180,1407,181,1952]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ancho sp. n. (B) zamora sp. n. (C) yantzaza sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892054" httpUri="https://zenodo.org/record/7892054/files/figure.png" pageId="2" pageNumber="93">Fig. 1B</figureCitation>
|
||
)............................................................................................
|
||
<emphasis bold="true" box="[1274,1436,738,759]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1274,1376,738,759]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="zamora" status="sp. n.">
|
||
<emphasis bold="true" box="[1274,1376,738,759]" italics="true" pageId="2" pageNumber="93">O. zamora</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,738,759]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- Elytral ground punctation not bearing setae; size smaller (
|
||
<quantity box="[781,901,768,789]" metricMagnitude="-3" metricUnit="m" metricValue="3.5999999999999996" metricValueMax="4.1" metricValueMin="3.1" pageId="2" pageNumber="93" unit="mm" value="3.5999999999999996" valueMax="4.1" valueMin="3.1">3.1–4.1 mm</quantity>
|
||
), ground punctation finer (
|
||
<figureCitation box="[1157,1231,767,789]" captionStart="FIGURE 4" captionStartId="13.[151,250,1523,1547]" captionTargetBox="[151,1436,181,1505]" captionTargetId="figure-20@13.[151,1436,181,1505]" captionTargetPageId="13" captionText="FIGURE 4. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) lepidus sp. n. (B) morgani/trujillo group" figureDoi="http://doi.org/10.5281/zenodo.7892068" httpUri="https://zenodo.org/record/7892068/files/figure.png" pageId="2" pageNumber="93">Fig. 4B</figureCitation>
|
||
)........................................................................................
|
||
<emphasis bold="true" box="[1105,1436,796,819]" pageId="2" pageNumber="93">
|
||
<taxonomicName authorityName="Garcia-Hernandez" authorityYear="2009" box="[1105,1218,797,818]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="morgani">
|
||
<emphasis bold="true" box="[1105,1218,797,818]" italics="true" pageId="2" pageNumber="93">O. morgani</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1225,1293,796,818]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="trujillo">
|
||
<emphasis bold="true" box="[1225,1293,796,818]" italics="true" pageId="2" pageNumber="93">trujillo</emphasis>
|
||
</taxonomicName>
|
||
species group
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">16(11) First row of elytral systematic punctures continuous, without short breaks. Lateral rows of elytral systematic punctures much coarser than surrounding ground punctation, making them easy to discern....................................... 18</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">- First row of elytral systematic punctures not continuous, interrupted by short breaks. Lateral rows of elytral systematic punctures similar in size to surrounding ground punctation, making them difficult to discern................................. 17</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,943,965]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,943,965]" pageId="2" pageNumber="93">
|
||
17(16’) Lateral rows of elytra systematic punctures coarser than surrounding punctures.............................
|
||
<taxonomicName authorityName="Short, Greene & Garcia" authorityYear="2013" box="[1347,1436,943,965]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="maluz">
|
||
<emphasis bold="true" box="[1347,1436,943,965]" italics="true" pageId="2" pageNumber="93">O. maluz</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- Lateral rows of elytra systematic punctures similar in size to surrounding ground punctation, making them difficult to discern..........................................................................................
|
||
<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1327,1436,1002,1024]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="andinus">
|
||
<emphasis bold="true" box="[1327,1436,1002,1024]" italics="true" pageId="2" pageNumber="93">O. andinus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
18(16) Aedeagus with parameres inwardly curved at apex, not sinuate along outer margin. Elytral margins distinctly explanate in posterior third. Dorsal ground punctation on pronotum and elytra fine but distinct.......................
|
||
<taxonomicName authorityName="Short & Garcia" authorityYear="2010" box="[1300,1437,1060,1082]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="meridensis">
|
||
<emphasis bold="true" box="[1300,1437,1060,1082]" italics="true" pageId="2" pageNumber="93">O. meridensis</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- Aedeagus with parameres slightly inwardly curved at apex, sinuate along outer margin. Elytral margins not to very weakly explanate in posterior third. Dorsal ground punctation on pronotum and elytra very fine; present but appearing nearly effaced.........................................................................................
|
||
<taxonomicName authorityName="Short & Perkins" authorityYear="2004" box="[1314,1436,1148,1170]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="substillus">
|
||
<emphasis bold="true" box="[1314,1436,1148,1170]" italics="true" pageId="2" pageNumber="93">O. substillus</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,1178,1200]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,1178,1200]" pageId="2" pageNumber="93">19(4) Mesoventral process set with thin hairs. Elytra bearing dense setae from ground punctures.......................... 20</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,1207,1229]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,1207,1229]" pageId="2" pageNumber="93">- Mesoventral process set with thickened spines. Elytra at most with setae arising in rows............................ 21</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,1237,1259]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,1237,1259]" pageId="2" pageNumber="93">
|
||
20(19) Anterolateral corners of pronotum with pale spot..............................................
|
||
<emphasis bold="true" box="[1260,1436,1237,1258]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1260,1376,1237,1258]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="rupestris" status="sp. n.">
|
||
<emphasis bold="true" box="[1260,1376,1237,1258]" italics="true" pageId="2" pageNumber="93">O. rupestris</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,1237,1258]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1437,1266,1288]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1437,1266,1288]" pageId="2" pageNumber="93">
|
||
- Anterolateral corners of pronotum without pale spot...................................................
