466 lines
67 KiB
XML
466 lines
67 KiB
XML
<document id="D2780DAFF8EC2AF718C928675000FF1B" ID-DOI="10.37520/aemnp.2021.006" ID-ISSN="1804-6487" ID-Zenodo-Dep="5037544" ID-ZooBank="7A927D20-F3B3-4611-BDC3-7EC99A131B69" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624896857361" checkinUser="carolina" docAuthor="Anichtchenko, Alexander, Minkina, Łukasz, Vasiljeva, Anastasija & Medina, Milton Norman D." docDate="2021" docId="03AA8121533CFFD253F36843B826F950" docLanguage="en" docName="ActaEntMusNatPra.61.1.99-111.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 61 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2021.006" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Rhyparus helophoroides Fairmaire 1893" docType="treatment" docVersion="3" lastPageNumber="108" masterDocId="FF93F9595338FFDB50746D24BA16FFBC" masterDocTitle="A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae)" masterLastPageNumber="111" masterPageNumber="99" pageNumber="103" updateTime="1698977438183" updateUser="plazi" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="94550F52DFFA706AB1E1CEBD63BF9F70">A review of the genus Rhyparus in the Philippines, with descriptions of two new species from Mindanao (Coleoptera: Scarabaeidae: Aphodiinae)</mods:title>
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<mods:namePart id="F939DD139165A1BC3F086512BD9EB235">Anichtchenko, Alexander</mods:namePart>
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<mods:affiliation id="740EA3F2A9F41AE6FC8C77F0E7D8AB57">Institute of Life Sciences and Technologies, Daugavpils University, Daugavpils, Latvia</mods:affiliation>
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<mods:namePart id="51DB019CB832C44A5BE5662C2612A902">Minkina, Łukasz</mods:namePart>
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<mods:affiliation id="C736B5FF2BD3082C2DD9C0C28F65635A">os. Polana Szalfarska</mods:affiliation>
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<mods:namePart id="F86D3FCF6238B20C7E285F30822A1B90">Vasiljeva, Anastasija</mods:namePart>
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<mods:affiliation id="201D5462953F16E224230027981C67D8">Institute of Life Sciences and Technologies, Daugavpils University, Daugavpils, Latvia</mods:affiliation>
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<mods:namePart id="C5F1F984864B7532FF9EF915743D7AF0">Medina, Milton Norman D.</mods:namePart>
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<mods:affiliation id="2D7138A7E8DDC2234EDCAA787C166EEF">Institute of Biodiversity and Environment, Coleoptera Research Center, University of Mindanao, Davao City, Philippines</mods:affiliation>
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<subSubSection id="C31963BC533CFFDF53F36843BF46FAC3" box="[903,1360,1383,1407]" pageId="4" pageNumber="103" type="nomenclature">
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<paragraph id="8BBC3037533CFFDF53F36843BF46FAC3" blockId="4.[903,1360,1383,1407]" box="[903,1360,1383,1407]" pageId="4" pageNumber="103">
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<heading id="D0F4875B533CFFDF53F36843BF46FAC3" box="[903,1360,1383,1407]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="103" reason="3">
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<taxonomicName id="4C034BB4533CFFDF53F36843BF46FAC3" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[903,1360,1383,1407]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="species" species="helophoroides">
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<emphasis id="B977EC25533CFFDF53F36843BF46FAC3" bold="true" box="[903,1360,1383,1407]" pageId="4" pageNumber="103">
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<emphasis id="B977EC25533CFFDF53F36843BE83FAC3" bold="true" box="[903,1173,1383,1407]" italics="true" pageId="4" pageNumber="103">Rhyparus helophoroides</emphasis>
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Fairmaire, 1893
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C31963BC533CFFDF539868A2BEFAFA25" box="[1004,1260,1414,1433]" pageId="4" pageNumber="103" type="description">
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<paragraph id="8BBC3037533CFFDF539868A2BEFAFA25" blockId="4.[828,1437,1414,1795]" box="[1004,1260,1414,1433]" pageId="4" pageNumber="103">
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(
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<figureCitation id="13382CB2533CFFDF538668A2BE51FA25" box="[1010,1095,1414,1433]" captionStart="Fig" captionStartId="5.[150,181,1312,1331]" captionTargetBox="[153,1408,195,1277]" captionTargetId="figure-325@5.[150,1435,178,1285]" captionTargetPageId="5" captionText="Fig. 4. Habitus of Rhyparus helophoroides Fairmaire, 1893: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037554" httpUri="https://zenodo.org/record/5037554/files/figure.png" pageId="4" pageNumber="103">Figs 4a–c</figureCitation>
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,
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<figureCitation id="13382CB2533CFFDF542468A2BE71FA25" box="[1104,1127,1414,1433]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="4" pageNumber="103">10</figureCitation>
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,
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<figureCitation id="13382CB2533CFFDF540468A2BE91FA25" box="[1136,1159,1414,1433]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="4" pageNumber="103">16</figureCitation>
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,
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<figureCitation id="13382CB2533CFFDF54E468A2BEB1FA25" box="[1168,1191,1414,1433]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi & Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="4" pageNumber="103">22</figureCitation>
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,
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<figureCitation id="13382CB2533CFFDF54C468A2BEF0FA25" box="[1200,1254,1414,1433]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="4" pageNumber="103">28a–b</figureCitation>
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)
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</paragraph>
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Material examined.
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: Lඎඓඈඇ:
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</emphasis>
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1 J, Camirines Sur, Lagonoy,
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<date id="FFBD16F7533CFFDF530D68EFB9A1FA62" box="[889,951,1483,1502]" pageId="4" pageNumber="103" value="2019-02">ii.2019</date>
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, local coll. (
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).
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<emphasis id="B977EC25533CFFDF541F68EFBEC0FA62" bold="true" box="[1131,1238,1483,1502]" pageId="4" pageNumber="103">Mංඇൽൺඇൺඈ:</emphasis>
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3JJ
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, Araibo, Pantukan,
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, Candalaga Mts.,
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, Anichtchenko A. leg. (
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); 4 JJ
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, New
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, Cagan,
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,
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<date id="FFBD16F7533CFFDF54D76B3FBF09F992" box="[1187,1311,1563,1582]" pageId="4" pageNumber="103" value="2019-10-18" valueMax="2019-10-21" valueMin="2019-10-18">18.–21.x.2019</date>
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,Anichtchenko A. leg.ʼ (
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).
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, Maragusan, Marangig Falls, Candalaga Mts.,
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,
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,
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<date id="FFBD16F7533CFFDF54036B74BECBF9DF" box="[1143,1245,1616,1635]" pageId="4" pageNumber="103" value="2019-05-03" valueMax="2019-05-04" valueMin="2019-05-03">3.–4.v.2019</date>
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, A. Anichtchenko leg. (
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).
