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<document id="40913FC0A2A89ABB40C4870D0821B7D3" ID-DOI="10.11646/phytotaxa.312.2.3" ID-ISSN="1179-3163" ID-Zenodo-Dep="13701413" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725576598284" checkinUser="felipe" docAuthor="Nguyen, Khang Sinh, Averyanov, Leonid V., Тanaka, Noriyuki, Konstantinov, Eugene L., Maisak, Tatiana V. &amp; Nguyen, Hiep Tien" docDate="2017" docId="03BD6B4BFF9ACD06FF43E48628E84231" docLanguage="en" docName="phytotaxa.312.2.3.pdf" docOrigin="Phytotaxa 312 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.312.2.3" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Peliosanthes hirsuta Averyanov &amp; N. Tanaka 2017, sp. nov." docType="treatment" docVersion="2" lastPageNumber="203" masterDocId="FF841333FF9BCD02FFCBE6302B6F4058" masterDocTitle="New taxa of Peliosanthes and Tupistra (Asparagaceae) in the flora of Laos and Vietnam and supplemental data for T. patula" masterLastPageNumber="212" masterPageNumber="199" pageNumber="200" updateTime="1725587358570" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="B902C06BD315337F8097008C72CFF9D5">New taxa of Peliosanthes and Tupistra (Asparagaceae) in the flora of Laos and Vietnam and supplemental data for T. patula</mods:title>
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<mods:namePart id="5987051771C459FC9346EABB55335FF5">Nguyen, Khang Sinh</mods:namePart>
<mods:affiliation id="91EB8339194CF24FE8B58FD38FF83AA7">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam.</mods:affiliation>
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<mods:namePart id="4DA64D668D428194BEBF2CD04710D7AA">Averyanov, Leonid V.</mods:namePart>
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<mods:namePart id="8E68FEFFA5DA0CED9DA559B35751920A">Тanaka, Noriyuki</mods:namePart>
<mods:affiliation id="6B06E473B985CBE1FAF2F64503521E4D">98 - 11, Otsuka, Hachioji, Tokyo 192 - 0352, Japan.</mods:affiliation>
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<mods:namePart id="D52930469DEB8023CC3F55DC99E361A2">Konstantinov, Eugene L.</mods:namePart>
<mods:affiliation id="775DB5833FAE643C2884725051183585">Kaluga State University, Kaluga, Stepan Razin str., 26, RF- 0248023, Russia.</mods:affiliation>
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<mods:namePart id="BBC5808ED99DC4A0CD80C5545BFDF361">Maisak, Tatiana V.</mods:namePart>
<mods:affiliation id="E198EE7F366C5C4522AA742EC26407AB">Institute of Ecology and Biological Resources, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Cau Giay, Ha Noi, Vietnam.</mods:affiliation>
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<mods:namePart id="41811E6DE91C82B339991B2C4522D42D">Nguyen, Hiep Tien</mods:namePart>
<mods:affiliation id="7DB56B2FF42A31750AD1C225924E4D7B">Center for Plant Conservation, 25 / 32, lane 191, Lac Long Quan, Nghia Do, Cau Giay District, Ha Noi, Vietnam.</mods:affiliation>
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<mods:title id="02EEA9B7CB82516C03172F6C189455CF">Phytotaxa</mods:title>
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<mods:date id="34136204AC46F34D265017B6E53E62A5">2017</mods:date>
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<subSubSection id="C30E89D6FF9ACD03FF43E486281D4297" box="[136,882,694,719]" pageId="1" pageNumber="200" type="nomenclature">
<paragraph id="8BABDA5DFF9ACD03FF43E486281D4297" blockId="1.[136,882,694,719]" box="[136,882,694,719]" pageId="1" pageNumber="200">
