275 lines
31 KiB
XML
275 lines
31 KiB
XML
<document ID-DOI="10.11646/zootaxa.5263.3.2" ID-ISSN="1175-5326" ID-Zenodo-Dep="7804268" ID-ZooBank="20CB3CA0-BEF9-474C-8931-6A7948B9CA61" approvalRequired="5" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="4" checkinTime="1680760231104" checkinUser="plazi" docAuthor="Řezáč, Milan, Cardoso, Pedro & Řezáčová, Veronika" docDate="2023" docId="039CE028E863FFD6FF12FA03FAF16441" docLanguage="en" docName="zootaxa.5263.3.2.pdf" docOrigin="Zootaxa 5263 (3)" docSource="http://dx.doi.org/10.11646/zootaxa.5263.3.2" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Harpactea pekari Rezac 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="355" masterDocId="FFA59850E870FFC2FF85FFA1FFD2667D" masterDocTitle="Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal" masterLastPageNumber="364" masterPageNumber="335" pageNumber="354" updateTime="1680760785065" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>Review of Harpactea ground-dwelling spiders (Araneae: Dysderidae) of Portugal</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Řezáč, Milan</mods:namePart>
|
||
<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Cardoso, Pedro</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0001-8119-9960</mods:nameIdentifier>
|
||
<mods:affiliation>Laboratory for Integrative Biodiversity Research (LIBRe), Finnish Museum of Natural History Luomus, University of Helsinki, P. O. Box 17, FI- 00014 Helsinki, Finland. pedro. cardoso @ helsinki. fi; https: // orcid. org / 0000 - 0001 - 8119 - 9960</mods:affiliation>
|
||
<mods:nameIdentifier type="email">pedro.cardoso@helsinki.fi</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Řezáčová, Veronika</mods:namePart>
|
||
<mods:nameIdentifier type="ORCID">0000-0002-1749-0355</mods:nameIdentifier>
|
||
<mods:affiliation>Crop Research Institute, Drnovská 507, CZ- 16106 Prague 6, Czechia & strver @ seznam. cz; https: // orcid. org / 0000 - 0002 - 1749 - 0355</mods:affiliation>
|
||
<mods:nameIdentifier type="email">strver@seznam.cz</mods:nameIdentifier>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2023</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2023-04-06</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>5263</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="issue">
|
||
<mods:number>3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>335</mods:start>
|
||
<mods:end>364</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location>
|
||
<mods:url>http://dx.doi.org/10.11646/zootaxa.5263.3.2</mods:url>
|
||
</mods:location>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.11646/zootaxa.5263.3.2</mods:identifier>
|
||
<mods:identifier type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">7804268</mods:identifier>
|
||
<mods:identifier type="ZooBank">20CB3CA0-BEF9-474C-8931-6A7948B9CA61</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7804308" ID-Zenodo-Dep="7804308" LSID="urn:lsid:plazi:treatment:039CE028E863FFD6FF12FA03FAF16441" httpUri="http://treatment.plazi.org/id/039CE028E863FFD6FF12FA03FAF16441" lastPageId="20" lastPageNumber="355" pageId="19" pageNumber="354">
|
||
<subSubSection box="[151,545,1441,1471]" pageId="19" pageNumber="354" type="nomenclature">
|
||
<paragraph blockId="19.[151,545,1441,1504]" box="[151,545,1441,1471]" pageId="19" pageNumber="354">
|
||
<heading bold="true" box="[151,545,1441,1471]" fontSize="11" level="1" pageId="19" pageNumber="354" reason="1">
|
||
<emphasis bold="true" box="[151,545,1441,1471]" pageId="19" pageNumber="354">
|
||
<taxonomicName authority="Rezac" authorityName="Rezac" authorityYear="2023" box="[151,445,1441,1471]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="pekari" status="sp. nov.">
|
||
<emphasis bold="true" box="[151,363,1441,1468]" italics="true" pageId="19" pageNumber="354">Harpactea pekari</emphasis>
|
||
Řezáč
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[452,545,1442,1468]" pageId="19" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[151,479,1478,1504]" pageId="19" pageNumber="354" type="description">
|
||
<paragraph blockId="19.[151,545,1441,1504]" box="[151,479,1478,1504]" pageId="19" pageNumber="354">
|
||
(
|
||
<figureCitation box="[159,251,1478,1504]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="19" pageNumber="354">Figs 1D</figureCitation>
|
||
,
|
||
<figureCitation box="[264,301,1478,1504]" captionStart="FIGURE 2" captionStartId="4.[151,250,1823,1847]" captionTargetBox="[250,1344,181,1799]" captionTargetId="figure-24@4.[244,1344,181,1799]" captionTargetPageId="4" captionText="FIGURE 2. Portuguese Harpactea spiders. Right male chelicera.Above anterior (G, J, L, M) or posterior view (A, B, C, D, E, F, H, I, K, N); below median view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804274" httpUri="https://zenodo.org/record/7804274/files/figure.png" pageId="19" pageNumber="354">2D</figureCitation>
|
||
,
|
||
<figureCitation box="[312,349,1478,1504]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="19" pageNumber="354">3D</figureCitation>
|
||
,
|
||
<figureCitation box="[360,397,1478,1504]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="19" pageNumber="354">4D</figureCitation>
|
||
,
|
||
<figureCitation box="[409,443,1478,1504]" captionStart="FIGURE 6" captionStartId="15.[151,250,1048,1072]" captionTargetBox="[151,1431,181,1020]" captionTargetId="figure-19@15.[151,1436,181,1024]" captionTargetPageId="15" captionText="FIGURE 6. Portuguese Harpactea spiders. Vulva. A. H. fageli. Portugal: Setúbal; B. H. henriquesi sp. nov. Portugal: Torres Novas; C. H.pekari sp. nov. Portugal:Corredoura.Views:dor.—dorsal,fro.—anterior,hin.—posterior, ven.—ventral.Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804286" httpUri="https://zenodo.org/record/7804286/files/figure.png" pageId="19" pageNumber="354">6C</figureCitation>
|
||
,
|
||
<figureCitation box="[455,470,1478,1504]" captionStart="FIGURE 9" captionStartId="22.[151,250,1892,1916]" captionTargetBox="[279,1342,195,1818]" captionTargetId="figure-24@22.[231,1356,181,1869]" captionTargetPageId="22" captionText="FIGURE 9. Known distribution of Portuguese Harpactea species. In each map the first species is represented by a transparent circle and the second species by a black diamond:A) H. adicensis & H. algarvensis; B) H. crespoi & H. dolanskyi; C) H. fageli & H. gaditana; D) H. henriquesi & H. hombergii; E) H. korenkoi & H. krejcii; F) H. magnibulbi & H. minoccii; G) H. pekari & H. proxima; H) H. stalitoides & H. subiasi; I) H. tavirensis." figureDoi="http://doi.org/10.5281/zenodo.7804296" httpUri="https://zenodo.org/record/7804296/files/figure.png" pageId="19" pageNumber="354">9</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="354" type="materials_examined">
|
||
<paragraph blockId="19.[151,1437,1544,2039]" pageId="19" pageNumber="354">
|
||
<emphasis bold="true" box="[151,325,1544,1571]" pageId="19" pageNumber="354">Type material.</emphasis>
|
||
<emphasis box="[333,443,1544,1570]" italics="true" pageId="19" pageNumber="354">
|
||
<typeStatus box="[333,439,1544,1570]" pageId="19" pageNumber="354">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
♁, Mertola, Corredoura,
|
||
<geoCoordinate box="[730,810,1544,1570]" degrees="37.746" direction="north" orientation="latitude" pageId="19" pageNumber="354" precision="55" value="37.746">37.746</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[821,896,1544,1570]" degrees="7.642" direction="west" orientation="longitude" pageId="19" pageNumber="354" precision="55" value="-7.642">-7.642</geoCoordinate>
|
||
,
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[908,1113,1545,1571]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="354" phylum="Tracheophyta" rank="species" species="coccifera">
|
||
<emphasis box="[908,1113,1545,1571]" italics="true" pageId="19" pageNumber="354">Quercus coccifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[1169,1243,1544,1571]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="354" phylum="Tracheophyta" rank="species" species="ilex">
|
||
<emphasis box="[1169,1243,1544,1571]" italics="true" pageId="19" pageNumber="354">Q. ilex</emphasis>
|
||
</taxonomicName>
|
||
wood,
|
||
<date pageId="19" pageNumber="354" value="2013-03-29">29 March 2013</date>
|
||
, leg. M. Řezáč, coll. National Natural History Museum,
|
||
<collectingRegion box="[844,925,1581,1607]" country="Czech Republic" name="Praha" pageId="19" pageNumber="354">Prague</collectingRegion>
|
||
, code P6A 7389.