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, S Cotobato, Lake Holon,
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, Anichtchenko A. leg.(
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).
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<specimenCount id="9D05FBBE533CFFDF53DB6BA1B9DBF924" box="[943,973,1663,1688]" pageId="4" pageNumber="103" type="female">1 ♀</specimenCount>
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,
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<collectingRegion id="49C7FED5533CFFDF53A26BA1BE3DF924" box="[982,1067,1669,1688]" country="Philippines" name="Bukidnon" pageId="4" pageNumber="103">Bukidnon</collectingRegion>
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, Kitanglad Mt.,
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<geoCoordinate id="EE3756F0533CFFDF556F6BA1BF8FF924" box="[1307,1433,1669,1688]" degrees="125" direction="east" minutes="00" orientation="longitude" pageId="4" pageNumber="103" precision="1" seconds="22.9" value="125.00636">125°00′22.9″E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFB9DD2533CFFDF53486B84B96BF90F" box="[828,893,1696,1715]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="4" pageNumber="103" unit="m" value="1200.0">1200 m</quantity>
|
||
,
|
||
<date id="FFBD16F7533CFFDF53F06B84BE16F90F" box="[900,1024,1696,1715]" pageId="4" pageNumber="103" value="2018-10-23" valueMax="2018-10-24" valueMin="2018-10-23">23.–24.x.2018</date>
|
||
,
|
||
<taxonomicName id="4C034BB4533CFFDF547C6B84BE93F90F" authority="MND." authorityName="MND." box="[1032,1157,1696,1715]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="genus">Medina MND.</taxonomicName>
|
||
leg.(
|
||
<collectionCode id="ED12A8F2533CFFDF54C56B84BF15F90F" box="[1201,1283,1696,1715]" pageId="4" pageNumber="103">UMCRC</collectionCode>
|
||
);
|
||
<specimenCount id="9D05FBBE533CFFDF55646B84BF39F90E" box="[1296,1327,1689,1715]" pageId="4" pageNumber="103" type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingRegion id="49C7FED5533CFFDF554C6B84BF66F90F" box="[1336,1392,1696,1715]" country="Philippines" name="Davao" pageId="4" pageNumber="103">Davao</collectingRegion>
|
||
City, Marilog Mts., Marilog,
|
||
<geoCoordinate id="EE3756F0533CFFDF547A6B9FBE6FF972" box="[1038,1145,1723,1742]" degrees="7" direction="north" minutes="27" orientation="latitude" pageId="4" pageNumber="103" precision="1" seconds="16.2" value="7.4544997">7°27′16.2″N</geoCoordinate>
|
||
<geoCoordinate id="EE3756F0533CFFDF54F56B9FBF17F972" box="[1153,1281,1723,1742]" degrees="125" direction="east" minutes="14" orientation="longitude" pageId="4" pageNumber="103" precision="1" seconds="52.7" value="125.24797">125°14′52.7″E</geoCoordinate>
|
||
,
|
||
<quantity id="4CFB9DD2533CFFDF55786B9FBF47F972" box="[1292,1361,1723,1742]" metricMagnitude="3" metricUnit="m" metricValue="1.3569999999999998" pageId="4" pageNumber="103" unit="m" value="1357.0">1357 m</quantity>
|
||
,
|
||
<date id="FFBD16F7533CFFDF55286B9FB992F954" pageId="4" pageNumber="103" value="2018-07-15" valueMax="2018-07-17" valueMin="2018-07-15">15.–17. vii.2018</date>
|
||
,
|
||
<taxonomicName id="4C034BB4533CFFDF53FB6BF1BE06F954" authority="MND." authorityName="MND." box="[911,1040,1749,1768]" class="Insecta" family="Tachinidae" genus="Medina" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="103" phylum="Arthropoda" rank="genus">Medina MND.</taxonomicName>
|
||
leg. (
|
||
<collectionCode id="ED12A8F2533CFFDF54376BF1BE83F954" box="[1091,1173,1749,1768]" pageId="4" pageNumber="103">UMCRC</collectionCode>
|
||
); 1 J, Dominorog,
|
||
<collectingRegion id="49C7FED5533CFFDF55376BF1BF8FF954" box="[1347,1433,1749,1768]" country="Philippines" name="Bukidnon" pageId="4" pageNumber="103">Bukidnon</collectingRegion>
|
||
,
|
||
<date id="FFBD16F7533CFFDF53486BD4B96BF8BF" box="[828,893,1776,1795]" pageId="4" pageNumber="103" value="2019-04">iv.2019</date>
|
||
, local coll.” (
|
||
<collectionCode id="ED12A8F2533CFFDF539A6BD4BE27F8BF" box="[1006,1073,1776,1795]" pageId="4" pageNumber="103">LMCN</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31963BC533CFFD253486A3FB81BF910" lastPageId="9" lastPageNumber="108" pageId="4" pageNumber="103" type="diagnosis">
|
||
<paragraph id="8BBC3037533CFFDE53486A3FBF8AF97D" blockId="4.[828,1437,1819,2066]" lastBlockId="5.[151,1437,1450,2081]" lastPageId="5" lastPageNumber="104" pageId="4" pageNumber="103">
|
||
<emphasis id="B977EC25533CFFDF53486A3FB9A6F88E" bold="true" box="[828,944,1819,1842]" pageId="4" pageNumber="103">Diagnosis.</emphasis>
|
||
Small species: body length
|
||
<quantity id="4CFB9DD2533CFFDF54AA6A3FBF48F88E" box="[1246,1374,1819,1842]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.5" metricValueMin="3.4" pageId="4" pageNumber="103" unit="mm" value="3.95" valueMax="4.5" valueMin="3.4">3.4–4.5 mm</quantity>
|
||
(
|
||
<figureCitation id="13382CB2533CFFDF55196A3FB964F8EE" captionStart="Fig" captionStartId="5.[150,181,1312,1331]" captionTargetBox="[153,1408,195,1277]" captionTargetId="figure-325@5.[150,1435,178,1285]" captionTargetPageId="5" captionText="Fig. 4. Habitus of Rhyparus helophoroides Fairmaire, 1893: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037554" httpUri="https://zenodo.org/record/5037554/files/figure.png" pageId="4" pageNumber="103">Figs 4a–c</figureCitation>
|
||
). Head as depicted in
|
||
<figureCitation id="13382CB2533CFFDF542B6A1FBEBBF8EE" box="[1119,1197,1851,1874]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="4" pageNumber="103">Fig. 10</figureCitation>
|
||
; epipharynx as in
|
||
<figureCitation id="13382CB2533CFFDF55066A1FB94CF8CE" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="4" pageNumber="103">Fig. 16</figureCitation>
|
||