<heading id="D0E36D31FF9ACD03FF43E486281D4297" box="[136,882,694,719]" fontSize="11" level="2" pageId="1" pageNumber="200" reason="3">
<taxonomicName id="4C14A1DEFF9ACD03FF43E48629E84297" ID-CoL="8X2KB" authority="Averyanov &amp; N.Tanaka" authorityName="Averyanov &amp; N. Tanaka" authorityYear="2017" box="[136,647,694,719]" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="1" pageNumber="200" phylum="Tracheophyta" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B960064FFF9ACD03FF43E4862A1C4297" bold="true" box="[136,371,694,719]" italics="true" pageId="1" pageNumber="200">Peliosanthes hirsuta</emphasis>
Averyanov &amp; N.Tanaka
</taxonomicName>
,
<emphasis id="B960064FFF9ACD03FD58E48629864297" bold="true" box="[659,745,694,719]" italics="true" pageId="1" pageNumber="200">
<taxonomicNameLabel id="A253BB34FF9ACD03FD58E48629864297" box="[659,745,694,719]" pageId="1" pageNumber="200" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="132FC6D8FF9ACD03FD32E48628294297" box="[761,838,694,719]" captionStart="FIGURE 1" captionStartId="2.[136,229,1919,1940]" captionTargetBox="[239,1342,207,1879]" captionTargetId="figure-17@2.[226,1362,187,1891]" captionTargetPageId="2" captionText="FIGURE 1. Peliosanthes hirsuta. A. Flowering plant. B. Inflorescence. C. Portion of rachis with floral bracts subtending a pedicel. D. Flattened floral bract and bracteole. E. Flower, side (left) and frontal (right) view. F. Flowers with flattened tepals, frontal (left) and rear view (right). G. Flower, sagittal section. H. Ovary, view from above. I. Ovary, side views. J. Ovary, transversal section at base (below) and in middle part (above). K, L. Separated singular carpels, adaxial views.All drawn from the type E. Konstantinov, K-305 by L.Averyanov and T. Maisak." figureDoi="http://doi.org/10.5281/zenodo.13701417" httpUri="https://zenodo.org/record/13701417/files/figure.png" pageId="1" pageNumber="200">Figs. 1</figureCitation>
,
<figureCitation id="132FC6D8FF9ACD03FC9FE486280A4297" box="[852,869,694,719]" captionStart="FIGURE 2" captionStartId="3.[136,229,1884,1905]" captionTargetBox="[237,1350,190,1859]" captionTargetId="figure-14@3.[237,1350,190,1859]" captionTargetPageId="3" captionText="FIGURE 2. Peliosanthes hirsuta sp. nov. A, B. Flowering plant. C. Inflorescence. D. Flower buds and flowers. E. Flattened plant. F. Inflorescence. G. Rachis with floral bracts (bract and bracteole), flowers removed. H. Bract and bracteole. I. Flowers, side view. J. Flowers, sagittal sections. K. Transversal section of perigone tube with intact ovary, side view (left) and view from above (right). L. Basal portion of flower in transversal section, showing basal portion (left; with ovules) and apical portion (right) of ovary. Photos of E. Konstantinov and L.Averyanov, correction and design by L.Averyanov [AE—photos from the type E. Konstantinov K-305; F-L—photos from paratype Averyanov et al, LA-VN 840]." figureDoi="http://doi.org/10.5281/zenodo.13701419" httpUri="https://zenodo.org/record/13701419/files/figure.png" pageId="1" pageNumber="200">2</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C30E89D6FF9ACD03FF43E5302EEC4773" pageId="1" pageNumber="200" type="materials_examined">
<paragraph id="8BABDA5DFF9ACD03FF43E53029494305" blockId="1.[136,1452,768,861]" pageId="1" pageNumber="200">
<emphasis id="B960064FFF9ACD03FF43E5302BD5434D" bold="true" box="[136,186,768,789]" pageId="1" pageNumber="200">Type</emphasis>
:—
<collectingCountry id="F3039ACDFF9ACD03FF13E5302A74434D" box="[216,283,768,789]" name="Laos" pageId="1" pageNumber="200">LAOS</collectingCountry>
.
