189 lines
24 KiB
XML
189 lines
24 KiB
XML
<document id="74D4F01A89CCFBF042D8DCE741B15F9C" ID-CLB-Dataset="2630" ID-DOI="10.5169/seals-787037" ID-GBIF-Dataset="d0b846f5-0d3d-4b6d-abb4-80bc84528180" ID-ISSN="2624-9170" ID-Zenodo-Dep="5842342" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1637934074083" checkinUser="carolina" docAuthor="Subai, Peter & Neubert, Eike" docDate="2014" docId="038C1808EF5EFFD259A5FDAC4D7D8917" docLanguage="en" docName="ContribNatHist.23.1-94.pdf.imf" docOrigin="Contributions to Natural History 23" docStyle="DocumentStyle:D319E9290ED630111D2A11900DAFC11F.4:ContribNatHist.2011-.journal_article" docStyleId="D319E9290ED630111D2A11900DAFC11F" docStyleName="ContribNatHist.2011-.journal_article" docStyleVersion="4" docTitle="Lindholmiola HESSE, Zoologica 1931" docType="treatment" docVersion="9" lastPageNumber="5" masterDocId="FFB56070EF5CFFD65913FFD24E798D1B" masterDocTitle="Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)" masterLastPageNumber="94" masterPageNumber="1" pageNumber="3" updateTime="1699470473833" updateUser="ExternalLinkService">
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<mods:title id="B2F6ABD3772F30B71C8F6A96D240FFF8">Revision of the genus Lindholmiola HESSE, 1931 (Gastropoda: Pulmonata: Helicodontidae)</mods:title>
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<mods:namePart id="CB8EF0AB6FB66898F6EA9E94EC9B62A5">Subai, Peter</mods:namePart>
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<mods:namePart id="ABC27E81923AA00EF64794908C7C825D">Neubert, Eike</mods:namePart>
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<mods:title id="84A32C47490EAB256CE65699D163AF53">Contributions to Natural History</mods:title>
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<mods:date id="18495EFD913C48AF20FCF6AB72F5207E">2014</mods:date>
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<treatment id="038C1808EF5EFFD259A5FDAC4D7D8917" ID-DOI="http://doi.org/10.5281/zenodo.5843617" ID-GBIF-Taxon="192044618" ID-Zenodo-Dep="5843617" LSID="urn:lsid:plazi:treatment:038C1808EF5EFFD259A5FDAC4D7D8917" httpUri="http://treatment.plazi.org/id/038C1808EF5EFFD259A5FDAC4D7D8917" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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<subSubSection id="C33FFA95EF5EFFD459A5FDAC4FA48F8D" box="[182,477,638,663]" pageId="2" pageNumber="3" type="nomenclature">
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<paragraph id="8B9AA91EEF5EFFD459A5FDAC4FA48F8D" blockId="2.[182,477,638,663]" box="[182,477,638,663]" pageId="2" pageNumber="3">
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<heading id="D0D21E72EF5EFFD459A5FDAC4FA48F8D" box="[182,477,638,663]" centered="true" fontSize="36" level="2" pageId="2" pageNumber="3" reason="4">
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<taxonomicName id="4C25D29DEF5EFFD459A5FDAC4FA48F8D" ID-CoL="7P7KD" authority="Hesse, 1931" authorityName="HESSE, Zoologica" authorityYear="1931" box="[182,477,638,663]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
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<emphasis id="B951750CEF5EFFD459A5FDAC4FA48F8D" bold="true" box="[182,477,638,663]" pageId="2" pageNumber="3">
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<emphasis id="B951750CEF5EFFD459A5FDAC4F358F8C" bold="true" box="[182,332,638,663]" italics="true" pageId="2" pageNumber="3">Lindholmiola</emphasis>
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<bibRefCitation id="EFB4D4EFEF5EFFD45842FDAC4FA48F8D" author="Hesse, P." box="[337,477,638,663]" pageId="2" pageNumber="3" pagination="1 - 118" refId="ref67366" refString="Hesse, P. (1931): Zur Anatomie und Systematik palaearktischer Stylommatophoren. - Zoologica, Original-Abhandlungen aus dem Gesamtgebiete der Zoologie 81: 1 - 118, Taf. 1 - 16." type="journal article" year="1931">Hesse, 1931</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FFA95EF5EFFD459A7FD1B4C518E1D" pageId="2" pageNumber="3" type="reference_group">
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<paragraph id="8B9AA91EEF5EFFD459A7FD1B4C518E1D" blockId="2.[180,1065,713,774]" pageId="2" pageNumber="3">
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<treatmentCitation id="0A848F0FEF5EFFD459A7FD1B4CED8FFA" box="[180,660,713,737]" pageId="2" pageNumber="3">