|
||
<taxonomicName authorityName="Short & Perkins" authorityYear="2004" box="[1344,1437,1266,1288]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="sharpi">
|
||
<emphasis bold="true" box="[1344,1437,1266,1288]" italics="true" pageId="2" pageNumber="93">O. sharpi</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
21(19’) Anterolateral corners of pronotum with pale spot (e.g.
|
||
<figureCitation box="[750,823,1295,1317]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1994]" captionTargetBox="[180,1407,181,1952]" captionTargetId="figure-20@3.[180,1407,181,1952]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ancho sp. n. (B) zamora sp. n. (C) yantzaza sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892054" httpUri="https://zenodo.org/record/7892054/files/figure.png" pageId="2" pageNumber="93">Fig. 1C</figureCitation>
|
||
); prosternal process with or without a pair of spines anteriorly. .................................................................................................. 22
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
- Anterolateral corners pronotum without pale spot (e.g.
|
||
<figureCitation box="[754,828,1354,1376]" captionStart="FIGURE 1" captionStartId="3.[151,250,1970,1994]" captionTargetBox="[180,1407,181,1952]" captionTargetId="figure-20@3.[180,1407,181,1952]" captionTargetPageId="3" captionText="FIGURE 1. Dorsal, ventral and lateral habitus views of Oocyclus species. (A) ancho sp. n. (B) zamora sp. n. (C) yantzaza sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892054" httpUri="https://zenodo.org/record/7892054/files/figure.png" pageId="2" pageNumber="93">Fig. 1B</figureCitation>
|
||
); prosternal process not set with a pair of spines anteriorly... .................................................................................................. 23
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<paragraph blockId="2.[151,1437,151,1523]" pageId="2" pageNumber="93">
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<keyLead pageId="2" pageNumber="93">
|
||
22(21) Prosternal process set with two spines anteriorly...................................................
|
||
<taxonomicName authority="Prosternal" authorityName="Prosternal" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="maculatus">
|
||
<emphasis bold="true" box="[1304,1437,1412,1434]" italics="true" pageId="2" pageNumber="93">O. maculatus</emphasis>
|
||
Prosternal
|
||
</taxonomicName>
|
||
process without spines...........................................................
|
||
<emphasis bold="true" box="[1262,1436,1442,1463]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1262,1376,1442,1463]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="yantzaza" status="sp. n.">
|
||
<emphasis bold="true" box="[1262,1376,1442,1463]" italics="true" pageId="2" pageNumber="93">O. yantzaza</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,1442,1463]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</keyStep>
|
||
</paragraph>
|
||
<keyStep pageId="2" pageNumber="93">
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,1471,1493]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1436,1471,1493]" pageId="2" pageNumber="93">
|
||
23(21’)Aedeagus with narrow median lobe (
|
||
<figureCitation box="[580,688,1471,1493]" captionStart="FIGURE 5" captionStartId="16.[151,250,1925,1949]" captionTargetBox="[208,1396,403,1876]" captionTargetId="figure-119@16.[155,1431,381,1908]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagi of Oocyclus species in Ecuador with rounded posterolateral corners of the pronotum. (A) rupestris sp. n. (B) sharpi, paratype from Costa Rica (C) sharpi, specimen from Ecuador (D) sharpi, specimen from Ecuador (E) kichwa sp. n. (F) kichwa sp. n. (G) yantzaza sp. n. (H) ancho sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892072" httpUri="https://zenodo.org/record/7892072/files/figure.png" pageId="2" pageNumber="93">Figs. 5E–F</figureCitation>
|
||
)...............................................
|
||
<emphasis bold="true" box="[1276,1436,1471,1493]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1276,1377,1471,1493]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="kichwa" status="sp. n.">
|
||
<emphasis bold="true" box="[1276,1377,1471,1493]" italics="true" pageId="2" pageNumber="93">O. kichwa</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,1472,1493]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph blockId="2.[151,1437,151,1523]" box="[151,1436,1500,1523]" pageId="2" pageNumber="93">
|
||
<keyLead box="[151,1432,1500,1523]" pageId="2" pageNumber="93">
|
||
- Aedeagus with broad median lobe (
|
||
<figureCitation box="[567,642,1501,1523]" captionStart="FIGURE 5" captionStartId="16.[151,250,1925,1949]" captionTargetBox="[208,1396,403,1876]" captionTargetId="figure-119@16.[155,1431,381,1908]" captionTargetPageId="16" captionText="FIGURE 5. Aedeagi of Oocyclus species in Ecuador with rounded posterolateral corners of the pronotum. (A) rupestris sp. n. (B) sharpi, paratype from Costa Rica (C) sharpi, specimen from Ecuador (D) sharpi, specimen from Ecuador (E) kichwa sp. n. (F) kichwa sp. n. (G) yantzaza sp. n. (H) ancho sp. n." figureDoi="http://doi.org/10.5281/zenodo.7892072" httpUri="https://zenodo.org/record/7892072/files/figure.png" pageId="2" pageNumber="93">Fig. 5H</figureCitation>
|
||
)....................................................
|
||
<emphasis bold="true" box="[1286,1436,1500,1522]" pageId="2" pageNumber="93">
|
||
<taxonomicName authority="Short & Torres-Gavosto & Hettinger, 2023" authorityName="Short & Torres-Gavosto & Hettinger" authorityYear="2023" box="[1286,1376,1500,1522]" class="Insecta" family="Hydrophilidae" genus="Oocyclus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="ancho" status="sp. n.">
|
||
<emphasis bold="true" box="[1286,1376,1500,1522]" italics="true" pageId="2" pageNumber="93">O. ancho</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1382,1436,1501,1522]" pageId="2" pageNumber="93" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |