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<document ID-DOI="http://doi.org/10.5281/zenodo.3571116" ID-GBIF-Dataset="a02f182b-32e4-4649-afe0-983cb21ceb9e" ID-GBIF-Taxon="162362554" ID-Zenodo-Dep="3571116" approvalRequired="5" approvalRequired_for_illustrations="3" approvalRequired_for_treatments="2" checkinTime="1576077498408" checkinUser="jeremy" docAuthor="M. A. Lieftinck" docDate="1940" docId="03B487FFFFBDFFD0FEF8BA0DFC2BAA1F" docLanguage="en" docName="Lieftinck - 1940 - Descriptions and records of South-east Asiatic Odonata (II) ABBYYed.pdf.imf" docOrigin="Treubia 17 (4)" docStyle="DocumentStyle{}" docTitle="Amphicnemis annae Lieftinck, 1940, sp. n." docType="treatment" docVersion="6" lastPageId="36" lastPageNumber="371" masterDocId="FF8DFF87FF9EFFF4FFF0BE36FF92AF52" masterDocTitle="Descriptions and records of South-East Asiatic Odonata (II)" masterLastPageNumber="392" masterPageNumber="337" pageId="35" pageNumber="370" updateTime="1632324461921" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Descriptions and records of South-East Asiatic Odonata (II)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>M. A. Lieftinck</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Treubia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>1940</mods:date>
<mods:detail type="pubDate">
<mods:number>1940-01-01</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>17</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>337</mods:start>
<mods:end>392</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://doi.org/10.5281/zenodo.3571116</mods:identifier>
<mods:identifier type="GBIF-Dataset">a02f182b-32e4-4649-afe0-983cb21ceb9e</mods:identifier>
<mods:identifier type="Zenodo-Dep">3571116</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="162362554" LSID="urn:lsid:plazi:treatment:03B487FFFFBDFFD0FEF8BA0DFC2BAA1F" httpUri="http://treatment.plazi.org/id/03B487FFFFBDFFD0FEF8BA0DFC2BAA1F" lastPageId="36" lastPageNumber="371" pageId="35" pageNumber="370">
<subSubSection box="[264,1356,1082,1122]" pageId="35" pageNumber="370" type="nomenclature">
<paragraph box="[264,1356,1082,1122]" pageId="35" pageNumber="370">
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[264,667,1083,1121]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="annae" status="sp . n .">
<emphasis bold="true" box="[264,667,1083,1121]" pageId="35" pageNumber="370">Amphicnemis annae</emphasis>
</taxonomicName>
,
<taxonomicNameLabel box="[710,800,1087,1121]" pageId="35" pageNumber="370">sp. n.</taxonomicNameLabel>
(
<figureCitation box="[839,1066,1082,1121]" captionStart="Pl" captionStartId="28.[749,785,1761,1789]" captionTargetBox="[510,2495,140,1742]" captionTargetPageId="28" captionText="Pl . 15 . — Structural characters of Malaysian species of Amphicnemis . 1 - 3 . Male anal appendages , dorsal view and right side . 4 . Right lateral view of prothorax ." figureDoi="http://doi.org/10.5281/zenodo.3571151" httpUri="https://zenodo.org/record/3571151/files/figure.png" pageId="35" pageNumber="370">pl. 15 fig. 4</figureCitation>
,
<figureCitation box="[1100,1327,1083,1122]" captionStart="Pl" captionStartId="29.[722,754,1768,1794]" captionTargetBox="[683,2440,144,1755]" captionTargetPageId="29" captionText="Pl. 16. — Male anal appendages of Malaysian species of Amphicnemis, dorsal view and right side" figureDoi="http://doi.org/10.5281/zenodo.3571153" httpUri="https://zenodo.org/record/3571153/files/figure.png" pageId="35" pageNumber="370">
pl. 16 fig.
<emphasis bold="true" box="[1308,1327,1085,1113]" pageId="35" pageNumber="370">4</emphasis>
</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="370" type="reference_group">
<paragraph pageId="35" pageNumber="370">
1911. RIS, Ann. Soc. ent. Belg. 55, p. 236 237 (key) fig-. 5 (♂ apps.). — ♂ (nec ♀!), W. Borneo (
<taxonomicName authorityName="Selys" authorityYear="1863" box="[589,785,1190,1222]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis bold="true" box="[589,611,1191,1216]" italics="true" pageId="35" pageNumber="370">A</emphasis>
.
<emphasis box="[643,785,1190,1222]" italics="true" pageId="35" pageNumber="370">wallacei</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph lastBlockId="35.[376,601,1280,1312]" pageId="35" pageNumber="370">
1913.
<emphasis box="[377,519,1235,1261]" italics="true" pageId="35" pageNumber="370">LAIDLAW</emphasis>
, P.
<emphasis box="[580,605,1235,1261]" italics="true" pageId="35" pageNumber="370">Z</emphasis>
. S. London, p. 70; 74 (key ♂), pl. 4
<figureCitation box="[1219,1308,1234,1265]" captionStart="Fig" captionStartId="17.[225,275,1195,1223]" captionTargetBox="[228,1073,591,1173]" captionTargetPageId="17" captionText="Fig. 7. Coeliccia arcuata, sp. n. ♂. Anal ap ­ pendages, dorsal view and right side ." figureDoi="http://doi.org/10.5281/zenodo.3571133" httpUri="https://zenodo.org/record/3571133/files/figure.png" pageId="35" pageNumber="370">fig. 7</figureCitation>
(♂ proth.). — ♂ Sarawak (
<taxonomicName authorityName="Selys" authorityYear="1863" box="[389,574,1280,1312]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis box="[389,411,1280,1305]" italics="true" pageId="35" pageNumber="370">A</emphasis>
. wallacei
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection pageId="35" pageNumber="370" type="materials_examined">
<paragraph pageId="35" pageNumber="370">
<materialsCitation ID-GBIF-Occurrence="2572187097" collectingCode="Buitenzorg Museum" collectingDate="1934-01-16" collectorName="L. COOMANS DE RUITER" country="Malaysia" location="Singkawang, forest-marsh near Tjapkala" pageId="35" pageNumber="370" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="W. Borneo">
Material studied: —
<specimenCount box="[754,811,1333,1363]" pageId="35" pageNumber="370" type="male">1 ♂</specimenCount>
(ad.),
<specimenCount box="[948,1006,1332,1363]" pageId="35" pageNumber="370" type="female">2 ♀</specimenCount>
(juv., red), W
<collectingRegion box="[1317,1497,1334,1364]" pageId="35" pageNumber="370">Borneo</collectingRegion>
,
<collectingMunicipality box="[1530,1748,1333,1372]" pageId="35" pageNumber="370">Singkawang</collectingMunicipality>
, forest-marsh near
<location LSID="urn:lsid:plazi:treatment:03B487FFFFBDFFD0FEF8BA0DFC2BAA1F:8EC26032FFBDFFD7FD91BB5DFC9AAAC0" box="[609,776,1387,1426]" country="Borneo" municipality="Singkawang" name="Tjapkala" pageId="35" pageNumber="370" stateProvince="Borneo">Tjapkala</location>
,
<date box="[808,1057,1387,1424]" pageId="35" pageNumber="370" value="1934-null-16">
<collectingDate box="[808,1057,1387,1424]" pageId="35" pageNumber="370" value="1934-null-16">Jan. 16, 1934</collectingDate>
</date>
,
<collectorName box="[1089,1521,1387,1418]" pageId="35" pageNumber="370">
L.
<emphasis box="[1147,1369,1387,1418]" italics="true" pageId="35" pageNumber="370">COOMANS DE</emphasis>
RUITER
</collectorName>
leg.
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2572187096" collectingCode="Buitenzorg Museum" collectingDate="1934-01-16" collectorName="L. COOMANS DE RUITER" country="Malaysia" location="Singkawang, forest-marsh near Tjapkala" pageId="35" pageNumber="370" specimenCount="1" specimenCount-male="1" stateProvince="W. Borneo" typeStatus="holotype">
<typeStatus pageId="35" pageNumber="370">Holo- type</typeStatus>
<specimenCount box="[398,420,1441,1471]" pageId="35" pageNumber="370" type="male">♂</specimenCount>
and
</materialsCitation>
<materialsCitation ID-GBIF-Occurrence="2572187098" box="[537,1339,1440,1480]" collectingCode="Buitenzorg Museum" collectingDate="1934-01-16" collectorName="L. COOMANS DE RUITER" country="Malaysia" location="Singkawang, forest-marsh near Tjapkala" pageId="35" pageNumber="370" specimenCount="1" specimenCount-female="1" stateProvince="W. Borneo" typeStatus="allotype">
<typeStatus box="[537,753,1441,1480]" pageId="35" pageNumber="370">allotype</typeStatus>
<specimenCount box="[777,796,1440,1471]" pageId="35" pageNumber="370" type="female">♀</specimenCount>
in the Buitenzorg Museum
</materialsCitation>
.
</paragraph>
<paragraph pageId="35" pageNumber="370">
This very distinct species has long been known as
<taxonomicName authorityName="Selys" authorityYear="1863" box="[1273,1426,1512,1551]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis box="[1273,1426,1512,1551]" italics="true" pageId="35" pageNumber="370">wallacei</emphasis>
</taxonomicName>
; but since SELYS’S type of
<taxonomicName authorityName="Selys" authorityYear="1863" box="[416,622,1565,1597]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis box="[416,441,1567,1597]" italics="true" pageId="35" pageNumber="370">A</emphasis>
.
<emphasis box="[475,622,1565,1597]" italics="true" pageId="35" pageNumber="370">wallacei</emphasis>
</taxonomicName>
is indistinguishable from the species described by LAIDLAW as louisae, this name unfortunately should be dropped as a synonym of
<taxonomicName authorityName="Selys" authorityYear="1863" box="[1603,1749,1620,1651]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">wallacei</taxonomicName>
.
</paragraph>
</subSubSection>
<paragraph pageId="35" pageNumber="370">
<subSubSection pageId="35" pageNumber="370" type="discussion">
Only few specimens. of
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[806,911,1683,1704]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="annae">annae</taxonomicName>
were known, and no descriptions of it are available in the literature. I believe its position to be closest to
<taxonomicName authorityName="Selys" authorityYear="1863" box="[1510,1658,1727,1759]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">wallacei</taxonomicName>
. The supposed allotype ♀ of
<taxonomicName authorityName="Selys" authorityYear="1863" box="[712,860,1779,1811]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">wallacei</taxonomicName>
, described by RIS, neither belongs to
<taxonomicName authorityName="Selys" authorityYear="1863" box="[1613,1760,1781,1812]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="wallacei">wallacei</taxonomicName>
nor even to
<taxonomicName authorityName="M.A. Lieftinck" authorityYear="1940" box="[512,616,1844,1865]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="370" phylum="Arthropoda" rank="species" species="annae">
<emphasis box="[512,616,1844,1865]" italics="true" pageId="35" pageNumber="370">annae</emphasis>
</taxonomicName>
, and I am unable at the present to identify it from the
</subSubSection>
<subSubSection lastPageId="36" lastPageNumber="371" pageId="35" pageNumber="370" type="description">
description alone. The following notes may be added to those given in the key to the species of both sexes:—
<emphasis box="[347,438,2048,2080]" italics="true" pageId="35" pageNumber="370">Male</emphasis>
(ad.). —- Labium pale yellow. Mandibles yellow, each with a large, slightly diffuse, triangular, brownish spot upon middle. Labrum shining black, its anterior margin bright yellow and fringed with yellow hairs. Anteclypeus yellow, postclypeus shining black. Frons in front black with two yellow' spots, one on each side of the middle. Dorsal surface of head metallic-green. Antennae brown, basal joints striped with yellow anteriorly. Rear of the head black.
<paragraph lastBlockId="35.[264,528,2479,2517]" pageId="35" pageNumber="370">Prothorax metallic-green above, anterior lobe with a pair of round orangish spots, one on each side of the middle; propleuron also metallic-green, its lateral border yellow.</paragraph>
<paragraph pageId="35" pageNumber="370">Synthorax metallic-green, as far down as the first lateral suture and the spiracle, including the dorsal two-thirds of the mesinfraepisternites and the dorsal fifth of the metepisternites. Sides and under surfaces pale yellow.</paragraph>
<paragraph box="[347,1715,208,252]" pageId="36" pageNumber="371">
M. A. LıEıfTıNck: Descriptions
<emphasis box="[925,1136,212,244]" italics="true" pageId="36" pageNumber="371">and records</emphasis>
0
<emphasis box="[1173,1227,214,252]" italics="true" pageId="36" pageNumber="371">ƒ S</emphasis>
.
<emphasis box="[1239,1266,216,245]" italics="true" pageId="36" pageNumber="371">E</emphasis>
.
<emphasis box="[1295,1591,215,248]" italics="true" pageId="36" pageNumber="371">
Asiatic
<taxonomicName authorityName="Fabricius" authorityYear="1793" box="[1439,1591,217,248]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="36" pageNumber="371" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
</emphasis>
.
<pageNumber ambiguity="12" box="[1661,1715,220,249]" fuzzyness="0" pageId="36" pageNumber="371" score="117.0" value="371">371</pageNumber>
</paragraph>
<paragraph pageId="36" pageNumber="371">Legs yellow, spines brown. Femora pale ochreous, with sharply defined black apical ring; a very diffuse brownish line runs along the distal half of their posterior surfaces (most marked on anterior pair of femora).</paragraph>
<paragraph pageId="36" pageNumber="371">
Wings hyaline;
<emphasis box="[607,643,471,501]" italics="true" pageId="36" pageNumber="371">M</emphasis>
3 arises at the subnodus or slightly distal to it,
<emphasis box="[1553,1599,483,514]" italics="true" pageId="36" pageNumber="371">Rs</emphasis>
about midway between subnodus and Px 1. Pterostigma greyish-brown with fine yellow margin. Postnodals 12 - 13 in fore wing, 10 - 11 in hind wing.
</paragraph>
<paragraph pageId="36" pageNumber="371">
Abdomen coloured as described for “
<taxonomicName authorityName="Laidlaw" authorityYear="1913" box="[1039,1161,635,666]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="36" pageNumber="371" phylum="Arthropoda" rank="species" species="louisae">louisae</taxonomicName>
”
<subSubSection pageId="36" pageNumber="371" type="description">
(=
<taxonomicName authorityName="Selys" authorityYear="1863" box="[1291,1439,638,670]" class="Insecta" family="Coenagrionidae" genus="Amphicnemis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="36" pageNumber="371" phylum="Arthropoda" rank="species" species="wallacei">
<emphasis box="[1291,1439,638,670]" italics="true" pageId="36" pageNumber="371">wallacei</emphasis>
</taxonomicName>
) by
<emphasis box="[1555,1709,643,674]" italics="true" pageId="36" pageNumber="371">LAIDLAW</emphasis>
. Tenth segment pale reddish-brown basally and along posterior margin. Anal appendages of the same colour, shaped as shown on
<figureCitation box="[1265,1498,745,784]" captionStart="Pl" captionStartId="29.[722,754,1768,1794]" captionTargetBox="[683,2440,144,1755]" captionTargetPageId="29" captionText="Pl. 16. — Male anal appendages of Malaysian species of Amphicnemis, dorsal view and right side" figureDoi="http://doi.org/10.5281/zenodo.3571153" httpUri="https://zenodo.org/record/3571153/files/figure.png" pageId="36" pageNumber="371">pl. 16 fig. 4</figureCitation>
.
<paragraph pageId="36" pageNumber="371">
<emphasis box="[311,442,793,826]" italics="true" pageId="36" pageNumber="371">Female</emphasis>
(red colour-phase). — Head coloured much as in the opposite sex, but pale areas slightly enlarged. Labrum with the distal half yellow, frons with a complete yellow anterior fascia, and a yellow line along margin of compound eyes.
</paragraph>
<paragraph pageId="36" pageNumber="371">Pro- and synthorax, and legs throughout scarlet-red. Spine of posterior lobe of prothorax equal in length to that of the male. Ante-alar triangles metallic blue-green. Postnodals 12 - 13 in fore wing, 11 in hind wing. Abdomen with segm. 10 and appendages whitish; valves also pale-coloured, slightly surpassing end of abdomen.</paragraph>
<paragraph box="[321,953,1305,1357]" pageId="36" pageNumber="371">Apparently a very scarce species.</paragraph>
</subSubSection>
</paragraph>
</subSubSection>
</paragraph>
</treatment>
</document>
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