|
||
</paragraph>
|
||
<paragraph blockId="19.[151,1437,1544,2039]" pageId="19" pageNumber="354">
|
||
<emphasis box="[199,320,1617,1642]" italics="true" pageId="19" pageNumber="354">
|
||
<typeStatus box="[199,316,1617,1642]" pageId="19" pageNumber="354">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
Mertola, Corredoura,
|
||
<geoCoordinate box="[568,648,1616,1642]" degrees="37.746" direction="north" orientation="latitude" pageId="19" pageNumber="354" precision="55" value="37.746">37.746</geoCoordinate>
|
||
,
|
||
<geoCoordinate box="[658,733,1616,1642]" degrees="7.642" direction="west" orientation="longitude" pageId="19" pageNumber="354" precision="55" value="-7.642">-7.642</geoCoordinate>
|
||
,
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1984" box="[743,947,1617,1643]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="354" phylum="Tracheophyta" rank="species" species="coccifera">
|
||
<emphasis box="[743,947,1617,1643]" italics="true" pageId="19" pageNumber="354">Quercus coccifera</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[1000,1073,1616,1643]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="19" pageNumber="354" phylum="Tracheophyta" rank="species" species="ilex">
|
||
<emphasis box="[1000,1073,1616,1643]" italics="true" pageId="19" pageNumber="354">Q. ilex</emphasis>
|
||
</taxonomicName>
|
||
wood, 9 ♁♁,
|
||
<specimenCount box="[1230,1271,1617,1643]" pageId="19" pageNumber="354" type="female">1 ♀</specimenCount>
|
||
, 1999, 13 ♁♁,
|
||
<specimenCount box="[151,216,1653,1679]" pageId="19" pageNumber="354" type="female">2 ♀♀</specimenCount>
|
||
, 2000, leg. P. Cardoso, coll. Finnish Museum of Natural History, Helsinki (http://id.luomus.fi/KN.23951);
|
||
<specimenCount pageId="19" pageNumber="354" type="female">1 ♀</specimenCount>
|
||
,
|
||
<specimenCount box="[187,253,1688,1715]" pageId="19" pageNumber="354" type="generic">1 juv.</specimenCount>
|
||
,
|
||
<date box="[264,471,1688,1715]" pageId="19" pageNumber="354" value="2005-11-08">8 November 2005</date>
|
||
,
|
||
<specimenCount box="[483,526,1689,1715]" pageId="19" pageNumber="354" type="female">1 ♀</specimenCount>
|
||
,
|
||
<date box="[540,686,1688,1715]" pageId="19" pageNumber="354" value="2008-04-03">3 April 2008</date>
|
||
,
|
||
<specimenCount box="[698,763,1689,1715]" pageId="19" pageNumber="354" type="female">3 ♀♀</specimenCount>
|
||
,
|
||
<specimenCount box="[777,843,1688,1715]" pageId="19" pageNumber="354" type="generic">3 juv.</specimenCount>
|
||
,
|
||
<date box="[854,1030,1688,1715]" pageId="19" pageNumber="354" value="2013-03-29">29 March 2013</date>
|
||
, leg. M. Řezáč, coll. Crop Research Institute,
|
||
<collectingRegion box="[257,338,1725,1751]" country="Czech Republic" name="Praha" pageId="19" pageNumber="354">Prague</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="354" type="etymology">
|
||
<paragraph blockId="19.[151,1437,1544,2039]" pageId="19" pageNumber="354">
|
||
<emphasis bold="true" box="[199,334,1761,1787]" pageId="19" pageNumber="354">Etymology.</emphasis>
|
||
Named after the Czechoslovak arachnologist Stano Pekár, our friend who helped us to collect material for this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="19" pageNumber="354" type="diagnosis">
|
||
<paragraph blockId="19.[151,1437,1544,2039]" pageId="19" pageNumber="354">
|
||
<emphasis bold="true" box="[199,320,1832,1858]" pageId="19" pageNumber="354">Diagnosis.</emphasis>
|
||
The smallest and more gracile Portuguese
|
||
<taxonomicName box="[792,1004,1832,1858]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[792,911,1833,1858]" italics="true" pageId="19" pageNumber="354">Harpactea</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
It resembles
|
||
<emphasis box="[1149,1390,1832,1858]" italics="true" pageId="19" pageNumber="354">
|
||
<taxonomicName authorityName="Pesarini" authorityYear="1988" box="[1149,1283,1832,1858]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="gaditana">H. gaditana</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Brignoli" authorityYear="1980" box="[1294,1390,1832,1858]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="fageli">H. fageli</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="Řezáč & Cardoso & Řezáčová, 2023" authorityName="Řezáč & Cardoso & Řezáčová" authorityYear="2023" box="[151,302,1868,1894]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="henriquesi" status="sp. nov.">
|
||
<emphasis box="[151,302,1868,1894]" italics="true" pageId="19" pageNumber="354">H. henriquesi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[308,397,1869,1895]" pageId="19" pageNumber="354">
|
||
<taxonomicNameLabel box="[308,397,1869,1895]" pageId="19" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
by slightly elongated unmodified cymbium (
|
||
<figureCitation box="[896,984,1868,1895]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="19" pageNumber="354">Fig. 3D</figureCitation>
|
||
). It can be distinguished from the former species by spherical tegulum, no conductor and long regularly curved embolus, transversal in respect to tegulum (
|
||
<figureCitation captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="19" pageNumber="354">Fig. 4D</figureCitation>
|
||
). The reduced (small, only slightly sclerotised and without spermathecae) vulva resembles
|
||
<emphasis italics="true" pageId="19" pageNumber="354">
|
||
<taxonomicName authorityName="Ferrandez" authorityYear="1990" box="[1229,1396,1940,1966]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="algarvensis">H. algarvensis</taxonomicName>
|
||
,
|
||
<taxonomicName authorityName="Wunderlich" authorityYear="2020" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="tavirensis">H. tavirensis</taxonomicName>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName authority="Řezáč & Cardoso & Řezáčová, 2023" authorityName="Řezáč & Cardoso & Řezáčová" authorityYear="2023" box="[313,428,1977,2002]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="crespoi" status="sp. nov.">
|
||
<emphasis box="[313,428,1977,2002]" italics="true" pageId="19" pageNumber="354">H. crespoi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[435,525,1977,2003]" pageId="19" pageNumber="354">
|
||
<taxonomicNameLabel box="[435,525,1977,2003]" pageId="19" pageNumber="354" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
but it can be distinguished from these species by narrow posterior transversal bar bearing two furrows (
|
||
<figureCitation box="[391,479,2012,2038]" captionStart="FIGURE 6" captionStartId="15.[151,250,1048,1072]" captionTargetBox="[151,1431,181,1020]" captionTargetId="figure-19@15.[151,1436,181,1024]" captionTargetPageId="15" captionText="FIGURE 6. Portuguese Harpactea spiders. Vulva. A. H. fageli. Portugal: Setúbal; B. H. henriquesi sp. nov. Portugal: Torres Novas; C. H.pekari sp. nov. Portugal:Corredoura.Views:dor.—dorsal,fro.—anterior,hin.—posterior, ven.—ventral.Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804286" httpUri="https://zenodo.org/record/7804286/files/figure.png" pageId="19" pageNumber="354">Fig. 6C</figureCitation>
|
||
). It co-occurs with
|
||
<emphasis box="[693,829,2013,2038]" italics="true" pageId="19" pageNumber="354">
|
||
<taxonomicName authorityName="Ferrandez" authorityYear="1982" box="[693,825,2013,2038]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="354" phylum="Arthropoda" rank="species" species="minoccii">H. minoccii</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
from which it differs by smaller and lighter body.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="355" type="description">
|
||
<paragraph blockId="20.[151,1437,150,573]" pageId="20" pageNumber="355">
|
||
<emphasis bold="true" box="[199,345,150,176]" pageId="20" pageNumber="355">Description.</emphasis>
|
||
<emphasis box="[351,537,150,176]" italics="true" pageId="20" pageNumber="355">
|
||
Male (
|
||
<typeStatus box="[423,525,150,176]" pageId="20" pageNumber="355">holotype</typeStatus>
|
||
).
|
||
</emphasis>
|
||
Carapace yellow-orange, head region darker, matting (
|
||
<figureCitation box="[1146,1234,150,176]" captionStart="FIGURE 1" captionStartId="3.[151,250,1773,1797]" captionTargetBox="[151,1436,667,1745]" captionTargetId="figure-280@3.[151,1436,666,1747]" captionTargetPageId="3" captionText="FIGURE 1. Portuguese Harpactea spiders. Males (dorsal view). A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804270" httpUri="https://zenodo.org/record/7804270/files/figure.png" pageId="20" pageNumber="355">Fig. 1D</figureCitation>
|
||
). Sternum yellow, matting. Chelicerae brown. Legs yellow, femora I and pedipalps darker. For measurements and leg spination see the
|
||
<tableCitation box="[192,276,222,248]" captionStart="TABLE 1" captionStartId="17.[152,244,151,175]" captionText="TABLE 1. Measurements and spination of holotypes of the new Portuguese Harpactea species and Harpactea tavirensis." pageId="20" pageNumber="355">Table 1</tableCitation>
|
||
. Relative leg length: IV>I>II>III. Cymbium with only slightly elongated distal part (
|
||
<figureCitation box="[1228,1318,222,248]" captionStart="FIGURE 3" captionStartId="7.[151,248,1802,1826]" captionTargetBox="[151,1428,345,1768]" captionTargetId="figure-52@7.[151,1436,334,1778]" captionTargetPageId="7" captionText="FIGURE 3. Portuguese Harpactea spiders. Male pedipalp. A, C, D, G, H, J, M, N—right pedipalp, prolateral view; B, E, F, I, K, L—left pedipalp, retrolateral view.A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal:Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira; I. H. adicensis sp. nov. Portugal: Sobral da Adiça; J. H. algarvensis. Portugal: Barranco do Velho; K. H. krejcii sp. nov. Portugal: Ameixial; L. H. dolanskyi sp. nov. Portugal: Mertola; M. H. tavirensis. Portugal: Barranco do Velho; N. H. crespoi sp. nov. Portugal: Sobral da Adiça." figureDoi="http://doi.org/10.5281/zenodo.7804278" httpUri="https://zenodo.org/record/7804278/files/figure.png" pageId="20" pageNumber="355">Fig. 3D</figureCitation>
|
||
). Tegulum large, longer than wide (
|
||
<figureCitation box="[430,523,258,284]" captionStart="FIGURE 4" captionStartId="8.[151,250,1860,1884]" captionTargetBox="[151,1436,741,1836]" captionTargetId="figure-289@8.[151,1436,741,1836]" captionTargetPageId="8" captionText="FIGURE 4. Portuguese Harpactea spiders. Right male bulbus. A. H. fageli. Portugal: Setúbal; B. H. gaditana. Portugal: Alcomgosta; C. H. henriquesi sp. nov. Portugal: Torres Novas; D. H. pekari sp. nov. Portugal: Corredoura; E. H. minoccii. Portugal: Mertola; F. H. subiasi. Portugal: Setúbal; G. H. magnibulbi. Portugal: Fóia; H. H. korenkoi sp. nov. Portugal: Carrapateira. Structures: c—conductor, e—embolus, t—tegulum. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804282" httpUri="https://zenodo.org/record/7804282/files/figure.png" pageId="20" pageNumber="355">Fig. 4D</figureCitation>
|
||
). Conductor atrophied. Embolus dark, long, regularly bent, directing to the side. Opisthosoma cylindrical, whitish.
|
||
</paragraph>
|
||
<paragraph blockId="20.[151,1437,150,573]" pageId="20" pageNumber="355">
|
||
<emphasis box="[199,291,330,356]" italics="true" pageId="20" pageNumber="355">Female.</emphasis>
|
||
All somatic characters as described for the male. The vulva is of the
|
||
<typeStatus box="[1066,1114,331,357]" pageId="20" pageNumber="355">type</typeStatus>
|
||
<taxonomicName authorityName="Rezac" authorityYear="2023" box="[1122,1194,330,356]" class="Arachnida" family="Dysderidae" genus="Harpactea" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="355" phylum="Arthropoda" rank="species" species="pekari">
|
||
<emphasis box="[1122,1194,330,356]" italics="true" pageId="20" pageNumber="355">pekari</emphasis>
|
||
</taxonomicName>
|
||
(see the Material and Methods). It is very reduced and does not possess any posterior diverticle (
|
||
<figureCitation box="[989,1078,366,392]" captionStart="FIGURE 6" captionStartId="15.[151,250,1048,1072]" captionTargetBox="[151,1431,181,1020]" captionTargetId="figure-19@15.[151,1436,181,1024]" captionTargetPageId="15" captionText="FIGURE 6. Portuguese Harpactea spiders. Vulva. A. H. fageli. Portugal: Setúbal; B. H. henriquesi sp. nov. Portugal: Torres Novas; C. H.pekari sp. nov. Portugal:Corredoura.Views:dor.—dorsal,fro.—anterior,hin.—posterior, ven.—ventral.Structures: aa—anterior arc, cb—copulatory bursa, kp—keel-like projection, mr—median rod, pd—posterior diverticle, s—spermatheca, tb—posterior transversal bar, th—transversal hinge, wt—whitish tissue. The photos in grey scale are made under the scanning electron microscope." figureDoi="http://doi.org/10.5281/zenodo.7804286" httpUri="https://zenodo.org/record/7804286/files/figure.png" pageId="20" pageNumber="355">Fig. 6C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="20.[151,1437,150,573]" pageId="20" pageNumber="355">
|
||
<emphasis box="[199,324,402,428]" italics="true" pageId="20" pageNumber="355">Variability.</emphasis>
|
||
Male carapace length
|
||
<quantity box="[589,755,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.585" metricValueMax="1.84" metricValueMin="1.33" pageId="20" pageNumber="355" unit="mm" value="1.585" valueMax="1.84" valueMin="1.33">1.33–1.84 mm</quantity>
|
||
(1.58±0.14, N=21), female carapace length
|
||
<quantity box="[1270,1437,402,429]" metricMagnitude="-3" metricUnit="m" metricValue="1.555" metricValueMax="1.6" metricValueMin="1.51" pageId="20" pageNumber="355" unit="mm" value="1.555" valueMax="1.6" valueMin="1.51">1.51–1.60 mm</quantity>
|
||
(1.54±0.04, N=4).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="20" pageNumber="355" type="biology_ecology">
|
||
<paragraph blockId="20.[151,1437,150,573]" pageId="20" pageNumber="355">
|
||
<emphasis bold="true" box="[199,300,474,500]" pageId="20" pageNumber="355">Ecology.</emphasis>
|
||
Adults are found in early spring in leaf litter and under stones on humid northern slopes with
|
||
<taxonomicName class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="20" pageNumber="355" phylum="Tracheophyta" rank="species" species="ilex">
|
||
<emphasis italics="true" pageId="20" pageNumber="355">Quercus ilex</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName box="[246,467,510,536]" class="Pinopsida" family="Cupressaceae" genus="Juniperus" kingdom="Plantae" order="Pinales" pageId="20" pageNumber="355" phylum="Tracheophyta" rank="species" species="turbinata">
|
||
<emphasis box="[246,467,510,536]" italics="true" pageId="20" pageNumber="355">Juniperus turbinata</emphasis>
|
||
</taxonomicName>
|
||
bush.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,1315,546,573]" pageId="20" pageNumber="355" type="distribution">
|
||
<paragraph blockId="20.[151,1437,150,573]" box="[199,1315,546,573]" pageId="20" pageNumber="355">
|
||
<emphasis bold="true" box="[199,353,546,572]" pageId="20" pageNumber="355">Distribution.</emphasis>
|
||
Known only from the valley of the river Guadiana near Mertola in southern
|
||
<collectingCountry box="[1212,1311,546,572]" name="Portugal" pageId="20" pageNumber="355">Portugal</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |