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<document id="F39A486C4E966336EF10FC58CF9C6121" ID-CLB-Dataset="168932" ID-DOI="10.5852/ejt.2022.844.1949" ID-GBIF-Dataset="3a08058c-14f8-4232-ad02-65ed5b28380f" ID-ISSN="2118-9773" ID-Zenodo-Dep="7222155" ID-ZooBank="193EDD91-B24D-455C-B8AA-8133586A00A1" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666126129383" checkinUser="felipe" docAuthor="Grzelak, Katarzyna &amp; Sørensen, Martin V." docDate="2022" docId="03B487885B092A3CFDDE3EB06147F9B7" docLanguage="en" docName="ejt-844_1-108.pdf" docOrigin="European Journal of Taxonomy 844" docSource="http://dx.doi.org/10.5852/ejt.2022.844.1949" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Echinoderes gandalfi Grzelak &amp; Sørensen 2022, sp. nov." docType="treatment" docUuid="84D3AEAA-391B-4C5C-81FC-380B198251F2" docUuidSource="ZooBank" docVersion="13" lastPageNumber="45" masterDocId="FF8DFFF05B2C2A10FFB63A55654FFFB8" masterDocTitle="Echinoderes (Kinorhyncha: Cyclorhagida) from the Hikurangi Margin, New Zealand" masterLastPageNumber="108" masterPageNumber="1" pageNumber="38" updateTime="1699407836451" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C1FA87DDD4BE3920D42541B419717E99">Echinoderes (Kinorhyncha: Cyclorhagida) from the Hikurangi Margin, New Zealand</mods:title>
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<mods:namePart id="12D8B807FC940E28F6E9BD80B74C1003">Grzelak, Katarzyna</mods:namePart>
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<mods:affiliation id="836578521A8E4AF79469619DA4C0C57D">Polish Academy of Sciences, Institute of Oceanology, Sopot, Poland. &amp; Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:namePart id="92DE4D2702B0521A60DF6B22A9919091">Sørensen, Martin V.</mods:namePart>
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<mods:affiliation id="B25999A097F4044EEF570DD08E4B7DC8">Natural History Museum of Denmark, University of Copenhagen, DK- 2100 Copenhagen, Denmark.</mods:affiliation>
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<mods:date id="E3CFDDD7202D0AD8A390637891141897">2022</mods:date>
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<subSubSection id="C30765155B092A35FDDE3EB06684FAB8" box="[616,971,1253,1280]" pageId="37" pageNumber="38" type="nomenclature">
<paragraph id="8BA2369E5B092A35FDDE3EB06684FAB8" blockId="37.[364,1223,1253,1353]" box="[616,971,1253,1280]" pageId="37" pageNumber="38">
<heading id="D0EA81F25B092A35FDDE3EB06684FAB8" box="[616,971,1253,1280]" centered="true" fontSize="11" level="2" pageId="37" pageNumber="38" reason="2">
<taxonomicName id="4C1D4D1D5B092A35FDDE3EB06625FAB8" ID-CoL="948d8539-132a-4440-886b-9384a8faa3b0" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[616,874,1253,1280]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B092A35FDDE3EB06625FAB8" bold="true" box="[616,874,1253,1280]" italics="true" pageId="37" pageNumber="38">Echinoderes gandalfi</emphasis>
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<taxonomicNameLabel id="A25A57F75B092A35FCC73EB36684FAB8" box="[881,971,1254,1280]" pageId="37" pageNumber="38" rank="species">sp. nov.</taxonomicNameLabel>
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urn:lsid:zoobank.org:act:
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</uri>
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<figureCitation id="13262A1B5B092A35FD353F7B6645FAF0" box="[643,778,1326,1352]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="39.[189,232,1927,1953]" captionStartId-1="40.[189,232,1857,1883]" captionStartId-2="41.[189,232,1793,1819]" captionTargetBox-0="[212,1377,269,1898]" captionTargetBox-1="[190,1398,766,1824]" captionTargetBox-2="[196,1392,267,1742]" captionTargetId-0="figure-60@39.[211,1377,265,1898]" captionTargetId-1="figure-235@40.[189,1399,766,1825]" captionTargetId-2="figure-0@41.[192,1402,265,1743]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionText-0="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." captionText-1="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." captionText-2="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.7222187" figureDoi-1="http://doi.org/10.5281/zenodo.7222189" figureDoi-2="http://doi.org/10.5281/zenodo.7222191" httpUri-0="https://zenodo.org/record/7222187/files/figure.png" httpUri-1="https://zenodo.org/record/7222189/files/figure.png" httpUri-2="https://zenodo.org/record/7222191/files/figure.png" pageId="37" pageNumber="38">Figs 1416</figureCitation>
;
<tableCitation id="C69F03255B092A35FCA33F7B66FFFAF1" box="[789,944,1326,1353]" captionStart-0="Table 10" captionStart-1="Table 11" captionStartId-0="42.[188,257,265,291]" captionStartId-1="43.[189,258,265,291]" captionTargetPageId-0="42" captionTargetPageId-1="43" captionText-0="Table 10. Measurements from light microscopy of Echinoderes gandalfi sp. nov. (in µm) from the Hikurangi Margin, including number of measured specimens (n) and standard deviation (SD)." captionText-1="Table 11. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes gandalfi sp. nov." pageId="37" pageNumber="38">Tables 1011</tableCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30765155B092A35FF0B3F2467F2F9F5" pageId="37" pageNumber="38" type="diagnosis">
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<heading id="D0EA81F25B092A35FF0B3F246479FA33" bold="true" box="[189,310,1393,1419]" fontSize="11" level="3" pageId="37" pageNumber="38" reason="3">
<emphasis id="B969EA8C5B092A35FF0B3F246479FA33" bold="true" box="[189,310,1393,1419]" pageId="37" pageNumber="38">Diagnosis</emphasis>
</heading>
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<taxonomicName id="4C1D4D1D5B092A35FF0B3FF46400FA03" authorityName="Claparede" authorityYear="1863" box="[189,335,1441,1467]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="37" pageNumber="38" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B092A35FF0B3FF46400FA03" box="[189,335,1441,1467]" italics="true" pageId="37" pageNumber="38">Echinoderes</emphasis>
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with spines in middorsal position on segments 6 and 8, and spines in lateroventral positions on segments 6 to 9. Tubes present in ventrolateral positions on segment 2, lateroventral positions on segment 5, lateral accessory positions on segment 8, and laterodorsal positions on segments 9 and 10. A protuberance-like structure emerges between segments 10 and
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middorsal position. Lateral terminal spines long, always exceeding trunk length.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B092A35FF0B3C236726F8B0" pageId="37" pageNumber="38" type="etymology">
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<emphasis id="B969EA8C5B092A35FF0B3C23640BF928" bold="true" box="[189,324,1654,1680]" pageId="37" pageNumber="38">Etymology</emphasis>
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The species name refers to Gandalf the Grey, one of the main characters in the novel “
<emphasis id="B969EA8C5B092A35FB123CF06450F95C" italics="true" pageId="37" pageNumber="38">The Fellowship of the Ring</emphasis>
”, the first volume of J.R.R. Tolkiens “
<emphasis id="B969EA8C5B092A35FD5B3C9F66A2F95C" box="[749,1005,1738,1764]" italics="true" pageId="37" pageNumber="38">The Lord of the Rings</emphasis>
”. He helped form the Fellowship of the Ring to destroy the One Ring.
</paragraph>
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<emphasis id="B969EA8C5B092A35FF0B3D6464E5F8F3" bold="true" box="[189,426,1841,1867]" pageId="37" pageNumber="38">Material examined</emphasis>
</heading>
</paragraph>
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</emphasis>
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<collectingCountry id="F30A760E5B092A35FF0B3DD364DDF818" box="[189,402,1926,1952]" name="New Zealand" pageId="37" pageNumber="38">NEW ZEALAND</collectingCountry>
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;
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, stn TAN1004/31;
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,
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;
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b.s.l.
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;
<date id="FFA3105E5B092A35FAF53DD365B4F87C" pageId="37" pageNumber="38" value="2010-04">
<collectingDate id="EFE7E9B65B092A35FAF53DD365B4F87C" pageId="37" pageNumber="38" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCode id="DBBB9EE55B092A35FD5B3DFF66D0F87D" box="[749,927,1962,1989]" pageId="37" pageNumber="38">NIWA-159407</specimenCode>
. Mounted for LM in Fluoromount G on HS slide.
</paragraph>
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<materialsCitation id="3B753CC35B0A2A36FF5C3B5C6428FE9B" ID-GBIF-Occurrence="3926368302" box="[234,359,265,291]" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="730" latitude="-41.4962" location="NEW ZEALAND" longLatPrecision="6" longitude="175.6828" pageId="38" pageNumber="39" specimenCode="NHMD-916356" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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<typeStatus id="54A6883C5B0A2A36FF5C3B5C6428FE9B" box="[234,359,265,291]" pageId="38" pageNumber="39" type="paratype">Paratypes</typeStatus>
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<collectingCountry id="F30A760E5B0A2A36FF0B3B7B64DEFEFF" box="[189,401,301,328]" name="New Zealand" pageId="38" pageNumber="39">NEW ZEALAND</collectingCountry>
• 2 ♀♀; same collection data as for
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;
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,
<materialsCitation id="3B753CC35B0A2A36FB153B7B64FAFED3" ID-GBIF-Occurrence="3926368310" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="730" latitude="-41.4962" location="Pahaua Canyon" longLatPrecision="6" longitude="175.6828" pageId="38" pageNumber="39" specimenCode="NIWA-159408" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FB153B7B61F6FEF0" box="[1187,1209,302,328]" pageId="38" pageNumber="39" type="female"></specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FB773B78603CFEF0" box="[1217,1395,301,328]" pageId="38" pageNumber="39">NIWA-159408</specimenCode>
. Mounted as holotype
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FE783B076058FE28" ID-GBIF-Occurrence="3926364377" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1350" latitude="-41.5508" location="Pahaua Canyon" longLatPrecision="6" longitude="175.725" municipality="Pahaua Canyon" pageId="38" pageNumber="39" specimenCode="NIWA-159409" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FE783B0764B4FED4" box="[462,507,338,364]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
;
<collectingMunicipality id="6BC6ACE45B0A2A36FDBF3B04678CFED4" box="[521,707,337,364]" pageId="38" pageNumber="39">Pahaua Canyon</collectingMunicipality>
, stn TAN1004/12;
<geoCoordinate id="EE2950595B0A2A36FC143B046169FED4" box="[930,1062,337,364]" degrees="41.5508" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-41.5508">41.5508° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FB853B046186FED4" box="[1075,1225,337,364]" degrees="175.7250" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="175.725">175.7250° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FB603B04603FFED3" box="[1238,1392,337,364]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="38" pageNumber="39" unit="m" value="1350.0">
<quantity id="4CE59B7B5B0A2A36FB603B046060FED4" box="[1238,1327,337,364]" metricMagnitude="3" metricUnit="m" metricValue="1.35" pageId="38" pageNumber="39" unit="m" value="1350.0">1350 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FF0B3B236473FE28" box="[189,316,374,400]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FF0B3B236473FE28" box="[189,316,374,400]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCode id="DBBB9EE55B0A2A36FCF93B23614DFE29" box="[847,1026,374,401]" pageId="38" pageNumber="39">NIWA-159409</specimenCode>
. Mounted as holotype
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FA8B3B236675FE62" ID-GBIF-Occurrence="3926368308" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1188" latitude="-41.51" location="Pahaua Canyon" longLatPrecision="6" longitude="175.7187" pageId="38" pageNumber="39" specimenCode="NHMD-916362" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FA8B3B236020FE29" box="[1341,1391,374,401]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
, 1 ♂;
<location id="8EC260455B0A2A36FEB73BCE64F0FE0D" LSID="urn:lsid:plazi:treatment:03B487885B092A3CFDDE3EB06147F9B7:8EC260455B0A2A36FEB73BCE64F0FE0D" box="[257,447,411,437]" country="New Zealand" latitude="-41.51" longLatPrecision="6" longitude="175.7187" name="Pahaua Canyon" pageId="38" pageNumber="39">Pahaua Canyon</location>
, stn TAN1004/22;
<geoCoordinate id="EE2950595B0A2A36FD1F3BCF667DFE0D" box="[681,818,410,437]" degrees="41.5100" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-41.51">41.5100° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FCF53BCF6692FE0D" box="[835,989,410,437]" degrees="175.7187" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="175.7187">175.7187° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FC5B3BCE61C0FE0D" box="[1005,1167,411,437]" metricMagnitude="3" metricUnit="m" metricValue="1.188" pageId="38" pageNumber="39" unit="m" value="1188.0">
<quantity id="4CE59B7B5B0A2A36FC5B3BCE6106FE0D" box="[1005,1097,411,437]" metricMagnitude="3" metricUnit="m" metricValue="1.188" pageId="38" pageNumber="39" unit="m" value="1188.0">1188 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FB293BCE6054FE0D" box="[1183,1307,411,437]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FB293BCE6054FE0D" box="[1183,1307,411,437]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCount id="9D1BFD175B0A2A36FDE13B956722FE62" box="[599,621,448,474]" pageId="38" pageNumber="39" type="female"></specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FDC13BEA667DFE61" box="[631,818,447,473]" pageId="38" pageNumber="39">NHMD-916362</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FCF23B95606EFE61" ID-GBIF-Occurrence="3926368304" box="[836,1313,447,474]" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1188" latitude="-41.51" longLatPrecision="6" longitude="175.7187" pageId="38" pageNumber="39" specimenCode="NIWA-159410" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FCF23B956615FE62" box="[836,858,448,474]" pageId="38" pageNumber="39" type="male"></specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FCD23BEA6159FE62" box="[868,1046,447,474]" pageId="38" pageNumber="39">NIWA-159410</specimenCode>
. Mounted as holotype
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FA893BEA66F2FD9A" ID-GBIF-Occurrence="3926364358" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1013" latitude="-41.4983" location="Pahaua Canyon" longLatPrecision="6" longitude="175.7043" pageId="38" pageNumber="39" specimenCode="NIWA-159411" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FA893BEA603EFE62" box="[1343,1393,447,474]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
;
<location id="8EC260455B0A2A36FF0B3BB16438FE46" LSID="urn:lsid:plazi:treatment:03B487885B092A3CFDDE3EB06147F9B7:8EC260455B0A2A36FF0B3BB16438FE46" box="[189,375,484,510]" country="New Zealand" latitude="-41.4983" longLatPrecision="6" longitude="175.7043" name="Pahaua Canyon" pageId="38" pageNumber="39">Pahaua Canyon</location>
, stn TAN1004/27;
<geoCoordinate id="EE2950595B0A2A36FDE23BB16797FE46" box="[596,728,484,510]" degrees="41.4983" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-41.4983">41.4983° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FD533BB16634FE46" box="[741,891,484,511]" degrees="175.7043" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="175.7043">175.7043° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FC313BB1616EFE46" box="[903,1057,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="38" pageNumber="39" unit="m" value="1013.0">
<quantity id="4CE59B7B5B0A2A36FC313BB166AFFE46" box="[903,992,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="38" pageNumber="39" unit="m" value="1013.0">1013 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FB9A3BB161EAFE46" box="[1068,1189,484,510]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FB9A3BB161EAFE46" box="[1068,1189,484,510]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCount id="9D1BFD175B0A2A36FE6A385C64BDFD9B" box="[476,498,521,547]" pageId="38" pageNumber="39" type="male"></specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FE48385D67E1FD9B" box="[510,686,520,547]" pageId="38" pageNumber="39">NIWA-159411</specimenCode>
. Mounted as holotype
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FC68385D6748FDD4" ID-GBIF-Occurrence="3926368309" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="670" latitude="-41.408" location="Honeycomb Canyon" longLatPrecision="6" longitude="175.8987" municipality="Honeycomb Canyon" pageId="38" pageNumber="39" specimenCode="NIWA-159412" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
1 ♀, 2 ♂♂;
<collectingMunicipality id="6BC6ACE45B0A2A36FBCF385D603DFD9A" box="[1145,1394,520,546]" pageId="38" pageNumber="39">Honeycomb Canyon</collectingMunicipality>
, stn TAN1004/58;
<geoCoordinate id="EE2950595B0A2A36FE213878676FFDFF" box="[407,544,557,583]" degrees="41.4080" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-41.408">41.4080° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FD8738786784FDFF" box="[561,715,557,584]" degrees="175.8987" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="175.8987">175.8987° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FD6D38786620FDFF" box="[731,879,557,583]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="38" pageNumber="39" unit="m" value="670.0">
<quantity id="4CE59B7B5B0A2A36FD6D38786666FDFF" box="[731,809,557,583]" metricMagnitude="2" metricUnit="m" metricValue="6.7" pageId="38" pageNumber="39" unit="m" value="670.0">670 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FCC8387866B4FDFF" box="[894,1019,557,583]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FCC8387866B4FDFF" box="[894,1019,557,583]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment;
<specimenCount id="9D1BFD175B0A2A36FE8338076404FDD4" box="[309,331,594,620]" pageId="38" pageNumber="39" type="female"></specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FEE43804674CFDD4" box="[338,515,593,620]" pageId="38" pageNumber="39">NIWA-159412</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FDB838076129FDD3" ID-GBIF-Occurrence="3926368306" box="[526,1126,593,620]" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="670" latitude="-41.408" longLatPrecision="6" longitude="175.8987" pageId="38" pageNumber="39" specimenCode="NHMD-916357-916358" specimenCount="1" specimenCount-female="2" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FDB838076775FDD4" box="[526,570,594,620]" pageId="38" pageNumber="39" type="male">♂♂</specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FDF63804662DFDD4" box="[576,866,593,620]" pageId="38" pageNumber="39">NHMD-916357916358</specimenCode>
. Mounted as holotype
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FBCA38046704FD0D" ID-GBIF-Occurrence="3926368305" collectedFrom="soft sediment" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1495" latitude="-42.1422" location="Campbell Canyon" longLatPrecision="6" longitude="174.5492" municipality="Campbell Canyon" pageId="38" pageNumber="39" specimenCode="NHMD-916360" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
1 ♀, 3 ♂♂;
<collectingMunicipality id="6BC6ACE45B0A2A36FAB238046454FD28" pageId="38" pageNumber="39">Campbell Canyon</collectingMunicipality>
, stn TAN1004/126;
<geoCoordinate id="EE2950595B0A2A36FDB4382367CBFD28" box="[514,644,630,656]" degrees="42.1422" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-42.1422">42.1422° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FD393823666CFD28" box="[655,803,630,657]" degrees="174.5492" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="174.5492">174.5492° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FC9B3823668BFD28" box="[813,964,630,657]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="38" pageNumber="39" unit="m" value="1495.0">
<quantity id="4CE59B7B5B0A2A36FC9B382366CBFD28" box="[813,900,630,657]" metricMagnitude="3" metricUnit="m" metricValue="1.4949999999999999" pageId="38" pageNumber="39" unit="m" value="1495.0">1495 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FC7B3823610BFD28" box="[973,1092,630,656]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FC7B3823610BFD28" box="[973,1092,630,656]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage;
<collectedFrom id="380175635B0A2A36FF0D38CF6412FD0C" box="[187,349,666,692]" pageId="38" pageNumber="39">soft sediment</collectedFrom>
;
<specimenCount id="9D1BFD175B0A2A36FEDD38CE64CEFD0D" box="[363,385,667,693]" pageId="38" pageNumber="39" type="female"></specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FE3F38CF670BFD0C" box="[393,580,666,692]" pageId="38" pageNumber="39">NHMD-916360</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FDE538CE6672FD0D" ID-GBIF-Occurrence="3926368303" box="[595,829,666,693]" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1495" latitude="-42.1422" longLatPrecision="6" longitude="174.5492" pageId="38" pageNumber="39" specimenCode="NIWA-159413" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FDE538CE67CFFD0D" box="[595,640,667,693]" pageId="38" pageNumber="39" type="male">1 ♂</specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FD3E38CF6676FD0D" box="[648,825,666,693]" pageId="38" pageNumber="39">NIWA-159413</specimenCode>
,
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FCF338CE6466FD61" ID-GBIF-Occurrence="3926368307" collectingDate="2010-04" collectionCode="NIWA" country="New Zealand" elevation="1495" latitude="-42.1422" longLatPrecision="6" longitude="174.5492" pageId="38" pageNumber="39" specimenCode="NHMD-916359" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D1BFD175B0A2A36FCF338CE66C7FD0D" box="[837,904,667,693]" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
<specimenCode id="DBBB9EE55B0A2A36FC2638CF6104FD0D" box="[912,1099,666,693]" pageId="38" pageNumber="39">NHMD-916359</specimenCode>
and 916361. Mounted as holotype
</materialsCitation>
.
</paragraph>
<paragraph id="8BA2369E5B0A2A36FF5C395464AEFCA3" blockId="38.[188,1399,769,1052]" box="[234,481,769,795]" pageId="38" pageNumber="39">
<emphasis id="B969EA8C5B0A2A36FF5C395464AEFCA3" bold="true" box="[234,481,769,795]" pageId="38" pageNumber="39">Additional material</emphasis>
</paragraph>
<paragraph id="8BA2369E5B0A2A36FF0B3972644FFBA4" blockId="38.[188,1399,769,1052]" pageId="38" pageNumber="39">
<materialsCitation id="3B753CC35B0A2A36FF0B39726490FC32" ID-GBIF-Occurrence="3926364315" collectingDate="2010-04" collectionCode="NIWA, SEM" country="New Zealand" elevation="1013" latitude="-41.4983" location="Pahaua Canyon" longLatPrecision="6" longitude="175.7043" pageId="38" pageNumber="39" specimenCode="TAN1004" specimenCount="8" specimenCount-female="5" specimenCount-male="3">
<collectingCountry id="F30A760E5B0A2A36FF0B397264D7FCF9" box="[189,408,807,833]" name="New Zealand" pageId="38" pageNumber="39">NEW ZEALAND</collectingCountry>
<specimenCount id="9D1BFD175B0A2A36FE7639736745FCF9" box="[448,522,806,833]" pageId="38" pageNumber="39" type="female">5 ♀♀</specimenCount>
,
<specimenCount id="9D1BFD175B0A2A36FDA939736724FCF9" box="[543,619,806,833]" pageId="38" pageNumber="39" type="male">3 ♂♂</specimenCount>
;
<location id="8EC260455B0A2A36FDC93973660FFCF8" LSID="urn:lsid:plazi:treatment:03B487885B092A3CFDDE3EB06147F9B7:8EC260455B0A2A36FDC93973660FFCF8" box="[639,832,806,832]" country="New Zealand" latitude="-41.4983" longLatPrecision="6" longitude="175.7043" name="Pahaua Canyon" pageId="38" pageNumber="39">Pahaua Canyon</location>
, stn TAN1004/27;
<geoCoordinate id="EE2950595B0A2A36FB823973618FFCF8" box="[1076,1216,806,832]" degrees="41.4983" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-41.4983">41.4983° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FB623973603EFCF9" box="[1236,1393,806,833]" degrees="175.7043" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="175.7043">175.7043° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FF0B391E642EFCDD" box="[189,353,843,869]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="38" pageNumber="39" unit="m" value="1013.0">
<quantity id="4CE59B7B5B0A2A36FF0B391E6454FCDD" box="[189,283,843,869]" metricMagnitude="3" metricUnit="m" metricValue="1.013" pageId="38" pageNumber="39" unit="m" value="1013.0">1013 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FEC7391E64A1FCDD" box="[369,494,843,869]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FEC7391E64A1FCDD" box="[369,494,843,869]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment; personal reference collection of MVS. Mounted for SEM
</materialsCitation>
<materialsCitation id="3B753CC35B0A2A36FE40392564DEFC6B" ID-GBIF-Occurrence="3926364325" collectingDate="2010-04" collectionCode="NIWA, SEM" country="New Zealand" elevation="1171" latitude="-41.476" location="Honeycomb Canyon" longLatPrecision="6" longitude="175.9477" pageId="38" pageNumber="39" specimenCode="TAN1004" specimenCount="3" specimenCount-female="1" specimenCount-male="2">
<specimenCount id="9D1BFD175B0A2A36FE403925676EFC32" box="[502,545,880,906]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
,
<specimenCount id="9D1BFD175B0A2A36FD993925673CFC32" box="[559,627,880,906]" pageId="38" pageNumber="39" type="male">2 ♂♂</specimenCount>
;
<location id="8EC260455B0A2A36FDC8393A663DFC32" LSID="urn:lsid:plazi:treatment:03B487885B092A3CFDDE3EB06147F9B7:8EC260455B0A2A36FDC8393A663DFC32" box="[638,882,879,906]" country="New Zealand" latitude="-41.476" longLatPrecision="6" longitude="175.9477" name="Honeycomb Canyon" pageId="38" pageNumber="39">Honeycomb Canyon</location>
, stn TAN1004/62;
<geoCoordinate id="EE2950595B0A2A36FBFB393A619FFC32" box="[1101,1232,879,906]" degrees="41.4760" direction="south" orientation="latitude" pageId="38" pageNumber="39" precision="5" value="-41.476">41.4760° S</geoCoordinate>
,
<geoCoordinate id="EE2950595B0A2A36FB6A393A603EFC32" box="[1244,1393,879,906]" degrees="175.9477" direction="east" orientation="longitude" pageId="38" pageNumber="39" precision="5" value="175.9477">175.9477° E</geoCoordinate>
;
<elevation id="0030D1AD5B0A2A36FF0B39C1641BFC16" box="[189,340,916,942]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="38" pageNumber="39" unit="m" value="1171.0">
<quantity id="4CE59B7B5B0A2A36FF0B39C1645BFC16" box="[189,276,916,942]" metricMagnitude="3" metricUnit="m" metricValue="1.1709999999999998" pageId="38" pageNumber="39" unit="m" value="1171.0">1171 m</quantity>
b.s.l.
</elevation>
;
<date id="FFA3105E5B0A2A36FEE839C16499FC16" box="[350,470,916,942]" pageId="38" pageNumber="39" value="2010-04">
<collectingDate id="EFE7E9B65B0A2A36FEE839C16499FC16" box="[350,470,916,942]" pageId="38" pageNumber="39" value="2010-04">Apr. 2010</collectingDate>
</date>
; NIWA TAN1004 Voyage; soft sediment; personal reference collection of MVS. Mounted for SEM
</materialsCitation>
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<specimenCount id="9D1BFD175B0A2A36FE1C39EC6498FC6B" box="[426,471,953,979]" pageId="38" pageNumber="39" type="female">1 ♀</specimenCount>
;
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, stn TAN1004/126;
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,
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;
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b.s.l.
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; NIWA TAN1004 Voyage; soft sediment; personal reference collection of MVS. Mounted for SEM
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B0A2A3BFF0B3E11664FF850" lastPageId="43" lastPageNumber="44" pageId="38" pageNumber="39" type="description">
<paragraph id="8BA2369E5B0A2A36FF0B3E116401FBE6" blockId="38.[189,334,1092,1118]" box="[189,334,1092,1118]" pageId="38" pageNumber="39">
<heading id="D0EA81F25B0A2A36FF0B3E116401FBE6" bold="true" box="[189,334,1092,1118]" fontSize="11" level="3" pageId="38" pageNumber="39" reason="3">
<emphasis id="B969EA8C5B0A2A36FF0B3E116401FBE6" bold="true" box="[189,334,1092,1118]" pageId="38" pageNumber="39">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B0A2A36FF0B3E21601CFB44" blockId="38.[187,1399,1140,1276]" pageId="38" pageNumber="39">
GENERAL. Adults with head, neck and eleven trunk segments (
<figureCitation id="13262A1B5B0A2A36FC223E216151FB36" box="[916,1054,1140,1166]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="39.[189,232,1927,1953]" captionStartId-1="40.[189,232,1857,1883]" captionStartId-2="41.[189,232,1793,1819]" captionTargetBox-0="[212,1377,269,1898]" captionTargetBox-1="[190,1398,766,1824]" captionTargetBox-2="[196,1392,267,1742]" captionTargetId-0="figure-60@39.[211,1377,265,1898]" captionTargetId-1="figure-235@40.[189,1399,766,1825]" captionTargetId-2="figure-0@41.[192,1402,265,1743]" captionTargetPageId-0="39" captionTargetPageId-1="40" captionTargetPageId-2="41" captionText-0="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." captionText-1="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." captionText-2="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi-0="http://doi.org/10.5281/zenodo.7222187" figureDoi-1="http://doi.org/10.5281/zenodo.7222189" figureDoi-2="http://doi.org/10.5281/zenodo.7222191" httpUri-0="https://zenodo.org/record/7222187/files/figure.png" httpUri-1="https://zenodo.org/record/7222189/files/figure.png" httpUri-2="https://zenodo.org/record/7222191/files/figure.png" pageId="38" pageNumber="39">Figs 1416</figureCitation>
). Overview of measurements and dimensions in
<tableCitation id="C69F03255B0A2A36FE233ECC64B3FB0B" box="[405,508,1177,1203]" captionStart="Table 10" captionStartId="42.[188,257,265,291]" captionTargetPageId="42" captionText="Table 10. Measurements from light microscopy of Echinoderes gandalfi sp. nov. (in µm) from the Hikurangi Margin, including number of measured specimens (n) and standard deviation (SD)." pageId="38" pageNumber="39">Table 10</tableCitation>
. Distribution of cuticular structures, i.e., sensory spots, glandular cell outlets, spines and tubes, summarized in
<tableCitation id="C69F03255B0A2A36FDFB3EE867F6FB6F" box="[589,697,1213,1239]" captionStart="Table 11" captionStartId="43.[189,258,265,291]" captionTargetPageId="43" captionText="Table 11. Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in Echinoderes gandalfi sp. nov." pageId="38" pageNumber="39">Table 11</tableCitation>
. No details regarding scalid arrangement and morphology could be provided, because introverts of all specimens mounted for SEM fully or partially retracted.
</paragraph>
<paragraph id="8BA2369E5B0A2A36FF0B3F7066CAFA30" blockId="38.[189,1399,1316,1416]" pageId="38" pageNumber="39">
NECK. With 16 placids. Midventral placid broadest, 8 µm in width and 15 µm in length, whereas all others narrower, measuring 6 µm in width at their bases and 13 µm in length, similar in size (
<figureCitation id="13262A1B5B0A2A36FAB63F1C659BFA30" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="38" pageNumber="39">Fig. 15B C</figureCitation>
). The trichoscalid plates are well-developed (
<figureCitation id="13262A1B5B0A2A36FD5F3F3B6637FA30" box="[745,888,1389,1416]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="38" pageNumber="39">Fig. 15BC</figureCitation>
).
</paragraph>
<paragraph id="8BA2369E5B0A2A36FF0B3FE46008F980" blockId="38.[189,1399,1456,1593]" pageId="38" pageNumber="39">
SEGMENT 1. Consists of complete cuticular ring. Sensory spots located on anterior half of segment, in subdorsal positions. Glandular cell outlet
<typeStatus id="54A6883C5B0A2A36FD6A3F836640FA48" box="[732,783,1494,1520]" pageId="38" pageNumber="39">type</typeStatus>
1 present in middorsal and ventrolateral positions (
<figureCitation id="13262A1B5B0A2A36FF733FAC642AF9AC" box="[197,357,1529,1556]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="38" pageNumber="39">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B0A2A36FEC83FAC649AF9AC" box="[382,469,1529,1556]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="38" pageNumber="39">15BC</figureCitation>
). Cuticular hairs relatively long, distributed around segment. Posterior segment margin almost straight, forming pectinate fringe with very short, sawtooth-like fringe tips.
</paragraph>
<paragraph id="8BA2369E5B0A2A36FF0B3C346798F851" blockId="38.[188,1400,1633,2025]" pageId="38" pageNumber="39">
SEGMENT 2. Consists of complete cuticular ring, with tubes located in ventrolateral positions (
<figureCitation id="13262A1B5B0A2A36FB4C3C34659CF918" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="38" pageNumber="39">Figs 14A B</figureCitation>
,
<figureCitation id="13262A1B5B0A2A36FF553CD06458F918" box="[227,279,1669,1696]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="38" pageNumber="39">15C</figureCitation>
,
<figureCitation id="13262A1B5B0A2A36FE903CD06414F927" box="[294,347,1669,1695]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="38" pageNumber="39">16C</figureCitation>
). Sensory spots present in middorsal, laterodorsal and ventromedial positions; sensory spots on this and following eight segments relatively large, with numerous micropapillae surrounding central pore and long marginal hair, giving them droplet-shaped appearance (
<figureCitation id="13262A1B5B0A2A36FBF83C9B61F5F950" box="[1102,1210,1742,1768]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="38" pageNumber="39">Fig. 16B</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B0A2A36FEA13CA16405F8B6" box="[279,330,1780,1806]" pageId="38" pageNumber="39">type</typeStatus>
1 present in middorsal and ventromedial positions (
<figureCitation id="13262A1B5B0A2A36FC7A3CA6612CF8B5" box="[972,1123,1779,1805]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="38" pageNumber="39">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B0A2A36FBC53CA661E8F8B6" box="[1139,1191,1779,1806]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="38" pageNumber="39">15B</figureCitation>
). Pachycyclus of anterior segment margin of regular thickness, without interruption. Secondary pectinate fringe present near anterior segment margin of this and following segments, but usually covered by preceding segment. Fairly long cuticular hairs evenly distributed across tergal plate; paraventral areas on this and following three segments with thinner and much shorter, non-bracteate hairs, emerging through perforation sites visible in SEM only. Posterior segment margin almost straight, but with rounded midventral extension; pectinate fringe tips as on preceding segment.
</paragraph>
<caption id="DF6266165B0B2A37FF0B3DD26004F850" ID-DOI="http://doi.org/10.5281/zenodo.7222187" ID-Zenodo-Dep="7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="39" pageNumber="40" startId="39.[189,232,1927,1953]" targetBox="[212,1377,269,1898]" targetPageId="39">
<paragraph id="8BA2369E5B0B2A37FF0B3DD26004F850" blockId="39.[189,1399,1927,2025]" pageId="39" pageNumber="40">
<emphasis id="B969EA8C5B0B2A37FF0B3DD26454F81A" bold="true" box="[189,283,1927,1954]" pageId="39" pageNumber="40">Fig. 14.</emphasis>
Line art illustrations of
<taxonomicName id="4C1D4D1D5B0B2A37FDF23DDD6672F81A" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[580,829,1928,1954]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="39" pageNumber="40" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B0B2A37FDF23DDD6672F81A" box="[580,829,1928,1954]" italics="true" pageId="39" pageNumber="40">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B0B2A37FCF13DDD66ECF81A" box="[839,931,1928,1954]" pageId="39" pageNumber="40" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B0B2A37FC183DD2668EF819" bold="true" box="[942,961,1927,1953]" pageId="39" pageNumber="40">A</emphasis>
. ♂, dorsal view.
<emphasis id="B969EA8C5B0B2A37FB243DDD61E9F81A" bold="true" box="[1170,1190,1928,1954]" pageId="39" pageNumber="40">B</emphasis>
. ♂, ventral view.
<emphasis id="B969EA8C5B0B2A37FF0B3DFE659DF87D" bold="true" box="[189,210,1963,1989]" pageId="39" pageNumber="40">C</emphasis>
. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view.
<emphasis id="B969EA8C5B0B2A37FB173DFE61F9F87D" bold="true" box="[1185,1206,1963,1989]" pageId="39" pageNumber="40">D</emphasis>
. ♀, segments 10 to 11, dorsal view.
<emphasis id="B969EA8C5B0B2A37FE2D3D9B64E0F850" bold="true" box="[411,431,1998,2024]" pageId="39" pageNumber="40">E</emphasis>
. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<paragraph id="8BA2369E5B042A38FF0B3B5C6771FE4F" blockId="40.[186,1399,265,503]" pageId="40" pageNumber="41">
SEGMENT 3. Present segment, and eight remaining ones, consist of one tergal and two sternal plates (
<figureCitation id="13262A1B5B042A38FF733B79641AFEFE" box="[197,341,300,326]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="40" pageNumber="41">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B042A38FEE93B7964F9FEFF" box="[351,438,300,327]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="40" pageNumber="41">15AC</figureCitation>
,
<figureCitation id="13262A1B5B042A38FE763B796758FEFE" box="[448,535,300,326]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="40" pageNumber="41">16BC</figureCitation>
). Pachycyclus of anterior segment margin of medium thickness, interrupted at tergosternal and midsternal junctions and middorsally, on this and following seven segments. Segment with middorsal and ventromedial glandular cell outlets
<typeStatus id="54A6883C5B042A38FC003B2666A6FE35" box="[950,1001,371,397]" pageId="40" pageNumber="41">type</typeStatus>
1; no sensory spots or other traits observed. Cuticular hairs interrupted by hairless midlateral area, otherwise as on preceding segment. Posterior segment margin straight, terminating in pectinate fringe with longer and more slender fringe tips than on preceding segments.
</paragraph>
<paragraph id="8BA2369E5B042A38FF0B384B670CFDE3" blockId="40.[189,1399,542,603]" pageId="40" pageNumber="41">
SEGMENT 4. With glandular cell outlets
<typeStatus id="54A6883C5B042A38FD04384B67AAFD80" box="[690,741,542,568]" pageId="40" pageNumber="41">type</typeStatus>
1 present in paradorsal and ventromedial positions. Segment otherwise as segment 3.
</paragraph>
<paragraph id="8BA2369E5B042A3AFF0B38D764EDF9B3" blockId="40.[189,1399,642,704]" lastBlockId="42.[188,1398,1415,1547]" lastPageId="42" lastPageNumber="43" pageId="40" pageNumber="41">
SEGMENT 5. With tubes in lateroventral positions (
<figureCitation id="13262A1B5B042A38FCAF38D766C3FD24" box="[793,908,642,668]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="40" pageNumber="41">Figs 14B</figureCitation>
,
<figureCitation id="13262A1B5B042A38FC2D38D76680FD25" box="[923,975,642,669]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="40" pageNumber="41">15C</figureCitation>
,
<figureCitation id="13262A1B5B042A38FC6938D76177FD24" box="[991,1080,642,668]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="40" pageNumber="41">16AB</figureCitation>
). Sensory spots present in midlateral and ventromedial positions; the latter, however, missing in some specimens examined under SEM (
<figureCitation id="13262A1B5B062A3AFEBF3FD2643EFA19" box="[265,369,1415,1441]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="42" pageNumber="43">Fig. 16B</figureCitation>
) and for some specimens (including
<typeStatus id="54A6883C5B062A3AFCA93FD266C7FA19" box="[799,904,1415,1441]" pageId="42" pageNumber="43" type="holotype">holotype</typeStatus>
) examined under LM presence/absence of ventromedial sensory spots could not be confirmed. Glandular cell outlets type 1 present in paradorsal and ventromedial positions. Pectinate fringe of posterior segment margin and cuticular hairs as on preceding segment.
</paragraph>
<caption id="DF6266165B042A38FF0B3D1467A4F850" ID-DOI="http://doi.org/10.5281/zenodo.7222189" ID-Zenodo-Dep="7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="40" pageNumber="41" startId="40.[189,232,1857,1883]" targetBox="[190,1398,766,1824]" targetPageId="40">
<paragraph id="8BA2369E5B042A38FF0B3D1467A4F850" blockId="40.[189,1399,1857,2025]" pageId="40" pageNumber="41">
<emphasis id="B969EA8C5B042A38FF0B3D146455F8E3" bold="true" box="[189,282,1857,1883]" pageId="40" pageNumber="41">Fig. 15.</emphasis>
Light micrographs showing overview and details of
<taxonomicName id="4C1D4D1D5B042A38FC3A3D1461CAF8E3" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[908,1157,1857,1883]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="40" pageNumber="41" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B042A38FC3A3D1461CAF8E3" box="[908,1157,1857,1883]" italics="true" pageId="40" pageNumber="41">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B042A38FB3B3D1761A7F8E4" box="[1165,1256,1858,1884]" pageId="40" pageNumber="41" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B042A38FB463D146005F8E3" bold="true" box="[1264,1354,1857,1883]" pageId="40" pageNumber="41">AD, F</emphasis>
. ♂, holotype (NIWA-159407).
<emphasis id="B969EA8C5B042A38FE4A3D306779F8C6" bold="true" box="[508,566,1892,1919]" pageId="40" pageNumber="41">E, G</emphasis>
. ♀, paratype (NHMD-916362).
<emphasis id="B969EA8C5B042A38FC053D316687F8C6" bold="true" box="[947,968,1892,1918]" pageId="40" pageNumber="41">A</emphasis>
. Ventral overview.
<emphasis id="B969EA8C5B042A38FB1B3D30618EF8C7" bold="true" box="[1197,1217,1893,1919]" pageId="40" pageNumber="41">B</emphasis>
. Segments 1 to 6, dorsal view.
<emphasis id="B969EA8C5B042A38FEC43DD264C9F819" bold="true" box="[370,390,1927,1953]" pageId="40" pageNumber="41">C</emphasis>
. Segments 1 to 9, ventral view.
<emphasis id="B969EA8C5B042A38FCB03DDD6655F81A" bold="true" box="[774,794,1928,1954]" pageId="40" pageNumber="41">D</emphasis>
. Segments 6 to 10, dorsal view.
<emphasis id="B969EA8C5B042A38FB283DDD61FEF81A" bold="true" box="[1182,1201,1928,1954]" pageId="40" pageNumber="41">E</emphasis>
. Segments 10 to 11 of female, dorsal view.
<emphasis id="B969EA8C5B042A38FE453DFE674AF87D" bold="true" box="[499,517,1963,1989]" pageId="40" pageNumber="41">F</emphasis>
. Segments 10 to 11 of male, dorsal view.
<emphasis id="B969EA8C5B042A38FC463DFE6148F87D" bold="true" box="[1008,1031,1963,1989]" pageId="40" pageNumber="41">G</emphasis>
. Segment 11 of female, ventral view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6266165B052A39FF0B3D546073F83C" ID-DOI="http://doi.org/10.5281/zenodo.7222191" ID-Zenodo-Dep="7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="41" pageNumber="42" startId="41.[189,232,1793,1819]" targetBox="[196,1392,267,1742]" targetPageId="41">
<paragraph id="8BA2369E5B052A39FF0B3D546073F83C" blockId="41.[188,1399,1793,1925]" pageId="41" pageNumber="42">
<emphasis id="B969EA8C5B052A39FF0B3D546456F8A3" bold="true" box="[189,281,1793,1819]" pageId="41" pageNumber="42">Fig. 16.</emphasis>
Scanning electron micrographs showing overviews and details of
<taxonomicName id="4C1D4D1D5B052A39FBAB3D546059F8A3" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1053,1302,1793,1819]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="41" pageNumber="42" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B052A39FBAB3D546059F8A3" box="[1053,1302,1793,1819]" italics="true" pageId="41" pageNumber="42">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B052A39FAAB3D576038F8A4" box="[1309,1399,1794,1820]" pageId="41" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="B969EA8C5B052A39FF0A3D71659EF886" bold="true" box="[188,209,1828,1854]" pageId="41" pageNumber="42">A</emphasis>
. Lateral overview.
<emphasis id="B969EA8C5B052A39FE723D716497F886" bold="true" box="[452,472,1828,1854]" pageId="41" pageNumber="42">B</emphasis>
. Ventral overview.
<emphasis id="B969EA8C5B052A39FD7D3D7167AFF886" bold="true" box="[715,736,1828,1854]" pageId="41" pageNumber="42">C</emphasis>
. Segments 1 to 4, ventral view.
<emphasis id="B969EA8C5B052A39FBCA3D7161DEF886" bold="true" box="[1148,1169,1828,1854]" pageId="41" pageNumber="42">D</emphasis>
. Lateral overview.
<emphasis id="B969EA8C5B052A39FF0B3D1D659EF8DA" bold="true" box="[189,209,1864,1890]" pageId="41" pageNumber="42">E</emphasis>
. Segments 4 to 10, dorsal view.
<emphasis id="B969EA8C5B052A39FDE93D1D6720F8DA" bold="true" box="[607,623,1864,1890]" pageId="41" pageNumber="42">F</emphasis>
. Segments 8 to 9, lateral view.
<emphasis id="B969EA8C5B052A39FC463D12614AF8D9" bold="true" box="[1008,1029,1863,1889]" pageId="41" pageNumber="42">G</emphasis>
. Segments 10 to 11 of female, dorsal view.
<emphasis id="B969EA8C5B052A39FEE43D3E6426F83D" bold="true" box="[338,361,1899,1925]" pageId="41" pageNumber="42">H</emphasis>
. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods.
</paragraph>
</caption>
<caption id="DF6266165B062A3AFF0A3B5C6044FEFE" pageId="42" pageNumber="43" startId="42.[188,257,265,291]" targetBox="[242,1315,376,1359]" targetIsTable="true" targetPageId="42">
<paragraph id="8BA2369E5B062A3AFF0A3B5C6044FEFE" blockId="42.[188,1399,265,327]" pageId="42" pageNumber="43">
<emphasis id="B969EA8C5B062A3AFF0A3B5C647EFE9B" bold="true" box="[188,305,265,291]" pageId="42" pageNumber="43">Table 10.</emphasis>
Measurements from light microscopy of
<taxonomicName id="4C1D4D1D5B062A3AFC873B5C6161FE9B" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[817,1070,265,291]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="42" pageNumber="43" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B062A3AFC873B5C6161FE9B" box="[817,1070,265,291]" italics="true" pageId="42" pageNumber="43">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B062A3AFB8C3B5C61D5FE9B" box="[1082,1178,265,291]" pageId="42" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
(in µm) from the Hikurangi Margin, including number of measured specimens (n) and standard deviation (SD).
</paragraph>
</caption>
<paragraph id="8BA2369E5B062A3AFF403B2D605FFAF7" pageId="42" pageNumber="43">
<table id="F91DC43E5B06D5EFFF443B2D606CFAF7" box="[242,1315,376,1359]" colsContinueFrom="26.[242,1308,378,1315]" colsContinueIn="49.[242,1308,391,1365]" gridcols="5" gridrows="26" pageId="42" pageNumber="43">
<tr id="352D34DC5B06D5EFFF443B2D606CFE2B" box="[242,1315,376,403]" gridrow="0" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443B2D6434FE2B" box="[242,379,376,403]" gridcol="0" gridrow="0" pageId="42" pageNumber="43">
<emphasis id="B969EA8C5B062A3AFF403B2D6438FE2A" bold="true" box="[246,375,376,402]" pageId="42" pageNumber="43">Character</emphasis>
</th>
<th id="76FC5DA05B06D5EFFDAF3B2D6779FE2B" box="[537,566,376,403]" gridcol="1" gridrow="0" pageId="42" pageNumber="43">
<emphasis id="B969EA8C5B062A3AFD963B2C677FFE2B" bold="true" box="[544,560,377,403]" pageId="42" pageNumber="43">n</emphasis>
</th>
<th id="76FC5DA05B06D5EFFD793B2D662BFE2B" box="[719,868,376,403]" gridcol="2" gridrow="0" pageId="42" pageNumber="43">
<emphasis id="B969EA8C5B062A3AFD443B2C660DFE2B" bold="true" box="[754,834,377,403]" pageId="42" pageNumber="43">Range</emphasis>
</th>
<th id="76FC5DA05B06D5EFFC683B2D6176FE2B" box="[990,1081,376,403]" gridcol="3" gridrow="0" pageId="42" pageNumber="43">
<emphasis id="B969EA8C5B062A3AFC5E3B2C617FFE2B" bold="true" box="[1000,1072,377,403]" pageId="42" pageNumber="43">Mean</emphasis>
</th>
<th id="76FC5DA05B06D5EFFB6E3B2D606CFE2B" box="[1240,1315,376,403]" gridcol="4" gridrow="0" pageId="42" pageNumber="43">
<emphasis id="B969EA8C5B062A3AFB5D3B2D605FFE2A" bold="true" box="[1259,1296,376,402]" pageId="42" pageNumber="43">SD</emphasis>
</th>
</tr>
<tr id="352D34DC5B06D5EFFF443BFF606CFE7D" box="[242,1315,426,453]" gridrow="1" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443BFF6434FE7D" box="[242,379,426,453]" gridcol="0" gridrow="1" pageId="42" pageNumber="43">TL</th>
<td id="76FC5DA05B06D5EFFDAF3BFF6779FE7D" box="[537,566,426,453]" gridcol="1" gridrow="1" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793BFF662BFE7D" box="[719,868,426,453]" gridcol="2" gridrow="1" pageId="42" pageNumber="43">140222</td>
<td id="76FC5DA05B06D5EFFC683BFF6176FE7D" box="[990,1081,426,453]" gridcol="3" gridrow="1" pageId="42" pageNumber="43">186</td>
<td id="76FC5DA05B06D5EFFB6E3BFF606CFE7D" box="[1240,1315,426,453]" gridcol="4" gridrow="1" pageId="42" pageNumber="43">24.5</td>
</tr>
<tr id="352D34DC5B06D5EFFF443B9A606CFE52" box="[242,1315,463,490]" gridrow="2" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443B9A6434FE52" box="[242,379,463,490]" gridcol="0" gridrow="2" pageId="42" pageNumber="43">MSW-6</th>
<td id="76FC5DA05B06D5EFFDAF3B9A6779FE52" box="[537,566,463,490]" gridcol="1" gridrow="2" pageId="42" pageNumber="43">7</td>
<td id="76FC5DA05B06D5EFFD793B9A662BFE52" box="[719,868,463,490]" gridcol="2" gridrow="2" pageId="42" pageNumber="43">4255</td>
<td id="76FC5DA05B06D5EFFC683B9A6176FE52" box="[990,1081,463,490]" gridcol="3" gridrow="2" pageId="42" pageNumber="43">50</td>
<td id="76FC5DA05B06D5EFFB6E3B9A606CFE52" box="[1240,1315,463,490]" gridcol="4" gridrow="2" pageId="42" pageNumber="43">4.8</td>
</tr>
<tr id="352D34DC5B06D5EFFF443BA0606CFDA8" box="[242,1315,501,528]" gridrow="3" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443BA06434FDA8" box="[242,379,501,528]" gridcol="0" gridrow="3" pageId="42" pageNumber="43">MSW-6/TL</th>
<td id="76FC5DA05B06D5EFFDAF3BA06779FDA8" box="[537,566,501,528]" gridcol="1" gridrow="3" pageId="42" pageNumber="43">7</td>
<td id="76FC5DA05B06D5EFFD793BA0662BFDA8" box="[719,868,501,528]" gridcol="2" gridrow="3" pageId="42" pageNumber="43">21.632.0%</td>
<td id="76FC5DA05B06D5EFFC683BA06176FDA8" box="[990,1081,501,528]" gridcol="3" gridrow="3" pageId="42" pageNumber="43">27.6%</td>
<td id="76FC5DA05B06D5EFFB6E3BA0606CFDA8" box="[1240,1315,501,528]" gridcol="4" gridrow="3" pageId="42" pageNumber="43">4.3%</td>
</tr>
<tr id="352D34DC5B06D5EFFF44384E606CFD8E" box="[242,1315,539,566]" gridrow="4" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF44384E6434FD8E" box="[242,379,539,566]" gridcol="0" gridrow="4" pageId="42" pageNumber="43">SW-10</th>
<td id="76FC5DA05B06D5EFFDAF384E6779FD8E" box="[537,566,539,566]" gridcol="1" gridrow="4" pageId="42" pageNumber="43">7</td>
<td id="76FC5DA05B06D5EFFD79384E662BFD8E" box="[719,868,539,566]" gridcol="2" gridrow="4" pageId="42" pageNumber="43">3742</td>
<td id="76FC5DA05B06D5EFFC68384E6176FD8E" box="[990,1081,539,566]" gridcol="3" gridrow="4" pageId="42" pageNumber="43">39</td>
<td id="76FC5DA05B06D5EFFB6E384E606CFD8E" box="[1240,1315,539,566]" gridcol="4" gridrow="4" pageId="42" pageNumber="43">1.9</td>
</tr>
<tr id="352D34DC5B06D5EFFF443814606CFDE4" box="[242,1315,577,604]" gridrow="5" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4438146434FDE4" box="[242,379,577,604]" gridcol="0" gridrow="5" pageId="42" pageNumber="43">SW-10/TL</th>
<td id="76FC5DA05B06D5EFFDAF38146779FDE4" box="[537,566,577,604]" gridcol="1" gridrow="5" pageId="42" pageNumber="43">7</td>
<td id="76FC5DA05B06D5EFFD793814662BFDE4" box="[719,868,577,604]" gridcol="2" gridrow="5" pageId="42" pageNumber="43">18.326.4%</td>
<td id="76FC5DA05B06D5EFFC6838146176FDE4" box="[990,1081,577,604]" gridcol="3" gridrow="5" pageId="42" pageNumber="43">21.6%</td>
<td id="76FC5DA05B06D5EFFB6E3814606CFDE4" box="[1240,1315,577,604]" gridcol="4" gridrow="5" pageId="42" pageNumber="43">3.2%</td>
</tr>
<tr id="352D34DC5B06D5EFFF443832606CFD39" box="[242,1315,615,641]" gridrow="6" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4438326434FD39" box="[242,379,615,641]" gridcol="0" gridrow="6" pageId="42" pageNumber="43">S1</th>
<td id="76FC5DA05B06D5EFFDAF38326779FD39" box="[537,566,615,641]" gridcol="1" gridrow="6" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD793832662BFD39" box="[719,868,615,641]" gridcol="2" gridrow="6" pageId="42" pageNumber="43">1821</td>
<td id="76FC5DA05B06D5EFFC6838326176FD39" box="[990,1081,615,641]" gridcol="3" gridrow="6" pageId="42" pageNumber="43">19</td>
<td id="76FC5DA05B06D5EFFB6E3832606CFD39" box="[1240,1315,615,641]" gridcol="4" gridrow="6" pageId="42" pageNumber="43">1.8</td>
</tr>
<tr id="352D34DC5B06D5EFFF4438D8606CFD1F" box="[242,1315,653,679]" gridrow="7" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4438D86434FD1F" box="[242,379,653,679]" gridcol="0" gridrow="7" pageId="42" pageNumber="43">S2</th>
<td id="76FC5DA05B06D5EFFDAF38D86779FD1F" box="[537,566,653,679]" gridcol="1" gridrow="7" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD7938D8662BFD1F" box="[719,868,653,679]" gridcol="2" gridrow="7" pageId="42" pageNumber="43">1723</td>
<td id="76FC5DA05B06D5EFFC6838D86176FD1F" box="[990,1081,653,679]" gridcol="3" gridrow="7" pageId="42" pageNumber="43">19</td>
<td id="76FC5DA05B06D5EFFB6E38D8606CFD1F" box="[1240,1315,653,679]" gridcol="4" gridrow="7" pageId="42" pageNumber="43">1.7</td>
</tr>
<tr id="352D34DC5B06D5EFFF4438E7606CFD75" box="[242,1315,690,717]" gridrow="8" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4438E76434FD75" box="[242,379,690,717]" gridcol="0" gridrow="8" pageId="42" pageNumber="43">S3</th>
<td id="76FC5DA05B06D5EFFDAF38E76779FD75" box="[537,566,690,717]" gridcol="1" gridrow="8" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD7938E7662BFD75" box="[719,868,690,717]" gridcol="2" gridrow="8" pageId="42" pageNumber="43">1525</td>
<td id="76FC5DA05B06D5EFFC6838E76176FD75" box="[990,1081,690,717]" gridcol="3" gridrow="8" pageId="42" pageNumber="43">21</td>
<td id="76FC5DA05B06D5EFFB6E38E7606CFD75" box="[1240,1315,690,717]" gridcol="4" gridrow="8" pageId="42" pageNumber="43">3.4</td>
</tr>
<tr id="352D34DC5B06D5EFFF44388D606CFD4A" box="[242,1315,728,754]" gridrow="9" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF44388D6434FD4A" box="[242,379,728,754]" gridcol="0" gridrow="9" pageId="42" pageNumber="43">S4</th>
<td id="76FC5DA05B06D5EFFDAF388D6779FD4A" box="[537,566,728,754]" gridcol="1" gridrow="9" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD79388D662BFD4A" box="[719,868,728,754]" gridcol="2" gridrow="9" pageId="42" pageNumber="43">1729</td>
<td id="76FC5DA05B06D5EFFC68388D6176FD4A" box="[990,1081,728,754]" gridcol="3" gridrow="9" pageId="42" pageNumber="43">23</td>
<td id="76FC5DA05B06D5EFFB6E388D606CFD4A" box="[1240,1315,728,754]" gridcol="4" gridrow="9" pageId="42" pageNumber="43">4.1</td>
</tr>
<tr id="352D34DC5B06D5EFFF4438AB606CFCA1" box="[242,1315,766,793]" gridrow="10" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4438AB6434FCA1" box="[242,379,766,793]" gridcol="0" gridrow="10" pageId="42" pageNumber="43">S5</th>
<td id="76FC5DA05B06D5EFFDAF38AB6779FCA1" box="[537,566,766,793]" gridcol="1" gridrow="10" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD7938AB662BFCA1" box="[719,868,766,793]" gridcol="2" gridrow="10" pageId="42" pageNumber="43">2231</td>
<td id="76FC5DA05B06D5EFFC6838AB6176FCA1" box="[990,1081,766,793]" gridcol="3" gridrow="10" pageId="42" pageNumber="43">26</td>
<td id="76FC5DA05B06D5EFFB6E38AB606CFCA1" box="[1240,1315,766,793]" gridcol="4" gridrow="10" pageId="42" pageNumber="43">2.9</td>
</tr>
<tr id="352D34DC5B06D5EFFF443971606CFC86" box="[242,1315,804,830]" gridrow="11" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4439716434FC86" box="[242,379,804,830]" gridcol="0" gridrow="11" pageId="42" pageNumber="43">S6</th>
<td id="76FC5DA05B06D5EFFDAF39716779FC86" box="[537,566,804,830]" gridcol="1" gridrow="11" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD793971662BFC86" box="[719,868,804,830]" gridcol="2" gridrow="11" pageId="42" pageNumber="43">2431</td>
<td id="76FC5DA05B06D5EFFC6839716176FC86" box="[990,1081,804,830]" gridcol="3" gridrow="11" pageId="42" pageNumber="43">27</td>
<td id="76FC5DA05B06D5EFFB6E3971606CFC86" box="[1240,1315,804,830]" gridcol="4" gridrow="11" pageId="42" pageNumber="43">2.1</td>
</tr>
<tr id="352D34DC5B06D5EFFF44391F606CFCDC" box="[242,1315,842,868]" gridrow="12" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF44391F6434FCDC" box="[242,379,842,868]" gridcol="0" gridrow="12" pageId="42" pageNumber="43">S7</th>
<td id="76FC5DA05B06D5EFFDAF391F6779FCDC" box="[537,566,842,868]" gridcol="1" gridrow="12" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD79391F662BFCDC" box="[719,868,842,868]" gridcol="2" gridrow="12" pageId="42" pageNumber="43">2433</td>
<td id="76FC5DA05B06D5EFFC68391F6176FCDC" box="[990,1081,842,868]" gridcol="3" gridrow="12" pageId="42" pageNumber="43">30</td>
<td id="76FC5DA05B06D5EFFB6E391F606CFCDC" box="[1240,1315,842,868]" gridcol="4" gridrow="12" pageId="42" pageNumber="43">2.2</td>
</tr>
<tr id="352D34DC5B06D5EFFF443925606CFC32" box="[242,1315,880,906]" gridrow="13" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4439256434FC32" box="[242,379,880,906]" gridcol="0" gridrow="13" pageId="42" pageNumber="43">S8</th>
<td id="76FC5DA05B06D5EFFDAF39256779FC32" box="[537,566,880,906]" gridcol="1" gridrow="13" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD793925662BFC32" box="[719,868,880,906]" gridcol="2" gridrow="13" pageId="42" pageNumber="43">2937</td>
<td id="76FC5DA05B06D5EFFC6839256176FC32" box="[990,1081,880,906]" gridcol="3" gridrow="13" pageId="42" pageNumber="43">32</td>
<td id="76FC5DA05B06D5EFFB6E3925606CFC32" box="[1240,1315,880,906]" gridcol="4" gridrow="13" pageId="42" pageNumber="43">2.3</td>
</tr>
<tr id="352D34DC5B06D5EFFF4439C0606CFC17" box="[242,1315,917,943]" gridrow="14" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4439C06434FC17" box="[242,379,917,943]" gridcol="0" gridrow="14" pageId="42" pageNumber="43">S9</th>
<td id="76FC5DA05B06D5EFFDAF39C06779FC17" box="[537,566,917,943]" gridcol="1" gridrow="14" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD7939C0662BFC17" box="[719,868,917,943]" gridcol="2" gridrow="14" pageId="42" pageNumber="43">2936</td>
<td id="76FC5DA05B06D5EFFC6839C06176FC17" box="[990,1081,917,943]" gridcol="3" gridrow="14" pageId="42" pageNumber="43">33</td>
<td id="76FC5DA05B06D5EFFB6E39C0606CFC17" box="[1240,1315,917,943]" gridcol="4" gridrow="14" pageId="42" pageNumber="43">2.3</td>
</tr>
<tr id="352D34DC5B06D5EFFF4439EE606CFC6D" box="[242,1315,955,981]" gridrow="15" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4439EE6434FC6D" box="[242,379,955,981]" gridcol="0" gridrow="15" pageId="42" pageNumber="43">S10</th>
<td id="76FC5DA05B06D5EFFDAF39EE6779FC6D" box="[537,566,955,981]" gridcol="1" gridrow="15" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD7939EE662BFC6D" box="[719,868,955,981]" gridcol="2" gridrow="15" pageId="42" pageNumber="43">2633</td>
<td id="76FC5DA05B06D5EFFC6839EE6176FC6D" box="[990,1081,955,981]" gridcol="3" gridrow="15" pageId="42" pageNumber="43">30</td>
<td id="76FC5DA05B06D5EFFB6E39EE606CFC6D" box="[1240,1315,955,981]" gridcol="4" gridrow="15" pageId="42" pageNumber="43">2.2</td>
</tr>
<tr id="352D34DC5B06D5EFFF4439B4606CFC44" box="[242,1315,993,1020]" gridrow="16" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF4439B46434FC44" box="[242,379,993,1020]" gridcol="0" gridrow="16" pageId="42" pageNumber="43">S11</th>
<td id="76FC5DA05B06D5EFFDAF39B46779FC44" box="[537,566,993,1020]" gridcol="1" gridrow="16" pageId="42" pageNumber="43">9</td>
<td id="76FC5DA05B06D5EFFD7939B4662BFC44" box="[719,868,993,1020]" gridcol="2" gridrow="16" pageId="42" pageNumber="43">1928</td>
<td id="76FC5DA05B06D5EFFC6839B46176FC44" box="[990,1081,993,1020]" gridcol="3" gridrow="16" pageId="42" pageNumber="43">23</td>
<td id="76FC5DA05B06D5EFFB6E39B4606CFC44" box="[1240,1315,993,1020]" gridcol="4" gridrow="16" pageId="42" pageNumber="43">5.1</td>
</tr>
<tr id="352D34DC5B06D5EFFF443E53606CFB99" box="[242,1315,1030,1057]" gridrow="17" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443E536434FB99" box="[242,379,1030,1057]" gridcol="0" gridrow="17" pageId="42" pageNumber="43">MD6 (ac)</th>
<td id="76FC5DA05B06D5EFFDAF3E536779FB99" box="[537,566,1030,1057]" gridcol="1" gridrow="17" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793E53662BFB99" box="[719,868,1030,1057]" gridcol="2" gridrow="17" pageId="42" pageNumber="43">3442</td>
<td id="76FC5DA05B06D5EFFC683E536176FB99" box="[990,1081,1030,1057]" gridcol="3" gridrow="17" pageId="42" pageNumber="43">39</td>
<td id="76FC5DA05B06D5EFFB6E3E53606CFB99" box="[1240,1315,1030,1057]" gridcol="4" gridrow="17" pageId="42" pageNumber="43">2.9</td>
</tr>
<tr id="352D34DC5B06D5EFFF443E79606CFBFF" box="[242,1315,1068,1095]" gridrow="18" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443E796434FBFF" box="[242,379,1068,1095]" gridcol="0" gridrow="18" pageId="42" pageNumber="43">MD8 (ac)</th>
<td id="76FC5DA05B06D5EFFDAF3E796779FBFF" box="[537,566,1068,1095]" gridcol="1" gridrow="18" pageId="42" pageNumber="43">8</td>
<td id="76FC5DA05B06D5EFFD793E79662BFBFF" box="[719,868,1068,1095]" gridcol="2" gridrow="18" pageId="42" pageNumber="43">4956</td>
<td id="76FC5DA05B06D5EFFC683E796176FBFF" box="[990,1081,1068,1095]" gridcol="3" gridrow="18" pageId="42" pageNumber="43">52</td>
<td id="76FC5DA05B06D5EFFB6E3E79606CFBFF" box="[1240,1315,1068,1095]" gridcol="4" gridrow="18" pageId="42" pageNumber="43">2.1</td>
</tr>
<tr id="352D34DC5B06D5EFFF443E07606CFBD4" box="[242,1315,1106,1132]" gridrow="19" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443E076434FBD4" box="[242,379,1106,1132]" gridcol="0" gridrow="19" pageId="42" pageNumber="43">LV6 (ac)</th>
<td id="76FC5DA05B06D5EFFDAF3E076779FBD4" box="[537,566,1106,1132]" gridcol="1" gridrow="19" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793E07662BFBD4" box="[719,868,1106,1132]" gridcol="2" gridrow="19" pageId="42" pageNumber="43">1723</td>
<td id="76FC5DA05B06D5EFFC683E076176FBD4" box="[990,1081,1106,1132]" gridcol="3" gridrow="19" pageId="42" pageNumber="43">20</td>
<td id="76FC5DA05B06D5EFFB6E3E07606CFBD4" box="[1240,1315,1106,1132]" gridcol="4" gridrow="19" pageId="42" pageNumber="43">2.3</td>
</tr>
<tr id="352D34DC5B06D5EFFF443E2D606CFB2B" box="[242,1315,1144,1171]" gridrow="20" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443E2D6434FB2B" box="[242,379,1144,1171]" gridcol="0" gridrow="20" pageId="42" pageNumber="43">LV7 (ac)</th>
<td id="76FC5DA05B06D5EFFDAF3E2D6779FB2B" box="[537,566,1144,1171]" gridcol="1" gridrow="20" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793E2D662BFB2B" box="[719,868,1144,1171]" gridcol="2" gridrow="20" pageId="42" pageNumber="43">1925</td>
<td id="76FC5DA05B06D5EFFC683E2D6176FB2B" box="[990,1081,1144,1171]" gridcol="3" gridrow="20" pageId="42" pageNumber="43">22</td>
<td id="76FC5DA05B06D5EFFB6E3E2D606CFB2B" box="[1240,1315,1144,1171]" gridcol="4" gridrow="20" pageId="42" pageNumber="43">2.7</td>
</tr>
<tr id="352D34DC5B06D5EFFF443ECB606CFB01" box="[242,1315,1182,1209]" gridrow="21" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443ECB6434FB01" box="[242,379,1182,1209]" gridcol="0" gridrow="21" pageId="42" pageNumber="43">LV8 (ac)</th>
<td id="76FC5DA05B06D5EFFDAF3ECB6779FB01" box="[537,566,1182,1209]" gridcol="1" gridrow="21" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793ECB662BFB01" box="[719,868,1182,1209]" gridcol="2" gridrow="21" pageId="42" pageNumber="43">2328</td>
<td id="76FC5DA05B06D5EFFC683ECB6176FB01" box="[990,1081,1182,1209]" gridcol="3" gridrow="21" pageId="42" pageNumber="43">25</td>
<td id="76FC5DA05B06D5EFFB6E3ECB606CFB01" box="[1240,1315,1182,1209]" gridcol="4" gridrow="21" pageId="42" pageNumber="43">2.5</td>
</tr>
<tr id="352D34DC5B06D5EFFF443E96606CFB66" box="[242,1315,1219,1246]" gridrow="22" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443E966434FB66" box="[242,379,1219,1246]" gridcol="0" gridrow="22" pageId="42" pageNumber="43">LV9 (ac)</th>
<td id="76FC5DA05B06D5EFFDAF3E966779FB66" box="[537,566,1219,1246]" gridcol="1" gridrow="22" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793E96662BFB66" box="[719,868,1219,1246]" gridcol="2" gridrow="22" pageId="42" pageNumber="43">2431</td>
<td id="76FC5DA05B06D5EFFC683E966176FB66" box="[990,1081,1219,1246]" gridcol="3" gridrow="22" pageId="42" pageNumber="43">26</td>
<td id="76FC5DA05B06D5EFFB6E3E96606CFB66" box="[1240,1315,1219,1246]" gridcol="4" gridrow="22" pageId="42" pageNumber="43">2.4</td>
</tr>
<tr id="352D34DC5B06D5EFFF443EBC606CFABC" box="[242,1315,1257,1284]" gridrow="23" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443EBC6434FABC" box="[242,379,1257,1284]" gridcol="0" gridrow="23" pageId="42" pageNumber="43">LTS</th>
<td id="76FC5DA05B06D5EFFDAF3EBC6779FABC" box="[537,566,1257,1284]" gridcol="1" gridrow="23" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793EBC662BFABC" box="[719,868,1257,1284]" gridcol="2" gridrow="23" pageId="42" pageNumber="43">191275</td>
<td id="76FC5DA05B06D5EFFC683EBC6176FABC" box="[990,1081,1257,1284]" gridcol="3" gridrow="23" pageId="42" pageNumber="43">241</td>
<td id="76FC5DA05B06D5EFFB6E3EBC606CFABC" box="[1240,1315,1257,1284]" gridcol="4" gridrow="23" pageId="42" pageNumber="43">26.2</td>
</tr>
<tr id="352D34DC5B06D5EFFF443F5A606CFA91" box="[242,1315,1295,1321]" gridrow="24" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443F5A6434FA91" box="[242,379,1295,1321]" gridcol="0" gridrow="24" pageId="42" pageNumber="43">LTS/TL</th>
<td id="76FC5DA05B06D5EFFDAF3F5A6779FA91" box="[537,566,1295,1321]" gridcol="1" gridrow="24" pageId="42" pageNumber="43">10</td>
<td id="76FC5DA05B06D5EFFD793F5A662BFA91" box="[719,868,1295,1321]" gridcol="2" gridrow="24" pageId="42" pageNumber="43">103169.3%</td>
<td id="76FC5DA05B06D5EFFC683F5A6176FA91" box="[990,1081,1295,1321]" gridcol="3" gridrow="24" pageId="42" pageNumber="43">132.0%</td>
<td id="76FC5DA05B06D5EFFB6E3F5A606CFA91" box="[1240,1315,1295,1321]" gridcol="4" gridrow="24" pageId="42" pageNumber="43">24.1%</td>
</tr>
<tr id="352D34DC5B06D5EFFF443F60606CFAF7" box="[242,1315,1333,1359]" gridrow="25" pageId="42" pageNumber="43">
<th id="76FC5DA05B06D5EFFF443F606434FAF7" box="[242,379,1333,1359]" gridcol="0" gridrow="25" pageId="42" pageNumber="43">LTAS</th>
<td id="76FC5DA05B06D5EFFDAF3F606779FAF7" box="[537,566,1333,1359]" gridcol="1" gridrow="25" pageId="42" pageNumber="43">3</td>
<td id="76FC5DA05B06D5EFFD793F60662BFAF7" box="[719,868,1333,1359]" gridcol="2" gridrow="25" pageId="42" pageNumber="43">2834</td>
<td id="76FC5DA05B06D5EFFC683F606176FAF7" box="[990,1081,1333,1359]" gridcol="3" gridrow="25" pageId="42" pageNumber="43">30</td>
<td id="76FC5DA05B06D5EFFB6E3F60606CFAF7" box="[1240,1315,1333,1359]" gridcol="4" gridrow="25" pageId="42" pageNumber="43">3.2</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA2369E5B062A3AFF0B3C676471F961" blockId="42.[188,1399,1586,1753]" pageId="42" pageNumber="43">
SEGMENT 6. With spines in middorsal and lateroventral positions (
<figureCitation id="13262A1B5B062A3AFC703C676115F9F4" box="[966,1114,1586,1612]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="42" pageNumber="43">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFBDE3C6761F0F9F5" box="[1128,1215,1586,1613]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="42" pageNumber="43">15CD</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFB7A3C676025F9F4" box="[1228,1386,1586,1612]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="42" pageNumber="43">16AB, DF</figureCitation>
). Sensory spots present in paradorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B062A3AFB993C0061F0F9D7" box="[1071,1215,1621,1647]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="42" pageNumber="43">Fig. 14AB</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B062A3AFEA23C2C6408F92B" box="[276,327,1657,1683]" pageId="42" pageNumber="43">type</typeStatus>
1 present in paradorsal and ventromedial positions (
<figureCitation id="13262A1B5B062A3AFC093C2D611BF92A" box="[959,1108,1656,1682]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="42" pageNumber="43">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFBD53C2D61D8F92B" box="[1123,1175,1656,1683]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="42" pageNumber="43">15C</figureCitation>
). Tips of pectinate fringe of posterior segment margin slightly longer than on preceding segments. Segment otherwise as segment 5.
</paragraph>
<paragraph id="8BA2369E5B062A3AFF0B3D556138F83C" blockId="42.[187,1399,1792,1924]" pageId="42" pageNumber="43">
SEGMENT 7. With spines in lateroventral positions, and sensory spots in paradorsal, midlateral and ventromedial positions (
<figureCitation id="13262A1B5B062A3AFE633D766729F885" box="[469,614,1827,1853]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="42" pageNumber="43">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFDC63D766788F886" box="[624,711,1827,1854]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="42" pageNumber="43">15CD</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFD643D766648F885" box="[722,775,1827,1853]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="42" pageNumber="43">16B</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B062A3AFBAE3D716104F886" box="[1048,1099,1828,1854]" pageId="42" pageNumber="43">type</typeStatus>
1 present in ventromedial positions, not observed on the dorsal side. Cuticular hairs covering as on preceding segment except for hairless paraventral and ventromedial areas, on this and following four segments.
</paragraph>
<paragraph id="8BA2369E5B062A3BFF0B3DFE64EDFBF9" blockId="42.[188,1399,1963,2025]" lastBlockId="43.[189,1398,1025,1089]" lastPageId="43" lastPageNumber="44" pageId="42" pageNumber="43">
SEGMENT 8. With spines in middorsal and lateroventral positions, and tubes in lateral accessory positions (
<figureCitation id="13262A1B5B062A3AFF723D9B6419F850" box="[196,342,1998,2024]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="42" pageNumber="43">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFED73D9B64F7F851" box="[353,440,1998,2025]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="42" pageNumber="43">15CD</figureCitation>
,
<figureCitation id="13262A1B5B062A3AFE753D9B6757F850" box="[451,536,1998,2024]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="42" pageNumber="43">16D, F</figureCitation>
). Sensory spots present in paradorsal positions only. Glandular cell outlets
<typeStatus id="54A6883C5B072A3BFF0B3E5765BFFBA4" box="[189,240,1026,1052]" pageId="43" pageNumber="44">type</typeStatus>
1 as on preceding segment. Pectinate fringe of posterior segment margin and cuticular hairs as on preceding segment.
</paragraph>
<caption id="DF6266165B072A3BFF0B3B5C665CFEFF" pageId="43" pageNumber="44" startId="43.[189,258,265,291]" targetBox="[206,1386,376,909]" targetIsTable="true" targetPageId="43">
<paragraph id="8BA2369E5B072A3BFF0B3B5C665CFEFF" blockId="43.[189,1398,265,327]" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFF0B3B5C647FFE9B" bold="true" box="[189,304,265,291]" pageId="43" pageNumber="44">Table 11.</emphasis>
Summary of nature and location of sensory spots, glandular cell outlets, tubes and spines arranged by series in
<taxonomicName id="4C1D4D1D5B072A3BFE0E3B7967FEFEFE" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[440,689,300,326]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="43" pageNumber="44" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B072A3BFE0E3B7967FEFEFE" box="[440,689,300,326]" italics="true" pageId="43" pageNumber="44">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B072A3BFD0E3B78665CFEFF" box="[696,787,301,327]" pageId="43" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BA2369E5B072A3BFF783B2D6124FC35" pageId="43" pageNumber="44">
<table id="F91DC43E5B07D5EFFF783B2D6025FC35" box="[206,1386,376,909]" gridcols="10" gridrows="12" pageId="43" pageNumber="44">
<tr id="352D34DC5B07D5EFFF783B2D6025FE2B" box="[206,1386,376,403]" gridrow="0" pageId="43" pageNumber="44">
<th id="76FC5DA05B07D5EFFF783B2D64EEFE2B" box="[206,417,376,403]" gridcol="0" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFF783B2D64EEFE2B" bold="true" box="[206,417,376,403]" pageId="43" pageNumber="44">Position segment</emphasis>
</th>
<th id="76FC5DA05B07D5EFFE0D3B2D677BFE2B" box="[443,564,376,403]" gridcol="1" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFE693B2C675FFE2B" bold="true" box="[479,528,377,403]" pageId="43" pageNumber="44">MD</emphasis>
</th>
<th id="76FC5DA05B07D5EFFDF13B2D67D1FE2B" box="[583,670,376,403]" gridcol="2" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFDE93B2C67C9FE2B" bold="true" box="[607,646,377,403]" pageId="43" pageNumber="44">PD</emphasis>
</th>
<th id="76FC5DA05B07D5EFFD003B2D67BFFE2B" box="[694,752,376,403]" gridcol="3" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFD763B2D67AAFE2A" bold="true" box="[704,741,376,402]" pageId="43" pageNumber="44">SD</emphasis>
</th>
<th id="76FC5DA05B07D5EFFCBB3B2D6679FE2B" box="[781,822,376,403]" gridcol="4" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFCBB3B2C6679FE2B" bold="true" box="[781,822,377,403]" pageId="43" pageNumber="44">LD</emphasis>
</th>
<th id="76FC5DA05B07D5EFFCE93B2D66F1FE2B" box="[863,958,376,403]" gridcol="5" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFCC23B2C66ECFE2B" bold="true" box="[884,931,377,403]" pageId="43" pageNumber="44">ML</emphasis>
</th>
<th id="76FC5DA05B07D5EFFC6A3B2D6161FE2B" box="[988,1070,376,403]" gridcol="6" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFC433B2D6151FE2A" bold="true" box="[1013,1054,376,402]" pageId="43" pageNumber="44">LA</emphasis>
</th>
<th id="76FC5DA05B07D5EFFBFC3B2D613FFE2B" box="[1098,1136,376,403]" gridcol="7" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFBFC3B2C613FFE2B" bold="true" box="[1098,1136,377,403]" pageId="43" pageNumber="44">LV</emphasis>
</th>
<th id="76FC5DA05B07D5EFFB173B2D6195FE2B" box="[1185,1242,376,403]" gridcol="8" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFB1C3B2C619CFE2B" bold="true" box="[1194,1235,377,403]" pageId="43" pageNumber="44">VL</emphasis>
</th>
<th id="76FC5DA05B07D5EFFAB23B2D6025FE2B" box="[1284,1386,376,403]" gridcol="9" gridrow="0" pageId="43" pageNumber="44">
<emphasis id="B969EA8C5B072A3BFAA93B2C601FFE2B" bold="true" box="[1311,1360,377,403]" pageId="43" pageNumber="44">VM</emphasis>
</th>
</tr>
<tr id="352D34DC5B07D5EFFF783BFB6025FE71" box="[206,1386,430,457]" gridrow="1" pageId="43" pageNumber="44" rowspan-2="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="76FC5DA05B07D5EFFF783BFB64EEFE71" box="[206,417,430,457]" gridcol="0" gridrow="1" pageId="43" pageNumber="44">1</th>
<td id="76FC5DA05B07D5EFFE0D3BFB677BFE71" box="[443,564,430,457]" gridcol="1" gridrow="1" pageId="43" pageNumber="44">gco1</td>
<td id="76FC5DA05B07D5EFFD003BFB67BFFE71" box="[694,752,430,457]" gridcol="3" gridrow="1" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFB173BFB6195FE71" box="[1185,1242,430,457]" gridcol="8" gridrow="1" pageId="43" pageNumber="44">gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF783B8E6025FE4E" box="[206,1386,475,502]" gridrow="2" pageId="43" pageNumber="44" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76FC5DA05B07D5EFFF783B8E64EEFE4E" box="[206,417,475,502]" gridcol="0" gridrow="2" pageId="43" pageNumber="44">2</th>
<td id="76FC5DA05B07D5EFFE0D3B8E677BFE4E" box="[443,564,475,502]" gridcol="1" gridrow="2" pageId="43" pageNumber="44">gco1,ss</td>
<td id="76FC5DA05B07D5EFFCBB3B8E6679FE4E" box="[781,822,475,502]" gridcol="4" gridrow="2" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFB173B8E6195FE4E" box="[1185,1242,475,502]" gridcol="8" gridrow="2" pageId="43" pageNumber="44">tu</td>
<td id="76FC5DA05B07D5EFFAB23B8E6025FE4E" box="[1284,1386,475,502]" gridcol="9" gridrow="2" pageId="43" pageNumber="44">gco1,ss</td>
</tr>
<tr id="352D34DC5B07D5EFFF78385D6025FD9A" box="[206,1386,520,546]" gridrow="3" pageId="43" pageNumber="44" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B07D5EFFF78385D64EEFD9A" box="[206,417,520,546]" gridcol="0" gridrow="3" pageId="43" pageNumber="44">3</th>
<td id="76FC5DA05B07D5EFFE0D385D677BFD9A" box="[443,564,520,546]" gridcol="1" gridrow="3" pageId="43" pageNumber="44">gco1</td>
<td id="76FC5DA05B07D5EFFAB2385D6025FD9A" box="[1284,1386,520,546]" gridcol="9" gridrow="3" pageId="43" pageNumber="44">gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7838636025FDE8" box="[206,1386,566,592]" gridrow="4" pageId="43" pageNumber="44" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1" rowspan-8="1">
<th id="76FC5DA05B07D5EFFF78386364EEFDE8" box="[206,417,566,592]" gridcol="0" gridrow="4" pageId="43" pageNumber="44">4</th>
<td id="76FC5DA05B07D5EFFDF1386367D1FDE8" box="[583,670,566,592]" gridcol="2" gridrow="4" pageId="43" pageNumber="44">gco1</td>
<td id="76FC5DA05B07D5EFFAB238636025FDE8" box="[1284,1386,566,592]" gridcol="9" gridrow="4" pageId="43" pageNumber="44">gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7838366025FDC6" box="[206,1386,611,638]" gridrow="5" pageId="43" pageNumber="44" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76FC5DA05B07D5EFFF78383664EEFDC6" box="[206,417,611,638]" gridcol="0" gridrow="5" pageId="43" pageNumber="44">5</th>
<td id="76FC5DA05B07D5EFFDF1383667D1FDC6" box="[583,670,611,638]" gridcol="2" gridrow="5" pageId="43" pageNumber="44">gco1</td>
<td id="76FC5DA05B07D5EFFCE9383666F1FDC6" box="[863,958,611,638]" gridcol="5" gridrow="5" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFBFC3836613FFDC6" box="[1098,1136,611,638]" gridcol="7" gridrow="5" pageId="43" pageNumber="44">tu</td>
<td id="76FC5DA05B07D5EFFAB238366025FDC6" box="[1284,1386,611,638]" gridcol="9" gridrow="5" pageId="43" pageNumber="44">ss*,gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7838C56025FD13" box="[206,1386,656,683]" gridrow="6" pageId="43" pageNumber="44" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76FC5DA05B07D5EFFF7838C564EEFD13" box="[206,417,656,683]" gridcol="0" gridrow="6" pageId="43" pageNumber="44">6</th>
<td id="76FC5DA05B07D5EFFE0D38C5677BFD13" box="[443,564,656,683]" gridcol="1" gridrow="6" pageId="43" pageNumber="44">ac</td>
<td id="76FC5DA05B07D5EFFDF138C567D1FD13" box="[583,670,656,683]" gridcol="2" gridrow="6" pageId="43" pageNumber="44">gco1,ss</td>
<td id="76FC5DA05B07D5EFFCE938C566F1FD13" box="[863,958,656,683]" gridcol="5" gridrow="6" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFBFC38C5613FFD13" box="[1098,1136,656,683]" gridcol="7" gridrow="6" pageId="43" pageNumber="44">ac</td>
<td id="76FC5DA05B07D5EFFAB238C56025FD13" box="[1284,1386,656,683]" gridcol="9" gridrow="6" pageId="43" pageNumber="44">ss, gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7838EB6025FD60" box="[206,1386,702,728]" gridrow="7" pageId="43" pageNumber="44" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-8="1">
<th id="76FC5DA05B07D5EFFF7838EB64EEFD60" box="[206,417,702,728]" gridcol="0" gridrow="7" pageId="43" pageNumber="44">7</th>
<td id="76FC5DA05B07D5EFFDF138EB67D1FD60" box="[583,670,702,728]" gridcol="2" gridrow="7" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFCE938EB66F1FD60" box="[863,958,702,728]" gridcol="5" gridrow="7" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFBFC38EB613FFD60" box="[1098,1136,702,728]" gridcol="7" gridrow="7" pageId="43" pageNumber="44">ac</td>
<td id="76FC5DA05B07D5EFFAB238EB6025FD60" box="[1284,1386,702,728]" gridcol="9" gridrow="7" pageId="43" pageNumber="44">ss,gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7838BE6025FCBE" box="[206,1386,747,774]" gridrow="8" pageId="43" pageNumber="44" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-8="1">
<th id="76FC5DA05B07D5EFFF7838BE64EEFCBE" box="[206,417,747,774]" gridcol="0" gridrow="8" pageId="43" pageNumber="44">8</th>
<td id="76FC5DA05B07D5EFFE0D38BE677BFCBE" box="[443,564,747,774]" gridcol="1" gridrow="8" pageId="43" pageNumber="44">ac</td>
<td id="76FC5DA05B07D5EFFDF138BE67D1FCBE" box="[583,670,747,774]" gridcol="2" gridrow="8" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFC6A38BE6161FCBE" box="[988,1070,747,774]" gridcol="6" gridrow="8" pageId="43" pageNumber="44">tu</td>
<td id="76FC5DA05B07D5EFFBFC38BE613FFCBE" box="[1098,1136,747,774]" gridcol="7" gridrow="8" pageId="43" pageNumber="44">ac</td>
<td id="76FC5DA05B07D5EFFAB238BE6025FCBE" box="[1284,1386,747,774]" gridcol="9" gridrow="8" pageId="43" pageNumber="44">gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF78394D6025FC8B" box="[206,1386,792,819]" gridrow="9" pageId="43" pageNumber="44" rowspan-1="1" rowspan-5="1">
<th id="76FC5DA05B07D5EFFF78394D64EEFC8B" box="[206,417,792,819]" gridcol="0" gridrow="9" pageId="43" pageNumber="44">9</th>
<td id="76FC5DA05B07D5EFFDF1394D67D1FC8B" box="[583,670,792,819]" gridcol="2" gridrow="9" pageId="43" pageNumber="44">gco1,ss</td>
<td id="76FC5DA05B07D5EFFD00394D67BFFC8B" box="[694,752,792,819]" gridcol="3" gridrow="9" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFCBB394D6679FC8B" box="[781,822,792,819]" gridcol="4" gridrow="9" pageId="43" pageNumber="44">tu</td>
<td id="76FC5DA05B07D5EFFC6A394D6161FC8B" box="[988,1070,792,819]" gridcol="6" gridrow="9" pageId="43" pageNumber="44">si</td>
<td id="76FC5DA05B07D5EFFBFC394D613FFC8B" box="[1098,1136,792,819]" gridcol="7" gridrow="9" pageId="43" pageNumber="44">ac</td>
<td id="76FC5DA05B07D5EFFB17394D6195FC8B" box="[1185,1242,792,819]" gridcol="8" gridrow="9" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFAB2394D6025FC8B" box="[1284,1386,792,819]" gridcol="9" gridrow="9" pageId="43" pageNumber="44">gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7839136025FCD8" box="[206,1386,838,864]" gridrow="10" pageId="43" pageNumber="44" rowspan-2="1" rowspan-3="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76FC5DA05B07D5EFFF78391364EEFCD8" box="[206,417,838,864]" gridcol="0" gridrow="10" pageId="43" pageNumber="44">10</th>
<td id="76FC5DA05B07D5EFFE0D3913677BFCD8" box="[443,564,838,864]" gridcol="1" gridrow="10" pageId="43" pageNumber="44">gco1,gco1</td>
<td id="76FC5DA05B07D5EFFCBB39136679FCD8" box="[781,822,838,864]" gridcol="4" gridrow="10" pageId="43" pageNumber="44">tu</td>
<td id="76FC5DA05B07D5EFFB1739136195FCD8" box="[1185,1242,838,864]" gridcol="8" gridrow="10" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFAB239136025FCD8" box="[1284,1386,838,864]" gridcol="9" gridrow="10" pageId="43" pageNumber="44">gco1</td>
</tr>
<tr id="352D34DC5B07D5EFFF7839266025FC35" box="[206,1386,883,909]" gridrow="11" pageId="43" pageNumber="44" rowspan-4="1" rowspan-8="1" rowspan-9="1">
<th id="76FC5DA05B07D5EFFF78392664EEFC35" box="[206,417,883,909]" gridcol="0" gridrow="11" pageId="43" pageNumber="44">11</th>
<td id="76FC5DA05B07D5EFFE0D3926677BFC35" box="[443,564,883,909]" gridcol="1" gridrow="11" pageId="43" pageNumber="44">pr</td>
<td id="76FC5DA05B07D5EFFDF1392667D1FC35" box="[583,670,883,909]" gridcol="2" gridrow="11" pageId="43" pageNumber="44">ss</td>
<td id="76FC5DA05B07D5EFFD00392667BFFC35" box="[694,752,883,909]" gridcol="3" gridrow="11" pageId="43" pageNumber="44">gco1</td>
<td id="76FC5DA05B07D5EFFCE9392666F1FC35" box="[863,958,883,909]" gridcol="5" gridrow="11" pageId="43" pageNumber="44">pe 3(♂)</td>
<td id="76FC5DA05B07D5EFFC6A39266161FC35" box="[988,1070,883,909]" gridcol="6" gridrow="11" pageId="43" pageNumber="44">ltas(♀)</td>
<td id="76FC5DA05B07D5EFFBFC3926613FFC35" box="[1098,1136,883,909]" gridcol="7" gridrow="11" pageId="43" pageNumber="44">lts</td>
</tr>
</table>
</paragraph>
<paragraph id="8BA2369E5B072A3BFF0B3E3D612CFA8B" blockId="43.[189,1400,1128,1331]" pageId="43" pageNumber="44">
SEGMENT 9. With spines in lateroventral positions and long (~18 µm) tubes in laterodorsal positions (
<figureCitation id="13262A1B5B072A3BFF733EDE6416FB1E" box="[197,345,1163,1190]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFED03EDE64D4FB1E" box="[358,411,1163,1190]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="43" pageNumber="44">15D</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFE1E3EDE64B1FB1D" box="[424,510,1163,1189]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="43" pageNumber="44">16DF</figureCitation>
). Sensory spots located in paradorsal, subdorsal and ventrolateral positions (
<figureCitation id="13262A1B5B072A3BFF703EFA6410FB71" box="[198,351,1199,1225]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFEC73EFB6488FB71" box="[369,455,1198,1225]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="43" pageNumber="44">15BC</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFE6F3EFA6744FB71" box="[473,523,1199,1225]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="43" pageNumber="44">16F</figureCitation>
). Glandular cell outlets
<typeStatus id="54A6883C5B072A3BFC8D3EFA6621FB71" box="[827,878,1199,1225]" pageId="43" pageNumber="44">type</typeStatus>
1 present in paradorsal and ventromedial positions. Small, rounded sieve plates located in lateral accessory positions. Cuticular hairs covering as on preceding segment, but less dense on dorsal side around sensory spots and tubes. Pectinate fringe of posterior segment margin with tips slightly shorter than on preceding segments.
</paragraph>
<paragraph id="8BA2369E5B072A3BFF0B3F0F61E9F9D3" blockId="43.[188,1399,1370,1643]" pageId="43" pageNumber="44">
SEGMENT 10. With laterodorsal tubes located near posterior segment margin; tubes well developed in both sexes (
<figureCitation id="13262A1B5B072A3BFEF03F286480FA20" box="[326,463,1405,1432]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="43" pageNumber="44">Figs 15EF</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFE6C3F286779FA2F" box="[474,566,1405,1431]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="43" pageNumber="44">16GH</figureCitation>
). Sensory spots present in ventrolateral positions. Glandular cell outlets
<typeStatus id="54A6883C5B072A3BFF0A3FF465A0FA03" box="[188,239,1441,1467]" pageId="43" pageNumber="44">type</typeStatus>
1 present as two middorsal ones, and in ventromedial positions. Cuticular hairs significantly scarcer than on preceding segment. Central part of tergal palate devoid of hairs; short cuticular hairs lightly scattered on lateral halves only. Hairs on sternal plates short and present mostly on lateral halves and near posterior segment margin. Posterior segment margin of tergal plate straight, with shorter fringe tips than those on preceding segment; margins of sternal plates with longer fringe tips than those on tergal plate and extend midventrally, almost reaching posterior margin of terminal segment.
</paragraph>
<paragraph id="8BA2369E5B072A3BFF0B3CC7664FF850" blockId="43.[187,1401,1681,2025]" pageId="43" pageNumber="44">
SEGMENT 11. With very long lateral terminal spines, always exceeding trunk length (
<figureCitation id="13262A1B5B072A3BFB3D3CC461B9F914" box="[1163,1270,1681,1708]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Fig. 14C</figureCitation>
;
<tableCitation id="C69F03255B072A3BFAB73CC46026F914" box="[1281,1385,1681,1708]" captionStart="Table 10" captionStartId="42.[188,257,265,291]" captionTargetPageId="42" captionText="Table 10. Measurements from light microscopy of Echinoderes gandalfi sp. nov. (in µm) from the Hikurangi Margin, including number of measured specimens (n) and standard deviation (SD)." pageId="43" pageNumber="44">Table 10</tableCitation>
). Males with three pairs of penile spines; dorsal and ventral ones are long, relatively thin tubes, whereas median ones markedly thicker, conical and stout (
<figureCitation id="13262A1B5B072A3BFCA93C8D66F9F94A" box="[799,950,1752,1778]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Figs 14AB</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFC703C8D66B3F94A" box="[966,1020,1752,1778]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="43" pageNumber="44">16H</figureCitation>
). Females with lateral terminal accessory spines (
<figureCitation id="13262A1B5B072A3BFE3D3CAE6757F8AD" box="[395,536,1787,1813]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Fig. 14DE</figureCitation>
). Sensory spots present in paradorsal positions (
<figureCitation id="13262A1B5B072A3BFB8C3CAE6185F8AE" box="[1082,1226,1787,1814]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Figs 14A, D</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFB653CAE6045F8AD" box="[1235,1290,1787,1813]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="43" pageNumber="44">16D</figureCitation>
); sensory spots smaller than on preceding segments, without long marginal hairs. Glandular cell outlets
<typeStatus id="54A6883C5B072A3BFA9F3D4A6013F881" box="[1321,1372,1823,1849]" pageId="43" pageNumber="44">type</typeStatus>
1 present in subdorsal positions. Middorsal protuberance-like structure extends from intersegmentary joint (
<figureCitation id="13262A1B5B072A3BFEB33D3064DBF8C7" box="[261,404,1893,1919]" captionStart="Fig" captionStartId="39.[189,232,1927,1953]" captionTargetBox="[212,1377,269,1898]" captionTargetId="figure-60@39.[211,1377,265,1898]" captionTargetPageId="39" captionText="Fig. 14. Line art illustrations of Echinoderes gandalfi sp. nov. A. ♂, dorsal view. B. ♂, ventral view. C. ♂, segments 10 to 11, with lateral terminal spines drawn in full length, dorsal view. D. ♀, segments 10 to 11, dorsal view. E. ♀, segments 10 to 11, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222187" httpUri="https://zenodo.org/record/7222187/files/figure.png" pageId="43" pageNumber="44">Figs 14A,D</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFE143D3064BEF838" box="[418,497,1893,1920]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="43" pageNumber="44">15EF</figureCitation>
,
<figureCitation id="13262A1B5B072A3BFE493D306779F8C7" box="[511,566,1893,1919]" captionStart="Fig" captionStartId="41.[189,232,1793,1819]" captionTargetBox="[196,1392,267,1742]" captionTargetId="figure-0@41.[192,1402,265,1743]" captionTargetPageId="41" captionText="Fig. 16. Scanning electron micrographs showing overviews and details of Echinoderes gandalfi sp. nov. A. Lateral overview. B. Ventral overview. C. Segments 1 to 4, ventral view. D. Lateral overview. E. Segments 4 to 10, dorsal view. F. Segments 8 to 9, lateral view. G. Segments 10 to 11 of female, dorsal view. H. Segments 10 to 11 of male, dorsal view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222191" httpUri="https://zenodo.org/record/7222191/files/figure.png" pageId="43" pageNumber="44">16G</figureCitation>
). Segment devoid of cuticular hairs, but with short cuticular hair-like structures covering paradorsal area and short fringes covering margins of tergal and sternal plates. Tergal extensions short and pointed, with small tooth at inner margin (
<figureCitation id="13262A1B5B072A3BFC033DFE616CF87E" box="[949,1059,1963,1990]" captionStart="Fig" captionStartId="40.[189,232,1857,1883]" captionTargetBox="[190,1398,766,1824]" captionTargetId="figure-235@40.[189,1399,766,1825]" captionTargetPageId="40" captionText="Fig. 15. Light micrographs showing overview and details of Echinoderes gandalfi sp. nov. AD, F. ♂, holotype (NIWA-159407). E, G. ♀, paratype (NHMD-916362). A. Ventral overview. B. Segments 1 to 6, dorsal view. C. Segments 1 to 9, ventral view. D. Segments 6 to 10, dorsal view. E. Segments 10 to 11 of female, dorsal view. F. Segments 10 to 11 of male, dorsal view. G. Segment 11 of female, ventral view. Abbreviations: see Material and methods." figureDoi="http://doi.org/10.5281/zenodo.7222189" httpUri="https://zenodo.org/record/7222189/files/figure.png" pageId="43" pageNumber="44">Fig. 15G</figureCitation>
). Sternal extensions broadly rounded, not extending beyond tergal extensions.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B002A3CFF0B3B5D6628FEC2" pageId="44" pageNumber="45" type="distribution">
<paragraph id="8BA2369E5B002A3CFF0B3B5D6419FE9A" blockId="44.[189,342,264,290]" box="[189,342,264,290]" pageId="44" pageNumber="45">
<heading id="D0EA81F25B002A3CFF0B3B5D6419FE9A" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B969EA8C5B002A3CFF0B3B5D6419FE9A" bold="true" box="[189,342,264,290]" pageId="44" pageNumber="45">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B002A3CFF0B3B6F6628FEC2" blockId="44.[189,1398,314,378]" pageId="44" pageNumber="45">
Canyons: Pahaua, Campbell, Honeycomb,
<quantity id="4CE59B7B5B002A3CFD0D3B6F661FFEEC" box="[699,848,314,341]" metricMagnitude="3" metricUnit="m" metricValue="1.1125" metricValueMax="1.4949999999999999" metricValueMin="0.73" pageId="44" pageNumber="45" unit="m" value="1112.5" valueMax="1495.0" valueMin="730.0">7301495 m</quantity>
b.s.l. See
<figureCitation id="13262A1B5B002A3CFC7B3B6F615BFEEC" box="[973,1044,314,340]" captionStart="Fig" captionStartId="2.[189,232,1961,1987]" captionTargetBox="[189,1399,1136,1922]" captionTargetId="figure-333@2.[189,1399,1136,1923]" captionTargetPageId="2" captionText="Fig. 1. Map showing the sampling stations. Colour codes indicate slopes (red), canyons (green), and seamounts (yellow). Inset shows New Zealand with a frame marking the sampling area." figureDoi="http://doi.org/10.5281/zenodo.7222159" httpUri="https://zenodo.org/record/7222159/files/figure.png" pageId="44" pageNumber="45">Fig. 1</figureCitation>
for a geographic overview of stations and
<tableCitation id="C69F03255B002A3CFEE73B3564E7FEC2" box="[337,424,352,378]" captionStart="Table 1" captionStartId="3.[185,254,549,577]" captionTargetBox="[192,1910,658,1734]" captionTargetPageId="3" captionText="Table 1 (continued on next four pages). Coordinates and basic data on sampling stations, and identities, type status and catalogue numbers of species of Echinoderes Claparède, 1863 from each station." httpUri="http://table.plazi.org/id/DF6266165B2F2A13FF0F38706644FDDC" pageId="44" pageNumber="45" tableUuid="DF6266165B2F2A13FF0F38706644FDDC">Table 1</tableCitation>
for station and specimen information.
</paragraph>
</subSubSection>
<subSubSection id="C30765155B002A3CFF0B3BF16147F9B7" pageId="44" pageNumber="45" type="discussion">
<paragraph id="8BA2369E5B002A3CFF0B3BF16603FE06" blockId="44.[189,844,419,446]" box="[189,844,419,446]" pageId="44" pageNumber="45">
<heading id="D0EA81F25B002A3CFF0B3BF16603FE06" bold="true" box="[189,844,419,446]" fontSize="11" level="3" pageId="44" pageNumber="45" reason="3">
<emphasis id="B969EA8C5B002A3CFF0B3BF16603FE06" bold="true" box="[189,844,419,446]" pageId="44" pageNumber="45">
Taxonomic remarks on
<taxonomicName id="4C1D4D1D5B002A3CFE533BF667A8FE06" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[485,743,419,446]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B002A3CFE533BF667A8FE06" bold="true" box="[485,743,419,446]" italics="true" pageId="44" pageNumber="45">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B002A3CFD583BF16603FE06" box="[750,844,420,446]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BA2369E5B002A3CFF0B3B80602BFCFD" blockId="44.[188,1400,469,837]" pageId="44" pageNumber="45">
The spine pattern with middorsal spines on segments 6 and 8 is rare among species of
<taxonomicName id="4C1D4D1D5B002A3CFB683B80603FFE57" authorityName="Claparede" authorityYear="1863" box="[1246,1392,469,495]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="genus">
<emphasis id="B969EA8C5B002A3CFB683B80603FFE57" box="[1246,1392,469,495]" italics="true" pageId="44" pageNumber="45">Echinoderes</emphasis>
</taxonomicName>
, and is shared with only three species, i.e.,
<taxonomicName id="4C1D4D1D5B002A3CFD643BA96631FDAD" authorityName="Grzelak &amp; Sorensen" authorityYear="2017" box="[722,894,507,533]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="daenerysae">
<emphasis id="B969EA8C5B002A3CFD643BA96631FDAD" box="[722,894,507,533]" italics="true" pageId="44" pageNumber="45">E. daenerysae</emphasis>
</taxonomicName>
,
<taxonomicName id="4C1D4D1D5B002A3CFC273BA9600FFDAD" authority="Sorensen et al., 2018" authorityName="Sorensen" authorityYear="2018" box="[913,1344,507,534]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="hviidarum">
<emphasis id="B969EA8C5B002A3CFC273BA96160FDAD" box="[913,1071,507,533]" italics="true" pageId="44" pageNumber="45">E. hviidarum</emphasis>
<bibRefCitation id="EF8C4B6F5B002A3CFB8D3BAE600FFDAD" author="Sorensen M. V. &amp; Rohal M. &amp; Thistle D." box="[1083,1344,507,534]" pageId="44" pageNumber="45" pagination="1 - 75" refId="ref53235" refString="Sorensen M. V., Rohal M. &amp; Thistle D. 2018. Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific. European Journal of Taxonomy 456: 1 - 75. https: // doi. org / 10.5852 / ejt. 2018.456" type="journal article" year="2018">
Sørensen
<emphasis id="B969EA8C5B002A3CFB043BA961A4FDAD" box="[1202,1259,507,533]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
., 2018
</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B002A3CFF0B387767A4FD83" authority="Adrianov &amp; Maiorova, 2019" authorityName="Adrianov &amp; Maiorova" authorityYear="2019" box="[189,747,545,571]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="ultraabyssalis">
<emphasis id="B969EA8C5B002A3CFF0B387764C9FD83" box="[189,390,545,571]" italics="true" pageId="44" pageNumber="45">E. ultraabyssalis</emphasis>
<bibRefCitation id="EF8C4B6F5B002A3CFE39387467A4FD83" author="Adrianov A. V. &amp; Maiorova A. S." box="[399,747,545,571]" pageId="44" pageNumber="45" pagination="102142" refId="ref49562" refString="Adrianov A. V. &amp; Maiorova A. S. 2019. Echinoderes ultraabyssalis sp. nov. from the Kuril-Kamchatka Trench - the first hadal representative of the Kinorhyncha (Kinorhyncha: Cyclorhagida). Progress in Oceanography 178: e 102142. https: // doi. org / 10.1016 / j. pocean. 2019.102142" type="journal article" year="2019">Adrianov &amp; Maiorova, 2019</bibRefCitation>
</taxonomicName>
, and an undescribed species,
<taxonomicName id="4C1D4D1D5B002A3CFBE338746063FD83" box="[1109,1324,545,572]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="undefined-1">
<emphasis id="B969EA8C5B002A3CFBE3387461A8FD83" box="[1109,1255,545,571]" italics="true" pageId="44" pageNumber="45">Echinoderes</emphasis>
sp. 1
</taxonomicName>
, from the
<collectingRegion id="49D9F87C5B002A3CFF5C3812641DFDD9" box="[234,338,583,609]" country="Chile" name="Atacama" pageId="44" pageNumber="45">Atacama</collectingRegion>
Trench (
<bibRefCitation id="EF8C4B6F5B002A3CFE773812664EFDD9" author="Grzelak K. &amp; Sorensen M. V." box="[449,769,583,609]" pageId="44" pageNumber="45" pagination="113 - 147" refId="ref50257" refString="Grzelak K. &amp; Sorensen M. V. 2018. New species of Echinoderes (Kinorhyncha: Cyclorhagida) from Spitsbergen, with additional information about known Arctic species. Marine Biology Research 14: 113 - 147. https: // doi. org / 10.1080 / 17451000.2017.1367096" type="journal article" year="2018">Grzelak &amp; Sørensen 2018</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B002A3CFCA638126141FDD9" author="Sorensen M. V. &amp; Rohal M. &amp; Thistle D." box="[784,1038,583,610]" pageId="44" pageNumber="45" pagination="1 - 75" refId="ref53235" refString="Sorensen M. V., Rohal M. &amp; Thistle D. 2018. Deep-sea Echinoderidae (Kinorhyncha: Cyclorhagida) from the Northwest Pacific. European Journal of Taxonomy 456: 1 - 75. https: // doi. org / 10.5852 / ejt. 2018.456" type="journal article" year="2018">
Sørensen
<emphasis id="B969EA8C5B002A3CFC31381D66F1FDD9" box="[903,958,583,609]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
. 2018
</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B002A3CFBAA3812603DFDD9" author="Adrianov A. V. &amp; Maiorova A. S." box="[1052,1394,583,609]" pageId="44" pageNumber="45" pagination="102142" refId="ref49562" refString="Adrianov A. V. &amp; Maiorova A. S. 2019. Echinoderes ultraabyssalis sp. nov. from the Kuril-Kamchatka Trench - the first hadal representative of the Kinorhyncha (Kinorhyncha: Cyclorhagida). Progress in Oceanography 178: e 102142. https: // doi. org / 10.1016 / j. pocean. 2019.102142" type="journal article" year="2019">Adrianov &amp; Maiorova 2019</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B002A3CFF0B383864EFFD3F" author="Grzelak K. &amp; Zeppilli D. &amp; Shimabukuro M. &amp; Sorensen M. V." box="[189,416,621,648]" pageId="44" pageNumber="45" pagination="670735" refId="ref50375" refString="Grzelak K., Zeppilli D., Shimabukuro M. &amp; Sorensen M. V. 2021. Hadal mud dragons: First insight into the diversity of Kinorhyncha from the Atacama Trench. Frontiers in Marine Science 8: e 670735. https: // doi. org / 10.3389 / fmars. 2021.670735" type="journal article" year="2021">
Grzelak
<emphasis id="B969EA8C5B002A3CFE96383B641CFD3F" box="[288,339,621,647]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
. 2021
</bibRefCitation>
). Furthermore,
<taxonomicName id="4C1D4D1D5B002A3CFDE5383B679DFD3F" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[595,722,621,647]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B002A3CFDE5383B679DFD3F" box="[595,722,621,647]" italics="true" pageId="44" pageNumber="45">E. gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B002A3CFD6C383B667BFD30" box="[730,820,622,648]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
possesses laterodorsal tubes on segment 9, which is another relatively rare trait, shared with all the abovementioned species, which suggests that these species might represent a group of closely related species. The only other species with tubes in this position on the dorsal side of segment 9 is
<taxonomicName id="4C1D4D1D5B002A3CFD0B38B56676FD41" box="[701,825,735,761]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="belenae">
<emphasis id="B969EA8C5B002A3CFD0B38B56676FD41" box="[701,825,735,761]" italics="true" pageId="44" pageNumber="45">E. belenae</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EF8C4B6F5B002A3CFCCF388A612FFD41" author="Pardos F. &amp; Herranz M. &amp; Sanchez N." box="[889,1120,735,761]" pageId="44" pageNumber="45" pagination="26 - 47" refId="ref52161" refString="Pardos F., Herranz M. &amp; Sanchez N. 2016 b. Two sides of a coin: The Phylum Kinorhyncha in Panama. II) Pacific Panama. Zoologischer Anzeiger 265: 26 - 47. https: // doi. org / 10.1016 / j. jcz. 2016.06.006" type="journal article" year="2016">
Pardos
<emphasis id="B969EA8C5B002A3CFC6438B56141FD41" box="[978,1038,735,761]" italics="true" pageId="44" pageNumber="45">et al.</emphasis>
2016b
</bibRefCitation>
) and
<taxonomicName id="4C1D4D1D5B002A3CFB1238B56042FD41" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1188,1293,735,761]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="frodoi" status="sp. nov.">
<emphasis id="B969EA8C5B002A3CFB1238B56042FD41" box="[1188,1293,735,761]" italics="true" pageId="44" pageNumber="45">E. frodoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B002A3CFAA2388A603DFD41" box="[1300,1394,735,761]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C1D4D1D5B002A3CFF083953640AFCA7" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[190,325,773,799]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="dalzottoi" status="sp. nov.">
<emphasis id="B969EA8C5B002A3CFF083953640AFCA7" box="[190,325,773,799]" italics="true" pageId="44" pageNumber="45">E. dalzottoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B002A3CFEFC395064ECFCA7" box="[330,419,773,799]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
and
<taxonomicName id="4C1D4D1D5B002A3CFE6F3953670EFCA7" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[473,577,773,799]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="leduci" status="sp. nov.">
<emphasis id="B969EA8C5B002A3CFE6F3953670EFCA7" box="[473,577,773,799]" italics="true" pageId="44" pageNumber="45">E. leduci</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B002A3CFDFE395067EBFCA7" box="[584,676,773,799]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
, all described in the present study. Nevertheless, none of these species can in any way be confused with
<taxonomicName id="4C1D4D1D5B002A3CFD173979666EFCFD" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[673,801,811,837]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFD173979666EFCFD" box="[673,801,811,837]" italics="true" pageId="44" pageNumber="45">E. gandalfi</emphasis>
</taxonomicName>
due to their significantly different spine patterns.
</paragraph>
<paragraph id="8BA2369E5B002A3CFF0B393A6147F9B7" blockId="44.[187,1401,879,1551]" pageId="44" pageNumber="45">
<taxonomicName id="4C1D4D1D5B002A3CFF0B393A64F3FC31" authority="Grzelak &amp; Sørensen, 2022" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[189,444,879,905]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi" status="sp. nov.">
<emphasis id="B969EA8C5B002A3CFF0B393A64F3FC31" box="[189,444,879,905]" italics="true" pageId="44" pageNumber="45">Echinoderes gandalfi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A25A57F75B002A3CFE7F39256765FC32" box="[457,554,880,906]" pageId="44" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
probably shows most resemblance to
<taxonomicName id="4C1D4D1D5B002A3CFBBB39256196FC31" authorityName="Adrianov &amp; Maiorova" authorityYear="2019" box="[1037,1241,879,905]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="ultraabyssalis">
<emphasis id="B969EA8C5B002A3CFBBB39256196FC31" box="[1037,1241,879,905]" italics="true" pageId="44" pageNumber="45">E. ultraabyssalis</emphasis>
</taxonomicName>
, the deepest kinorhynch species described so far (&gt;
<quantity id="4CE59B7B5B002A3CFD2C39C067B9FC08" box="[666,758,917,944]" metricMagnitude="3" metricUnit="m" metricValue="9.0" pageId="44" pageNumber="45" unit="m" value="9000.0">9000 m</quantity>
water depth) from the Kuril-Kamchatka Trench, and
<taxonomicName id="4C1D4D1D5B002A3CFF0B39EE64DCFC6D" box="[189,403,955,981]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="undefined-1">
<emphasis id="B969EA8C5B002A3CFF0B39EE6400FC6D" box="[189,335,955,981]" italics="true" pageId="44" pageNumber="45">Echinoderes</emphasis>
sp. 1
</taxonomicName>
, found in the
<collectingRegion id="49D9F87C5B002A3CFD8939EE67E8FC6D" box="[575,679,955,981]" country="Chile" name="Atacama" pageId="44" pageNumber="45">Atacama</collectingRegion>
Trench at a water depth of
<quantity id="4CE59B7B5B002A3CFC4A39EE6118FC6D" box="[1020,1111,955,981]" metricMagnitude="3" metricUnit="m" metricValue="7.7" pageId="44" pageNumber="45" unit="m" value="7700.0">7700 m</quantity>
(
<bibRefCitation id="EF8C4B6F5B002A3CFBDF39EE65B4FC43" author="Adrianov A. V. &amp; Maiorova A. S." pageId="44" pageNumber="45" pagination="102142" refId="ref49562" refString="Adrianov A. V. &amp; Maiorova A. S. 2019. Echinoderes ultraabyssalis sp. nov. from the Kuril-Kamchatka Trench - the first hadal representative of the Kinorhyncha (Kinorhyncha: Cyclorhagida). Progress in Oceanography 178: e 102142. https: // doi. org / 10.1016 / j. pocean. 2019.102142" type="journal article" year="2019">Adrianov &amp; Maiorova 2019</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B002A3CFEBA39B464B0FC43" author="Grzelak K. &amp; Zeppilli D. &amp; Shimabukuro M. &amp; Sorensen M. V." box="[268,511,993,1019]" pageId="44" pageNumber="45" pagination="670735" refId="ref50375" refString="Grzelak K., Zeppilli D., Shimabukuro M. &amp; Sorensen M. V. 2021. Hadal mud dragons: First insight into the diversity of Kinorhyncha from the Atacama Trench. Frontiers in Marine Science 8: e 670735. https: // doi. org / 10.3389 / fmars. 2021.670735" type="journal article" year="2021">
Grzelak
<emphasis id="B969EA8C5B002A3CFEC339B764E2FC43" box="[373,429,993,1019]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
. 2021
</bibRefCitation>
).
<taxonomicName id="4C1D4D1D5B002A3CFDAE39B46659FC43" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[536,790,993,1019]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFDAE39B46659FC43" box="[536,790,993,1019]" italics="true" pageId="44" pageNumber="45">Echinoderes gandalfi</emphasis>
</taxonomicName>
shares several features with these hadal/deep-sea trench species, including the presence of tubes in ventrolateral positions on segment 2, lateroventral positions on segment 5, and in lateral accessory positions on segment 8, the presence of a middorsal protuberance between segments 10 and 11, as well as very long lateral terminal spines, the shape of the tergal extensions and the lack of glandular cell outlets
<typeStatus id="54A6883C5B002A3CFC983E2C662EFB2B" box="[814,865,1145,1171]" pageId="44" pageNumber="45">type</typeStatus>
2. Nevertheless, despite the overall similarity,
<taxonomicName id="4C1D4D1D5B002A3CFF0D3ECA6470FB01" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[187,319,1183,1209]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFF0D3ECA6470FB01" box="[187,319,1183,1209]" italics="true" pageId="44" pageNumber="45">E. gandalfi</emphasis>
</taxonomicName>
can be distinguished from both congeners by its morphometrics.
<taxonomicName id="4C1D4D1D5B002A3CFBEB3ECA6016FB01" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[1117,1369,1183,1209]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFBEB3ECA6016FB01" box="[1117,1369,1183,1209]" italics="true" pageId="44" pageNumber="45">Echinoderes gandalfi</emphasis>
</taxonomicName>
is markedly smaller in trunk length than both species (TL: 186 µm vs 267 µm and 257 µm, respectively) but has proportionally longer lateral terminal spines, which always exceed the trunk length, resulting in a markedly higher LTS/TL ratio in
<taxonomicName id="4C1D4D1D5B002A3CFDC03F4467B9FA92" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[630,758,1296,1322]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFDC03F4467B9FA92" box="[630,758,1296,1322]" italics="true" pageId="44" pageNumber="45">E. gandalfi</emphasis>
</taxonomicName>
in comparison with
<taxonomicName id="4C1D4D1D5B002A3CFC513F4461E2FA92" authorityName="Adrianov &amp; Maiorova" authorityYear="2019" box="[999,1197,1296,1322]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="ultraabyssalis">
<emphasis id="B969EA8C5B002A3CFC513F4461E2FA92" box="[999,1197,1296,1322]" italics="true" pageId="44" pageNumber="45">E. ultraabyssalis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C1D4D1D5B002A3CFB533F4565BBFAE9" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="undefined-1">
<emphasis id="B969EA8C5B002A3CFB533F456038FA92" box="[1253,1399,1296,1322]" italics="true" pageId="44" pageNumber="45">Echinoderes</emphasis>
sp. 1
</taxonomicName>
(LTS/TL: 132% vs 59% and 91%, respectively) (
<bibRefCitation id="EF8C4B6F5B002A3CFC873F636133FAE9" author="Adrianov A. V. &amp; Maiorova A. S." box="[817,1148,1334,1361]" pageId="44" pageNumber="45" pagination="102142" refId="ref49562" refString="Adrianov A. V. &amp; Maiorova A. S. 2019. Echinoderes ultraabyssalis sp. nov. from the Kuril-Kamchatka Trench - the first hadal representative of the Kinorhyncha (Kinorhyncha: Cyclorhagida). Progress in Oceanography 178: e 102142. https: // doi. org / 10.1016 / j. pocean. 2019.102142" type="journal article" year="2019">Adrianov &amp; Maiorova 2019</bibRefCitation>
;
<bibRefCitation id="EF8C4B6F5B002A3CFB313F636025FAE9" author="Grzelak K. &amp; Zeppilli D. &amp; Shimabukuro M. &amp; Sorensen M. V." box="[1159,1386,1334,1361]" pageId="44" pageNumber="45" pagination="670735" refId="ref50375" refString="Grzelak K., Zeppilli D., Shimabukuro M. &amp; Sorensen M. V. 2021. Hadal mud dragons: First insight into the diversity of Kinorhyncha from the Atacama Trench. Frontiers in Marine Science 8: e 670735. https: // doi. org / 10.3389 / fmars. 2021.670735" type="journal article" year="2021">
Grzelak
<emphasis id="B969EA8C5B002A3CFB5D3F626052FAE8" box="[1259,1309,1334,1360]" italics="true" pageId="44" pageNumber="45">et al</emphasis>
. 2021
</bibRefCitation>
). It appears that also the arrangement and number of sensory spots show some differences. Unique for
<taxonomicName id="4C1D4D1D5B002A3CFF093FD6640FFA24" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[191,320,1410,1436]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFF093FD6640FFA24" box="[191,320,1410,1436]" italics="true" pageId="44" pageNumber="45">E. gandalfi</emphasis>
</taxonomicName>
is the presence of laterodorsal sensory spots on segment 2 but the lack of these structures in ventrolateral positions on segment 1, and in middorsal position on segment 10 as found in the other two species. In addition,
<taxonomicName id="4C1D4D1D5B002A3CFE1A3F9A673EFA50" authorityName="Adrianov &amp; Maiorova" authorityYear="2019" box="[428,625,1486,1512]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="ultraabyssalis">
<emphasis id="B969EA8C5B002A3CFE1A3F9A673EFA50" box="[428,625,1486,1512]" italics="true" pageId="44" pageNumber="45">E. ultraabyssalis</emphasis>
</taxonomicName>
lacks well-developed laterodorsal tubes on segment 10, which are present in both sexes and easy to visualize even with LM in
<taxonomicName id="4C1D4D1D5B002A3CFC363FA0614EF9B6" authorityName="Grzelak &amp; Sørensen" authorityYear="2022" box="[896,1025,1524,1550]" class="Cyclorhagida" family="Echinoderidae" genus="Echinoderes" kingdom="Animalia" order="Echinorhagata" pageId="44" pageNumber="45" phylum="Kinorhyncha" rank="species" species="gandalfi">
<emphasis id="B969EA8C5B002A3CFC363FA0614EF9B6" box="[896,1025,1524,1550]" italics="true" pageId="44" pageNumber="45">E. gandalfi</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>