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<document id="F179DF855E12C6F4A8A9608720D6E038" ID-DOI="10.5281/zenodo.13244519" ID-ISSN="2345-7600" ID-Zenodo-Dep="13244519" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722970045800" checkinUser="felipe" docAuthor="Ng, Peter K. L. &amp; Liu, H. - C." docDate="2004" docId="039C6C2F691DFFE804D3F9A9FC7A43A2" docLanguage="en" docName="RafflesBZool.52.1.239-249.pdf" docOrigin="Raffles Bulletin of Zoology 52 (1)" docStyle="DocumentStyle:19380AF0FBA5B2F218DD253A34ABE57F.5:RafflesBZool.2003-2005.journal_article" docStyleId="19380AF0FBA5B2F218DD253A34ABE57F" docStyleName="RafflesBZool.2003-2005.journal_article" docStyleVersion="5" docTitle="Geosesarma De Man 1892" docType="treatment" docVersion="2" lastPageNumber="240" masterDocId="FFA51457691DFFE9073EFFDFFF82411A" masterDocTitle="Geosesarma Hednon, A New Species Of Terrestrial Crab (Crustacea: Decapoda: Brachyura: Sesarmidae) From Taiwan And The Philippines" masterLastPageNumber="249" masterPageNumber="239" pageNumber="239" updateTime="1722974637517" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="9EFAAE5AD46419F002EE3DA6BE8125C3">Geosesarma Hednon, A New Species Of Terrestrial Crab (Crustacea: Decapoda: Brachyura: Sesarmidae) From Taiwan And The Philippines</mods:title>
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<mods:namePart id="3839C75A3AA083EAB2655A023937B921">Ng, Peter K. L.</mods:namePart>
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<mods:namePart id="A066D7733FE5D7B117AE9E0F4FCDA19A">Liu, H. - C.</mods:namePart>
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<mods:date id="E01D2681BFD2CCFA5559109B5059FA44">2004</mods:date>
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<treatment id="039C6C2F691DFFE804D3F9A9FC7A43A2" LSID="urn:lsid:plazi:treatment:039C6C2F691DFFE804D3F9A9FC7A43A2" httpUri="http://treatment.plazi.org/id/039C6C2F691DFFE804D3F9A9FC7A43A2" lastPageId="1" lastPageNumber="240" pageId="0" pageNumber="239">
<subSubSection id="C32F8EB2691DFFE904D3F9A9FA9A4794" box="[1005,1304,1654,1678]" pageId="0" pageNumber="239" type="nomenclature">
<paragraph id="8B8ADD39691DFFE904D3F9A9FA9A4794" blockId="0.[1005,1304,1654,1678]" box="[1005,1304,1654,1678]" pageId="0" pageNumber="239">
<heading id="D0C26A55691DFFE904D3F9A9FA9A4794" bold="true" box="[1005,1304,1654,1678]" centered="true" fontSize="10" level="2" pageId="0" pageNumber="239" reason="2">
<taxonomicName id="4C35A6BA691DFFE904D3F9A9FA9A4794" ID-CoL="7P3H8" authority="De Man 1892" authorityName="De Man" authorityYear="1892" box="[1005,1304,1654,1678]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="B941012B691DFFE904D3F9A9FA9A4794" bold="true" box="[1005,1304,1654,1678]" pageId="0" pageNumber="239">
<emphasis id="B941012B691DFFE904D3F9A9FBF64794" bold="true" box="[1005,1140,1654,1678]" italics="true" pageId="0" pageNumber="239">Geosesarma</emphasis>
De
<bibRefCitation id="EFA4A0C8691DFFE9039BF9A9FA9A4794" author="Man, J" box="[1189,1304,1654,1678]" pageId="0" pageNumber="239" pagination="265 - 527" refId="ref5526" refString="Man, J. G., De, 1892. Decapoden des Indischen Archipels. In: Weber, M. (ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 2: 265 - 527, pls. 15 - 29." type="book chapter" year="1892">Man 1892</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F8EB2691DFFE8047DF969FC7A43A2" lastPageId="1" lastPageNumber="240" pageId="0" pageNumber="239" type="discussion">
<paragraph id="8B8ADD39691DFFE8047DF969FD294342" blockId="0.[835,1475,1718,2094]" lastBlockId="1.[113,753,193,600]" lastPageId="1" lastPageNumber="240" pageId="0" pageNumber="239">
<emphasis id="B941012B691DFFE9047DF969FC4147D4" bold="true" box="[835,963,1718,1742]" italics="true" pageId="0" pageNumber="239">Remarks. –</emphasis>
The genus
<taxonomicName id="4C35A6BA691DFFE90374F968FAF147D7" authority="De Man 1892" authorityName="De Man" authorityYear="1892" box="[1098,1395,1718,1742]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="B941012B691DFFE90374F968FB5247D4" box="[1098,1232,1719,1742]" italics="true" pageId="0" pageNumber="239">Geosesarma</emphasis>
De
<bibRefCitation id="EFA4A0C8691DFFE9023DF969FAF147D7" author="Man, J" box="[1283,1395,1718,1741]" pageId="0" pageNumber="239" pagination="265 - 527" refId="ref5526" refString="Man, J. G., De, 1892. Decapoden des Indischen Archipels. In: Weber, M. (ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 2: 265 - 527, pls. 15 - 29." type="book chapter" year="1892">Man 1892</bibRefCitation>
</taxonomicName>
(sensu Serène &amp; Soh, 1970) is clearly not homogeneous, and
<bibRefCitation id="EFA4A0C8691DFFE9029FF909FC3F4617" author="Ng, P" pageId="0" pageNumber="239" refId="ref5693" refString="Ng, P. K. L., 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, pp. i-viii, 1 - 156, 4 colour pls." type="journal volume" year="1988">Ng (1988: 119)</bibRefCitation>
recognised three separate groups for convenience. As has been discussed by some authors (see
<bibRefCitation id="EFA4A0C8691DFFE90224F8C9FA374637" author="Ng, P" box="[1306,1461,1814,1837]" pageId="0" pageNumber="239" refId="ref5693" refString="Ng, P. K. L., 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, pp. i-viii, 1 - 156, 4 colour pls." type="journal volume" year="1988">Ng, 1988: 118</bibRefCitation>
), the genus
<taxonomicName id="4C35A6BA691DFFE90494F8E8FBAF4654" authorityName="De Man" authorityYear="1892" box="[938,1069,1847,1870]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="B941012B691DFFE90494F8E8FBAF4654" box="[938,1069,1847,1870]" italics="true" pageId="0" pageNumber="239">Geosesarma</emphasis>
</taxonomicName>
is not well defined, and is characterised mainly by their wholly freshwater habits, swollen basal antennular segment and large eggs (ca. 1.0 mm in diameter or larger). Many other sesarmid genera, however, also have representatives living in fresh water habitats, sometimes great distances from the sea. The swollen antennular segment is a useful character, but is not always reliable as it is also evident in members of other genera (e.g.
<taxonomicName id="4C35A6BA691DFFE90392F7C8FAA74934" authorityName="Serene &amp; Soh" authorityYear="1970" box="[1196,1317,2071,2094]" class="Malacostraca" family="Sesarmidae" genus="Labuanium" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="239" phylum="Arthropoda" rank="genus">
<emphasis id="B941012B691DFFE90392F7C8FAA74934" box="[1196,1317,2071,2094]" italics="true" pageId="0" pageNumber="239">Labuanium</emphasis>
</taxonomicName>
) (unpublished data). The egg sizes of many species currently referred to
<taxonomicName id="4C35A6BA691CFFE8074FFF3EFF7541E2" authorityName="De Man" authorityYear="1892" box="[113,247,225,248]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis id="B941012B691CFFE8074FFF3EFF7541E2" box="[113,247,225,248]" italics="true" pageId="1" pageNumber="240">Geosesarma</emphasis>
</taxonomicName>
are still not known, and it is doubtful if this character alone is sufficient to diagnose a genus. The development of only two species is known for certain,
<taxonomicName id="4C35A6BA691CFFE805E8FEFEFF514042" baseAuthorityName="Nobili" baseAuthorityYear="1903" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="peraccae">
<emphasis id="B941012B691CFFE805E8FEFEFF514042" italics="true" pageId="1" pageNumber="240">G. peraccae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C35A6BA691CFFE80639FE9EFE284042" authorityName="PKL Ng &amp; CGS Tan" authorityYear="1995" box="[263,426,321,344]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="notophorum">
<emphasis id="B941012B691CFFE80639FE9EFE284042" box="[263,426,321,344]" italics="true" pageId="1" pageNumber="240">G. notophorum</emphasis>
</taxonomicName>
.
<taxonomicName id="4C35A6BA691CFFE80686FE9EFD244042" baseAuthorityName="Nobili" baseAuthorityYear="1903" box="[440,678,321,344]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="peraccae">
<emphasis id="B941012B691CFFE80686FE9EFD244042" box="[440,678,321,344]" italics="true" pageId="1" pageNumber="240">Geosesarma peraccae</emphasis>
</taxonomicName>
has an abbreviated development, with the advanced zoeae possessing their own food reserves and developing rapidly in the burrows (
<bibRefCitation id="EFA4A0C8691CFFE80747FE7EFF6E40A2" author="Soh, C" box="[121,236,417,440]" pageId="1" pageNumber="240" pagination="357 - 370" refId="ref6129" refString="Soh, C. L., 1969. Abbreviated development of a non-marine crab, Sesarma (Geosesarma) perracae (Brachyura; Grapsidae) from Singapore. Journal of Zoology, 158: 357 - 370." type="journal article" year="1969">Soh, 1969</bibRefCitation>
), while
<taxonomicName id="4C35A6BA691CFFE80678FE7EFE6940A2" authorityName="PKL Ng &amp; CGS Tan" authorityYear="1995" box="[326,491,417,440]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="notophorum">
<emphasis id="B941012B691CFFE80678FE7EFE6940A2" box="[326,491,417,440]" italics="true" pageId="1" pageNumber="240">G. notophorum</emphasis>
</taxonomicName>
has direct development with the young crabs staying on the mother’s back for a short time (Ng &amp; Tan, 1995). Large eggs and abbreviated development have also been reported for other inland and coastal sesarmid taxa (
<bibRefCitation id="EFA4A0C8691CFFE80654FDFEFD8F4322" author="Hartnoll, R" box="[362,525,545,568]" pageId="1" pageNumber="240" pagination="145 - 169" refId="ref5496" refString="Hartnoll, R. G., 1964. The freshwater grapsid crabs of Jamaica. Proceedings of the Linnaean Society, London, 175: 145 - 169." type="journal article" year="1964">Hartnoll, 1964</bibRefCitation>
; Schubart &amp; Cuesta, 1998;
<bibRefCitation id="EFA4A0C8691CFFE80789FD9EFE074342" author="Cuesta, J &amp; Schuh, R" box="[183,389,577,600]" pageId="1" pageNumber="240" pagination="26 - 41" refId="ref5329" refString="Cuesta, J. A., M. Schuh, R. Diesel &amp; C. D. Schubart, 1999. Abbreviated larval development of Armases miersii (Grapsidae: Sesarminae), a crab that breeds in supralittoral rock pools. Journal of Crustacean Biology, 19 (1): 26 - 41." type="journal article" year="1999">Cuesta et al., 1999</bibRefCitation>
; Schubart &amp; Ng, in prep.).
</paragraph>
<paragraph id="8B8ADD39691CFFE8074FFD5EFC7A43A2" blockId="1.[113,753,641,920]" lastBlockId="1.[813,1452,193,696]" pageId="1" pageNumber="240">
The present description of a new species,
<taxonomicName id="4C35A6BA691CFFE80502FD5EFD2C4382" authorityName="Ng &amp; Liu" authorityYear="2004" box="[572,686,641,664]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="hednon">
<emphasis id="B941012B691CFFE80502FD5EFD2C4382" box="[572,686,641,664]" italics="true" pageId="1" pageNumber="240">G. hednon</emphasis>
</taxonomicName>
, from the central
<collectingCountry id="F3229DA9691CFFE807D6FD7EFEE243A2" box="[232,352,673,696]" name="Philippines" pageId="1" pageNumber="240">Philippines</collectingCountry>
and
<collectingCountry id="F3229DA9691CFFE806AAFD7EFE6643A2" box="[404,484,673,696]" name="Taiwan" pageId="1" pageNumber="240">Taiwan</collectingCountry>
is noteworthy as the two sites are some distance apart. However, many ovigerous females of
<taxonomicName id="4C35A6BA691CFFE807D0FD3EFEE243E2" authorityName="Ng &amp; Liu" authorityYear="2004" box="[238,352,737,760]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="hednon">
<emphasis id="B941012B691CFFE807D0FD3EFEE243E2" box="[238,352,737,760]" italics="true" pageId="1" pageNumber="240">G. hednon</emphasis>
</taxonomicName>
had been obtained and the eggs are small, much smaller than those known for other
<taxonomicName id="4C35A6BA691CFFE80555FCDEFF3C4222" pageId="1" pageNumber="240">
<emphasis id="B941012B691CFFE80555FCDEFD724202" box="[619,752,769,792]" italics="true" pageId="1" pageNumber="240">Geosesarma</emphasis>
species
</taxonomicName>
(see
<bibRefCitation id="EFA4A0C8691CFFE807C2FCFEFE1A4222" author="Ng, P" box="[252,408,801,824]" pageId="1" pageNumber="240" refId="ref5693" refString="Ng, P. K. L., 1988. The Freshwater Crabs of Peninsular Malaysia and Singapore. Department of Zoology, National University of Singapore, Shinglee Press, Singapore, pp. i-viii, 1 - 156, 4 colour pls." type="journal volume" year="1988">Ng 1988: 118</bibRefCitation>
). The eggs from some of these ovigerous females hatched into normal planktonic zoeae and they will be described at a later date. As such, this species has a rather wide distribution compared to more typical
<taxonomicName id="4C35A6BA691CFFE80413FF1EFB8F41C2" box="[813,1037,193,216]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B941012B691CFFE80413FF1EFC3041C2" box="[813,946,193,216]" italics="true" pageId="1" pageNumber="240">Geosesarma</emphasis>
species.
</taxonomicName>
<taxonomicName id="4C35A6BA691CFFE8032AFF1EFB7241C2" authorityName="Ng &amp; Liu" authorityYear="2004" box="[1044,1264,193,216]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="hednon">
<emphasis id="B941012B691CFFE8032AFF1EFB7241C2" box="[1044,1264,193,216]" italics="true" pageId="1" pageNumber="240">Geosesarma hednon</emphasis>
</taxonomicName>
thus lacks one of the key attributes of the genus, i.e. the possession of large eggs with abbreviated or direct development. The authors have on hand, ovigerous females of
<taxonomicName id="4C35A6BA691CFFE80397FEFEFA0A4022" authority="Ng, 1986" authorityName="Ng" authorityYear="1986" box="[1193,1416,289,312]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="nemesis">
<emphasis id="B941012B691CFFE80397FEFEFA9F4022" box="[1193,1309,289,312]" italics="true" pageId="1" pageNumber="240">G. nemesis</emphasis>
<bibRefCitation id="EFA4A0C8691CFFE8021DFEFEFA0A4022" author="Ng, P" box="[1315,1416,289,312]" pageId="1" pageNumber="240" pagination="36 - 44" refId="ref5639" refString="Ng, P. K. L., 1986. Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia. Journal of the Singapore National Academy of Science, 15: 36 - 44." type="journal article" year="1986">Ng, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C35A6BA691CFFE802ACFEFEFC784042" authority="Ng, 1986" authorityName="Ng" authorityYear="1986" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="cataracta">
<emphasis id="B941012B691CFFE802ACFEFEFC104042" italics="true" pageId="1" pageNumber="240">G. cataracta</emphasis>
<bibRefCitation id="EFA4A0C8691CFFE804A9FE9EFC784042" author="Ng, P" box="[919,1018,321,344]" pageId="1" pageNumber="240" pagination="36 - 44" refId="ref5639" refString="Ng, P. K. L., 1986. Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia. Journal of the Singapore National Academy of Science, 15: 36 - 44." type="journal article" year="1986">Ng, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C35A6BA691CFFE8033DFE9EFA854042" authority="Ng, 1986" authorityName="Ng" authorityYear="1986" box="[1027,1287,321,344]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="tiomanicum">
<emphasis id="B941012B691CFFE8033DFE9EFB1D4042" box="[1027,1183,321,344]" italics="true" pageId="1" pageNumber="240">G. tiomanicum</emphasis>
<bibRefCitation id="EFA4A0C8691CFFE8039AFE9EFA854042" author="Ng, P" box="[1188,1287,321,344]" pageId="1" pageNumber="240" pagination="36 - 44" refId="ref5639" refString="Ng, P. K. L., 1986. Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia. Journal of the Singapore National Academy of Science, 15: 36 - 44." type="journal article" year="1986">Ng, 1986</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C35A6BA691CFFE8022EFE9EFC4D4062" authority="Tweedie, 1940" authorityName="Tweedie" authorityYear="1940" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="penangense">
<emphasis id="B941012B691CFFE8022EFE9EFA2E4042" box="[1296,1452,321,344]" italics="true" pageId="1" pageNumber="240">G. penangense</emphasis>
<bibRefCitation id="EFA4A0C8691CFFE80413FEBEFC4D4062" author="Tweedie, M" box="[813,975,353,376]" pageId="1" pageNumber="240" pagination="88 - 113" refId="ref6168" refString="Tweedie, M. W. F., 1940. New and interesting Malaysian species of Sesarma and Utica (Crustacea Brachyura). Bulletin of the Raffles Museum, Singapore, 16: 88 - 113, pl. 14." type="journal article" year="1940">Tweedie, 1940</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C35A6BA691CFFE804E5FEBEFB774062" authority="Ng &amp; Lim" authorityName="Ng &amp; Lim" box="[987,1269,353,376]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="malayanum">
<emphasis id="B941012B691CFFE804E5FEBEFBFB4062" box="[987,1145,353,376]" italics="true" pageId="1" pageNumber="240">G. malayanum</emphasis>
Ng &amp; Lim
</taxonomicName>
, in
<bibRefCitation id="EFA4A0C8691CFFE80220FEBEFA074062" author="Ng, P" box="[1310,1413,353,376]" pageId="1" pageNumber="240" pagination="36 - 44" refId="ref5639" refString="Ng, P. K. L., 1986. Preliminary descriptions of 17 new freshwater crabs of the genera Geosesarma, Parathelphusa, Johora and Stoliczia (Crustacea Decapoda, Brachyura) from South East Asia. Journal of the Singapore National Academy of Science, 15: 36 - 44." type="journal article" year="1986">Ng, 1986</bibRefCitation>
,
<taxonomicName id="4C35A6BA691CFFE802ACFEBEFB8E4082" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="aurantium">
<emphasis id="B941012B691CFFE802ACFEBEFC1E4082" italics="true" pageId="1" pageNumber="240">G. aurantium</emphasis>
Ng, 1995
</taxonomicName>
, and
<taxonomicName id="4C35A6BA691CFFE80376FE5EFA254082" authority="(De Man, 1892)" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[1096,1447,385,408]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="noduliferum">
<emphasis id="B941012B691CFFE80376FE5EFB6F4082" box="[1096,1261,385,408]" italics="true" pageId="1" pageNumber="240">G. noduliferum</emphasis>
(De
<bibRefCitation id="EFA4A0C8691CFFE80218FE5EFA224082" author="Man, J" box="[1318,1440,385,408]" pageId="1" pageNumber="240" pagination="265 - 527" refId="ref5526" refString="Man, J. G., De, 1892. Decapoden des Indischen Archipels. In: Weber, M. (ed.), Zoologische Ergebnisse einer Reise in Niederlandisch Ost-Indien, 2: 265 - 527, pls. 15 - 29." type="book chapter" year="1892">Man, 1892</bibRefCitation>
)
</taxonomicName>
, all of which have large eggs. We expect some of the other
<taxonomicName id="4C35A6BA691CFFE80413FE1EFB8440C2" box="[813,1030,449,472]" pageId="1" pageNumber="240">
<emphasis id="B941012B691CFFE80413FE1EFC3040C2" box="[813,946,449,472]" italics="true" pageId="1" pageNumber="240">Geosesarma</emphasis>
species
</taxonomicName>
(e.g.
<taxonomicName id="4C35A6BA691CFFE8037DFE1EFB5840C2" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[1091,1242,449,472]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="maculatum">
<emphasis id="B941012B691CFFE8037DFE1EFB5840C2" box="[1091,1242,449,472]" italics="true" pageId="1" pageNumber="240">G. maculatum</emphasis>
</taxonomicName>
) to also have small eggs like
<taxonomicName id="4C35A6BA691CFFE804A4FE3EFB8E40E2" authorityName="Ng &amp; Liu" authorityYear="2004" box="[922,1036,481,504]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="hednon">
<emphasis id="B941012B691CFFE804A4FE3EFB8E40E2" box="[922,1036,481,504]" italics="true" pageId="1" pageNumber="240">G. hednon</emphasis>
</taxonomicName>
. While we have seriously considered establishing a new genus for
<taxonomicName id="4C35A6BA691CFFE80355FDDEFB5E4302" authorityName="Ng &amp; Liu" authorityYear="2004" box="[1131,1244,513,536]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="hednon">
<emphasis id="B941012B691CFFE80355FDDEFB5E4302" box="[1131,1244,513,536]" italics="true" pageId="1" pageNumber="240">G. hednon</emphasis>
</taxonomicName>
and its close allies (the
<typeStatus id="548E639B691CFFE80462FDFEFC084322" box="[860,906,545,568]" pageId="1" pageNumber="240">type</typeStatus>
species of
<taxonomicName id="4C35A6BA691CFFE804C2FDFEFB034322" authorityName="De Man" authorityYear="1892" box="[1020,1153,545,568]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="genus">
<emphasis id="B941012B691CFFE804C2FDFEFB034322" box="[1020,1153,545,568]" italics="true" pageId="1" pageNumber="240">Geosesarma</emphasis>
</taxonomicName>
is
<taxonomicName id="4C35A6BA691CFFE803A0FDFEFAC24322" baseAuthorityName="De Man" baseAuthorityYear="1892" box="[1182,1344,545,568]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="240" phylum="Arthropoda" rank="species" species="noduliferum">
<emphasis id="B941012B691CFFE803A0FDFEFAC24322" box="[1182,1344,545,568]" italics="true" pageId="1" pageNumber="240">G. noduliferum</emphasis>
</taxonomicName>
), we have deferred from doing so as the affinities of too many of the species in the genus are still poorly understood. Most of the species have not even been reported or redescribed since their original discovery.
</paragraph>
</subSubSection>
</treatment>
</document>
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