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<document id="5F23EA5FCD0C011B33D5B1C0208BE562" ID-CLB-Dataset="32408" ID-DOI="10.11646/zootaxa.4306.1.11" ID-GBIF-Dataset="ed19bdba-db4a-4caa-98ea-75450b624e21" ID-ISSN="1175-5326" ID-Zenodo-Dep="843081" ID-ZooBank="7C11D4DC-7A47-40C6-A5FF-358667B56AD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1502782628146" checkinUser="plazi" docAuthor="Král, David, Hrůzová, Lucie, Lu, Yuanyuan &amp; Bai, Ming" docDate="2017" docId="039C7918FFD4D134E9BAC446FA7DA4EB" docLanguage="en" docName="zootaaxa.4306.1.11.pdf" docOrigin="Zootaxa 4306 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Glaresis Král &amp; Hrůzová &amp; Lu &amp; Bai 2017" docType="key" docVersion="7" lastPageNumber="148" masterDocId="FFA50160FFD7D137E92DC12BFFEBA31F" masterDocTitle="First records of Glaresidae (Coleoptera) in China, with the description of a new species from Inner Mongolia and Shaanxi" masterLastPageNumber="150" masterPageNumber="145" pageNumber="148" updateTime="1698466543568" updateUser="ExternalLinkService">
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<mods:title id="8C666995F1447E557DBB59EAF99DC8C6">First records of Glaresidae (Coleoptera) in China, with the description of a new species from Inner Mongolia and Shaanxi</mods:title>
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<mods:namePart id="6131A3C072C6D8233E4C94938BA93A84">Král, David</mods:namePart>
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<mods:namePart id="B13ED728E489BDD5BBCD0AA6E6C10703">Hrůzová, Lucie</mods:namePart>
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<mods:namePart id="5059CEA6C6FE73E2C1D2B399AED5CE9F">Lu, Yuanyuan</mods:namePart>
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<mods:namePart id="5104A043166F7A2AE5FD70259D600380">Bai, Ming</mods:namePart>
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<treatment id="039C7918FFD4D134E9BAC446FA7DA4EB" ID-DOI="http://doi.org/10.5281/zenodo.6033679" ID-GBIF-Taxon="132650243" ID-Zenodo-Dep="6033679" LSID="urn:lsid:plazi:treatment:039C7918FFD4D134E9BAC446FA7DA4EB" httpUri="http://treatment.plazi.org/id/039C7918FFD4D134E9BAC446FA7DA4EB" lastPageNumber="148" pageId="3" pageNumber="148">
<subSubSection id="C32F9B85FFD4D134E9BAC446FC0CA6B8" pageId="3" pageNumber="148" type="nomenclature">
<paragraph id="8B8AC80EFFD4D134E9BAC446FC0CA6B8" blockId="3.[151,1436,1389,2036]" pageId="3" pageNumber="148">
<heading id="D0C27F62FFD4D134E9BAC446FC51A698" bold="true" box="[151,954,1389,1415]" fontSize="11" level="1" pageId="3" pageNumber="148" reason="1">
<emphasis id="B941141CFFD4D134E9BAC446FC51A698" bold="true" box="[151,954,1389,1415]" pageId="3" pageNumber="148">
Key for identifying Middle and Central Asian species of
<taxonomicName id="4C35B38DFFD4D134EA7AC446FC51A698" ID-CoL="4NT7" authorityName="Král &amp; Hrůzová &amp; Lu &amp; Bai" authorityYear="2017" box="[855,954,1389,1415]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="genus">
<emphasis id="B941141CFFD4D134EA7AC446FC51A698" bold="true" box="[855,954,1389,1415]" italics="true" pageId="3" pageNumber="148">Glaresis</emphasis>
</taxonomicName>
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Species distribution adopted from
<bibRefCitation id="EFA4B5FFFFD4D134EB2BC4BBFD56A6B8" author="Nikolajev" box="[518,701,1424,1447]" pageId="3" pageNumber="148" refString="Nikolajev, G. V. (1987) Plastinchatousye zhuki (Coleoptera, Scarabaeoidea) Kazakhstana i Srednei Azii [Scarabaeoidea (Coleoptera) of Kazakhstan and Middle Asia]. Izdatelstvo Nauka KazSSR, Alma-Ata, Kazakhstan, 232 pp. [in Russian]" type="book" year="1987">Nikolajev (1987)</bibRefCitation>
and
<bibRefCitation id="EFA4B5FFFFD4D134EBDDC4BBFC08A6B8" author="Kral" box="[752,995,1424,1447]" pageId="3" pageNumber="148" refString="Kral, D. &amp; Bezdek, A. (2016) Family Glaresidae. In: Lobl, I. &amp; Lobl, D. (Eds.), Catalogue of Palaearctic Coleoptera. Folume 3. Scarabaeoidea - Scirtoidea - Dasciloidea - Buprestoidea - Byrrhoidea. Revised and Updated Edition. E. J. Brill, Leiden, p. 58" type="journal article" year="2016">Král &amp; Bezděk (2016)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32F9B85FFD4D134E9BAC499FA7DA4EB" pageId="3" pageNumber="148" type="key">
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1 Anterior margin of clypeus with four teeth; body length 4.55.0 mm; Middle Asia and
<collectingCountry id="F322889EFFD4D134ED24C499FB90A6D8" box="[1033,1147,1458,1479]" name="Kazakhstan" pageId="3" pageNumber="148">Kazakhstan</collectingCountry>
,
<collectingCountry id="F322889EFFD4D134EDA8C499FB47A6D8" box="[1157,1196,1458,1479]" name="Iran" pageId="3" pageNumber="148">Iran</collectingCountry>
....................................... ............................................................................................................................................................
<taxonomicName id="4C35B38DFFD4D134ED50C4E4FA77A6FB" ID-CoL="3G5XS" authority="Solsky, 1870" authorityName="Solsky" authorityYear="1870" box="[1149,1436,1487,1508]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="beckeri">
<emphasis id="B941141CFFD4D134ED50C4E4FAF1A6FB" box="[1149,1306,1487,1508]" italics="true" pageId="3" pageNumber="148">Glaresis beckeri</emphasis>
Solsky, 1870
</taxonomicName>
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</paragraph>
<paragraph id="8B8AC80EFFD4D134E9BAC4C6FA77A51D" blockId="3.[151,1436,1389,2036]" box="[151,1436,1517,1538]" pageId="3" pageNumber="148">
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- Anterior margin of clypeus without teeth; body length 3.0
<quantity id="4CCD65EBFFD4D134EBD2C4C6FCACA51D" box="[767,839,1517,1538]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="3" pageNumber="148" unit="mm" value="4.5">4.5 mm</quantity>
............................................................................................... 2
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<paragraph id="8B8AC80EFFD4D134E9BAC721FA77A523" blockId="3.[151,1436,1389,2036]" pageId="3" pageNumber="148">
<materialsCitation id="3B5DC253FFD4D134E9BAC721FA77A523" ID-GBIF-Occurrence="1586033252" country="Kazakhstan" location="central Europe" pageId="3" pageNumber="148" specimenCount="1" stateProvince="Balkan">
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2 Pronotum with four shallow grooves beside medial longitudinal groove; mesotibial fossae on meso-metaventrum absent; body length 3.0
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; from
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and the
<collectingRegion id="49F106ECFFD4D134EBB6C70CFD34A523" box="[667,735,1575,1596]" country="Turkmenistan" name="Balkan Welayaty" pageId="3" pageNumber="148">Balkan</collectingRegion>
Peninsula to
<collectingCountry id="F322889EFFD4D134EA4CC70CFC3FA523" box="[865,980,1575,1596]" name="Kazakhstan" pageId="3" pageNumber="148">Kazakhstan</collectingCountry>
. ...........................
<taxonomicName id="4C35B38DFFD4D134EDABC70CFA77A523" ID-CoL="3G5ZY" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[1158,1436,1575,1596]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="rufa">
<emphasis id="B941141CFFD4D134EDABC70CFAEEA523" box="[1158,1285,1575,1596]" italics="true" pageId="3" pageNumber="148">Glaresis rufa</emphasis>
Erichson, 1848
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<paragraph id="8B8AC80EFFD4D134E9BAC76EFA70A568" blockId="3.[151,1436,1389,2036]" pageId="3" pageNumber="148">
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- Pronotum almost flat, only with indistinct, shallow medial longitudinal groove (
<figureCitation id="130ED48BFFD4D134EACBC76EFBC7A545" box="[998,1068,1605,1626]" captionStart="FIGURES 1 10" captionStartId="2.[151,258,1913,1935]" captionTargetBox="[176,1413,290,1861]" captionTargetId="figure@2.[151,1436,277,1891]" captionTargetPageId="2" captionText="FIGURES 1 10. Glaresis ordosensis Král, Hrůzová, Lu, &amp; Bai, new species. 1, Habitus, holotype, ♂, dorsal view; 2, habitus, allotype, ♀, ventral view; 3, aedeagus, dorsal view; 4, head, dorsal view; 5, meso-metaventral oblique grooves, ventral view; 6, detail of left elytron, dorsal view; 7, left mesothoracic leg, dorsal view; 8, left mesothoracic leg, ventral view; 9, right metathoracic leg, dorsal view; 10, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843083/files/figure.png" pageId="3" pageNumber="148">Figs. 1</figureCitation>
,
<figureCitation id="130ED48BFFD4D134ED16C76EFB92A545" box="[1083,1145,1605,1626]" captionStart="FIGURES 11 20" captionStartId="4.[151,258,1753,1775]" captionTargetBox="[176,1418,206,1720]" captionTargetId="figure@4.[152,1435,194,1732]" captionTargetPageId="4" captionText="FIGURES 11 20. Glaresis orientalis Medvedev, 1976 ♂ from Alxa, China (11) and Bayankhongor Province, Mongolia (12 20). 11, habitus, dorsal view; 12, habitus, dorsal view; 13, aedeagus, dorsal view; 14, head, dorsal view; 15, meso-metaventral oblique grooves, ventral view; 16, detail of left elytron, dorsal view; 17, left mesothoracic leg, dorsal view; 18, left mesothoracic leg, ventral view; 19, right metathoracic leg, dorsal view; 20, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843085/files/figure.png" pageId="3" pageNumber="148">1112</figureCitation>
); mesotibial fossae on mesometaventrum present (
<figureCitation id="130ED48BFFD4D134E8B0C749FE0AA568" box="[413,481,1634,1655]" captionStart="FIGURES 1 10" captionStartId="2.[151,258,1913,1935]" captionTargetBox="[176,1413,290,1861]" captionTargetId="figure@2.[151,1436,277,1891]" captionTargetPageId="2" captionText="FIGURES 1 10. Glaresis ordosensis Král, Hrůzová, Lu, &amp; Bai, new species. 1, Habitus, holotype, ♂, dorsal view; 2, habitus, allotype, ♀, ventral view; 3, aedeagus, dorsal view; 4, head, dorsal view; 5, meso-metaventral oblique grooves, ventral view; 6, detail of left elytron, dorsal view; 7, left mesothoracic leg, dorsal view; 8, left mesothoracic leg, ventral view; 9, right metathoracic leg, dorsal view; 10, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843083/files/figure.png" pageId="3" pageNumber="148">Figs. 5</figureCitation>
,
<figureCitation id="130ED48BFFD4D134E8C1C749FDEBA568" box="[492,512,1634,1655]" captionStart="FIGURES 11 20" captionStartId="4.[151,258,1753,1775]" captionTargetBox="[176,1418,206,1720]" captionTargetId="figure@4.[152,1435,194,1732]" captionTargetPageId="4" captionText="FIGURES 11 20. Glaresis orientalis Medvedev, 1976 ♂ from Alxa, China (11) and Bayankhongor Province, Mongolia (12 20). 11, habitus, dorsal view; 12, habitus, dorsal view; 13, aedeagus, dorsal view; 14, head, dorsal view; 15, meso-metaventral oblique grooves, ventral view; 16, detail of left elytron, dorsal view; 17, left mesothoracic leg, dorsal view; 18, left mesothoracic leg, ventral view; 19, right metathoracic leg, dorsal view; 20, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843085/files/figure.png" pageId="3" pageNumber="148">15</figureCitation>
)..................................................................................................................................................... 3
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30C1D0ABFFD4D134E9BAC754FA70A45B" pageId="3" pageNumber="148">
<paragraph id="8B8AC80EFFD4D134E9BAC754FA77A5D0" blockId="3.[151,1436,1389,2036]" pageId="3" pageNumber="148">
<keyLead id="30C46B3BFFD4D134E9BAC754FA77A5D0" pageId="3" pageNumber="148">
3 Anterior margin of clypeus broadly rounded; posterior angles of pronotum not protruding as spines; mesotibia shorter with distal part of outer edge distinctly emarginate; mesotibial fossae on meso-metaventrum present, shallowly impressed, without sharp edges; body length
<quantity id="4CCD65EBFFD4D134E8AFC791FE13A5D0" box="[386,504,1722,1743]" metricMagnitude="-3" metricUnit="m" metricValue="3.95" metricValueMax="4.5" metricValueMin="3.4" pageId="3" pageNumber="148" unit="mm" value="3.95" valueMax="4.5" valueMin="3.4">3.44.5 mm</quantity>
; Middle Asia and
<collectingCountry id="F322889EFFD4D134EB8AC791FCF2A5D0" box="[679,793,1722,1743]" name="Kazakhstan" pageId="3" pageNumber="148">Kazakhstan</collectingCountry>
,
<collectingCountry id="F322889EFFD4D134EA0EC791FC70A5D0" box="[803,923,1722,1743]" name="Afghanistan" pageId="3" pageNumber="148">Afghanistan</collectingCountry>
,
<collectingCountry id="F322889EFFD4D134EA89C791FC26A5D0" box="[932,973,1722,1743]" name="Iraq" pageId="3" pageNumber="148">Iraq</collectingCountry>
,
<collectingCountry id="F322889EFFD4D134EAFAC791FBC0A5D0" box="[983,1067,1722,1743]" name="Pakistan" pageId="3" pageNumber="148">Pakistan</collectingCountry>
.........
<taxonomicName id="4C35B38DFFD4D134ED46C791FA77A5D0" ID-CoL="3G5ZQ" authority="Semenov, 1892" authorityName="Semenov" authorityYear="1892" box="[1131,1436,1722,1743]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="148" phylum="Arthropoda" rank="species" species="oxiana">
<emphasis id="B941141CFFD4D134ED46C791FAE8A5D0" box="[1131,1283,1722,1743]" italics="true" pageId="3" pageNumber="148">Glaresis oxiana</emphasis>
Semenov, 1892
</taxonomicName>
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</paragraph>
<paragraph id="8B8AC80EFFD4D134E9BAC7FCFA70A45B" blockId="3.[151,1436,1389,2036]" pageId="3" pageNumber="148">
<keyLead id="30C46B3BFFD4D134E9BAC7FCFA70A45B" pageId="3" pageNumber="148">
- Anterior margin of clypeus straight to shallowly emarginate (
<figureCitation id="130ED48BFFD4D134EA22C7FCFCB8A5F3" box="[783,851,1751,1772]" captionStart="FIGURES 1 10" captionStartId="2.[151,258,1913,1935]" captionTargetBox="[176,1413,290,1861]" captionTargetId="figure@2.[151,1436,277,1891]" captionTargetPageId="2" captionText="FIGURES 1 10. Glaresis ordosensis Král, Hrůzová, Lu, &amp; Bai, new species. 1, Habitus, holotype, ♂, dorsal view; 2, habitus, allotype, ♀, ventral view; 3, aedeagus, dorsal view; 4, head, dorsal view; 5, meso-metaventral oblique grooves, ventral view; 6, detail of left elytron, dorsal view; 7, left mesothoracic leg, dorsal view; 8, left mesothoracic leg, ventral view; 9, right metathoracic leg, dorsal view; 10, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843083/files/figure.png" pageId="3" pageNumber="148">Figs. 4</figureCitation>
,
<figureCitation id="130ED48BFFD4D134EA70C7FCFC9CA5F3" box="[861,887,1751,1772]" captionStart="FIGURES 11 20" captionStartId="4.[151,258,1753,1775]" captionTargetBox="[176,1418,206,1720]" captionTargetId="figure@4.[152,1435,194,1732]" captionTargetPageId="4" captionText="FIGURES 11 20. Glaresis orientalis Medvedev, 1976 ♂ from Alxa, China (11) and Bayankhongor Province, Mongolia (12 20). 11, habitus, dorsal view; 12, habitus, dorsal view; 13, aedeagus, dorsal view; 14, head, dorsal view; 15, meso-metaventral oblique grooves, ventral view; 16, detail of left elytron, dorsal view; 17, left mesothoracic leg, dorsal view; 18, left mesothoracic leg, ventral view; 19, right metathoracic leg, dorsal view; 20, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843085/files/figure.png" pageId="3" pageNumber="148">14</figureCitation>
); posterior angles of pronotum protruding more or less as spines (
<figureCitation id="130ED48BFFD4D134E83EC7DEFEB3A415" box="[275,344,1781,1802]" captionStart="FIGURES 1 10" captionStartId="2.[151,258,1913,1935]" captionTargetBox="[176,1413,290,1861]" captionTargetId="figure@2.[151,1436,277,1891]" captionTargetPageId="2" captionText="FIGURES 1 10. Glaresis ordosensis Král, Hrůzová, Lu, &amp; Bai, new species. 1, Habitus, holotype, ♂, dorsal view; 2, habitus, allotype, ♀, ventral view; 3, aedeagus, dorsal view; 4, head, dorsal view; 5, meso-metaventral oblique grooves, ventral view; 6, detail of left elytron, dorsal view; 7, left mesothoracic leg, dorsal view; 8, left mesothoracic leg, ventral view; 9, right metathoracic leg, dorsal view; 10, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843083/files/figure.png" pageId="3" pageNumber="148">Figs. 1</figureCitation>
,
<figureCitation id="130ED48BFFD4D134E845C7DEFE4DA415" box="[360,422,1781,1802]" captionStart="FIGURES 11 20" captionStartId="4.[151,258,1753,1775]" captionTargetBox="[176,1418,206,1720]" captionTargetId="figure@4.[152,1435,194,1732]" captionTargetPageId="4" captionText="FIGURES 11 20. Glaresis orientalis Medvedev, 1976 ♂ from Alxa, China (11) and Bayankhongor Province, Mongolia (12 20). 11, habitus, dorsal view; 12, habitus, dorsal view; 13, aedeagus, dorsal view; 14, head, dorsal view; 15, meso-metaventral oblique grooves, ventral view; 16, detail of left elytron, dorsal view; 17, left mesothoracic leg, dorsal view; 18, left mesothoracic leg, ventral view; 19, right metathoracic leg, dorsal view; 20, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843085/files/figure.png" pageId="3" pageNumber="148">1112</figureCitation>
); mesotibia longer with distal part of outer edge straight or shallowly emarginate (
<figureCitation id="130ED48BFFD4D134EDC9C7DEFAA8A415" box="[1252,1347,1781,1802]" captionStart="FIGURES 1 10" captionStartId="2.[151,258,1913,1935]" captionTargetBox="[176,1413,290,1861]" captionTargetId="figure@2.[151,1436,277,1891]" captionTargetPageId="2" captionText="FIGURES 1 10. Glaresis ordosensis Král, Hrůzová, Lu, &amp; Bai, new species. 1, Habitus, holotype, ♂, dorsal view; 2, habitus, allotype, ♀, ventral view; 3, aedeagus, dorsal view; 4, head, dorsal view; 5, meso-metaventral oblique grooves, ventral view; 6, detail of left elytron, dorsal view; 7, left mesothoracic leg, dorsal view; 8, left mesothoracic leg, ventral view; 9, right metathoracic leg, dorsal view; 10, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843083/files/figure.png" pageId="3" pageNumber="148">Figs. 78</figureCitation>
,
<figureCitation id="130ED48BFFD4D134EC7CC7DEFA7BA415" box="[1361,1424,1781,1802]" captionStart="FIGURES 11 20" captionStartId="4.[151,258,1753,1775]" captionTargetBox="[176,1418,206,1720]" captionTargetId="figure@4.[152,1435,194,1732]" captionTargetPageId="4" captionText="FIGURES 11 20. Glaresis orientalis Medvedev, 1976 ♂ from Alxa, China (11) and Bayankhongor Province, Mongolia (12 20). 11, habitus, dorsal view; 12, habitus, dorsal view; 13, aedeagus, dorsal view; 14, head, dorsal view; 15, meso-metaventral oblique grooves, ventral view; 16, detail of left elytron, dorsal view; 17, left mesothoracic leg, dorsal view; 18, left mesothoracic leg, ventral view; 19, right metathoracic leg, dorsal view; 20, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843085/files/figure.png" pageId="3" pageNumber="148">1718</figureCitation>
); mesotibial fossae on meso-metaventrum present, distinctly impressed, with sharp edges, divergent posteriad; body length 4.0
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(
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,
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) ................................................................................................................................................................................ 4
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4 Distal part of outer edge of mesotibiae straight, bearing at least 79 short, stout spines (
<figureCitation id="130ED48BFFD4D134ED19C666FB4DA47D" box="[1076,1190,1869,1890]" captionStart="FIGURES 11 20" captionStartId="4.[151,258,1753,1775]" captionTargetBox="[176,1418,206,1720]" captionTargetId="figure@4.[152,1435,194,1732]" captionTargetPageId="4" captionText="FIGURES 11 20. Glaresis orientalis Medvedev, 1976 ♂ from Alxa, China (11) and Bayankhongor Province, Mongolia (12 20). 11, habitus, dorsal view; 12, habitus, dorsal view; 13, aedeagus, dorsal view; 14, head, dorsal view; 15, meso-metaventral oblique grooves, ventral view; 16, detail of left elytron, dorsal view; 17, left mesothoracic leg, dorsal view; 18, left mesothoracic leg, ventral view; 19, right metathoracic leg, dorsal view; 20, right metathoracic leg, ventral view." httpUri="https://zenodo.org/record/843085/files/figure.png" pageId="3" pageNumber="148">Figs 1718</figureCitation>
); basal external tooth on mesotibia straight basally; aedeagus as in
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; body length
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- Distal part of outer edge of mesotibiae shallowly emarginate, bearing 46 short, stout spines (
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); basal external tooth on mesotibia emarginate basally; aedeagus as in
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; body length 4.0
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.........................................................................................................................................................
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