<collectingRegion id="49D014BFFF9ACD03FEE8E5302AB3434D" box="[291,476,768,789]" country="Laos" name="Viangchan" pageId="1" pageNumber="200">Vientiane province</collectingRegion>
, Kasi town area, about
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to the north by old road Kasi—
<collectingRegion id="49D014BFFF9ACD03FBE6E5302FA5434D" box="[1069,1226,768,789]" country="Laos" name="Louangphabang" pageId="1" pageNumber="200">Louangphabang</collectingRegion>
, evergreen shady forest on slope of Nam Kean River at lower western slope of Ph. Phaday Mountain at elevation about
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a.s.l.,
<date id="FFAAFC9DFF9ACD03FB19E5142EE24361" box="[1234,1421,804,825]" pageId="1" pageNumber="200" value="2014-11-30">30 November 2014</date>
,
<emphasis id="B960064FFF9ACD03FA5CE5142AEB4305" italics="true" pageId="1" pageNumber="200">
<collectionCode id="ED054298FF9ACD03FA5CE5142EC64361" box="[1431,1449,804,825]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15670" name="Royal Botanic Garden Edinburgh" pageId="1" pageNumber="200" type="Herbarium">E</collectionCode>
. Konstantinov, K-305
</emphasis>
(
<typeStatus id="54AF64FFFF9ACD03FE5BE5782A874305" box="[400,488,840,861]" pageId="1" pageNumber="200" type="holotype">holotype</typeStatus>
,
<collectionCode id="ED054298FF9ACD03FE39E57829784305" box="[498,535,840,861]" country="Brazil" name="Servico de Microbiologia e Imunologia" pageId="1" pageNumber="200">LE</collectionCode>
!).
</paragraph>
<paragraph id="8BABDA5DFF9ACD03FF43E5BE2EEC4773" blockId="1.[136,1452,910,2015]" pageId="1" pageNumber="200">
Terrestrial or lithophytic perennial herb. Stem erect or oblique, to (2)34(5) cm high, covered with many whitish scales that are irregularly imbricate, cuneate, papyraceous and incised. Roots several, rigid, wiry, straight or slightly flexuose,
<collectionCode id="ED054298FF9ACD03FF3EE5E62A6643B7" box="[245,265,982,1007]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="200" type="Herbarium">Ø</collectionCode>
<quantity id="4CEC77B8FF9ACD03FEDBE5E62AEB43B7" box="[272,388,982,1007]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="1" pageNumber="200" unit="mm" value="1.75" valueMax="2.0" valueMin="1.5">1.52 mm</quantity>
. Leaves proteranthous, petiolate; petiole rigid, often curved or suberect, (3)48(10) cm long; leaf blade elliptic to broadly elliptic, acute to shortly acuminate, (6)913(15) cm long, (2.5)3.55.5(6.5) cm wide, leathery, glabrous, rugulose, often finely undulate along margin, adaxially uniformly dark green and glossy, abaxially light green to whitish green; longitudinal veins (5)69(11), prominent; secondary transverse veinlets almost invisible. Floriferous stem bearing a terminal raceme; peduncle erect, (3.5)48(9) cm long,
<collectionCode id="ED054298FF9ACD03FAD8E2562E484427" box="[1299,1319,1126,1151]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="200" type="Herbarium">Ø</collectionCode>
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, herbaceous, white, finely ridged longitudinally, hirsutulous to almost glabrous, bracteate; sterile bracts (1)23(4), loose, ensiform, acuminate, papyraceous, whitish, (1.5)25(6) cm long, (3)46(8) mm wide (when flattened for measuring); raceme ovoid, densely many-flowered, (1.5)23.5(4) cm long,
<collectionCode id="ED054298FF9ACD03FBCAE2E22F7A44B3" box="[1025,1045,1234,1259]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="1" pageNumber="200" type="Herbarium">Ø</collectionCode>
(1.2)1.51.8(2) cm; rachis thick, straight. Floral bracts 2 per flower, 2-whorled, whitish, scarious to papyraceous, narrowly triangular-ovate, acute, finely ciliate along margin and hirsutulous on abaxial surface, often somewhat descendent, slightly concave to almost flat; outer bract lying below flower, (4)610(12) mm long, (1.5)23(3.5) mm wide; inner one (bracteole) lying lateral to flower, elongate, twice smaller, oblique. Flowers solitary on very short, knob-like cylindric pedicel, not widely open, broadly campanulate, (5.5)67(8) mm across. Perigone adaxially dark dirty brown-purple, distally 6- lobed; proximal syntepalous (tubular) part broadly obconical, 6-ribbed, (2)2.53(3.5) mm long, (3.8)44.2(4.4) mm wide, abaxially white, sparsely hirsutulous; segments subsimilar, almost triangular, (2.2)2.5(2.7) mm long and broad, blunt to rounded at apex, adaxially thickened medially, abaxially densely hirsute, greenish, rimmed with purple along ciliate margin. Corona slightly convex to almost flat, to
<quantity id="4CEC77B8FF9ACD03FCF7E02628FE4677" box="[828,913,1558,1583]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="200" unit="mm" value="0.5">0.5 mm</quantity>
high, obscurely hexagonal,
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(3.3)3.54(4.2) mm, apical opening indistinctly hexagonal to irregularly circular,
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(1.4)1.61.8(2) mm. Anthers 6, nearly vertically attached to orifice of corona, sessile, ovoid to broadly ovoid, (0.7)0.8(1) mm long and wide, introrse, almost black; pollen dull pale yellow. Pistil 1, tricarpellary, dirty purplish, deeper in color distally; carpels hemispheric, distally shortly attenuate into style, adjoining carpels fused at base, at middle and apex closely juxtaposed or loosely connate along ventral sutures, ventral suture of each carpel slightly open in middle portion; ovary almost superior, broadly ovoid to almost globular,
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high and broad; style
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long and wide, deep dirty purple-violet; stigmas narrowly obovate, finely papillose; ovules 4 per locule, borne on basal placentae, narrowly oblong-ovoid.
</paragraph>
</subSubSection>
<subSubSection id="C30E89D6FF9ACD03FF76E1062AE947E3" pageId="1" pageNumber="200" type="etymology">
<paragraph id="8BABDA5DFF9ACD03FF76E1062E214717" blockId="1.[136,1452,910,2015]" box="[189,1358,1846,1871]" pageId="1" pageNumber="200">
<emphasis id="B960064FFF9ACD03FF76E1062A514717" bold="true" box="[189,318,1846,1871]" pageId="1" pageNumber="200">Etymology</emphasis>
:—The specific epithet refers to the hairy state of the perigones, bracts and floriferous stems.
</paragraph>
<paragraph id="8BABDA5DFF9ACD03FF76E16A2AE947E3" blockId="1.[136,1452,910,2015]" pageId="1" pageNumber="200">
<emphasis id="B960064FFF9ACD03FF76E16A2AB4472B" bold="true" box="[189,475,1882,1907]" pageId="1" pageNumber="200">Habitat and phenology</emphasis>
:—Primary and secondary broad-leaved evergreen forests on rocky, highly eroded, crystalline limestone at elevation
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a.s.l. Terrestrial or lithophytic herb on shady rocky slopes. Flowering in NovemberDecember.
</paragraph>
</subSubSection>
<subSubSection id="C30E89D6FF9ACD06FF76E1F628E84231" lastPageId="4" lastPageNumber="203" pageId="1" pageNumber="200" type="distribution">
<paragraph id="8BABDA5DFF9ACD03FF76E1F62F9F4787" blockId="1.[136,1452,910,2015]" box="[189,1264,1990,2015]" pageId="1" pageNumber="200">
<emphasis id="B960064FFF9ACD03FF76E1F62A204787" bold="true" box="[189,335,1990,2015]" pageId="1" pageNumber="200">Distribution</emphasis>
:—Central
<collectingCountry id="F3039ACDFF9ACD03FE06E1F6296C4787" box="[461,515,1990,2015]" name="Laos" pageId="1" pageNumber="200">Laos</collectingCountry>
(
<collectingRegion id="49D014BFFF9ACD03FDD9E1F629844787" box="[530,747,1990,2015]" country="Laos" name="Viangchan" pageId="1" pageNumber="200">Vientiane province</collectingRegion>
, Kasi district). Local endemic to central
<collectingCountry id="F3039ACDFF9ACD03FB79E1F62F834787" box="[1202,1260,1990,2015]" name="Laos" pageId="1" pageNumber="200">Laos</collectingCountry>
.
</paragraph>
<caption id="DF6B8AD5FF99CD00FF43E14F2A78487C" ID-DOI="http://doi.org/10.5281/zenodo.13701417" ID-Zenodo-Dep="13701417" httpUri="https://zenodo.org/record/13701417/files/figure.png" pageId="2" pageNumber="201" startId="2.[136,229,1919,1940]" targetBox="[239,1342,207,1879]" targetPageId="2" targetType="figure">
<paragraph id="8BABDA5DFF99CD00FF43E14F2A78487C" blockId="2.[136,1452,1919,2084]" pageId="2" pageNumber="201">
<emphasis id="B960064FFF99CD00FF43E14F2B9047CC" bold="true" box="[136,255,1919,1940]" pageId="2" pageNumber="201">FIGURE 1.</emphasis>
<taxonomicName id="4C14A1DEFF99CD00FECDE14F2AA347CC" authorityName="Averyanov &amp; N. Tanaka" authorityYear="2017" box="[262,460,1919,1940]" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="2" pageNumber="201" phylum="Tracheophyta" rank="species" species="hirsuta">
<emphasis id="B960064FFF99CD00FECDE14F2AA347CC" box="[262,460,1919,1940]" italics="true" pageId="2" pageNumber="201">Peliosanthes hirsuta</emphasis>
</taxonomicName>
. A. Flowering plant. B. Inflorescence. C. Portion of rachis with floral bracts subtending a pedicel. D. Flattened floral bract and bracteole. E. Flower, side (left) and frontal (right) view. F. Flowers with flattened tepals, frontal (left) and rear view (right). G. Flower, sagittal section. H. Ovary, view from above. I. Ovary, side views. J. Ovary, transversal section at base (below) and in middle part (above). K, L. Separated singular carpels, adaxial views. All drawn from the type
<emphasis id="B960064FFF99CD00FBECE1DB2E684858" box="[1063,1287,2027,2048]" italics="true" pageId="2" pageNumber="201">E. Konstantinov, K-305</emphasis>
by L. Averyanov and T. Maisak.
</paragraph>
</caption>
<caption id="DF6B8AD5FF98CD01FF43E16C295E487D" ID-DOI="http://doi.org/10.5281/zenodo.13701419" ID-Zenodo-Dep="13701419" httpUri="https://zenodo.org/record/13701419/files/figure.png" pageId="3" pageNumber="202" startId="3.[136,229,1884,1905]" targetBox="[237,1350,190,1859]" targetPageId="3" targetType="figure">
<paragraph id="8BABDA5DFF98CD01FF43E16C295E487D" blockId="3.[136,1452,1884,2085]" pageId="3" pageNumber="202">
<emphasis id="B960064FFF98CD01FF43E16C2A6E4729" bold="true" box="[136,257,1884,1905]" pageId="3" pageNumber="202">FIGURE 2.</emphasis>
<taxonomicName id="4C14A1DEFF98CD01FEC1E16C2ABD4729" authorityName="Nguyen &amp; Averyanov &amp; Тanaka &amp; Konstantinov &amp; Maisak &amp; Nguyen" authorityYear="2017" box="[266,466,1884,1905]" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="3" pageNumber="202" phylum="Tracheophyta" rank="species" species="hirsuta" status="sp. nov.">
<emphasis id="B960064FFF98CD01FEC1E16C2ABD4729" box="[266,466,1884,1905]" italics="true" pageId="3" pageNumber="202">Peliosanthes hirsuta</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A253BB34FF98CD01FE17E16C29454729" box="[476,554,1884,1905]" pageId="3" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