. Intermediate lobes of pronotum slightly larger than anterior one; median intercosta with dense punctation medially; second and third intercostae without or only with simple punctation basally. All intercostae of elytra with two rows of large punctures. Internal and median protrusion of caudal bulbs connected (
|
||
<figureCitation id="13382CB2533CFFDF54AB6ADFBF27F7AE" box="[1247,1329,2043,2066]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi & Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="4" pageNumber="103">Fig. 22</figureCitation>
|
||
); external protrusion weakly developed (see also TIJඣ*©ං & Bඈ*©ൽൺඍ intervals 1–4; anterior part of pronotal costae with minute 2012). Aedeagus as in
|
||
<figureCitation id="13382CB2533DFFDE51FD68EEB814FA5D" box="[393,514,1482,1505]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="5" pageNumber="104">Figs 28a–b</figureCitation>
|
||
. appressed microsetae as described below, but otherwise
|
||
<emphasis id="B977EC25533DFFDE50E368CEBB3CF9BD" bold="true" box="[151,298,1514,1537]" pageId="5" pageNumber="104">Distribution.</emphasis>
|
||
<collectingCountry id="F31470A7533DFFDE514468CEBB84F9BD" box="[304,402,1514,1537]" name="Australia" pageId="5" pageNumber="104">Australia</collectingCountry>
|
||
(introduced):
|
||
<collectingRegion id="49C7FED5533DFFDE525B68CEB8E5F9BD" box="[559,755,1514,1537]" country="Australia" name="New South Wales" pageId="5" pageNumber="104">New South Wales</collectingRegion>
|
||
, glabrous; black, except for dark brown antennae and mouth
|
||
<collectingRegion id="49C7FED5533DFFDE50E36B2EBB0EF99D" box="[151,280,1546,1569]" country="Australia" name="Queensland" pageId="5" pageNumber="104">Queensland</collectingRegion>
|
||
;
|
||
<collectingCountry id="F31470A7533DFFDE51526B2EBB84F99D" box="[294,402,1546,1569]" name="Indonesia" pageId="5" pageNumber="104">Indonesia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C7FED5533DFFDE51EB6B2EB83FF99D" box="[415,553,1546,1569]" country="Indonesia" name="Jawa Tengah" pageId="5" pageNumber="104">Central Java</collectingRegion>
|
||
,
|
||
<collectingRegion id="49C7FED5533DFFDE52426B2EB8E2F99D" box="[566,756,1546,1569]" country="Indonesia" name="Kalimantan Barat" pageId="5" pageNumber="104">West Kalimantan</collectingRegion>
|
||
, parts (
|
||
<figureCitation id="13382CB2533DFFDE53086B2EB9F0F99D" box="[892,998,1546,1569]" captionStart="Fig" captionStartId="6.[151,182,1388,1407]" captionTargetBox="[183,1426,194,1357]" captionTargetId="figure-337@6.[151,1436,183,1375]" captionTargetPageId="6" captionText="Fig. 5. Habitus of Rhyparus mindanaoensis sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037556" httpUri="https://zenodo.org/record/5037556/files/figure.png" pageId="5" pageNumber="104">Figs 5a–c</figureCitation>
|
||
).
|
||
<collectingRegion id="49C7FED5533DFFDE50E36B0EBB01F9FD" box="[151,279,1578,1601]" country="Indonesia" name="Papua Barat" pageId="5" pageNumber="104">West Papua</collectingRegion>
|
||
;
|
||
<collectingCountry id="F31470A7533DFFDE51556B0EBB74F9FD" box="[289,354,1578,1601]" name="Japan" pageId="5" pageNumber="104">Japan</collectingCountry>
|
||
: Honshu, Kyushu, Shikoku, Ryukyus; Head, as observed dorsally and frontally (
|
||
<figureCitation id="13382CB2533DFFDE55766B0EBF5BF9FD" box="[1282,1357,1578,1601]" captionStart="Fig" captionStartId="6.[151,182,1388,1407]" captionTargetBox="[183,1426,194,1357]" captionTargetId="figure-337@6.[151,1436,183,1375]" captionTargetPageId="6" captionText="Fig. 5. Habitus of Rhyparus mindanaoensis sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037556" httpUri="https://zenodo.org/record/5037556/files/figure.png" pageId="5" pageNumber="104">Figs 5c</figureCitation>
|
||
and
|
||
<figureCitation id="13382CB2533DFFDE55086B0EBF81F9FD" box="[1404,1431,1578,1601]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="5" pageNumber="104">11</figureCitation>
|
||
,
|
||
<collectingCountry id="F31470A7533DFFDE50E36B6EBAE8F9DD" box="[151,254,1610,1633]" name="Malaysia" pageId="5" pageNumber="104">Malaysia</collectingCountry>
|
||
:
|
||
<collectingRegion id="49C7FED5533DFFDE517F6B6EBB46F9DD" box="[267,336,1610,1633]" country="Malaysia" name="Sabah" pageId="5" pageNumber="104">Sabah</collectingRegion>
|
||
;
|
||
<collectingCountry id="F31470A7533DFFDE512B6B6EB81AF9DD" box="[351,524,1610,1633]" name="New Caledonia" pageId="5" pageNumber="104">New Caledonia</collectingCountry>
|
||
;
|
||
<collectingCountry id="F31470A7533DFFDE526E6B6EB8E3F9DD" box="[538,757,1610,1633]" name="Papua New Guinea" pageId="5" pageNumber="104">Papua New Guinea</collectingCountry>
|
||
: respectively), with anterior clypeal emargination with weak, Central Province,
|
||
<collectingRegion id="49C7FED5533DFFDE51246B4EB807F93D" box="[336,529,1642,1665]" country="Papua New Guinea" name="Eastern Highlands" pageId="5" pageNumber="104">Eastern Highlands</collectingRegion>
|
||
,
|
||
<collectingRegion id="49C7FED5533DFFDE526D6B4EB866F93D" box="[537,624,1642,1665]" country="Papua New Guinea" name="Morobe" pageId="5" pageNumber="104">Morobe</collectingRegion>
|
||
: the Philippi- very obtuse tooth on each side, distinct lateral emargination nes: Luzon, Negros;
|
||
<collectingCountry id="F31470A7533DFFDE51036BAEBBC4F91D" box="[375,466,1674,1697]" name="Vanuatu" pageId="5" pageNumber="104">Vanuatu</collectingCountry>
|
||
;
|
||
<collectingCountry id="F31470A7533DFFDE51A86BAEB838F91D" box="[476,558,1674,1697]" name="Taiwan" pageId="5" pageNumber="104">Taiwan</collectingCountry>
|
||
; ʻBornéoʼ (Mൾඇർඅ sinuate before widely rounded lateral angle; clypeal margin et al. 2013). First record from Mindanao Island. with upper and lower edges: upper edge concave, sharp and
|
||
</paragraph>
|
||
<caption id="DF7C60BF533DFFDE50E26804BF6DFA8F" ID-DOI="http://doi.org/10.5281/zenodo.5037554" ID-Zenodo-Dep="5037554" box="[150,1403,1312,1331]" httpUri="https://zenodo.org/record/5037554/files/figure.png" pageId="5" pageNumber="104" startId="5.[150,181,1312,1331]" targetBox="[153,1408,195,1277]" targetPageId="5">
|
||
<paragraph id="8BBC3037533DFFDE50E26804BF6DFA8F" blockId="5.[150,1403,1312,1331]" box="[150,1403,1312,1331]" pageId="5" pageNumber="104">
|
||
Fig. 4. Habitus of
|
||
<taxonomicName id="4C034BB4533DFFDE51466804B885FA8F" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[306,659,1312,1331]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="helophoroides">
|
||
<emphasis id="B977EC25533DFFDE51466804B814FA8F" box="[306,514,1312,1331]" italics="true" pageId="5" pageNumber="104">Rhyparus helophoroides</emphasis>
|
||
Fairmaire, 1893
|
||
</taxonomicName>
|
||
: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC3037533DFFDE53486BEEBF8BF95D" blockId="5.[151,1437,1450,2081]" box="[828,1437,1738,1761]" pageId="5" pageNumber="104">distinct, lower edge obsolete and not observable from above;</paragraph>
|
||
<paragraph id="8BBC3037533DFFDE517A6BCDBF8BF8BD" blockId="5.[151,1437,1450,2081]" box="[270,1437,1769,1793]" pageId="5" pageNumber="104">
|
||
<emphasis id="B977EC25533DFFDE517A6BCDB897F8BD" bold="true" box="[270,641,1769,1793]" pageId="5" pageNumber="104">
|
||
<taxonomicName id="4C034BB4533DFFDE517A6BCDB833F8BD" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[270,549,1769,1793]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="104" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
|
||
<emphasis id="B977EC25533DFFDE517A6BCDB833F8BD" bold="true" box="[270,549,1769,1793]" italics="true" pageId="5" pageNumber="104">Rhyparus mindanaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E533DFFDE52586BCEB897F8BD" box="[556,641,1770,1793]" pageId="5" pageNumber="104" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
clypeal disc very convex, ringed by deep groove, with pair
|
||
</paragraph>
|
||
<paragraph id="8BBC3037533DFFDE513C6A2DB8E1F80C" blockId="5.[151,1437,1450,2081]" pageId="5" pageNumber="104">
|
||
(
|
||
<figureCitation id="13382CB2533DFFDE513A6A2DBBB2F8A0" box="[334,420,1801,1820]" captionStart="Fig" captionStartId="6.[151,182,1388,1407]" captionTargetBox="[183,1426,194,1357]" captionTargetId="figure-337@6.[151,1436,183,1375]" captionTargetPageId="6" captionText="Fig. 5. Habitus of Rhyparus mindanaoensis sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037556" httpUri="https://zenodo.org/record/5037556/files/figure.png" pageId="5" pageNumber="104">Figs 5a–c</figureCitation>
|
||
,
|
||
<figureCitation id="13382CB2533DFFDE51D86A2DBBD4F8A0" box="[428,450,1801,1820]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="5" pageNumber="104">11</figureCitation>
|
||
,
|
||
<figureCitation id="13382CB2533DFFDE51BF6A2DBBF4F8A0" box="[459,482,1801,1820]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="5" pageNumber="104">17</figureCitation>
|
||
,
|
||
<figureCitation id="13382CB2533DFFDE519F6A2DB814F8A0" box="[491,514,1801,1820]" captionStart="Figs 19–24" captionStartId="10.[151,188,1324,1343]" captionTargetBox="[164,1415,209,1277]" captionTargetId="figure-243@10.[151,1436,189,1299]" captionTargetPageId="10" captionText="Figs 19–24. Elytra of Rhyparus species, oblique apico-dorsolateral view: 19 – R. barclayi Masumoto, Ochi & Ho, 2019; 20 – R. flavohirtus sp. nov.; 21 – R. gracilis Arrow, 1905; 22 – R. helophoroides Fairmaire, 1893; 23 – R. mindanaoensis sp. nov.; 24 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037564" httpUri="https://zenodo.org/record/5037564/files/figure.png" pageId="5" pageNumber="104">23</figureCitation>
|
||
,
|
||
<figureCitation id="13382CB2533DFFDE527F6A2DB857F8A0" box="[523,577,1801,1820]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="5" pageNumber="104">29a–b</figureCitation>
|
||
) of small rounded tubercles. Frons with four relatively long,
|
||
<emphasis id="B977EC25533DFFDE50E36A16BB0EF8F9" bold="true" box="[151,280,1842,1861]" pageId="5" pageNumber="104">
|
||
<typeStatus id="54B88E95533DFFDE50E36A16BAD5F8F9" box="[151,195,1842,1861]" pageId="5" pageNumber="104">Type</typeStatus>
|
||
material.
|
||
</emphasis>
|
||
Hඈඅඈඍඒඉൾ:J, labelled: ʻPhilippines, Mindanao, Maragu- sharp, longitudinal ridges of equal length; lateral ridges san,
|
||
<quantity id="4CFB9DD2533DFFDE50C86A69BAF8F8DC" box="[188,238,1869,1888]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="5" pageNumber="104" unit="m" value="700.0">700m</quantity>
|
||
, Marangig Falls, Candalaga Mts.,
|
||
<geoCoordinate id="EE3756F0533DFFDE527D6A69B865F8DC" box="[521,627,1869,1888]" degrees="7" direction="north" minutes="20" orientation="latitude" pageId="5" pageNumber="104" precision="1" seconds="28.4" value="7.3412223">7°20′28.4″N</geoCoordinate>
|
||
<geoCoordinate id="EE3756F0533DFFDE52026A69B8E3F8DC" box="[630,757,1869,1888]" degrees="126" direction="east" minutes="10" orientation="longitude" pageId="5" pageNumber="104" precision="1" seconds="19.8" value="126.172165">126°10′19.8″E</geoCoordinate>
|
||
, slightly less elevated; distance between median and lateral
|
||
<date id="FFBD16F7533DFFDE50E36A4CBAEEF8C7" box="[151,248,1896,1915]" pageId="5" pageNumber="104" value="2019-05-03" valueMax="2019-05-04" valueMin="2019-05-03">3–4.v.2019</date>
|
||
, A. Anichtchenko leg.ʼ (DUBC). Pൺ*©ൺඍඒඉൾඌ: 1J, labelled: ridges slightly larger than that between two median ones. ʻPhilippines, Mindanao,
|
||
<collectingRegion id="49C7FED5533DFFDE51006AA6BBDAF829" box="[372,460,1922,1941]" country="Philippines" name="Bukidnon" pageId="5" pageNumber="104">Bukidnon</collectingRegion>
|
||
, Kalatungan,
|
||
<date id="FFBD16F7533DFFDE523C6AA6B89AF829" box="[584,652,1922,1941]" pageId="5" pageNumber="104" value="2016-09">ix.2016</date>
|
||
, local coll.ʼ Head between genae almost as wide as pronotum; genae (LMCN).
|
||
<specimenCount id="9D05FBBE533DFFDE50866AB9BB03F80C" box="[242,277,1943,1968]" pageId="5" pageNumber="104" type="female">1 ♀</specimenCount>
|
||
, labelled: ʻPalawan (Central), Sabang,
|
||
<quantity id="4CFB9DD2533DFFDE520F6AB9B8D8F80C" box="[635,718,1949,1968]" metricMagnitude="1" metricUnit="m" metricValue="7.5" metricValueMax="10.0" metricValueMin="5.0" pageId="5" pageNumber="104" unit="m" value="75.0" valueMax="100.0" valueMin="50.0">
|
||
50-
|
||
<specimenCount id="9D05FBBE533DFFDE52ED6AB9B8D8F80C" box="[665,718,1949,1968]" pageId="5" pageNumber="104" type="generic">100m</specimenCount>
|
||
</quantity>
|
||
., via
|
||
</paragraph>
|
||
<paragraph id="8BBC3037533DFFDE53486A8EB850F877" blockId="5.[151,1437,1450,2081]" pageId="5" pageNumber="104">
|
||
very large and strongly protruding. Head irregularly punc-
|
||
<superScript id="7C769D7F533DFFDE50E36A9CB850F877" attach="left" box="[151,582,1976,1995]" fontSize="8" pageId="5" pageNumber="104">Underground River, I. Lobl, 30.xi.[19]95ʼ (ISEA).</superScript>
|
||
</paragraph>
|
||
<paragraph id="8BBC3037533DFFDE53486AEEBF86F79D" blockId="5.[151,1437,1450,2081]" pageId="5" pageNumber="104">
|
||
tate, more sparsely so in frons, punctures large, separated
|
||
<emphasis id="B977EC25533DFFDE50E36AC7BB35F846" bold="true" box="[151,291,2019,2042]" pageId="5" pageNumber="104">Description.</emphasis>
|
||
Male
|
||
<typeStatus id="54B88E95533DFFDE511A6AC7BBC6F846" box="[366,464,2019,2042]" pageId="5" pageNumber="104" type="holotype">holotype</typeStatus>
|
||
: Moderately large species, by about 1.5–2.0 times their diameter and bearing minute, body length
|
||
<quantity id="4CFB9DD2533DFFDE516E6527BB7BF7A6" box="[282,365,2051,2074]" metricMagnitude="-3" metricUnit="m" metricValue="5.5" pageId="5" pageNumber="104" unit="mm" value="5.5">5.5 mm</quantity>
|
||
; robust, matt, except for shiny tops of pale, relatively thin microsetae. Epipharynx as in
|
||
<figureCitation id="13382CB2533DFFDE5534652EBF9AF79D" box="[1344,1420,2058,2081]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="5" pageNumber="104">Fig. 17</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF7C60BF533EFFDD50E36848BF88FAC3" ID-DOI="http://doi.org/10.5281/zenodo.5037556" ID-Zenodo-Dep="5037556" box="[151,1438,1388,1407]" httpUri="https://zenodo.org/record/5037556/files/figure.png" pageId="6" pageNumber="105" startId="6.[151,182,1388,1407]" targetBox="[183,1426,194,1357]" targetPageId="6">
|
||
<paragraph id="8BBC3037533EFFDD50E36848BF88FAC3" blockId="6.[151,1438,1388,1407]" box="[151,1438,1388,1407]" pageId="6" pageNumber="105">
|
||
Fig. 5. Habitus of
|
||
<taxonomicName id="4C034BB4533EFFDD51596848B814FAC3" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[301,514,1388,1407]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="105" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
|
||
<emphasis id="B977EC25533EFFDD51596848B814FAC3" box="[301,514,1388,1407]" italics="true" pageId="6" pageNumber="105">Rhyparus mindanaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E533EFFDD52726848B85FFAC3" box="[518,585,1388,1407]" pageId="6" pageNumber="105" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC3037533EFFDD50C168CDBEF5F9DC" blockId="6.[151,760,1513,2080]" lastBlockId="6.[828,1437,1513,2080]" pageId="6" pageNumber="105">
|
||
Pronotum transverse, 1.26 times as wide as long; opaque, convex, anterior margin weakly bisinuate; disc with eight costae, six central costae sharp and well delimited. Costae 1 (paramedian pair) continuous, very highly elevated anteriorly and posteriorly, in anterior part forming elevated, elongate tubercles (
|
||
<figureCitation id="13382CB2533EFFDD51BA6BADB80AF91C" box="[462,540,1673,1696]" captionStart="Figs 7–12" captionStartId="8.[151,188,1611,1630]" captionTargetBox="[165,1410,204,1585]" captionTargetId="figure-191@8.[151,1436,189,1595]" captionTargetPageId="8" captionText="Figs 7–12. Head of Rhyparus species, frontal view: 7 – R. barclayi Masumoto, Ochi & Ho, 2019; 8 – R. flavohirtus sp. nov.; 9 – R. gracilis Arrow, 1905; 10 – R. helophoroides Fairmaire, 1893; 11 – R. mindanaoensis sp. nov.; 12 – R. philippinensis Arrow, 1905. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="6" pageNumber="105">Fig. 11</figureCitation>
|
||
); costae slightly and evenly convergent in central part. Costae 2 high and well delimited, flattened in transverse depression, anterior part short but very high; fossae of transverse depression small and caviform, with two coracoid processes hanging over fossae. Costae 3 continuous, distinctly elevated, weakly sinuous in anterior third. Costae 4 incomplete, flattened and low but distinct in median part, interrupted in apical third by two deep punctiform fossae, and effaced in basal third; one deep punctiform fossa situated at base of anterolateral pronotal process. Median intercosta densely covered with large punctures, punctures not confluent in median line and sparser in anterior half. Lateral intercostae 1 and 2 each with four large punctures in apical part; intercostae 2 and 3 with only few large punctures. Lateral margin with two lobes, anterior lobe large, relatively narrow and with pointed apex, median lobe subangulate.
