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<mods:title id="6539A2C30E751001C40189192AB6071C">The Cephalozygoptera, a new, extinct suborder of Odonata with new taxa from the early Eocene Okanagan Highlands, western North America</mods:title>
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<mods:namePart id="4D0768611469A060098E124BFC0CE0C8">Archibald, Bruce</mods:namePart>
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<mods:namePart id="A2F995CC24678CF7B5A021F887E460F7">Cannings, Robert A.</mods:namePart>
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<mods:namePart id="1F8FCA70900C59648938F2C2355F48AA">Erickson, Robert J.</mods:namePart>
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<mods:namePart id="4CD9485B3F46E94F991AF3A1087704A1">Bybee, Seth M.</mods:namePart>
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<mods:namePart id="90D8A34CF7C27AFAB312DF15D515859D">Mathewes, Rolf W.</mods:namePart>
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<mods:date id="79AAAC9E9992FB58829A5449C101DD88">2021</mods:date>
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<treatment id="03B487C20030FFB3FF5BFEC2FF4B12EC" ID-DOI="http://doi.org/10.5281/zenodo.4672715" ID-GBIF-Taxon="180881811" ID-Zenodo-Dep="4672715" LSID="urn:lsid:plazi:treatment:03B487C20030FFB3FF5BFEC2FF4B12EC" httpUri="http://treatment.plazi.org/id/03B487C20030FFB3FF5BFEC2FF4B12EC" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35">
<subSubSection id="C307655F0030FFB5FF5BFEC2FE751354" box="[151,475,295,321]" pageId="34" pageNumber="35" type="nomenclature">
<paragraph id="8BA236D40030FFB5FF5BFEC2FE751354" blockId="34.[151,475,295,357]" box="[151,475,295,321]" pageId="34" pageNumber="35">
<emphasis id="B969EAC60030FFB5FF5BFEC2FE751354" bold="true" box="[151,475,295,321]" pageId="34" pageNumber="35">
Genus
<taxonomicName id="4C1D4D570030FFB5FF22FEC2FE751354" authority="Scudder" authorityName="Scudder" box="[238,475,295,321]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF22FEC2FEC51354" bold="true" box="[238,363,295,321]" italics="true" pageId="34" pageNumber="35">Dysagrion</emphasis>
Scudder
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<subSubSection id="C307655F0030FFB5FF5BFEAEFEF51370" box="[151,347,331,357]" pageId="34" pageNumber="35" type="description">
<paragraph id="8BA236D40030FFB5FF5BFEAEFEF51370" blockId="34.[151,475,295,357]" box="[151,347,331,357]" pageId="34" pageNumber="35">
<figureCitation id="13262A510030FFB5FF5BFEAEFEAC1370" box="[151,258,331,357]" captionStart-0="FIGURE 4" captionStart-1="FIGURE 5" captionStart-2="FIGURE 6" captionStartId-0="11.[151,250,1923,1947]" captionStartId-1="12.[151,250,1943,1967]" captionStartId-2="13.[151,250,1943,1967]" captionTargetBox-0="[343,1242,181,1891]" captionTargetBox-1="[250,1324,200,1919]" captionTargetBox-2="[262,1315,184,1919]" captionTargetId-0="figure-17@11.[332,1255,181,1899]" captionTargetId-1="figure-19@12.[250,1338,181,1919]" captionTargetId-2="figure-17@13.[262,1326,181,1919]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionText-0="FIGURE 4. Dysagrion packardii cotype MCZ 4656 (Packard number 252): A, drawing; B, photograph; C, holotype (Packard number 146) redrawn from Zittel (1885: his Fig. 979), also redrawn by Scudder (1890: his plate 6, Fig. 3). A, B drawn from a photograph supplied by MCZ personnel. All to scale, 5 mm." captionText-1="FIGURE 5. Dysagrion lakesii Scudder type specimen MCZ 4101 (Packard number 259, not previously illustrated): A, photograph; B, drawing from a photograph supplied by MCZ personnel. Both to scale, 5 mm." captionText-2="FIGURE 6. Dysagrion fredericii Scudders number 4167/4168 (now MCZ numbers 381A, B “type”): A, photograph; B, drawing from a photograph supplied by MCZ personnel. Both to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558805" figureDoi-1="http://doi.org/10.5281/zenodo.4558807" figureDoi-2="http://doi.org/10.5281/zenodo.4558809" httpUri-0="https://zenodo.org/record/4558805/files/figure.png" httpUri-1="https://zenodo.org/record/4558807/files/figure.png" httpUri-2="https://zenodo.org/record/4558809/files/figure.png" pageId="34" pageNumber="35">Figs. 46</figureCitation>
,
<figureCitation id="13262A510030FFB5FEC1FEAEFEF81370" box="[269,342,331,357]" captionStart-0="FIGURE 14" captionStart-1="FIGURE 15" captionStart-2="FIGURE 16" captionStartId-0="35.[151,250,1193,1217]" captionStartId-1="35.[151,250,1929,1953]" captionStartId-2="37.[151,250,1946,1970]" captionTargetBox-0="[156,1427,245,1166]" captionTargetBox-1="[161,1428,1323,1898]" captionTargetBox-2="[225,1349,191,1914]" captionTargetId-0="figure-47@35.[151,1436,237,1170]" captionTargetId-1="figure-102@35.[151,1436,1314,1906]" captionTargetId-2="figure-17@37.[225,1363,181,1922]" captionTargetPageId-0="35" captionTargetPageId-1="35" captionTargetPageId-2="37" captionText-0="FIGURE 14. Tree resulting from parsimony-based analyses for Analysis 1: Tarsophlebiidae, the suborders of Odonata and former members of Dysagrionidae (Burmadysagrion, Electrodysagrion, Paleodysagrion). One single most-parsimonious tree (MPT) based on 23 continuous and 79 discrete characters. Symmetrical resampling values Ξ70 are labelled at the nodes." captionText-1="FIGURE 15. Tree resulting from parsimony-based analyses for Analysis 2 (relationship Whetwhetaksidae to confirmed families of Cephalozygoptera). One single most-parsimonious tree (MPT) based on 23 continuous and 79 discrete characters. Symmetrical resampling values Ξ70 are labeled at the nodes." captionText-2="FIGURE 16. Dysagrion pruettae, new species holotype, SR 13-005-012. A, photograph; B, drawing. Wings and labels (FW, forewing; HW, hind wing) colour-coded to match Figs. 17 and 18. Both to scale, 5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4558832" figureDoi-1="http://doi.org/10.5281/zenodo.4558834" figureDoi-2="http://doi.org/10.5281/zenodo.4558836" httpUri-0="https://zenodo.org/record/4558832/files/figure.png" httpUri-1="https://zenodo.org/record/4558834/files/figure.png" httpUri-2="https://zenodo.org/record/4558836/files/figure.png" pageId="34" pageNumber="35">1416</figureCitation>
.
</paragraph>
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<subSubSection id="C307655F0030FFB5FF5BFE76FB0414F4" pageId="34" pageNumber="35" type="diagnosis">
<paragraph id="8BA236D40030FFB5FF5BFE76FB0414F4" blockId="34.[151,1437,403,2014]" pageId="34" pageNumber="35">
<emphasis id="B969EAC60030FFB5FF5BFE76FE3613B8" bold="true" box="[151,408,403,429]" pageId="34" pageNumber="35">Diagnosis (emended).</emphasis>
Wings distinguished from those of other
<taxonomicName id="4C1D4D570030FFB5FCB4FE76FBB513B8" box="[888,1051,403,429]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
genera by subnodus with reverse obliquity [all others and tentative
<taxonomicName id="4C1D4D570030FFB5FDC3FE52FD1C13C4" box="[527,690,439,465]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
as below (where known), normal]; and from all
<taxonomicName id="4C1D4D570030FFB5FB03FE52FADC13C4" box="[1231,1394,439,465]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Dysagrioninae">Dysagrioninae</taxonomicName>
by: from
<taxonomicName id="4C1D4D570030FFB5FF1BFE3EFEC613E0" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[215,360,475,501]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF1BFE3EFEC613E0" box="[215,360,475,501]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FEB4FE39FE4313E0" box="[376,493,476,501]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FEB4FE39FE4313E0" box="[376,493,476,501]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
by pterostigma long (47 times its width) [
<taxonomicName id="4C1D4D570030FFB5FC29FE3EFBD813E0" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[997,1142,475,501]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC29FE3EFBD813E0" box="[997,1142,475,501]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
: 2.5 times;
<taxonomicName id="4C1D4D570030FFB5FACDFE39FAD813E0" box="[1281,1398,476,501]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FACDFE39FAD813E0" box="[1281,1398,476,501]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
: 3; not from
<taxonomicName id="4C1D4D570030FFB5FEC8FE1AFE25100C" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[260,395,511,537]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FEC8FE1AFE25100C" box="[260,395,511,537]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
: 34;
<taxonomicName id="4C1D4D570030FFB5FE1BFE1AFDD8100C" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[471,630,511,537]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE1BFE1AFDD8100C" box="[471,630,511,537]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
: 4;
<taxonomicName id="4C1D4D570030FFB5FD6AFE1AFC93100C" box="[678,829,511,537]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD6AFE1AFC93100C" box="[678,829,511,537]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
: 3.54;
<taxonomicName id="4C1D4D570030FFB5FC52FE1AFB90100C" authorityName="Cockerell" authorityYear="1908" box="[926,1086,511,537]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC52FE1AFB90100C" box="[926,1086,511,537]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
: 5;
<taxonomicName id="4C1D4D570030FFB5FBA3FDE5FAA2100C" box="[1135,1292,512,537]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FBA3FDE5FAA2100C" box="[1135,1292,512,537]" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
species: unknown]; with oblique brace vein joining posterior-proximal corner of pterostigma in forewing, hind wing [not present in
<taxonomicName id="4C1D4D570030FFB5FF12FDA2FECB1074" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[222,357,583,609]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF12FDA2FECB1074" box="[222,357,583,609]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FEB8FDA2FDBA1074" authorityName="Cockerell" authorityYear="1908" box="[372,532,583,609]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FEB8FDA2FDBA1074" box="[372,532,583,609]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
(both known from fore-, hind wings), not known in
<taxonomicName id="4C1D4D570030FFB5FBAAFDA2FB531074" box="[1126,1277,583,609]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FBAAFDA2FB531074" box="[1126,1277,583,609]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
(fore- or hind wings?); known in
<taxonomicName id="4C1D4D570030FFB5FEB8FD89FD381090" authority=", Okanoptyeryx" authorityName="Okanoptyeryx" box="[372,662,619,646]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FEB8FD89FE471090" box="[372,489,620,645]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
,
<emphasis id="B969EAC60030FFB5FE35FD8EFD381090" box="[505,662,619,645]" italics="true" pageId="34" pageNumber="35">Okanoptyeryx</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FD69FD8EFCEA1090" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[677,836,619,645]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD69FD8EFCEA1090" box="[677,836,619,645]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
; wing region not preserved in
<taxonomicName id="4C1D4D570030FFB5FB6BFD8EFA3B1090" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[1191,1429,619,645]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FB6BFD8EFA3B1090" box="[1191,1429,619,645]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FF5BFD75FE9A10BC" box="[151,308,656,681]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFD75FE9A10BC" box="[151,308,656,681]" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
species]; costal space distal to pterostigma 1 cell wide [distinct from
<taxonomicName id="4C1D4D570030FFB5FB91FD6AFB5A10BC" box="[1117,1268,655,681]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FB91FD6AFB5A10BC" box="[1117,1268,655,681]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FAC9FD6AFA3810BC" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1285,1430,655,681]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FAC9FD6AFA3810BC" box="[1285,1430,655,681]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FF5BFD56FE9810D8" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[151,310,691,717]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFD56FE9810D8" box="[151,310,691,717]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
: all 2;
<taxonomicName id="4C1D4D570030FFB5FE45FD56FDBE10D8" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[393,528,691,717]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE45FD56FDBE10D8" box="[393,528,691,717]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
: 35; shared with
<taxonomicName id="4C1D4D570030FFB5FD2BFD51FCF210D8" box="[743,860,692,717]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD2BFD51FCF210D8" box="[743,860,692,717]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FCA0FD56FBA210D8" authorityName="Cockerell" authorityYear="1908" box="[876,1036,691,717]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FCA0FD56FBA210D8" box="[876,1036,691,717]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
: 1;
<taxonomicName id="4C1D4D570030FFB5FBF7FD56FA8710D8" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[1083,1321,691,717]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FBF7FD56FA8710D8" box="[1083,1321,691,717]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FAF5FD51FF7410E4" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FAF5FD51FF7410E4" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
species: not known]; IR2 origin between RP1-2, RP3-4, closer to RP3-4, connected to RP1-2 notably more distally than to RP3-4 [other genera: origin close to or on RP1-2 (some
<taxonomicName id="4C1D4D570030FFB5FC19FD1EFBF21100" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[981,1116,763,789]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC19FD1EFBF21100" box="[981,1116,763,789]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
), or between RP1-2, RP3-4 as in
<taxonomicName id="4C1D4D570030FFB5FF15FCC5FEE1112C" box="[217,335,800,825]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF15FCC5FEE1112C" box="[217,335,800,825]" italics="true" pageId="34" pageNumber="35">Dysagrion</emphasis>
</taxonomicName>
(all others), but connected with RP1-2 similar level as RP3-4]; MA linear from quadrangle to zigzagged roughly mid-wing or further, linear again near, to margin [
<taxonomicName id="4C1D4D570030FFB5FC77FCA1FB9E1148" box="[955,1072,836,861]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC77FCA1FB9E1148" box="[955,1072,836,861]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
: similar (but differs by distinct curve beyond mid-wing);
<taxonomicName id="4C1D4D570030FFB5FE75FC82FDF71194" authorityName="Cockerell" authorityYear="1908" box="[441,601,871,897]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE75FC82FDF71194" box="[441,601,871,897]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
: very slightly zigzagged or not in mid-portion;
<taxonomicName id="4C1D4D570030FFB5FBA5FC82FAAE1194" box="[1129,1280,871,897]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FBA5FC82FAAE1194" box="[1129,1280,871,897]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FAC1FC82FA3A1194" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1293,1428,871,897]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FAC1FC82FA3A1194" box="[1293,1428,871,897]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
: not zigzagged;
<taxonomicName id="4C1D4D570030FFB5FE88FC6EFE7B11B0" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[324,469,907,933]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE88FC6EFE7B11B0" box="[324,469,907,933]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FE2FFC6EFD2C11B0" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[483,642,907,933]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE2FFC6EFD2C11B0" box="[483,642,907,933]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FD5CFC69FC8311B0" box="[656,813,908,933]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD5CFC69FC8311B0" box="[656,813,908,933]" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
species: linear to mid-wing, somewhat to strongly zigzagged to margin (in portions known); MA, MP close, 12 cells apart at margin [distinct from
<taxonomicName id="4C1D4D570030FFB5FB6BFC4AFA3B11DC" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[1191,1429,943,969]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FB6BFC4AFA3B11DC" box="[1191,1429,943,969]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FF5BFC31FEA211F8" box="[151,268,980,1005]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFC31FEA211F8" box="[151,268,980,1005]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FED1FC36FE1A11F8" box="[285,436,979,1005]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FED1FC36FE1A11F8" box="[285,436,979,1005]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FE09FC36FDE211F8" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[453,588,979,1005]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE09FC36FDE211F8" box="[453,588,979,1005]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FD92FC36FD5011F8" authorityName="Cockerell" authorityYear="1908" box="[606,766,979,1005]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD92FC36FD5011F8" box="[606,766,979,1005]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
: widely separated; not clearly distinct from
<taxonomicName id="4C1D4D570030FFB5FAC9FC36FA3811F8" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1285,1430,979,1005]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FAC9FC36FA3811F8" box="[1285,1430,979,1005]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FF5BFC12FE981604" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[151,310,1015,1041]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFC12FE981604" box="[151,310,1015,1041]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
, difficult to be sure of identity of MA near margin by strong zigzag, probably separated by two cells;
<taxonomicName id="4C1D4D570030FFB5FF5BFBF9FE9A1620" box="[151,308,1052,1077]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFBF9FE9A1620" box="[151,308,1052,1077]" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
species also difficult, perhaps one cell]; CuA increasingly zigzagged distal to level of nodus [distinct from
<taxonomicName id="4C1D4D570030FFB5FF1AFBA5FEE5164C" box="[214,331,1088,1113]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF1AFBA5FEE5164C" box="[214,331,1088,1113]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
: slightly zigzagged distal to about level of origin of RP2;
<taxonomicName id="4C1D4D570030FFB5FC28FBDAFB2A164C" authorityName="Cockerell" authorityYear="1908" box="[996,1156,1087,1113]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC28FBDAFB2A164C" box="[996,1156,1087,1113]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
: very slightly zigzagged in parts;
<taxonomicName id="4C1D4D570030FFB5FF30FB86FE441668" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[252,490,1123,1149]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF30FB86FE441668" box="[252,490,1123,1149]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FE36FB86FD3F1668" box="[506,657,1123,1149]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE36FB86FD3F1668" box="[506,657,1123,1149]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FD6CFB86FC891668" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[672,807,1123,1149]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD6CFB86FC891668" box="[672,807,1123,1149]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
: linear or only very slightly zigzagged; not clearly distinct from
<emphasis id="B969EAC60030FFB5FEDCFB62FE0316B4" box="[272,429,1159,1185]" italics="true" pageId="34" pageNumber="35">Okanoptyeryx</emphasis>
,
<taxonomicName id="4C1D4D570030FFB5FE70FB62FDF516B4" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[444,603,1159,1185]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE70FB62FDF516B4" box="[444,603,1159,1185]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
: zigzagged to various degrees distal to about mid-wing]; CuAA space 4 cells wide [separated from
<taxonomicName id="4C1D4D570030FFB5FE01FB49FDEC16D0" box="[461,578,1196,1221]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE01FB49FDEC16D0" box="[461,578,1196,1221]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
: 2;
<taxonomicName id="4C1D4D570030FFB5FDBCFB4EFCA116D0" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[624,783,1195,1221]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FDBCFB4EFCA116D0" box="[624,783,1195,1221]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
: 2 (most likely, wing incomplete);
<taxonomicName id="4C1D4D570030FFB5FB6FFB49FA3A16D0" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1187,1428,1195,1221]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="delinei">
<emphasis id="B969EAC60030FFB5FB6FFB49FA3A16D0" box="[1187,1428,1195,1221]" italics="true" pageId="34" pageNumber="35">Dysagrionites delinei</emphasis>
</taxonomicName>
: 3;
<taxonomicName id="4C1D4D570030FFB5FF79FB35FEFC16FC" box="[181,338,1232,1257]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF79FB35FEFC16FC" box="[181,338,1232,1257]" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
sp. A: 2;
<taxonomicName id="4C1D4D570030FFB5FE0FFB35FDCE16FC" box="[451,608,1232,1257]" class="Insecta" family="Megapodagrionidae" genus="Dysagrionites" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE0FFB35FDCE16FC" box="[451,608,1232,1257]" italics="true" pageId="34" pageNumber="35">Dysagrionites</emphasis>
</taxonomicName>
sp. B: probably 3 but unclear; not separated from
<taxonomicName id="4C1D4D570030FFB5FB6DFB2AFAEF16FC" authorityName="Cockerell" authorityYear="1908" box="[1185,1345,1231,1257]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FB6DFB2AFAEF16FC" box="[1185,1345,1231,1257]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
: 4;
<taxonomicName id="4C1D4D570030FFB5FAA3FB35FEA41718" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FAA3FB35FEA41718" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
: 36;
<taxonomicName id="4C1D4D570030FFB5FE9BFB16FE701718" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[343,478,1267,1293]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE9BFB16FE701718" box="[343,478,1267,1293]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
: 47;
<taxonomicName id="4C1D4D570030FFB5FDE7FB16FD121718" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[555,700,1267,1293]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FDE7FB16FD121718" box="[555,700,1267,1293]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
: 24; not distinct from
<taxonomicName id="4C1D4D570030FFB5FC1EFB16FB6E1718" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[978,1216,1267,1293]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC1EFB16FB6E1718" box="[978,1216,1267,1293]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
: 4 except
<quantity id="4CE59B310030FFB5FAF1FB16FAC41718" box="[1341,1386,1267,1294]" metricMagnitude="-1" metricUnit="m" metricValue="1.27" pageId="34" pageNumber="35" unit="in" value="5.0">5 in</quantity>
one column]; further from
<taxonomicName id="4C1D4D570030FFB5FE5BFAF2FD2B1724" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[407,645,1303,1329]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE5BFAF2FD2B1724" box="[407,645,1303,1329]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
by shorter petiole; further from
<taxonomicName id="4C1D4D570030FFB5FC3AFAF2FB231724" box="[1014,1165,1303,1329]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC3AFAF2FB231724" box="[1014,1165,1303,1329]" italics="true" pageId="34" pageNumber="35">Primorilestes</emphasis>
</taxonomicName>
by no linear intercalary veins in CuAA space. Postnodal, postsubnodal crossveins not aligned at least in proximal half [separated from
<taxonomicName id="4C1D4D570030FFB5FF5BFABAFE98176C" authorityName="Archibald &amp; Cannings" authorityYear="2021" box="[151,310,1375,1401]" class="Insecta" family="Dysagrionidae" genus="Stenodiafanus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFABAFE98176C" box="[151,310,1375,1401]" italics="true" pageId="34" pageNumber="35">Stenodiafanus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FE89FABAFE78176C" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[325,470,1375,1401]" class="Insecta" family="Thaumatoneuridae" genus="Okanopteryx" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FE89FABAFE78176C" box="[325,470,1375,1401]" italics="true" pageId="34" pageNumber="35">Okanopteryx</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FE28FABAFD72176C" box="[484,732,1375,1401]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="violetae">
<emphasis id="B969EAC60030FFB5FE28FABAFD72176C" box="[484,732,1375,1401]" italics="true" pageId="34" pageNumber="35">Primorilestes violetae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FD27FABAFC77176C" authorityName="Nel &amp; Arillo" authorityYear="2006" box="[747,985,1375,1401]" class="Insecta" family="Thaumatoneuridae" genus="Electrophenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FD27FABAFC77176C" box="[747,985,1375,1401]" italics="true" pageId="34" pageNumber="35">Electrophenacolestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FC24FA85FBF3176C" box="[1000,1117,1376,1401]" class="Insecta" family="Megapodagrionidae" genus="Furagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC24FA85FBF3176C" box="[1000,1117,1376,1401]" italics="true" pageId="34" pageNumber="35">Furagrion</emphasis>
</taxonomicName>
: all aligned or somewhat so in proximal half; not distinct from
<taxonomicName id="4C1D4D570030FFB5FDE3FA66FD611788" authorityName="Cockerell" authorityYear="1908" box="[559,719,1411,1437]" class="Insecta" family="Dysagrionidae" genus="Phenacolestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FDE3FA66FD611788" box="[559,719,1411,1437]" italics="true" pageId="34" pageNumber="35">Phenacolestes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FD2CFA66FC701788" box="[736,990,1411,1437]" class="Insecta" family="Thaumatoneuridae" genus="Primorilestes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="madseni">
<emphasis id="B969EAC60030FFB5FD2CFA66FC701788" box="[736,990,1411,1437]" italics="true" pageId="34" pageNumber="35">Primorilestes madseni</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570030FFB5FC3CFA66FBD91788" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1008,1143,1411,1437]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC3CFA66FBD91788" box="[1008,1143,1411,1437]" italics="true" pageId="34" pageNumber="35">Okanagrion</emphasis>
</taxonomicName>
: all not]. Separated from
<taxonomicName id="4C1D4D570030FFB5FF5BFA42FF4217D4" box="[151,236,1447,1473]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF5BFA42FF4217D4" box="[151,236,1447,1473]" italics="true" pageId="34" pageNumber="35">Valerea</emphasis>
</taxonomicName>
(tentative
<taxonomicName id="4C1D4D570030FFB5FEAFFA42FDA617D7" box="[355,520,1447,1474]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="subFamily" subFamily="Dysagrioninae">Dysagrioninae</taxonomicName>
) by no linear supplementary sector between RP1, IR1 [
<taxonomicName id="4C1D4D570030FFB5FBBFFA42FB6917D4" box="[1139,1223,1447,1473]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FBBFFA42FB6917D4" box="[1139,1223,1447,1473]" italics="true" pageId="34" pageNumber="35">Valerea</emphasis>
</taxonomicName>
: with such sector]; by one cell wide in costal space distal to pterostigma [
<taxonomicName id="4C1D4D570030FFB5FCC8FA2EFCF617F0" box="[772,856,1483,1509]" class="Insecta" genus="Valerea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FCC8FA2EFCF617F0" box="[772,856,1483,1509]" italics="true" pageId="34" pageNumber="35">Valerea</emphasis>
</taxonomicName>
: up to 5]. Most easily separated from other genera regarded here as possible
<taxonomicName id="4C1D4D570030FFB5FE78FA0AFDF9141C" box="[436,599,1519,1545]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
as follows: from the Chickaloon specimen (
<bibRefCitation id="EF8C4B250030FFB5FBF3FA0AFA94141C" author="Garrouste, R. &amp; Nel, A." box="[1087,1338,1519,1545]" pageId="34" pageNumber="35" pagination="1939 - 1946" refId="ref57867" refString="Garrouste, R. &amp; Nel, A. (2019) Alaskan Palaeogene insects: a challenge for a better knowledge of the Beringian ' route' (Odonata: Aeshnidae, Dysagrionidae). Journal of Systematic Palaeontology, 17, 1939 - 1946. https: // doi. org / 10.1080 / 14772019.2019.1572235" type="journal article" year="2019">Garrouste &amp; Nel 2019</bibRefCitation>
) by CuA increasingly zigzagged from level of nodus [linear at least to well past level of origin of IR1 (distal-most portion not preserved)]; MA, MP close [widening in distal-most preserved portion]; from
<taxonomicName id="4C1D4D570030FFB5FBDDF9D2FB7B1444" box="[1041,1237,1591,1617]" class="Insecta" genus="Thanetophilosina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FBDDF9D2FB7B1444" box="[1041,1237,1591,1617]" italics="true" pageId="34" pageNumber="35">Thanetophilosina</emphasis>
</taxonomicName>
by proximal side of quadrangle more than half length of distal side [less than half], IR2 originates between RP1-2, RP3-4 [originates on RP1-2]; from NHMUK I.9866/I.9718 by origin of RP3-4 [distinctly closer to subnodus]; from
<emphasis id="B969EAC60030FFB5FB1BF99AFB45148C" box="[1239,1259,1663,1689]" italics="true" pageId="34" pageNumber="35">cf</emphasis>
.
<taxonomicName id="4C1D4D570030FFB5FB35F99AFA32148C" box="[1273,1436,1663,1689]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
genus A, species A (
<bibRefCitation id="EF8C4B250030FFB5FEBBF946FD2514A8" author="Petrulevicius, J. F. &amp; Wappler, T. &amp; Wedmann, S. &amp; Rust, J. &amp; Nel, A." box="[375,651,1699,1726]" pageId="34" pageNumber="35" pagination="1173 - 1181" refId="ref61019" refString="Petrulevicius, J. F., Wappler, T., Wedmann, S., Rust, J. &amp; Nel, A. (2008) New Megapodagrionid Damselflies (Odonata: Zygop- tera) from the Paleogene of Europe. Journal of Paleontology, 82, 1173 - 1181. https: // doi. org / 10.1666 / 07 - 091.1" type="journal article" year="2008">
Petrulevičius
<emphasis id="B969EAC60030FFB5FDDEF941FDED14A8" box="[530,579,1699,1725]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2008
</bibRefCitation>
) by hyaline membrane [preserved portion infuscate except apex distal to pterostigma], by MA zigzagged mid-wing, CuA zigzagged distally [both linear to margin].
</paragraph>
</subSubSection>
<subSubSection id="C307655F0030FFB3FF0BF90EFF4B12EC" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="discussion">
<paragraph id="8BA236D40030FFB5FF0BF90EFE5B1580" blockId="34.[151,1437,403,2014]" pageId="34" pageNumber="35">
<emphasis id="B969EAC60030FFB5FF0BF90EFEE21510" bold="true" box="[199,332,1771,1797]" pageId="34" pageNumber="35">Discussion.</emphasis>
<taxonomicName id="4C1D4D570030FFB5FE9AF909FD901510" authorityName="Scudder" authorityYear="1878" box="[342,574,1771,1797]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="fredericii">
<emphasis id="B969EAC60030FFB5FE9AF909FD901510" box="[342,574,1771,1797]" italics="true" pageId="34" pageNumber="35">Dysagrion fredericii</emphasis>
</taxonomicName>
was designated the
<typeStatus id="54A688760030FFB5FCE0F909FCF21513" box="[812,860,1772,1798]" pageId="34" pageNumber="35">type</typeStatus>
species of
<taxonomicName id="4C1D4D570030FFB5FC2FF909FBF71510" box="[995,1113,1772,1797]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FC2FF909FBF71510" box="[995,1113,1772,1797]" italics="true" pageId="34" pageNumber="35">Dysagrion</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF8C4B250030FFB5FB46F90EFA911510" author="Scudder, S. H." box="[1162,1343,1771,1797]" pageId="34" pageNumber="35" pagination="519 - 543" refId="ref61543" refString="Scudder, S. H. (1878) An Account of some insects of unusual interest from the Tertiary rocks of Colorado and Wyoming. Bulletin of the United States Geological and Geographical Survey of the Territories, 4, 519 - 543." type="journal article" year="1878">Scudder (1878)</bibRefCitation>
. His diagnosis of the genus distinguished it from
<taxonomicName id="4C1D4D570030FFB5FDB6F8EAFC92153C" authority="Selys" authorityName="Selys" authorityYear="1862" box="[634,828,1807,1833]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FDB6F8EAFD5B153C" box="[634,757,1807,1833]" italics="true" pageId="34" pageNumber="35">Philogenia</emphasis>
Selys
</taxonomicName>
, a genus currently in need of revision (
<bibRefCitation id="EF8C4B250030FFB5FB3BF8EAFF7A1558" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." pageId="34" pageNumber="35" refId="ref57766" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly genera of the New World: an illustrated and annotated key to the Zygoptera. Johns Hopkins University Press, Baltimore, 490 pp." type="book" year="2010">
Garrison
<emphasis id="B969EAC60030FFB5FAAFF8F5FA3B153C" box="[1379,1429,1807,1833]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2010
</bibRefCitation>
).
<taxonomicName id="4C1D4D570030FFB5FF2AF8D6FECF1558" authorityName="Selys" authorityYear="1862" box="[230,353,1843,1869]" class="Insecta" family="Megapodagrionidae" genus="Philogenia" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FF2AF8D6FECF1558" box="[230,353,1843,1869]" italics="true" pageId="34" pageNumber="35">Philogenia</emphasis>
</taxonomicName>
had been placed in the polyphyletic
<taxonomicName id="4C1D4D570030FFB5FD3BF8D6FC741558" box="[759,986,1843,1869]" class="Insecta" family="Megapodagrionidae" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Megapodagrionidae</taxonomicName>
, which was recently divided into several families (
<bibRefCitation id="EF8C4B250030FFB5FF32F8B2FE771564" author="Dijkstra, K. - D. B. &amp; Kalkman, V. J. &amp; Dow, R. A. &amp; Stokvis, F. R. &amp; van Tol, J." box="[254,473,1879,1906]" pageId="34" pageNumber="35" pagination="68 - 96" refId="ref57269" refString="Dijkstra, K. - D. B., Kalkman, V. J., Dow, R. A., Stokvis, F. R. &amp; van Tol, J. (2014) Redefining the damselfly families: the first comprehensive molecular phylogeny of Zygoptera (Odonata). Systematic Entomology, 39, 68 - 96. https: // doi. org / 10.1111 / syen. 12035" type="journal article" year="2014">
Dijkstra
<emphasis id="B969EAC60030FFB5FEACF8BDFE3E1564" box="[352,400,1879,1905]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2014
</bibRefCitation>
), including the
<taxonomicName id="4C1D4D570030FFB5FD4EF8B2FCB11564" box="[642,799,1879,1905]" class="Insecta" family="Philogeniidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="family">Philogeniidae</taxonomicName>
, where it now resides. We, therefore, provide an emended diagnosis of
<taxonomicName id="4C1D4D570030FFB5FEEAF899FE321580" box="[294,412,1916,1941]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60030FFB5FEEAF899FE321580" box="[294,412,1916,1941]" italics="true" pageId="34" pageNumber="35">Dysagrion</emphasis>
</taxonomicName>
, above.
</paragraph>
<paragraph id="8BA236D40030FFB4FF0BF87AFD2912C0" blockId="34.[151,1437,403,2014]" lastBlockId="35.[151,1436,151,213]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
An oblique brace vein between RA and RP1 at the posterior-proximal corner of pterostigma is seen in the fore- and hind wings of
<taxonomicName id="4C1D4D570030FFB5FEA4F821FE4915C8" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[360,487,1988,2013]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="pruettae">
<emphasis id="B969EAC60030FFB5FEA4F821FE4915C8" box="[360,487,1988,2013]" italics="true" pageId="34" pageNumber="35">D. pruettae</emphasis>
</taxonomicName>
(
<figureCitation id="13262A510030FFB5FE3BF826FDFA15C8" box="[503,596,1987,2013]" captionStart="FIGURE 17" captionStartId="38.[151,250,1971,1995]" captionTargetBox="[332,1253,184,1939]" captionTargetId="figure-19@38.[319,1267,181,1947]" captionTargetPageId="38" captionText="FIGURE 17. Dysagrion pruettae holotype, SR 13-005-012.A, drawing of forewing (FW); B, drawing of hind wing (HW). FW, HW label colours refer to those in Fig. 16. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558840" httpUri="https://zenodo.org/record/4558840/files/figure.png" pageId="34" pageNumber="35">Figs. 17</figureCitation>
,
<figureCitation id="13262A510030FFB5FDACF826FDD015C8" box="[608,638,1987,2013]" captionStart="FIGURE 18" captionStartId="39.[151,250,1943,1967]" captionTargetBox="[433,1156,191,1908]" captionTargetId="figure-17@39.[418,1170,181,1919]" captionTargetPageId="39" captionText="FIGURE 18. Dysagrion pruettae holotype, SR 13-005-012.A, drawing of forewing (FW); B, drawing of hind wing (HW). FW, HW label colours refer to those in Fig. 16. Small, blue arrows indicate supplementary sectors. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558842" httpUri="https://zenodo.org/record/4558842/files/figure.png" pageId="34" pageNumber="35">18</figureCitation>
), and in the wings of
<taxonomicName id="4C1D4D570030FFB5FCBEF821FC7315C8" authorityName="Scudder" authorityYear="1890" box="[882,989,1987,2013]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="lakesii">
<emphasis id="B969EAC60030FFB5FCBEF821FC7315C8" box="[882,989,1987,2013]" italics="true" pageId="34" pageNumber="35">D. lakesii</emphasis>
</taxonomicName>
(
<figureCitation id="13262A510030FFB5FC21F826FB9F15CB" box="[1005,1073,1987,2014]" captionStart="FIGURE 5" captionStartId="12.[151,250,1943,1967]" captionTargetBox="[250,1324,200,1919]" captionTargetId="figure-19@12.[250,1338,181,1919]" captionTargetPageId="12" captionText="FIGURE 5. Dysagrion lakesii Scudder type specimen MCZ 4101 (Packard number 259, not previously illustrated): A, photograph; B, drawing from a photograph supplied by MCZ personnel. Both to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558807" httpUri="https://zenodo.org/record/4558807/files/figure.png" pageId="34" pageNumber="35">Fig. 5</figureCitation>
, slightly, but distinctly oblique),
<taxonomicName id="4C1D4D570031FFB4FF5BFF72FE8E12A4" authorityName="Scudder" authorityYear="1878" box="[151,288,151,177]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="fredericii">
<emphasis id="B969EAC60031FFB4FF5BFF72FF0512A5" box="[151,171,151,176]" italics="true" pageId="35" pageNumber="36">D</emphasis>
.
<emphasis id="B969EAC60031FFB4FF75FF72FE8E12A4" box="[185,288,151,177]" italics="true" pageId="35" pageNumber="36">fredericii</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF8C4B250031FFB4FE97FF72FE5812A4" author="Scudder, S. H." box="[347,502,151,177]" pageId="35" pageNumber="36" pagination="1 - 734" refId="ref61619" refString="Scudder, S. H. (1890) The Tertiary insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734, 28 pls. https: // doi. org / 10.5962 / bhl. title. 44698" type="journal article" year="1890">Scudder 1890</bibRefCitation>
plate 6
<figureCitation id="13262A510031FFB4FD82FF72FD3C12A4" box="[590,658,151,177]" captionStart="FIGURE 9" captionStartId="18.[151,250,1817,1841]" captionTargetBox="[210,1368,188,1783]" captionTargetId="figure-19@18.[206,1381,181,1793]" captionTargetPageId="18" captionText="FIGURE 9. Wing drawings of: Valerea multicellulata redrawn from Fig. 1 of Garrouste et al. (2017); Chickaloon Formation (Alaska) specimen redrawn from Fig. 3 of Garrouste &amp; Nel (2017); cf. Dysagrionidae gen. et. sp. A redrawn from Fig. 4 of Petrulevičius et al. (2008) (there as “Megapodagrionidae gen. et. sp. A”); Thanetophilosina menatensis redrawn from Fig. 21 of Nel et al. (1997); NHMUK I.9866 (Isle of Wight) redrawn from Fig. 28 of Nel &amp; Fleck (2014); Palaeodysagrion youlini redrawn from Fig. 2 of Zheng et al. (2018); Electrodysagrion neli redrawn from Fig. 2 of Zheng et al. (2019); Burmadysagrion zhangi redrawn from Fig. 3 of Zheng et al. (2016a). To associated scales, all 5 mm. Asterisks indicate quadrangles." figureDoi="http://doi.org/10.5281/zenodo.4558818" httpUri="https://zenodo.org/record/4558818/files/figure.png" pageId="35" pageNumber="36">Fig. 9</figureCitation>
),
<taxonomicName id="4C1D4D570031FFB4FD6BFF72FC9D12A4" authorityName="Scudder Lutetian Grube Messel" authorityYear="1885" box="[679,819,151,177]" class="Insecta" family="Dysagrionidae" genus="Dysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="packardii">
<emphasis id="B969EAC60031FFB4FD6BFF72FC9D12A4" box="[679,819,151,177]" italics="true" pageId="35" pageNumber="36">D. packardii</emphasis>
</taxonomicName>
(a portion preserved, but distinct:
<figureCitation id="13262A510031FFB4FB78FF72FB5912A4" box="[1204,1271,151,177]" captionStart="FIGURE 4" captionStartId="11.[151,250,1923,1947]" captionTargetBox="[343,1242,181,1891]" captionTargetId="figure-17@11.[332,1255,181,1899]" captionTargetPageId="11" captionText="FIGURE 4. Dysagrion packardii cotype MCZ 4656 (Packard number 252): A, drawing; B, photograph; C, holotype (Packard number 146) redrawn from Zittel (1885: his Fig. 979), also redrawn by Scudder (1890: his plate 6, Fig. 3). A, B drawn from a photograph supplied by MCZ personnel. All to scale, 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558805" httpUri="https://zenodo.org/record/4558805/files/figure.png" pageId="35" pageNumber="36">Fig. 4</figureCitation>
, redrawn from the original by
<bibRefCitation id="EF8C4B250031FFB4FE8EFF5EFE7212C0" author="Scudder, S. H." box="[322,476,187,213]" pageId="35" pageNumber="36" pagination="1 - 734" refId="ref61619" refString="Scudder, S. H. (1890) The Tertiary insects of North America. Report of the United States Geological Survey of the Territories, 13, 1 - 734, 28 pls. https: // doi. org / 10.5962 / bhl. title. 44698" type="journal article" year="1890">Scudder 1890</bibRefCitation>
plate 6
<figureCitation id="13262A510031FFB4FDFAFF5EFDD712C0" box="[566,633,187,213]" captionStart="FIGURE 3" captionStartId="10.[151,250,1805,1829]" captionTargetBox="[151,1436,181,1781]" captionTargetId="figure-19@10.[151,1436,181,1781]" captionTargetPageId="10" captionText="FIGURE 3. Phenacolestes parallelus. A, specimen YPMIP-220974 of the Yale Peabody Museum; B, drawing from A, photograph provided by Peabody Museum personnel; C, holotype, UCM-4503 of the University of Colorado (Boulder) paleontology collections; D, drawing of C, photograph provided by University of Colorado personnel; E, UCM-4545, University of Colorado (Boulder). AD to scale, E to scale, both scale bars 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4558803" httpUri="https://zenodo.org/record/4558803/files/figure.png" pageId="35" pageNumber="36">Fig. 3</figureCitation>
).
</paragraph>
<caption id="DF62665C0031FFB4FF5BFB4CFAF4171C" ID-DOI="http://doi.org/10.5281/zenodo.4558832" ID-Zenodo-Dep="4558832" httpUri="https://zenodo.org/record/4558832/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1193,1217]" targetBox="[156,1427,245,1166]" targetPageId="35">
<paragraph id="8BA236D40031FFB4FF5BFB4CFAF4171C" blockId="35.[151,1437,1193,1290]" pageId="35" pageNumber="36">
<emphasis id="B969EAC60031FFB4FF5BFB4CFE8C16D4" bold="true" box="[151,290,1193,1217]" pageId="35" pageNumber="36">FIGURE 14.</emphasis>
Tree resulting from parsimony-based analyses for Analysis 1:
<taxonomicName id="4C1D4D570031FFB4FC7AFB4CFBF416D4" box="[950,1114,1193,1217]" class="Insecta" family="Tarsophlebiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Tarsophlebiidae</taxonomicName>
, the suborders of
<taxonomicName id="4C1D4D570031FFB4FAD5FB4CFAC116D4" box="[1305,1391,1193,1217]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="order">Odonata</taxonomicName>
and former members of
<taxonomicName id="4C1D4D570031FFB4FEA9FB28FE5716F0" box="[357,505,1229,1253]" class="Insecta" family="Dysagrionidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="family">Dysagrionidae</taxonomicName>
(
<taxonomicName id="4C1D4D570031FFB4FDC5FB28FD1D16F0" box="[521,691,1229,1253]" class="Insecta" family="Dysagrionidae" genus="Burmadysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60031FFB4FDC5FB28FD1D16F0" box="[521,691,1229,1253]" italics="true" pageId="35" pageNumber="36">Burmadysagrion</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D570031FFB4FD0EFB28FBB116F0" authority=", Paleodysagrion" authorityName="Paleodysagrion" box="[706,1055,1229,1253]" class="Insecta" family="Dysagrionidae" genus="Electrodysagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B969EAC60031FFB4FD0EFB28FCDF16F0" box="[706,881,1229,1253]" italics="true" pageId="35" pageNumber="36">Electrodysagrion</emphasis>
,
<emphasis id="B969EAC60031FFB4FCB3FB28FBB116F0" box="[895,1055,1229,1253]" italics="true" pageId="35" pageNumber="36">Paleodysagrion</emphasis>
</taxonomicName>
). One single most-parsimonious tree (MPT) based on 23 continuous and 79 discrete characters. Symmetrical resampling values Ξ70 are labelled at the nodes.
</paragraph>
</caption>
<caption id="DF62665C0031FFB4FF5BF86CFD7F15FC" ID-DOI="http://doi.org/10.5281/zenodo.4558834" ID-Zenodo-Dep="4558834" httpUri="https://zenodo.org/record/4558834/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1929,1953]" targetBox="[161,1428,1323,1898]" targetPageId="35">
<paragraph id="8BA236D40031FFB4FF5BF86CFD7F15FC" blockId="35.[151,1436,1929,2025]" pageId="35" pageNumber="36">
<emphasis id="B969EAC60031FFB4FF5BF86CFE8E15B4" bold="true" box="[151,288,1929,1953]" pageId="35" pageNumber="36">FIGURE 15.</emphasis>
Tree resulting from parsimony-based analyses for Analysis 2 (relationship
<taxonomicName id="4C1D4D570031FFB4FBEDF86CFB7B15B4" authority="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes, 2021" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[1057,1237,1929,1953]" family="Whetwhetaksidae" kingdom="Animalia" order="Odonata" pageId="35" pageNumber="28" rank="family" status="fam. nov.">Whetwhetaksidae</taxonomicName>
to confirmed families of
<taxonomicName id="4C1D4D570031FFB4FF10F848FE3715D0" box="[220,409,1965,1989]" kingdom="Animalia" pageId="35" pageNumber="27" rank="subOrder" subOrder="Cephalozygoptera">Cephalozygoptera</taxonomicName>
). One single most-parsimonious tree (MPT) based on 23 continuous and 79 discrete characters. Symmetrical resampling values Ξ70 are labeled at the nodes.
</paragraph>
</caption>
<paragraph id="8BA236D40036FFB3FF0BFF72FF4B12EC" blockId="36.[151,1436,151,249]" pageId="36" pageNumber="37">
The subnodus appears to be of reverse obliquity in one specimen of
<taxonomicName id="4C1D4D570036FFB3FC0BFF72FB0A12A4" authorityName="Archibald &amp; Cannings &amp; Erickson &amp; Bybee &amp; Mathewes" authorityYear="2021" box="[967,1188,151,177]" class="Insecta" family="Dysagrionidae" genus="Okanagrion" higherTaxonomySource="GBIF" kingdom="Animalia" order="Odonata" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="hobani">
<emphasis id="B969EAC60036FFB3FC0BFF72FB0A12A4" box="[967,1188,151,177]" italics="true" pageId="36" pageNumber="37">Okanagrion hobani</emphasis>
</taxonomicName>
,
<typeStatus id="54A688760036FFB3FB7EFF72FABC12A4" box="[1202,1298,151,177]" pageId="36" pageNumber="37" type="paratype">paratype</typeStatus>
1, however, given that all other of its numerous specimens have normal obliquity, we consider this adventitious in this individual.
</paragraph>
</subSubSection>
</treatment>
</document>