A, B. Flowering plant. C. Inflorescence. D. Flower buds and flowers. E. Flattened plant. F. Inflorescence. G. Rachis with floral bracts (bract and bracteole), flowers removed. H. Bract and bracteole. I. Flowers, side view. J. Flowers, sagittal sections. K. Transversal section of perigone tube with intact ovary, side view (left) and view from above (right). L. Basal portion of flower in transversal section, showing basal portion (left; with ovules) and apical portion (right) of ovary. Photos of E. Konstantinov and L. Averyanov, correction and design by L. Averyanov [AE—photos from the type
<emphasis id="B960064FFF98CD01FB82E1DC2E4B4859" box="[1097,1316,2028,2049]" italics="true" pageId="3" pageNumber="202">E. Konstantinov K-305</emphasis>
; F-L—photos from paratype
<emphasis id="B960064FFF98CD01FEDEEE20294C487D" box="[277,547,2064,2085]" italics="true" pageId="3" pageNumber="202">Averyanov et al, LA-VN 840</emphasis>
].
</paragraph>
</caption>
<paragraph id="8BABDA5DFF9FCD06FF76E6902A2141C9" blockId="4.[136,1452,160,617]" pageId="4" pageNumber="203">
<emphasis id="B960064FFF9FCD06FF76E6902AB040E1" bold="true" box="[189,479,160,185]" pageId="4" pageNumber="203">Taxonomic relationships</emphasis>
:—
<taxonomicName id="4C14A1DEFF9FCD06FDC9E690298B40E1" authorityName="Nguyen &amp; Averyanov &amp; Тanaka &amp; Konstantinov &amp; Maisak &amp; Nguyen" authorityYear="2017" box="[514,740,160,185]" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="species" species="hirsuta">
<emphasis id="B960064FFF9FCD06FDC9E690298B40E1" box="[514,740,160,185]" italics="true" pageId="4" pageNumber="203">Peliosanthes hirsuta</emphasis>
</taxonomicName>
is superficially similar to
<taxonomicName id="4C14A1DEFF9FCD06FC37E6902A644085" authority="Averyanov &amp; Tanaka (2012: 153)" authorityName="Averyanov &amp; Tanaka" authorityPageNumber="153" authorityYear="2012" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="species" species="argenteostriata">
<emphasis id="B960064FFF9FCD06FC37E6902FAE40E1" box="[1020,1217,160,185]" italics="true" pageId="4" pageNumber="203">P. argenteostriata</emphasis>
<bibRefCitation id="EF85A7ACFF9FCD06FB08E6902A644085" author="Averyanov, L. V. &amp; Tanaka, N." pageId="4" pageNumber="203" pagination="153 - 167" refId="ref8623" refString="Averyanov, L. V. &amp; Tanaka, N. (2012) New species of Peliosanthes and Tupistra (Asparagaceae) from eastern Indochina. Taiwania 57: 153 - 167." type="journal article" year="2012">Averyanov &amp; Tanaka (2012: 153)</bibRefCitation>
</taxonomicName>
occurring in calcareous regions of central and northern
<collectingCountry id="F3039ACDFF9FCD06FCB2E6F428B74085" box="[889,984,196,221]" name="Vietnam" pageId="4" pageNumber="203">Vietnam</collectingCountry>
(
<bibRefCitation id="EF85A7ACFF9FCD06FC2EE6F42E784085" author="Averyanov, L. V. &amp; Tanaka, N." box="[997,1303,196,221]" pageId="4" pageNumber="203" pagination="153 - 167" refId="ref8623" refString="Averyanov, L. V. &amp; Tanaka, N. (2012) New species of Peliosanthes and Tupistra (Asparagaceae) from eastern Indochina. Taiwania 57: 153 - 167." type="journal article" year="2012">Averyanov &amp; Tanaka 2012</bibRefCitation>