|
||
</paragraph>
|
||
<paragraph id="8BBC3037533EFFDC532E6B4DB8EFF903" blockId="6.[828,1437,1513,2080]" lastBlockId="7.[151,761,1384,2079]" lastPageId="7" lastPageNumber="106" pageId="6" pageNumber="105">
|
||
Elytra (
|
||
<figureCitation id="13382CB2533EFFDD53DF6B4DBE01F93C" box="[939,1047,1641,1664]" captionStart="Fig" captionStartId="6.[151,182,1388,1407]" captionTargetBox="[183,1426,194,1357]" captionTargetId="figure-337@6.[151,1436,183,1375]" captionTargetPageId="6" captionText="Fig. 5. Habitus of Rhyparus mindanaoensis sp. nov., male, holotype: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.5037556" httpUri="https://zenodo.org/record/5037556/files/figure.png" pageId="6" pageNumber="105">Figs 5a–c</figureCitation>
|
||
;
|
||
<figureCitation id="13382CB2533EFFDD54506B4DBE29F93C" box="[1060,1087,1641,1664]" captionStart="Figs 13–18" captionStartId="9.[150,187,1526,1545]" captionTargetBox="[158,1411,197,1485]" captionTargetId="figure-259@9.[150,1435,161,1515]" captionTargetPageId="9" captionText="Figs 13–18. Epipharynx of Rhyparus species: 13 – R. barclayi Masumoto, Ochi & Ho, 2019; 14 – R. flavohirtus sp. nov.; 15 – R. gracilis Arrow, 1905; 16 – R. helophoroides Fairmaire, 1893; 17– R. mindanaoensis sp. nov.; 18 – R. philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="6" pageNumber="105">17</figureCitation>
|
||
) elongate, 1.65 times as long as wide. Humeral callus with large punctures separated by about one puncture diameter. Each elytron with five strong costae. First costa (sutural) straight, flattened on disc and slightly elevated in apical third, without apical tooth. Second costa sharp, very strongly elevated from base to apical bulbous area, and terminated as strong swelling with dense brown trichome at apex. Third costa slightly incurved, sharp, extending from base to a little before end of costa 2, apex slightly swollen and with brown elongate tubercle. Fourth costa high, of almost same length and width as costa 3, shortly curved at base and slightly sinuous near apex, with relatively long brown tubercle and dense trichome. Fifth costa sinuous, elevated, extending from base to apex, with apical portion strongly swollen and forming bisinuate transverse protrusion with three large tubercles in dorsal view. Elytral intercostae very weakly and transversally corrugated, matt, except for very shiny and impunctate apical parts of intercostae 1–4. Intercostae 1–4 with two longitudinal rows of large and deep punctures of almost equal size, extending from base to near apex, distances between neighboring punctures relatively uniform, about 1.5–2.0 times puncture diameter. Caudal bulbs distinctly developed. Internal and median protrusion of caudal bulbs well developed; external protrusion moderately developed.
|
||
</paragraph>
|
||
<caption id="DF7C60BF533FFFDC50E269CFBF88FB42" ID-DOI="http://doi.org/10.5281/zenodo.5037558" ID-Zenodo-Dep="5037558" box="[150,1438,1259,1278]" httpUri="https://zenodo.org/record/5037558/files/figure.png" pageId="7" pageNumber="106" startId="7.[150,181,1259,1278]" targetBox="[172,1389,192,1237]" targetPageId="7">
|
||
<paragraph id="8BBC3037533FFFDC50E269CFBF88FB42" blockId="7.[150,1438,1259,1278]" box="[150,1438,1259,1278]" pageId="7" pageNumber="106">
|
||
Fig. 6. Habitus of
|
||
<taxonomicName id="4C034BB4533FFFDC515A69CFB87AFB42" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[302,620,1259,1278]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="106" phylum="Arthropoda" rank="species" species="philippinensis">
|
||
<emphasis id="B977EC25533FFFDC515A69CFBBEBFB42" box="[302,509,1259,1278]" italics="true" pageId="7" pageNumber="106">Rhyparus philippinensis</emphasis>
|
||
<bibRefCitation id="EF924DC6533FFFDC527569CFB87AFB42" author="ARROW G. J." box="[513,620,1259,1278]" pageId="7" pageNumber="106" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
, syntypes: a – dorsal view; b – right dorsolateral view; c – right lateral view. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC3037533FFFDC50C16BECBB70F883" blockId="7.[151,761,1384,2079]" pageId="7" pageNumber="106">Pygidium with semicircular elevated area at base connected by narrow ridge with pygidial apex and by deep groove with basal margin, with areas on each side of ridge strongly depressed.</paragraph>
|
||
<paragraph id="8BBC3037533FFFDC50C16A6CBF3AF963" blockId="7.[151,761,1384,2079]" lastBlockId="7.[828,1437,1384,2079]" pageId="7" pageNumber="106">Legs. Profemora wide, with large punctures throughout. Protibiae thin at base; with slightly sinuate inner margin up to relatively large and slightly inward bent inner apical tooth, weakly dilated apically, with moderately arcuate and then emarginate inner margin in front of large outer tooth. Mesofemora with 4–5 tough, fairly long, sharp, pale microsetae along apical half of anterior edge; posterior margin bilobed, with deep groove, its surface punctate throughout, punctures of half size of those on profemoral surface. Mesotibiae narrow at base, inner margin strongly widened from base to about third of its length and then almost parallel to apex, with short, pale, decumbent setae on inner surface and rows of longer, pale, semierect setae on edges; apex with backward directed inferior spine and inward bent tooth. Metafemora long, narrow, posterior margin slightly bilobed, punctation smaller and sparser than on mesofemora. Metatibiae with outer margin concave, setation less developed than on mesotibiae; inferior apical tooth large and directed inward in male.</paragraph>
|
||
<paragraph id="8BBC3037533FFFD3532E6BCCBB75F941" blockId="7.[828,1437,1384,2079]" lastBlockId="8.[151,760,1734,2077]" lastPageId="8" lastPageNumber="107" pageId="7" pageNumber="106">Venter matt, black. Prosternum with long anterior process with rounded apex and hastate posterior process. Mesoventrite with shallow setigerous punctures. Meso- -metaventral plate with distinct, moderately deep medi- an impression, surface with small setigerous punctures medially and larger punctures laterally. Abdomen with ventrites 1–5 with small, shallow and sparse setiferous punctures, punctures along apical and basal margin crenate; ventrites 3–5 triangularly grooved on each lateral broadened area, groove distinctly larger on ventrite 5; ventrites 5 and 6 fused, ventrite 6 with large transverse groove with 16 large punctures.</paragraph>
|
||
<caption id="DF7C60BF5330FFD350E36B6FBEF4F9C5" ID-DOI="http://doi.org/10.5281/zenodo.5037560" ID-Zenodo-Dep="5037560" httpUri="https://zenodo.org/record/5037560/files/figure.png" pageId="8" pageNumber="107" startId="8.[151,188,1611,1630]" targetBox="[165,1410,204,1585]" targetPageId="8">
|
||
<paragraph id="8BBC30375330FFD350E36B6FBEF4F9C5" blockId="8.[151,1437,1611,1657]" pageId="8" pageNumber="107">
|
||
Figs 7–12. Head of
|
||
<taxonomicName id="4C034BB45330FFD351496B6FBBC5F9E2" box="[317,467,1611,1630]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B977EC255330FFD351496B6FBB99F9E2" box="[317,399,1611,1630]" italics="true" pageId="8" pageNumber="107">Rhyparus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, frontal view: 7 –
|
||
<taxonomicName id="4C034BB45330FFD3521F6B6FB9D2F9E2" authority="Masumoto, Ochi & Ho, 2019" authorityName="Masumoto, Ochi & Ho" authorityYear="2019" box="[619,964,1611,1630]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="barclayi">
|
||
<emphasis id="B977EC255330FFD3521F6B6FB8DEF9E2" box="[619,712,1611,1630]" italics="true" pageId="8" pageNumber="107">R. barclayi</emphasis>
|
||
<bibRefCitation id="EF924DC65330FFD352B96B6FB9D2F9E2" author="MASUMOTO K. & OCHI T. & HO B. H." box="[717,964,1611,1630]" pageId="8" pageNumber="107" pagination="13 - 18" refId="ref6667" refString="MASUMOTO K., OCHI T. & HO B. H. 2019: A new Rhyparus species from Taiwan (Scarabaeidae, Aphodiinae, Rhyparini). Kogane 22: 13 - 18." type="journal article" year="2019">Masumoto, Ochi & Ho, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
; 8 –
|
||
<taxonomicName id="4C034BB45330FFD3539A6B6FBE77F9E2" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[1006,1121,1611,1630]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="flavohirtus" status="sp. nov.">
|
||
<emphasis id="B977EC255330FFD3539A6B6FBE77F9E2" box="[1006,1121,1611,1630]" italics="true" pageId="8" pageNumber="107">R. flavohirtus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E5330FFD354116B6FBEBFF9E2" box="[1125,1193,1611,1630]" pageId="8" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 9 –
|
||
<taxonomicName id="4C034BB45330FFD354BB6B6FBF8FF9E2" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[1231,1433,1611,1630]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B977EC255330FFD354BB6B6FBF31F9E2" box="[1231,1319,1611,1630]" italics="true" pageId="8" pageNumber="107">R. gracilis</emphasis>
|
||
<bibRefCitation id="EF924DC65330FFD3555F6B6FBF8FF9E2" author="ARROW G. J." box="[1323,1433,1611,1630]" pageId="8" pageNumber="107" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
; 10 –
|
||
<taxonomicName id="4C034BB45330FFD350B66B42BBF5F9C5" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[194,483,1638,1657]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="helophoroides">
|
||
<emphasis id="B977EC255330FFD350B66B42BB44F9C5" box="[194,338,1638,1657]" italics="true" pageId="8" pageNumber="107">R. helophoroides</emphasis>
|
||
Fairmaire, 1893
|
||
</taxonomicName>
|
||
; 11 –
|
||
<taxonomicName id="4C034BB45330FFD352616B42B8BBF9C5" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[533,685,1638,1657]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
|
||
<emphasis id="B977EC255330FFD352616B42B8BBF9C5" box="[533,685,1638,1657]" italics="true" pageId="8" pageNumber="107">R. mindanaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E5330FFD352C76B42B8EEF9C5" box="[691,760,1638,1657]" pageId="8" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 12 –
|
||
<taxonomicName id="4C034BB45330FFD3535E6B42BE3BF9C5" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[810,1069,1638,1657]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="philippinensis">
|
||
<emphasis id="B977EC255330FFD3535E6B42B9ACF9C5" box="[810,954,1638,1657]" italics="true" pageId="8" pageNumber="107">R. philippinensis</emphasis>
|
||
<bibRefCitation id="EF924DC65330FFD353CB6B42BE3BF9C5" author="ARROW G. J." box="[959,1069,1638,1657]" pageId="8" pageNumber="107" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
. Scale bar = 1.0 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BBC30375330FFD350C16A22B896F8A1" blockId="8.[151,760,1734,2077]" box="[181,640,1798,1821]" pageId="8" pageNumber="107">
|
||
Male genitalia. Aedeagus as in
|
||
<figureCitation id="13382CB25330FFD352776A22B86AF8A1" box="[515,636,1798,1821]" captionStart="Figs 25–30" captionStartId="11.[150,187,1486,1505]" captionTargetBox="[207,1392,193,1454]" captionTargetId="figure-302@11.[151,1436,180,1472]" captionTargetPageId="11" captionText="Figs 25–30.Aedeagus of Rhyparus species, a – lateral view, b – dorsal view. 25 – R.barclayi Masumoto, Ochi & Ho, 2019; 26 – R. flavohirtus sp. nov.; 27 – R. gracilis Arrow, 1905; 28 – R. helophoroides Fairmaire, 1893; 29 – R.mindanaoensis sp. nov.; 30 – R.philippinensis Arrow, 1905. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5037566" httpUri="https://zenodo.org/record/5037566/files/figure.png" pageId="8" pageNumber="107">Figs 29a–b</figureCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBC30375330FFD250E36A02B81BF910" blockId="8.[151,760,1734,2077]" lastBlockId="9.[151,760,1653,1772]" lastPageId="9" lastPageNumber="108" pageId="8" pageNumber="107">
|
||
<emphasis id="B977EC255330FFD350E36A02BB99F881" bold="true" box="[151,399,1830,1853]" pageId="8" pageNumber="107">Differential diagnosis.</emphasis>
|
||
Based on the moderately elongate habitus with elytra widest in basal third, body length 5.5–6.0 mm, anterior lateral lobes of pronotum wider than intermediate ones, very high costae of pronotum and elytra with deep intercostae, presence of two rows of punctures in all intercostae of elytra, distinctly divided median and internal protrusion of caudal bulb of elytra with distinct external protrusion, the new species is very similar to
|
||
<taxonomicName id="4C034BB45330FFD353DF6BE2BF5EF961" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[939,1352,1734,1757]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="denticollis">
|
||
<emphasis id="B977EC255330FFD353DF6BE2BE98F961" box="[939,1166,1734,1757]" italics="true" pageId="8" pageNumber="107">Rhyparus denticollis</emphasis>
|
||
Fairmaire, 1893
|
||
</taxonomicName>
|
||
. However,
|
||
<taxonomicName id="4C034BB45330FFD3531E6BC2BE3FF941" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[874,1065,1766,1789]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="107" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
|
||
<emphasis id="B977EC255330FFD3531E6BC2BE3FF941" box="[874,1065,1766,1789]" italics="true" pageId="8" pageNumber="107">R. mindanaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E5330FFD3545B6BC2BE94F941" box="[1071,1154,1766,1789]" pageId="8" pageNumber="107" rank="species">sp. nov.</taxonomicNameLabel>
|
||
can be distinguished from the former species by median intercosta of pronotum somewhat narrower and with denser punctation, second and third intercostae of pronotum with sparse/very sparse, irregularly distributed punctation in basal part (versus moderately dense punctation, rather evenly distributed, with similar density to punctation in median furrow), sinuation between internal and median protrusion of caudal bulb of elytra more distinct and tibiae relatively slender.
|
||
<emphasis id="B977EC255330FFD353486522B9D6F7A1" bold="true" box="[828,960,2054,2077]" pageId="8" pageNumber="107">Etymology.</emphasis>
|
||
The specific epithet is derived from Min- danao
|
||
<collectingCountry id="F31470A75331FFD250AA6B51BB34F930" box="[222,290,1653,1676]" name="Iceland" pageId="9" pageNumber="108">Island</collectingCountry>
|
||
, where the
|
||
<typeStatus id="54B88E955331FFD251E96B51BBECF930" box="[413,506,1653,1676]" pageId="9" pageNumber="108" type="holotype">holotype</typeStatus>
|
||
of the new species was collected; the name is an adjective.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7C60BF5331FFD250E268D2BECBF998" ID-DOI="http://doi.org/10.5281/zenodo.5037562" ID-Zenodo-Dep="5037562" httpUri="https://zenodo.org/record/5037562/files/figure.png" pageId="9" pageNumber="108" startId="9.[150,187,1526,1545]" targetBox="[158,1411,197,1485]" targetPageId="9">
|
||
<paragraph id="8BBC30375331FFD250E268D2BECBF998" blockId="9.[150,1436,1526,1572]" pageId="9" pageNumber="108">
|
||
Figs 13–18. Epipharynx of
|
||
<taxonomicName id="4C034BB45331FFD251F468D2B80EF9B5" box="[384,536,1526,1545]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B977EC255331FFD251F468D2BBC4F9B5" box="[384,466,1526,1545]" italics="true" pageId="9" pageNumber="108">Rhyparus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
: 13 –
|
||
<taxonomicName id="4C034BB45331FFD2523E68D2B9BFF9B5" authority="Masumoto, Ochi & Ho, 2019" authorityName="Masumoto, Ochi & Ho" authorityYear="2019" box="[586,937,1526,1545]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="barclayi">
|
||
<emphasis id="B977EC255331FFD2523E68D2B8BEF9B5" box="[586,680,1526,1545]" italics="true" pageId="9" pageNumber="108">R. barclayi</emphasis>
|
||
<bibRefCitation id="EF924DC65331FFD252D968D2B9BFF9B5" author="MASUMOTO K. & OCHI T. & HO B. H." box="[685,937,1526,1545]" pageId="9" pageNumber="108" pagination="13 - 18" refId="ref6667" refString="MASUMOTO K., OCHI T. & HO B. H. 2019: A new Rhyparus species from Taiwan (Scarabaeidae, Aphodiinae, Rhyparini). Kogane 22: 13 - 18." type="journal article" year="2019">Masumoto, Ochi & Ho, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
; 14 –
|
||
<taxonomicName id="4C034BB45331FFD253A968D2BE46F9B5" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[989,1104,1526,1545]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="flavohirtus" status="sp. nov.">
|
||
<emphasis id="B977EC255331FFD253A968D2BE46F9B5" box="[989,1104,1526,1545]" italics="true" pageId="9" pageNumber="108">R. flavohirtus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E5331FFD2542168D2BE8CF9B5" box="[1109,1178,1526,1545]" pageId="9" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 15 –
|
||
<taxonomicName id="4C034BB45331FFD254B868D2BF8EF9B5" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[1228,1432,1526,1545]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B977EC255331FFD254B868D2BF33F9B5" box="[1228,1317,1526,1545]" italics="true" pageId="9" pageNumber="108">R. gracilis</emphasis>
|
||
<bibRefCitation id="EF924DC65331FFD2555E68D2BF8EF9B5" author="ARROW G. J." box="[1322,1432,1526,1545]" pageId="9" pageNumber="108" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
; 16 –
|
||
<taxonomicName id="4C034BB45331FFD250B56B35BBF4F998" authority="Fairmaire, 1893" authorityName="Fairmaire" authorityYear="1893" box="[193,482,1553,1572]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="helophoroides">
|
||
<emphasis id="B977EC255331FFD250B56B35BB44F998" box="[193,338,1553,1572]" italics="true" pageId="9" pageNumber="108">R. helophoroides</emphasis>
|
||
Fairmaire, 1893
|
||
</taxonomicName>
|
||
; 17–
|
||
<taxonomicName id="4C034BB45331FFD252646B35B8BEF998" authority="Anichtchenko & Minkina & Vasiljeva & Medina, 2021" authorityName="Anichtchenko & Minkina & Vasiljeva & Medina" authorityYear="2021" box="[528,680,1553,1572]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
|
||
<emphasis id="B977EC255331FFD252646B35B8BEF998" box="[528,680,1553,1572]" italics="true" pageId="9" pageNumber="108">R. mindanaoensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A244515E5331FFD252D96B35B8E4F998" box="[685,754,1553,1572]" pageId="9" pageNumber="108" rank="species">sp. nov.</taxonomicNameLabel>
|
||
; 18 –
|
||
<taxonomicName id="4C034BB45331FFD253516B35BE31F998" authority="Arrow, 1905" authorityName="Arrow" authorityYear="1905" box="[805,1063,1553,1572]" class="Insecta" family="Aphodiidae" genus="Rhyparus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="108" phylum="Arthropoda" rank="species" species="philippinensis">
|
||
<emphasis id="B977EC255331FFD253516B35B9A2F998" box="[805,948,1553,1572]" italics="true" pageId="9" pageNumber="108">R. philippinensis</emphasis>
|
||
<bibRefCitation id="EF924DC65331FFD253CD6B35BE31F998" author="ARROW G. J." box="[953,1063,1553,1572]" pageId="9" pageNumber="108" pagination="534 - 540" refId="ref6410" refString="ARROW G. J. 1905: On some Oriental aphodiid Coleoptera of the Rhyparus Group, with description of a new genus. Annals and Magazine of Natural History, Series 7 15: 534 - 540." type="journal article" year="1905">Arrow, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
. Scale bar = 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31963BC5331FFD250E36B91B826F950" pageId="9" pageNumber="108" type="distribution">
|
||
<paragraph id="8BBC30375331FFD250E36B91B826F950" blockId="9.[151,760,1653,1772]" pageId="9" pageNumber="108">
|
||
<emphasis id="B977EC255331FFD250E36B91BB3CF970" bold="true" box="[151,298,1717,1740]" pageId="9" pageNumber="108">Distribution.</emphasis>
|
||
Species known so far only from the
|
||
<collectingCountry id="F31470A75331FFD252DB6B91BAC5F950" name="Philippines" pageId="9" pageNumber="108">Philippines</collectingCountry>
|
||
: Mindanao and
|
||
<collectingRegion id="49C7FED55331FFD251096BF1BBC1F950" box="[381,471,1749,1772]" country="Philippines" name="Palawan" pageId="9" pageNumber="108">Palawan</collectingRegion>
|
||
islands.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |