treatments-xml/data/03/B4/24/03B42472FF84FF9EB38A41AF932EBCC3.xml

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<mods:title id="9DD48EADACBF9B98FECF7904D28C0A56">Two new species of Curculionichthys (Siluriformes: Loricariidae) from the rio Amazonas basin, Brazil</mods:title>
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<mods:namePart id="F6616248DF5D48F84DD1C2261999C8E3">Roxo, Fábio F.</mods:namePart>
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<mods:namePart id="61F27251A1A3F496622B59E6DCAE7753">Dias, Angelica C.</mods:namePart>
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<mods:namePart id="47F31CBAFF2D5A1B5FF30FCACA1D3345">Silva, Gabriel S. C.</mods:namePart>
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<emphasis id="B9694976FF84FF9BB38A41AF97D3BFEE" bold="true" box="[151,460,1238,1264]" italics="true" pageId="1">
<taxonomicName id="4C1DEEE7FF84FF9BB38A41AF97D9BFEE" authority="Roxo &amp; Dias &amp; Silva &amp; Oliveira, 2017" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[151,454,1238,1264]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="1" phylum="Chordata" rank="species" species="tukana" status="sp. nov.">Curculionichthys tukana</taxonomicName>
,
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<paragraph id="8BA29564FF84FF9BB38A41809725BE0C" blockId="1.[151,611,1238,1298]" box="[151,314,1273,1298]" pageId="1">
<figureCitation id="132689E1FF84FF9BB38A418096C7BE0C" box="[151,216,1273,1298]" captionStart="FIGURE 1" captionStartId="2.[151,250,1952,1974]" captionTargetBox="[204,1417,799,1930]" captionTargetId="figure@2.[204,1417,799,1930]" captionTargetPageId="2" captionText="FIGURE 1. Curculionichthys tukana, MZUSP 123010, 23.4 mm SL, holotype, Brazil, state of Maranhão, municipality of Estreito, ribeirão Bofe, rio Tocantins basin. Dorsal, lateral and ventral views." httpUri="https://zenodo.org/record/1039484/files/figure.png" pageId="1">Fig. 1</figureCitation>
, 
<tableCitation id="C69FA0DFFF84FF9BB3FB41809725BE0C" box="[230,314,1273,1298]" captionStart="TABLE 1" captionStartId="7.[151,239,584,607]" captionText="TABLE 1. Morphometric characters of C. tukana and C. itaim. SD = Standard deviation. N = number of analyzed specimens." pageId="1">Table 1</tableCitation>
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<emphasis id="B9694976FF84FF9BB38A4038975BBE48" box="[151,324,1345,1366]" italics="true" pageId="1">Curculionichthys</emphasis>
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<taxonomicNameLabel id="A25AF40DFF84FF9BB256403B979EBE49" box="[331,385,1346,1367]" pageId="1" rank="species">sp. n.</taxonomicNameLabel>
4: 
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<bibRefCitation id="EF8CE895FF84FF9BB2BC403B9451BE49" author="Roxo" box="[417,590,1345,1367]" pageId="1" refString="Roxo, F. F., Silva, G. S. C, Ochoa, L. E. &amp; Oliveira, C. (2015) Description of a new genus and three new species of Otothyrinae (Siluriformes, Loricariidae). Zookeys, 534, 103 - 134. https: // doi. org / 10.3897 / zookeys. 534.6169" type="journal article" year="2015">
Roxo 
<emphasis id="B9694976FF84FF9BB2C3403B940DBE48" box="[478,530,1345,1367]" italics="true" pageId="1">et al.</emphasis>
2015
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: 116
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(Fig. 8)
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</paragraph>
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<typeStatus id="54A62BC6FF84FF9BB38A40FD9719BE83" box="[151,262,1412,1437]" pageId="1" type="holotype">Holotype</typeStatus>
.
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<collectionCode id="ED0C0DA1FF84FF9BB20940FD976EBE82" box="[276,369,1412,1436]" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1">MZUSP</collectionCode>
<date id="FFA3B3A4FF84FF9BB26640FD97CDBE82" box="[379,466,1412,1436]" pageId="1" value="10-12-30">123010</date>
, 
<quantity id="4CE53881FF84FF9BB2FD40FD9459BE82" box="[480,582,1412,1436]" metricMagnitude="-2" metricUnit="m" metricValue="2.34" pageId="1" unit="mm" value="23.4">23.4 mm</quantity>
SL, 
<collectingCountry id="F30AD5F4FF84FF9BB19C40FD94D7BE83" box="[641,712,1412,1437]" name="Brazil" pageId="1">Brazil</collectingCountry>
, state of 
<collectingRegion id="49D95B86FF84FF9BB02940FD95B7BE83" box="[820,936,1412,1437]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
, municipality of Estreito, ribeirão Bofe, rio 
<collectingRegion id="49D95B86FF84FF9BB38A40DE971CBEDE" box="[151,259,1447,1472]" country="Brazil" name="Tocantins" pageId="1">Tocantins</collectingRegion>
basin, 
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, 
<geoCoordinate id="EE29F3A3FF84FF9BB2F040DE949BBEA1" box="[493,644,1447,1472]" direction="west" orientation="longitude" pageId="1" precision="15" value="-46.95">46°57’00” W</geoCoordinate>
, 
<date id="FFA3B3A4FF84FF9BB19240D19522BEDE" box="[655,829,1447,1472]" pageId="1" value="2005-03-22">22 March 2005</date>
, A. Akama &amp; E. Baena.
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<typeStatus id="54A62BC6FF84FF9BB3DA40B5975EBEFB" box="[199,321,1484,1509]" pageId="1" type="paratype">Paratypes</typeStatus>
.
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All from 
<collectingCountry id="F30AD5F4FF84FF9BB2A740B5941EBEFB" box="[442,513,1484,1509]" name="Brazil" pageId="1">Brazil</collectingCountry>
, rio 
<collectingRegion id="49D95B86FF84FF9BB12B40B594BDBEFB" box="[566,674,1484,1509]" country="Brazil" name="Tocantins" pageId="1">Tocantins</collectingRegion>
basin. LBP 22288, 
<specimenCode id="DBBB3D1FFF84FF9BB09540B59589BEFA" box="[904,918,1484,1508]" pageId="1">1</specimenCode>
, 
<quantity id="4CE53881FF84FF9BB0BB40B59213BEFA" box="[934,1036,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="2.2399999999999998" pageId="1" unit="mm" value="22.4">
<specimenCode id="DBBB3D1FFF84FF9BB0BB40B595C8BEFA" box="[934,983,1484,1508]" pageId="1">22.4</specimenCode>
mm
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SL, 1 c&amp;s, 
<quantity id="4CE53881FF84FF9BB78540B592E1BEFA" box="[1176,1278,1484,1508]" metricMagnitude="-2" metricUnit="m" metricValue="2.78" pageId="1" unit="mm" value="27.8">27.8 mm</quantity>
SL, collected with holotype
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB25E40899219BD16" ID-GBIF-Occurrence="1676480072" box="[323,1030,1519,1544]" collectionCode="MZUSP" country="Brazil" pageId="1" specimenCode="MZUSP 87452, 2, 20.2-22.6" specimenCount="1" stateProvince="Tocantins" typeStatus="paratype">
<specimenCode id="DBBB3D1FFF84FF9BB25E40899493BD16" box="[323,652,1519,1544]" pageId="1">
MZUSP 87452, 2, 
<quantity id="4CE53881FF84FF9BB10640969493BD16" box="[539,652,1519,1544]" metricMagnitude="-2" metricUnit="m" metricValue="2.1399999999999997" metricValueMax="2.26" metricValueMin="2.02" pageId="1" unit="mm" value="21.4" valueMax="22.6" valueMin="20.2">20.2–22.6</quantity>
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mm SL, collected with holotype
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB70C40899581BD51" ID-GBIF-Occurrence="1676480070" collectingDate="2005-03-25" collectionCode="MZUSP" collectorName="A. Akama &amp; E. G. Baena" country="Brazil" latitude="-7.045833" location="Feira Nova do Maranhao" longLatPrecision="21" longitude="-46.602497" municipality="rio Caraibas" pageId="1" specimenCode="MZUSP 87545, 24, 17.2-20.7" specimenCount="1" stateProvince="Maranhao" typeStatus="paratype">
<specimenCode id="DBBB3D1FFF84FF9BB70C40899377BD16" box="[1041,1384,1519,1544]" pageId="1">
MZUSP 87545, 24, 
<quantity id="4CE53881FF84FF9BB7E540899377BD16" box="[1272,1384,1520,1544]" metricMagnitude="-2" metricUnit="m" metricValue="1.895" metricValueMax="2.07" metricValueMin="1.72" pageId="1" unit="mm" value="18.95" valueMax="20.7" valueMin="17.2">17.2–20.7</quantity>
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mm SL, small unnamed tributary of 
<collectingMunicipality id="6BC60F1EFF84FF9BB101436D94B2BD33" box="[540,685,1556,1581]" pageId="1">rio Caraíbas</collectingMunicipality>
, municipality of 
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Feira Nova do 
<collectingRegion id="49D95B86FF84FF9BB725436D92B2BD33" box="[1080,1197,1556,1581]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB639436D9387BD33" box="[1316,1432,1556,1581]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
, 
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, 
<geoCoordinate id="EE29F3A3FF84FF9BB22D434E97D9BD51" box="[304,454,1591,1616]" direction="west" orientation="longitude" pageId="1" precision="15" value="-46.602497">46°36’09” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB2CC434E9462BD4E" box="[465,637,1591,1616]" pageId="1" value="2005-03-25">
<date id="FFA3B3A4FF84FF9BB2CC434E9462BD4E" box="[465,637,1591,1616]" pageId="1" value="2005-03-25">25 March 2005</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB194434194E3BD4E" box="[649,764,1591,1616]" pageId="1">A. Akama</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB00243419581BD51" box="[799,926,1592,1616]" pageId="1">E.G. Baena</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB0B443419227BD89" ID-GBIF-Occurrence="1676480069" collectingDate="2005-03-25" collectionCode="MZUSP" collectorName="A. Akama &amp; E. G. Baena" country="Brazil" county="Bom Jardim" latitude="-7.0680556" location="Feira Nova do Maranhao" longLatPrecision="21" longitude="-46.59722" municipality="rio Caraibas" pageId="1" specimenCode="MZUSP 87553, 7, 18.5-27.6" specimenCount="1" stateProvince="Maranhao" typeStatus="paratype">
<specimenCode id="DBBB3D1FFF84FF9BB0B4434192F3BD4E" box="[937,1260,1591,1616]" pageId="1">
MZUSP 87553, 7, 
<quantity id="4CE53881FF84FF9BB761434E92F3BD4E" box="[1148,1260,1591,1616]" metricMagnitude="-2" metricUnit="m" metricValue="2.305" metricValueMax="2.76" metricValueMin="1.85" pageId="1" unit="mm" value="23.05" valueMax="27.6" valueMin="18.5">18.5–27.6</quantity>
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mm SL, 1 c&amp;s, 
<quantity id="4CE53881FF84FF9BB38A432596E1BD6A" box="[151,254,1628,1652]" metricMagnitude="-2" metricUnit="m" metricValue="2.13" pageId="1" unit="mm" value="21.3">21.3 mm</quantity>
SL, córrego 
<collectingCounty id="62C3EDE8FF84FF9BB28543249438BD6B" box="[408,551,1628,1653]" pageId="1">Bom Jardim</collectingCounty>
, tributary of 
<collectingMunicipality id="6BC60F1EFF84FF9BB1DC43259550BD6B" box="[705,847,1628,1653]" pageId="1">rio Caraíbas</collectingMunicipality>
, municipality of 
<location id="8EC2C3BFFF84FF9BB70843259326BD6B" LSID="urn:lsid:plazi:treatment:03B42472FF84FF9EB38A41AF932EBCC3:8EC2C3BFFF84FF9BB70843259326BD6B" box="[1045,1337,1628,1653]" country="Brazil" county="Bom Jardim" latitude="-7.0680556" longLatPrecision="21" longitude="-46.59722" municipality="rio Caraibas" name="Feira Nova do Maranhao" pageId="1" stateProvince="Maranhao">
Feira Nova do 
<collectingRegion id="49D95B86FF84FF9BB7D843259326BD6B" box="[1221,1337,1628,1653]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB38A43069713BD86" box="[151,268,1663,1688]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
, 
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, 
<geoCoordinate id="EE29F3A3FF84FF9BB2A843069451BD89" box="[437,590,1663,1688]" direction="west" orientation="longitude" pageId="1" precision="15" value="-46.59722">46°35’50” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB14643069513BD86" box="[603,780,1663,1688]" pageId="1" value="2005-03-25">
<date id="FFA3B3A4FF84FF9BB14643069513BD86" box="[603,780,1663,1688]" pageId="1" value="2005-03-25">25 March 2005</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB00443F99591BD86" box="[793,910,1663,1688]" pageId="1">A. Akama</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB0AB43F99227BD89" box="[950,1080,1664,1688]" pageId="1">E.G. Baena</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB75943F99410BDFE" ID-GBIF-Occurrence="1676480080" collectingDate="2010-01-18" collectionCode="MZUSP" collectorName="M. V. Loeb &amp; H. R. Varella" country="Brazil" county="rio Parauapebas" latitude="-6.411389" location="Carajas" longLatPrecision="21" longitude="-50.195835" municipality="Canaa" pageId="1" specimenCode="MZUSP 105178, 6, 19.4-21.4" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<specimenCode id="DBBB3D1FFF84FF9BB75943F99383BD86" box="[1092,1436,1663,1688]" pageId="1">
MZUSP 105178, 6, 
<quantity id="4CE53881FF84FF9BB63143F99383BD86" box="[1324,1436,1664,1688]" metricMagnitude="-2" metricUnit="m" metricValue="2.04" metricValueMax="2.1399999999999997" metricValueMin="1.94" pageId="1" unit="mm" value="20.4" valueMax="21.4" valueMin="19.4">19.4–21.4</quantity>
</specimenCode>
mm SL, 
<collectingCounty id="62C3EDE8FF84FF9BB3E443DD97B1BDA3" box="[249,430,1700,1725]" pageId="1">rio Parauapebas</collectingCounty>
, municipality of 
<collectingMunicipality id="6BC60F1EFF84FF9BB17343DD94ABBDA2" box="[622,692,1700,1724]" pageId="1">Canaã</collectingMunicipality>
dos 
<location id="8EC2C3BFFF84FF9BB1F443DD955FBDA3" LSID="urn:lsid:plazi:treatment:03B42472FF84FF9EB38A41AF932EBCC3:8EC2C3BFFF84FF9BB1F443DD955FBDA3" box="[745,832,1700,1725]" country="Brazil" county="rio Parauapebas" latitude="-6.411389" longLatPrecision="21" longitude="-50.195835" municipality="Canaa" name="Carajas" pageId="1" stateProvince="Para">Carajás</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB0B843DD95C5BDA2" box="[933,986,1700,1724]" country="Brazil" name="Para" pageId="1">Pará</collectingRegion>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB0FB43DD926CBDA2" box="[998,1139,1700,1725]" direction="south" orientation="latitude" pageId="1" precision="15" value="-6.411389">06°24’41” S</geoCoordinate>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB79D43DD9309BDA2" box="[1152,1302,1700,1725]" direction="west" orientation="longitude" pageId="1" precision="15" value="-50.195835">50°11’45” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB63C43DD96CDBDFE" pageId="1" value="2010-01-18">
<date id="FFA3B3A4FF84FF9BB63C43DD96CDBDFE" pageId="1" value="2010-01-18">18 January 2010</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB3C343B19747BDFE" box="[222,344,1735,1760]" pageId="1">M.V. Loeb</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB26343B19410BDFE" box="[382,527,1735,1760]" pageId="1">H.R. Varella</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB10743B192B5BC1B" ID-GBIF-Occurrence="1676480071" collectingDate="2010-06-21" collectionCode="MZUSP" collectorName="M. Loeb &amp; H. R. Varella" country="Brazil" county="rio Parauapebas" latitude="-6.412778" location="Carajas" longLatPrecision="21" longitude="-50.026665" municipality="Canaa" pageId="1" specimenCode="MZUSP 106572, 3, 14.7-20.7" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<specimenCode id="DBBB3D1FFF84FF9BB10743B1956EBDFE" box="[538,881,1735,1760]" pageId="1">
MZUSP 106572, 3, 
<quantity id="4CE53881FF84FF9BB01C43B1956EBDFE" box="[769,881,1736,1760]" metricMagnitude="-2" metricUnit="m" metricValue="1.77" metricValueMax="2.07" metricValueMin="1.47" pageId="1" unit="mm" value="17.7" valueMax="20.7" valueMin="14.7">14.7–20.7</quantity>
</specimenCode>
mm SL, 
<collectingCounty id="62C3EDE8FF84FF9BB0C343BE928BBDFE" box="[990,1172,1735,1760]" pageId="1">rio Parauapebas</collectingCounty>
, municipality of 
<collectingMunicipality id="6BC60F1EFF84FF9BB64B43B19383BDFE" box="[1366,1436,1736,1760]" pageId="1">Canaã</collectingMunicipality>
dos 
<location id="8EC2C3BFFF84FF9BB3D943959704BC1B" LSID="urn:lsid:plazi:treatment:03B42472FF84FF9EB38A41AF932EBCC3:8EC2C3BFFF84FF9BB3D943959704BC1B" box="[196,283,1772,1797]" country="Brazil" county="rio Parauapebas" latitude="-6.412778" longLatPrecision="21" longitude="-50.026665" municipality="Canaa" name="Carajas" pageId="1" stateProvince="Para">Carajás</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB260439597ADBC1A" box="[381,434,1772,1796]" country="Brazil" name="Para" pageId="1">Pará</collectingRegion>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB2A043959456BC1A" box="[445,585,1772,1797]" direction="south" orientation="latitude" pageId="1" precision="15" value="-6.412778">06°24’46” S</geoCoordinate>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB14B439594F3BC1A" box="[598,748,1772,1797]" direction="west" orientation="longitude" pageId="1" precision="15" value="-50.026665">50°01’36” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB1EB43959591BC1A" box="[758,910,1772,1796]" pageId="1" value="2010-06-21">
<date id="FFA3B3A4FF84FF9BB1EB43959591BC1A" box="[758,910,1772,1796]" pageId="1" value="2010-06-21">21 June 2010</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB085439495E6BC1B" box="[920,1017,1772,1797]" pageId="1">M. Loeb</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB706439492B5BC1B" box="[1051,1194,1772,1797]" pageId="1">H.R. Varella</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB7A94395944DBC53" ID-GBIF-Occurrence="1676480078" collectingDate="2010-06-19" collectionCode="MZUSP" collectorName="M. V. Loeb &amp; H. R. Varella" country="Brazil" county="rio Sossego" latitude="-6.4066668" location="Carajas" longLatPrecision="21" longitude="-50.045555" municipality="Canaa" pageId="1" specimenCode="MZUSP 106584, 14, 14.8-20.8" specimenCount="1" stateProvince="Para" typeStatus="paratype">
<specimenCode id="DBBB3D1FFF84FF9BB7A943959718BC36" pageId="1">
MZUSP 106584, 14, 
<quantity id="4CE53881FF84FF9BB38A42699718BC36" box="[151,263,1808,1832]" metricMagnitude="-2" metricUnit="m" metricValue="1.78" metricValueMax="2.08" metricValueMin="1.48" pageId="1" unit="mm" value="17.8" valueMax="20.8" valueMin="14.8">14.8–20.8</quantity>
</specimenCode>
mm SL, 
<collectingCounty id="62C3EDE8FF84FF9BB269427697E5BC36" box="[372,506,1807,1832]" pageId="1">rio Sossego</collectingCounty>
, municipality of 
<collectingMunicipality id="6BC60F1EFF84FF9BB1A34269951BBC36" box="[702,772,1808,1832]" pageId="1">Canaã</collectingMunicipality>
dos 
<location id="8EC2C3BFFF84FF9BB0214276958CBC36" LSID="urn:lsid:plazi:treatment:03B42472FF84FF9EB38A41AF932EBCC3:8EC2C3BFFF84FF9BB0214276958CBC36" box="[828,915,1807,1832]" country="Brazil" county="rio Sossego" latitude="-6.4066668" longLatPrecision="21" longitude="-50.045555" municipality="Canaa" name="Carajas" pageId="1" stateProvince="Para">Carajás</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB0E14269922EBC36" box="[1020,1073,1808,1832]" country="Brazil" name="Para" pageId="1">Pará</collectingRegion>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB723427692D3BC36" box="[1086,1228,1807,1832]" direction="south" orientation="latitude" pageId="1" precision="15" value="-6.4066668">06°24’24” S</geoCoordinate>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB7C64276936CBC39" box="[1243,1395,1807,1832]" direction="west" orientation="longitude" pageId="1" precision="15" value="-50.045555">50°02’44” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB69D42699710BC52" pageId="1" value="2010-06-19">
<date id="FFA3B3A4FF84FF9BB69D42699710BC52" pageId="1" value="2010-06-19">19 June 2010</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB200424C9787BC53" box="[285,408,1844,1869]" pageId="1">M.V. Loeb</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB2DD424C944DBC53" box="[448,594,1844,1869]" pageId="1">H.R. Varella</collectorName>
</materialsCitation>
. 
<materialsCitation id="3B759F39FF84FF9BB142424C97ADBC8A" ID-GBIF-Occurrence="1676480079" collectingDate="2005-03-25" collectorName="A. Akama &amp; E. G. Baena." country="Brazil" county="Bom Jardim" latitude="-7.0680556" location="Feira Nova do Maranhao" longLatPrecision="21" longitude="-46.59722" municipality="rio Caraibas" pageId="1" specimenCode="2, 23.7" specimenCount="1" stateProvince="Maranhao" typeStatus="paratype">
NUP 19280, 
<specimenCode id="DBBB3D1FFF84FF9BB1EA424D9518BC52" box="[759,775,1844,1868]" pageId="1">2</specimenCode>
, 
<specimenCode id="DBBB3D1FFF84FF9BB008424D955ABC52" box="[789,837,1844,1868]" pageId="1">23.7</specimenCode>
–24.0 mm SL, córrego 
<collectingCounty id="62C3EDE8FF84FF9BB748424C92FBBC53" box="[1109,1252,1844,1869]" pageId="1">Bom Jardim</collectingCounty>
, tributary of 
<collectingMunicipality id="6BC60F1EFF84FF9BB660424D96E3BC6E" pageId="1">rio Caraíbas</collectingMunicipality>
, municipality of 
<location id="8EC2C3BFFF84FF9BB2A5422E94CEBC6E" LSID="urn:lsid:plazi:treatment:03B42472FF84FF9EB38A41AF932EBCC3:8EC2C3BFFF84FF9BB2A5422E94CEBC6E" box="[440,721,1879,1904]" country="Brazil" county="Bom Jardim" latitude="-7.0680556" longLatPrecision="21" longitude="-46.59722" municipality="rio Caraibas" name="Feira Nova do Maranhao" pageId="1" stateProvince="Maranhao">
Feira Nova do 
<collectingRegion id="49D95B86FF84FF9BB140422E94CEBC6E" box="[605,721,1879,1904]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB02E422E95B7BC6E" box="[819,936,1879,1904]" country="Brazil" name="Maranhao" pageId="1">Maranhão</collectingRegion>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB0AF422E9220BC6E" box="[946,1087,1879,1904]" direction="south" orientation="latitude" pageId="1" precision="15" value="-7.0680556">07°04’05” S</geoCoordinate>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB756422E92FEBC71" box="[1099,1249,1879,1904]" direction="west" orientation="longitude" pageId="1" precision="15" value="-46.59722">46°35’50” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB7F6422E9387BC6E" box="[1259,1432,1879,1904]" pageId="1" value="2005-03-25">
<date id="FFA3B3A4FF84FF9BB7F6422E9387BC6E" box="[1259,1432,1879,1904]" pageId="1" value="2005-03-25">25 March 2005</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB38A42059715BC8B" box="[151,266,1916,1941]" pageId="1">A. Akama</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB233420597ADBC8A" box="[302,434,1916,1940]" pageId="1">E.G. Baena.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C307C6EFFF84FF9BB3DA42D99379BCC3" pageId="1" type="description">
<paragraph id="8BA29564FF84FF9BB3DA42D99379BCC3" blockId="1.[151,1437,1412,2013]" pageId="1">
<materialsCitation id="3B759F39FF84FF9BB3DA42D99379BCC3" ID-GBIF-Occurrence="1676480075" collectingDate="2011-11-29" collectionCode="MZUSP" collectorName="A. M. Zanata &amp; P. C. A. Cardoso &amp; F. A. G. Melo &amp; O. T. Oyakawa." latitude="-13.856945" location="Cavalcante" longLatPrecision="21" longitude="-47.600277" municipality="rio Montes Claros" pageId="1" specimenCode="MZUSP 114384, 1, 20.5" specimenCount="1" stateProvince="Goias">
<emphasis id="B9694976FF84FF9BB3DA42D99758BCA7" bold="true" box="[199,327,1952,1977]" pageId="1">Non-types.</emphasis>
<specimenCode id="DBBB3D1FFF84FF9BB24D42D99413BCA6" box="[336,524,1952,1976]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1">MZUSP 114384</specimenCode>
, 
<specimenCode id="DBBB3D1FFF84FF9BB10442D99438BCA6" box="[537,551,1952,1976]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1">1</specimenCode>
, 
<quantity id="4CE53881FF84FF9BB12A42D99482BCA6" box="[567,669,1952,1977]" metricMagnitude="-2" metricUnit="m" metricValue="2.05" pageId="1" unit="mm" value="20.5">
<specimenCode id="DBBB3D1FFF84FF9BB12A42D99477BCA7" box="[567,616,1952,1977]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1">20.5</specimenCode>
mm
</quantity>
SL, 
<collectingMunicipality id="6BC60F1EFF84FF9BB1CB42D995B8BCA7" box="[726,935,1952,1977]" pageId="1">rio Montes Claros</collectingMunicipality>
, municipality of 
<location id="8EC2C3BFFF84FF9BB77642D992F5BCA7" LSID="urn:lsid:plazi:treatment:03B42472FF84FF9EB38A41AF932EBCC3:8EC2C3BFFF84FF9BB77642D992F5BCA7" box="[1131,1258,1952,1977]" latitude="-13.856945" longLatPrecision="21" longitude="-47.600277" municipality="rio Montes Claros" name="Cavalcante" pageId="1" stateProvince="Goias">Cavalcante</location>
, state of 
<collectingRegion id="49D95B86FF84FF9BB64942D99387BCA7" box="[1364,1432,1952,1977]" country="Brazil" name="Goias" pageId="1">Goiás</collectingRegion>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB38A42BD973BBCC2" box="[151,292,1988,2013]" direction="south" orientation="latitude" pageId="1" precision="15" value="-13.856945">13°51’25” S</geoCoordinate>
, 
<geoCoordinate id="EE29F3A3FF84FF9BB22D42BD97D8BCC2" box="[304,455,1988,2013]" direction="west" orientation="longitude" pageId="1" precision="15" value="-47.600277">47°36’01” W</geoCoordinate>
, 
<collectingDate id="EFE74A4CFF84FF9BB2CF42BD94B3BCC2" box="[466,684,1988,2013]" pageId="1" value="2011-11-29">
<date id="FFA3B3A4FF84FF9BB2CF42BD94B3BCC2" box="[466,684,1988,2013]" pageId="1" value="2011-11-29">29 November 2011</date>
</collectingDate>
, 
<collectorName id="26E8F0B2FF84FF9BB1AA42BD9557BCC2" box="[695,840,1988,2012]" pageId="1">A.M. Zanata</collectorName>
, 
<collectorName id="26E8F0B2FF84FF9BB04942BD921CBCC3" box="[852,1027,1988,2013]" pageId="1">P.C.A. Cardoso</collectorName>
, 
<collectorName id="26E8F0B2FF84FF9BB71342BD9287BCC3" box="[1038,1176,1988,2013]" pageId="1">F.A.G. Melo</collectorName>
&amp; 
<collectorName id="26E8F0B2FF84FF9BB7A142BD9379BCC3" box="[1212,1382,1988,2013]" pageId="1">O.T. Oyakawa.</collectorName>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C307C6EFFF87FF98B3DA45EE97E7B9F9" pageId="2" type="diagnosis">
<paragraph id="8BA29564FF87FF98B3DA45EE97E7B9F9" blockId="2.[151,1436,151,743]" pageId="2">
<emphasis id="B9694976FF87FF98B3DA45EE975EBBAE" bold="true" box="[199,321,151,176]" pageId="2">Diagnosis.</emphasis>
<taxonomicName id="4C1DEEE7FF87FF98B25545E19443BBAE" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[328,604,152,176]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF87FF98B25545E19443BBAE" box="[328,604,152,176]" italics="true" pageId="2">Curculionichthys tukana</emphasis>
</taxonomicName>
differs from all congeners, except 
<taxonomicName id="4C1DEEE7FF87FF98B0FB45E19272BBAE" authorityName="Silva, Roxo, Melo &amp; Oliveira" authorityYear="2016" box="[998,1133,152,176]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="karipuna">
<emphasis id="B9694976FF87FF98B0FB45E19272BBAE" box="[998,1133,152,176]" italics="true" pageId="2">C. karipuna</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1DEEE7FF87FF98B7B945E1933BBBAE" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" box="[1188,1316,152,176]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="oliveirai">
<emphasis id="B9694976FF87FF98B7B945E1933BBBAE" box="[1188,1316,152,176]" italics="true" pageId="2">C. oliveirai</emphasis>
</taxonomicName>
by having one abdominal median plate series, 
<figureCitation id="132689E1FF87FF98B13845C09465BBCF" box="[549,634,185,210]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="2">Fig. 2A</figureCitation>
(
<emphasis id="B9694976FF87FF98B19445C394B7BBCF" box="[649,680,186,209]" italics="true" pageId="2">vs.</emphasis>
three or more abdominal medial plates series); from 
<taxonomicName id="4C1DEEE7FF87FF98B7E945C09366BBCF" authorityName="Silva, Roxo, Melo &amp; Oliveira" authorityYear="2016" box="[1268,1401,185,209]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="karipuna">
<emphasis id="B9694976FF87FF98B7E945C09366BBCF" box="[1268,1401,185,209]" italics="true" pageId="2">C. karipuna</emphasis>
</taxonomicName>
by the absence of an irregular concentration of chromatophores that entirely cover the anal-fin origin and adjacent region, and distal portion of the first unbranched anal-fin ray (
<emphasis id="B9694976FF87FF98B04A45849569BA0A" box="[855,886,253,276]" italics="true" pageId="2">vs.</emphasis>
presence of such pigmentation pattern); from 
<taxonomicName id="4C1DEEE7FF87FF98B69F458596E9BA2B" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="oliveirai">
<emphasis id="B9694976FF87FF98B69F458596E9BA2B" italics="true" pageId="2">C. oliveirai</emphasis>
</taxonomicName>
, and also from 
<taxonomicName id="4C1DEEE7FF87FF98B2B344679459BA2B" baseAuthorityName="Britski &amp; Garavello" baseAuthorityYear="2003" box="[430,582,286,309]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="insperatus">
<emphasis id="B9694976FF87FF98B2B344679459BA2B" box="[430,582,286,309]" italics="true" pageId="2">C. insperatus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1DEEE7FF87FF98B14B446794A0BA2B" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" box="[598,703,286,309]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="paresi">
<emphasis id="B9694976FF87FF98B14B446794A0BA2B" box="[598,703,286,309]" italics="true" pageId="2">C. paresi</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1DEEE7FF87FF98B1E744679598BA2B" authorityName="Roxo, Silva, Ochoa &amp; Oliveira" authorityYear="2015" box="[762,903,286,309]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="sagarana">
<emphasis id="B9694976FF87FF98B1E744679598BA2B" box="[762,903,286,309]" italics="true" pageId="2">C. sagarana</emphasis>
</taxonomicName>
, by having seven to eight plates in abdominal lateral plates series, 
<figureCitation id="132689E1FF87FF98B265444797D2BA49" box="[376,461,318,343]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="2">Fig. 2A</figureCitation>
(
<emphasis id="B9694976FF87FF98B2C1444697E4BA48" box="[476,507,319,342]" italics="true" pageId="2">vs.</emphasis>
four to six plates in abdominal lateral plates series); from 
<taxonomicName id="4C1DEEE7FF87FF98B79F444692C4BA48" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[1154,1243,319,342]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="itaim">
<emphasis id="B9694976FF87FF98B79F444692C4BA48" box="[1154,1243,319,342]" italics="true" pageId="2">C. itaim</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C1DEEE7FF87FF98B7F444469383BA48" baseAuthorityName="Britski &amp; Garavello" baseAuthorityYear="2007" box="[1257,1436,318,342]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="luteofrenatus">
<emphasis id="B9694976FF87FF98B7F444469383BA48" box="[1257,1436,318,342]" italics="true" pageId="2">C. luteofrenatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1DEEE7FF87FF98B3DA4419976EBA66" baseAuthorityName="Martins &amp; Langeani" baseAuthorityYear="2012" box="[199,369,352,376]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="piracanjuba">
<emphasis id="B9694976FF87FF98B3DA4419976EBA66" box="[199,369,352,376]" italics="true" pageId="2">C. piracanjuba</emphasis>
</taxonomicName>
by having papillae aligned in series that extends from the distal portion of lower lip to dentary (
<emphasis id="B9694976FF87FF98B3BD44FB96A8BA87" box="[160,183,386,409]" italics="true" pageId="2">vs</emphasis>
. all papillae randomly distributed throughout lower lip, see figure 4 of Silva 
<emphasis id="B9694976FF87FF98B70944FB925BBA87" box="[1044,1092,385,409]" italics="true" pageId="2">et al</emphasis>
. [2016] for illustration of both character states). The new species can be further diagnosed from 
<taxonomicName id="4C1DEEE7FF87FF98B06044DA920BBAA4" baseAuthorityName="Britski &amp; Garavello" baseAuthorityYear="2003" box="[893,1044,419,442]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="insperatus">
<emphasis id="B9694976FF87FF98B06044DA920BBAA4" box="[893,1044,419,442]" italics="true" pageId="2">C. insperatus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1DEEE7FF87FF98B75144DA92D2BAA5" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" box="[1100,1229,419,443]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="oliveirai">
<emphasis id="B9694976FF87FF98B75144DA92D2BAA5" box="[1100,1229,419,443]" italics="true" pageId="2">C. oliveirai</emphasis>
</taxonomicName>
by the absence of large conspicuous odontodes forming rows on head and trunk (
<emphasis id="B9694976FF87FF98B09744BC95BEBAC2" box="[906,929,453,476]" italics="true" pageId="2">vs</emphasis>
. presence of large, conspicuous odontodes forming rows on the head); from 
<taxonomicName id="4C1DEEE7FF87FF98B117449F9496BAE3" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" box="[522,649,485,509]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="oliveirai">
<emphasis id="B9694976FF87FF98B117449F9496BAE3" box="[522,649,485,509]" italics="true" pageId="2">C. oliveirai</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C1DEEE7FF87FF98B1A3449F955BBAE3" authorityName="Roxo, Silva, Ochoa &amp; Oliveira" authorityYear="2015" box="[702,836,486,509]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="coxipone">
<emphasis id="B9694976FF87FF98B1A3449F955BBAE3" box="[702,836,486,509]" italics="true" pageId="2">C. coxipone</emphasis>
</taxonomicName>
by having the anterior profile of the head pointed (
<emphasis id="B9694976FF87FF98B660449F938BBAE3" box="[1405,1428,486,509]" italics="true" pageId="2">vs</emphasis>
. anterior profile of head rounded); from 
<taxonomicName id="4C1DEEE7FF87FF98B14F477E94A8B900" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" box="[594,695,519,542]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="paresi">
<emphasis id="B9694976FF87FF98B14F477E94A8B900" box="[594,695,519,542]" italics="true" pageId="2">C. paresi</emphasis>
</taxonomicName>
by having 12–18 premaxillary teeth (
<emphasis id="B9694976FF87FF98B743477E926AB900" box="[1118,1141,519,542]" italics="true" pageId="2">vs</emphasis>
. 6–10) and 10–16 dentary teeth (
<emphasis id="B9694976FF87FF98B3C0475096EBB95E" box="[221,244,553,576]" italics="true" pageId="2">vs</emphasis>
. 4–7); from 
<taxonomicName id="4C1DEEE7FF87FF98B29D475197FBB95E" authorityName="Roxo, Silva, Ochoa &amp; Oliveira" authorityYear="2015" box="[384,484,552,576]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="sabaji">
<emphasis id="B9694976FF87FF98B29D475197FBB95E" box="[384,484,552,576]" italics="true" pageId="2">C. sabaji</emphasis>
</taxonomicName>
by the absence of dark-brown spots scattered over the body (
<emphasis id="B9694976FF87FF98B790475092BBB95E" box="[1165,1188,553,576]" italics="true" pageId="2">vs</emphasis>
. the presence of darkbrown spots); from 
<taxonomicName id="4C1DEEE7FF87FF98B29D473097F5B97F" baseAuthorityName="Roxo, Zawadzki &amp; Troy" baseAuthorityYear="2014" box="[384,490,585,609]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="paresi">
<emphasis id="B9694976FF87FF98B29D473097F5B97F" box="[384,490,585,609]" italics="true" pageId="2">C. paresi</emphasis>
</taxonomicName>
by the lack of contrasting dark spots at the anterodorsal region of body (
<emphasis id="B9694976FF87FF98B64D47339370B97F" box="[1360,1391,586,609]" italics="true" pageId="2">vs.</emphasis>
the presence of such pigmentation pattern). 
<taxonomicName id="4C1DEEE7FF87FF98B16347129587B99D" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[638,920,619,643]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF87FF98B16347129587B99D" box="[638,920,619,643]" italics="true" pageId="2">Curculionichthys tukana</emphasis>
</taxonomicName>
can additionally be distinguished from 
<taxonomicName id="4C1DEEE7FF87FF98B69F4712973FB9BA" baseAuthorityName="Martins &amp; Langeani" baseAuthorityYear="2012" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="piracanjuba">
<emphasis id="B9694976FF87FF98B69F4712973FB9BA" italics="true" pageId="2">C. piracanjuba</emphasis>
</taxonomicName>
by presence of odontodes forming aligned rows, more evident in the dorsal portion of head and in the lateral portion of caudal peduncle (
<emphasis id="B9694976FF87FF98B10147D79424B9DB" box="[540,571,686,709]" italics="true" pageId="2">vs.</emphasis>
odontodes not forming rows on the supraoccipital, on the compound pterotic, and on the lateral plates series).
</paragraph>
</subSubSection>
<caption id="DF62C5ECFF87FF98B38A42D995BEBCC6" httpUri="https://zenodo.org/record/1039484/files/figure.png" pageId="2" targetBox="[204,1417,799,1930]" targetPageId="2">
<paragraph id="8BA29564FF87FF98B38A42D995BEBCC6" blockId="2.[151,1435,1952,2008]" pageId="2">
<emphasis id="B9694976FF87FF98B38A42D99709BCAB" bold="true" box="[151,278,1952,1974]" pageId="2">FIGURE 1.</emphasis>
<taxonomicName id="4C1DEEE7FF87FF98B20242D89404BCA8" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[287,539,1953,1974]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="2" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF87FF98B20242D89404BCA8" box="[287,539,1953,1974]" italics="true" pageId="2">Curculionichthys tukana</emphasis>
</taxonomicName>
, MZUSP 123010, 23.4 mm SL, holotype, Brazil, state of Maranhão, municipality of Estreito, ribeirão Bofe, rio Tocantins basin. Dorsal, lateral and ventral views.
</paragraph>
</caption>
<subSubSection id="C307C6EFFF86FF9EB3DA45EE9482BC36" lastPageId="4" pageId="3" type="description">
<paragraph id="8BA29564FF86FF99B3DA45EE970DB906" blockId="3.[151,1437,151,896]" pageId="3">
<emphasis id="B9694976FF86FF99B3DA45EE9746BBAE" bold="true" box="[199,345,151,176]" pageId="3">Description.</emphasis>
Morphometric and meristic data are given in 
<tableCitation id="C69FA0DFFF86FF99B06945EE95D4BBAE" box="[884,971,151,176]" captionStart="TABLE 1" captionStartId="7.[151,239,584,607]" captionText="TABLE 1. Morphometric characters of C. tukana and C. itaim. SD = Standard deviation. N = number of analyzed specimens." pageId="3">Table 1</tableCitation>
. Small sized loricariid (
<quantity id="4CE53881FF86FF99B7FE45E1937EBBB1" box="[1251,1377,152,176]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" metricValueMax="2.4" metricValueMin="1.8" pageId="3" unit="mm" value="21.0" valueMax="24.0" valueMin="18.0">18–24 mm</quantity>
SL). Head and snout slightly rounded, snout tip pointed anteriorly (48–56% HL). Area around tip of snout completely covered by odontodes. Snout elongate (48–57% HL); anterior region slightly depressed. Nostril small, almost reaching same diameter of eyes. Dorsal profile of head ascending approximately 45° to parieto-supraoccipital; slightly concavity between nostrils. Eyes relatively small (12–17% HL), dorsolaterally positioned, just posterior of midpoint of head. Superior margin of orbits not elevated. Presence of inconspicuous odontodes on suboperculum. Mouth moderate in size; oral disk ellipsoid with papillae randomly distributed. Lips without odontodes; lower lip larger than upper lip, almost reaching cleithrum; its border fringed; lower lip inner surface covered with small papillae, similar in size. Maxillary barbel short, slender and not adnate to lower lip. Teeth slender with two symmetrical cusps; central cusp larger than lateral cusps. Premaxillary teeth 10–18 (mode 12). Dentary teeth 12–18 (mode 12).
</paragraph>
<paragraph id="8BA29564FF86FF99B3DA475C96E0B89E" blockId="3.[151,1437,151,896]" pageId="3">Lower surface of head naked. Head lacking ridges. Supraoccipital process not elevated and without conspicuous tuft of odontodes in specimens of all sizes. Predorsal region without ridges. In dorsal view, body elongated and slightly depressed on caudal peduncle. Greatest body width at cleithral region (23–26% SL), progressively narrowing towards snout tip and caudal-fin insertion. Dorsal profile of head and trunk covered by dermal plates, except on area around dorsal-fin insertion. In lateral view, body depressed. Body convex from snout tip to posterior margin of parieto-supraoccipital; slight concave from that point to dorsal-fin origin; slightly concave and descending from dorsal-fin origin to first upper procurrent caudal-fin ray origin, elevating posteriorly to insertion of caudal fin. Greatest body depth at unbranched dorsal-fin ray insertion (16–19% SL). Cleithrum and coracoid exposed in ventral view, covered by odontodes. Arrector fossa partially enclosed by ventral lamina of coracoid.</paragraph>
<caption id="DF62C5ECFF86FF99B38A4222946CBCF5" httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="3" targetBox="[180,1404,939,1867]" targetPageId="3">
<paragraph id="8BA29564FF86FF99B38A4222946CBCF5" blockId="3.[151,1437,1882,2027]" pageId="3">
<emphasis id="B9694976FF86FF99B38A42229709BC6E" bold="true" box="[151,278,1883,1905]" pageId="3">FIGURE 2.</emphasis>
<taxonomicName id="4C1DEEE7FF86FF99B23D42229403BC6E" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[288,540,1883,1904]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="3" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF86FF99B23D42229403BC6E" box="[288,540,1883,1904]" italics="true" pageId="3">Curculionichthys tukana</emphasis>
</taxonomicName>
, LBP 22288, 27.8 mm SL. 
<emphasis id="B9694976FF86FF99B0544223957BBC6F" bold="true" box="[841,868,1882,1905]" pageId="3">A)</emphasis>
Ventral view of abdominal plates; 
<emphasis id="B9694976FF86FF99B7C1422392E9BC6F" bold="true" box="[1244,1270,1882,1905]" pageId="3">B)</emphasis>
Spinelet shape. Cranial bones and dermal plates of the head in 
<emphasis id="B9694976FF86FF99B18B420194AEBC91" bold="true" box="[662,689,1912,1935]" pageId="3">C)</emphasis>
dorsal and 
<emphasis id="B9694976FF86FF99B02E42019552BC91" bold="true" box="[819,845,1912,1935]" pageId="3">D)</emphasis>
dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io1–4 = infraorbitals; pr1–4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp1–2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D).
</paragraph>
</caption>
<paragraph id="8BA29564FF81FF9EB3DA45EE9219BA7B" blockId="4.[151,1437,151,2013]" pageId="4">Ventral profile straight and descending from snout tip to opercular region; convex from opercular region to anal-fin origin; slightly concave from that point to lower procurrent caudal-fin ray origin. Lateral surface of body entirely covered by plates; mid-dorsal plate series poorly developed (six plates), reaching middle of dorsal-fin base; median plate series uninterrupted in median portion of body and perforated by lateral line 24–26 plates (mode 25); mid-ventral series of lateral plates developed and almost reaching middle of caudal peduncle (13 plates). Body plates covered with minute, uniformly sized and evenly distributed odontodes.</paragraph>
<paragraph id="8BA29564FF81FF9EB3DA44169503B906" blockId="4.[151,1437,151,2013]" pageId="4">
Ventral portion of body totally covered by plates, except on ventral part of head and around pectoral- and pelvic-fin insertions, and urogenital opening. Abdomen entirely covered by bony plates. Lateral plates series larger and forming regular series of seven to eight elongate plates on each side; median plate series smaller, arranged in single series from distal portion of cleithrum to anal plate series on anterior portion of urogenital opening; anal plates series formed by four to five large plates (
<figureCitation id="132689E1FF81FF9EB1A844869510B906" box="[693,783,511,536]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="4">Fig. 2A</figureCitation>
).
</paragraph>
<paragraph id="8BA29564FF81FF9EB3DA475D9254B826" blockId="4.[151,1437,151,2013]" pageId="4">
Dorsal-fin rays ii,7; its origin slightly anterior through pelvic-fin origin. Dorsal-fin unbranched ray slightly convex. Tip of adpressed dorsal-fin rays surpassing anal fin unbranched ray insertion. Dorsal-fin spinelet small and 
<emphasis id="B9694976FF81FF9EB38A471496BAB99A" box="[151,165,621,644]" italics="true" pageId="4">V</emphasis>
-shaped, 
<figureCitation id="132689E1FF81FF9EB21047159777B99A" box="[269,360,620,645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="4">Fig. 2B</figureCitation>
; lock mechanism functional. Pectoral-fin rays i,6; its tip almost reaching pelvic-fin insertion when adpressed. Pectoral-fin axillary slit present above pectoral-fin insertion. Pelvic-fin rays i,5; its distal margin straight to slightly convex; tip of adpressed pelvic fin reaching anal-fin origin in males, but not in females. Absent of adipose fin. Anal-fin rays i,5; distal margin slightly convex. Caudal-fin rays i, 
<date id="FFA3B3A4FF81FF9EB74647AE929BB9EE" box="[1115,1156,727,752]" bridgedPair="-" pageId="4">7–i</date>
,7; slightly emarginated; unbranched rays of same size. Rays of all fins covered with pointed odontodes. Least body depth at caudal peduncle. Caudal peduncle ellipsoid in cross section, rounded dorsally and ventrally.
</paragraph>
<paragraph id="8BA29564FF81FF9EB3DA463D9765BF46" blockId="4.[151,1437,151,2013]" pageId="4">
Neurocranium presented in 
<figureCitation id="132689E1FF81FF9EB10C463D9489B842" box="[529,662,836,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="4">Fig. 2C, D</figureCitation>
. In dorsal view (
<figureCitation id="132689E1FF81FF9EB074463D95D7B842" box="[873,968,836,861]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="4">Fig. 2C</figureCitation>
), parieto-supraoccipital (soc) octagonal, bordered by compound pterotic (cpt) and sphenotic (sp) laterally and by frontal (f) anteriorly. Snout composed by two rostral square-shaped plates (r); two prenasals square-shape, and two rectangular-shape plates (pn) between nares, and two nasal bones (n) slightly chevron-shape, forming superior margin of orbit, bordered by infra-orbital (io2) and posteriorly by frontals (f). Upper region of neurocranium composed by three large bones: compound pterotic (cpt), parieto-supraoccipital (soc) and by frontal (f), largest bones of head; and one small bone: sphenotic (sp), bordering anteriorly by frontal (f), posteriorly by compound pterotic (cpt) and laterally by parietosupraoccipital (soc).
</paragraph>
<paragraph id="8BA29564FF81FF9EB3DA411C9482BE2E" blockId="4.[151,1437,151,2013]" pageId="4">
In lateral view (
<figureCitation id="132689E1FF81FF9EB261411D97C9BF62" box="[380,470,1124,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1883,1905]" captionTargetBox="[180,1404,939,1867]" captionTargetId="figure@3.[180,1404,939,1867]" captionTargetPageId="3" captionText="FIGURE 2. Curculionichthys tukana, LBP 22288, 27.8 mm SL. A) Ventral view of abdominal plates; B) Spinelet shape. Cranial bones and dermal plates of the head in C) dorsal and D) dorsolateral views. Abbreviations are: cpt = compound pterotic; f = frontal; io 1 – 4 = infraorbitals; pr 1 – 4 postrostral plates; n = nasal; op = opercle; pf = prefrontal; pop = preopercle; pn = prenasal; r = rostral plate; cp 1 – 2 = subocular cheek plate; sp = sphenotic; soc = parieto-supraoccipital; spop = supraopercular plate. Scale bars: 0.5 mm (B); 1 mm (A, C, D)." httpUri="https://zenodo.org/record/1039486/files/figure.png" pageId="4">Fig. 2D</figureCitation>
), posterior rostrum plates large, pr1 triangular-shaped, pr2 small, rectangular-shaped, pr3 larger, pentagon-shaped and pr4 triangular-shaped. Infraorbital plate series incomplete (io1-io4), io1 and io2 largest; io3 and io4 smallest, forming inferior orbital margin of eyes. Opercular series complete, preopercle (pop) and supraopercular (spop) reduced, rectangular-shaped, covered by laterosensory canal; opercle (op) rounded; two subocular cheek plates, cp1 largest, triangular-shaped; cp2, smaller; bordering by cp1, preopercle (pop) and opercle (op), forming posterior lateral margin of head.
</paragraph>
<paragraph id="8BA29564FF81FF9EB3DA40459405BD86" blockId="4.[151,1437,151,2013]" pageId="4">
<emphasis id="B9694976FF81FF9EB3DA40459787BE4B" bold="true" box="[199,408,1340,1365]" pageId="4">Color in Alcohol.</emphasis>
In dorsal view, light brown background; trunk with two longitudinal light bars from predorsal region to caudal peduncle. Three inconspicuous light dark bars along dorsal portion of body: one at end of dorsal-fin base; second at middle of caudal peduncle; third at lower procurrent caudal-fin ray origin. Unpigmented portion of lateral cleithral region and latero-anterior portion of head. Two hyaline parallel strips on snout, from rostral plate to nares. Pectoral fins hyaline, without chromatophores, except in region of its insertions. In lateral view, light brown head and hyaline trunk. Dark bar inconspicuous in median region of body flowing lateral line. Few chromatophores irregularly distributed on cleithral region. Dorsal-fin hyaline, without chromatophores. Caudal-fin hyaline, except by a dark spot at anterior to middle region of rays and two small bars of chromatophores on posterior portion of caudal-fin rays. Body hyaline in ventral view; chromatophores irregularly distributed on anal-fin base and caudal peduncle.
</paragraph>
<paragraph id="8BA29564FF81FF9EB3DA43DD9482BC36" blockId="4.[151,1437,151,2013]" pageId="4">
<emphasis id="B9694976FF81FF9EB3DA43DD97A7BDA3" bold="true" box="[199,440,1700,1725]" pageId="4">Sexual dimorphism.</emphasis>
Males with a urogenital papilla in ventral view, positioned anteriorly to anal opening; a dermal flap along dorsal margin of unbranched pelvic-fin ray; a tip of pelvic-fin rays extending beyond anal-fin origin. All features are absent in females. Males and females have a membrane covering almost entire urogenital opening, a feature more developed in females.
</paragraph>
</subSubSection>
<subSubSection id="C307C6EFFF81FF9EB3DA424D939FBC6E" pageId="4" type="distribution">
<paragraph id="8BA29564FF81FF9EB3DA424D939FBC6E" blockId="4.[151,1437,151,2013]" pageId="4">
<emphasis id="B9694976FF81FF9EB3DA424D977FBC53" bold="true" box="[199,352,1844,1869]" pageId="4">Distribution.</emphasis>
<taxonomicName id="4C1DEEE7FF81FF9EB276424D949EBC52" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[363,641,1844,1868]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF81FF9EB276424D949EBC52" box="[363,641,1844,1868]" italics="true" pageId="4">Curculionichthys tukana</emphasis>
</taxonomicName>
is known from five small rivers of the rio 
<collectingRegion id="49D95B86FF81FF9EB79D424D92F3BC53" box="[1152,1260,1844,1869]" country="Brazil" name="Tocantins" pageId="4">Tocantins</collectingRegion>
basin (
<figureCitation id="132689E1FF81FF9EB658424D9392BC52" box="[1349,1421,1844,1869]" captionStart="FIGURE 3" captionStartId="5.[151,250,1093,1115]" captionTargetBox="[153,1424,197,1057]" captionTargetId="figure@5.[151,1436,195,1071]" captionTargetPageId="5" captionText="FIGURE 3. Map of north South America showing the distribution of Curculionichthys itaim— Red Diamond = type locality at Igarapé Nambuaí, municipality of Itaituba, state of Pará, Black Circles = paratypes specimens; and Curculionichthys tukana — Red Star = type locality at ribeirão Bofe, municipality of Estreito, state of Maranhão, Black Square = paratypes and non-type specimens. The green color in the map shows the rio Tapajós basin and the purple color the rio Tocantins-Araguaia basin." httpUri="https://zenodo.org/record/1039488/files/figure.png" pageId="4">Fig. 3</figureCitation>
): ribeirão Bofe, córrego Bom Jardim, rio Parauapebas, rio Sossego and a small unnamed tributary of rio Caraíbas.
</paragraph>
</subSubSection>
<subSubSection id="C307C6EFFF81FF9EB3DA4205932EBCC3" pageId="4" type="etymology">
<paragraph id="8BA29564FF81FF9EB3DA4205932EBCC3" blockId="4.[151,1437,151,2013]" pageId="4">
<emphasis id="B9694976FF81FF9EB3DA42059751BC8B" bold="true" box="[199,334,1916,1941]" pageId="4">Etymology.</emphasis>
The specific name, 
<taxonomicName id="4C1DEEE7FF81FF9EB12D42059463BC8A" authorityName="Roxo &amp; Dias &amp; Silva &amp; Oliveira" authorityYear="2017" box="[560,636,1916,1940]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF81FF9EB12D42059463BC8A" box="[560,636,1916,1940]" italics="true" pageId="4">tukana</emphasis>
</taxonomicName>
, is from the Tupi-Guarani language, and it is related to the name of the rio 
<collectingRegion id="49D95B86FF81FF9EB3DF42D9972EBCA7" box="[194,305,1952,1977]" country="Brazil" name="Tocantins" pageId="4">Tocantins</collectingRegion>
, where this new species is distributed. 
<collectingRegion id="49D95B86FF81FF9EB01442D9956ABCA7" box="[777,885,1952,1977]" country="Brazil" name="Tocantins" pageId="4">Tocantins</collectingRegion>
came from Tupi-Guarani language and means “Tucano beak” with the junction of the two words 
<taxonomicName id="4C1DEEE7FF81FF9EB1D342BD9598BCC2" authority="(Tucano)" authorityName="Tucano" box="[718,903,1988,2012]" class="Actinopterygii" family="Loricariidae" genus="Curculionichthys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Siluriformes" pageId="4" phylum="Chordata" rank="species" species="tukana">
<emphasis id="B9694976FF81FF9EB1D342BD9504BCC2" box="[718,795,1988,2012]" italics="true" pageId="4">tukana</emphasis>
(Tucano)
</taxonomicName>
and 
<emphasis id="B9694976FF81FF9EB0A042BC95FEBCC2" box="[957,993,1989,2012]" italics="true" pageId="4">tim</emphasis>
(beak). A noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>