). However, it differs by the hairy flowers and the erect or oblique stem without accompanying any markedly developed plagiotropic rhizome. The new species is also unusual in having an almost apocarpous gynoecium. To our knowledge, there has been no report of such a gynoecium and hairy flowers in other members of
<taxonomicName id="4C14A1DEFF9FCD06FC2AE7002F1F4111" authorityName="Andrews" authorityYear="1808" box="[993,1136,304,329]" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="genus">
<emphasis id="B960064FFF9FCD06FC2AE7002F1F4111" box="[993,1136,304,329]" italics="true" pageId="4" pageNumber="203">Peliosanthes</emphasis>
</taxonomicName>
, or even of
<taxonomicName id="4C14A1DEFF9FCD06FB31E7002EC44111" box="[1274,1451,304,329]" pageId="4" pageNumber="199" rank="tribe" tribe="Ophiopogoneae">Ophiopogoneae</taxonomicName>
(
<bibRefCitation id="EF85A7ACFF9FCD06FF5BE7642A184135" author="Dahlgren, R. M. N. &amp; Clifford, H. T. &amp; Yeo, P. F." box="[144,375,340,365]" pageId="4" pageNumber="203" refId="ref9151" refString="Dahlgren, R. M. N., Clifford, H. T. &amp; Yeo, P. F. (1985) The Families of the Monocotyledons. Springer-Verlag, Berlin, 520 pp. https: // doi. org / 10.1007 / 978 - 3 - 642 - 61663 - 1" type="book" year="1985">
Dahlgren
<emphasis id="B960064FFF9FCD06FF34E7642A404135" box="[255,303,340,365]" italics="true" pageId="4" pageNumber="203">et al</emphasis>
. 1985
</bibRefCitation>
,
<bibRefCitation id="EF85A7ACFF9FCD06FE4BE76429E84135" author="Conran, J. G. &amp; Tamura, M. N." box="[384,647,340,365]" pageId="4" pageNumber="203" pagination="186 - 198" refId="ref9008" refString="Conran, J. G. &amp; Tamura, M. N. (1998) Convallariaceae. In: Kubitzki, K. (Ed.) The Families and Genera of Vascular Plants III. Flowering Plants, Monocotyledons, Lilianae (except Orchidaceae). Springer, Berlin, Heidelberg &amp; New York, pp. 186 - 198. https: // doi. org / 10.1007 / 978 - 3 - 662 - 03533 - 7 _ 25" type="book chapter" year="1998">Conran &amp; Tamura 1998</bibRefCitation>
) or
<taxonomicName id="4C14A1DEFF9FCD06FD79E76428E84135" box="[690,903,340,365]" class="Liliopsida" family="Asparagaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="199" phylum="Tracheophyta" rank="subFamily" subFamily="Ophiopogonoideae">Ophiopogonoideae</taxonomicName>
(
<bibRefCitation id="EF85A7ACFF9FCD06FC5FE7642F2B4135" author="Takhtajan, A." box="[916,1092,340,365]" pageId="4" pageNumber="203" refId="ref9403" refString="Takhtajan, A. (2009) Flowering plants. 2 nd ed .. Springer, Berlin etc., 871 pp. http: // dx. doi. org / 10.1007 / 978 - 1 - 4020 - 9609 - 9" type="book" year="2009">Takhtajan 2009</bibRefCitation>
) including
<taxonomicName id="4C14A1DEFF9FCD06FB0BE7642E7D4135" box="[1216,1298,340,365]" class="Liliopsida" family="Asparagaceae" genus="Liriope" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="genus">
<emphasis id="B960064FFF9FCD06FB0BE7642E7D4135" box="[1216,1298,340,365]" italics="true" pageId="4" pageNumber="203">Liriope</emphasis>
</taxonomicName>
,
<taxonomicName id="4C14A1DEFF9FCD06FAD4E7642EC44135" box="[1311,1451,340,365]" class="Liliopsida" family="Asparagaceae" genus="Ophiopogon" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="genus">
<emphasis id="B960064FFF9FCD06FAD4E7642EC44135" box="[1311,1451,340,365]" italics="true" pageId="4" pageNumber="203">Ophiopogon</emphasis>
</taxonomicName>
and
<taxonomicName id="4C14A1DEFF9FCD06FF7CE7482A2941C9" authorityName="Andrews" authorityYear="1808" box="[183,326,376,401]" class="Liliopsida" family="Asparagaceae" genus="Peliosanthes" kingdom="Plantae" order="Asparagales" pageId="4" pageNumber="203" phylum="Tracheophyta" rank="genus">
<emphasis id="B960064FFF9FCD06FF7CE7482A2941C9" box="[183,326,376,401]" italics="true" pageId="4" pageNumber="203">Peliosanthes</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BABDA5DFF9FCD06FF76E7AC28E84231" blockId="4.[136,1452,160,617]" pageId="4" pageNumber="203">
<emphasis id="B960064FFF9FCD06FF76E7AC29D741ED" bold="true" box="[189,696,412,437]" pageId="4" pageNumber="203">
Additional specimens studied (
<typeStatus id="54AF64FFFF9FCD06FDFDE7AC29DF41ED" box="[566,688,412,437]" pageId="4" pageNumber="203" type="paratype">paratypes</typeStatus>
)
</emphasis>
:—
<collectingCountry id="F3039ACDFF9FCD06FD11E7AC284741ED" box="[730,808,412,437]" name="Laos" pageId="4" pageNumber="203">LAOS</collectingCountry>
.
<date id="FFAAFC9DFF9FCD06FCFCE7AC289441ED" box="[823,1019,412,437]" pageId="4" pageNumber="203" value="2016-10-21">21 October 2016</date>
and 1 November,
<emphasis id="B960064FFF9FCD06FB11E7AC2B814181" italics="true" pageId="4" pageNumber="203">L. Averyanov, LA-VN 840a</emphasis>
(LE!). Specimen prepared from cultivated plant originally collected in central Laos (
<collectingRegion id="49D014BFFF9FCD06FB00E7F02EC74181" box="[1227,1448,448,473]" country="Laos" name="Viangchan" pageId="4" pageNumber="203">Vientiane province</collectingRegion>
, Kasi district, Namken village, Phachao Mountain, around point
<geoCoordinate id="EE20BC9AFF9FCD06FC9AE7D4289941A5" box="[849,1014,484,509]" degrees="19" direction="north" minutes="18" orientation="latitude" pageId="4" pageNumber="203" precision="1" seconds="45.5" value="19.312637">19°1845.5” N</geoCoordinate>
<geoCoordinate id="EE20BC9AFF9FCD06FC36E7D42FC141A5" box="[1021,1198,484,509]" degrees="102" direction="east" minutes="22" orientation="longitude" pageId="4" pageNumber="203" precision="1" seconds="31.4" value="102.37539">102°2231.4” E</geoCoordinate>
, primary broad-leaved evergreen forest on very steep rocky slopes of mountain composed of highly eroded solid crystalline limestone at elevations
<quantity id="4CEC77B8FF9FCD06FEC9E41C2AFB421D" box="[258,404,556,581]" metricMagnitude="2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="4" pageNumber="203" unit="m" value="850.0" valueMax="1000.0" valueMin="700.0">7001000 m</quantity>
. a.s.l., terrestrial herb on shady place, leaves uniformly dark green and glossy, common,
<date id="FFAAFC9DFF9FCD06FA5BE41C2A7C4231" pageId="4" pageNumber="203" value="2013-03-24">24 March 2013</date>
,
<emphasis id="B960064FFF9FCD06FED5E46028184231" box="[286,887,592,617]" italics="true" pageId="4" pageNumber="203">L. Averyanov, N.S. Khang, S. Lorphengsy, LA-VN 840</emphasis>
).
</paragraph>
</subSubSection>
</treatment>
</document>