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<bibRefCitation id="EFB4D4EFEF5EFFD459A7FD1B4E928FFA" author="Hesse, P." box="[180,235,713,737]" pageId="2" pageNumber="3" pagination="1 - 118" refId="ref67366" refString="Hesse, P. (1931): Zur Anatomie und Systematik palaearktischer Stylommatophoren. - Zoologica, Original-Abhandlungen aus dem Gesamtgebiete der Zoologie 81: 1 - 118, Taf. 1 - 16." type="journal article" year="1931">1931</bibRefCitation>
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<taxonomicName id="4C25D29DEF5EFFD459E2FD1B4FB38FFA" ID-CoL="7P7KD" authority="HESSE, Zoologica" authorityName="HESSE, Zoologica" authorityYear="1931" box="[241,458,713,737]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="genus">
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<emphasis id="B951750CEF5EFFD459E2FD1B4FF98FFA" box="[241,384,713,737]" italics="true" pageId="2" pageNumber="3">Lindholmiola</emphasis>
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HESSE
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</taxonomicName>
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, Zoologica, 18: 50
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</treatmentCitation>
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[
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<typeStatus id="549E17BCEF5EFFD45BB3FD1B4CAB8FFA" box="[672,722,713,737]" pageId="2" pageNumber="3">type</typeStatus>
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species by original designation
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<taxonomicName id="4C25D29DEF5EFFD459C8FD3C4C6E8E1D" authority="A. FeRUSSAC, 1832" authorityName="A. FeRUSSAC" authorityYear="1832" box="[219,535,750,774]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="lens">
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<emphasis id="B951750CEF5EFFD459C8FD3C4F3F8E1D" box="[219,326,750,774]" italics="true" pageId="2" pageNumber="3">Helix lens</emphasis>
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A. FéRUSSAC, 1832
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</taxonomicName>
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].
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</paragraph>
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</subSubSection>
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<subSubSection id="C33FFA95EF5EFFD259C8FCEB4C478C77" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
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<paragraph id="8B9AA91EEF5EFFD459C8FCEB4C2F89BA" blockId="2.[182,1072,825,1596]" pageId="2" pageNumber="3">
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Description: shell flat to lentiform (H:
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<quantity id="4CDD04FBEF5EFFD45B87FCEB4D638E4A" box="[660,794,825,849]" metricMagnitude="-3" metricUnit="m" metricValue="5.3" metricValueMax="7.5" metricValueMin="3.1" pageId="2" pageNumber="3" unit="mm" value="5.3" valueMax="7.5" valueMin="3.1">3.1–7.5 mm</quantity>
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, D:
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<quantity id="4CDD04FBEF5EFFD45A56FCEB4DA68E4A" box="[837,991,825,849]" metricMagnitude="-2" metricUnit="m" metricValue="1.185" metricValueMax="1.65" metricValueMin="0.72" pageId="2" pageNumber="3" unit="mm" value="11.85" valueMax="16.5" valueMin="7.2">7.2–16.5 mm</quantity>
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); spire slightly elevated (exception:
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<taxonomicName id="4C25D29DEF5EFFD45B14FC8C4C278E6D" baseAuthorityName="ROTH" baseAuthorityYear="1839" box="[519,606,862,886]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="gyria">
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<emphasis id="B951750CEF5EFFD45B14FC8C4C278E6D" box="[519,606,862,886]" italics="true" pageId="2" pageNumber="3">L. gyria</emphasis>
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</taxonomicName>
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with a concave spire); colour brown to yellowish-brown; teleoconch surface usually with hairs; teleoconch sculpture of fine to coarse radial riblets; whorls regularly increasing, 5¼–7¾ whorls; periphery of last whorl rounded, bluntly shouldered or sharply keeled; last whorl strongly bends downwards before reaching the aperture; umbilicus narrow to moderately wide, cylindrical to perspective, usually only slightly covered by the peristome (exception:
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<taxonomicName id="4C25D29DEF5EFFD45B5BFBEC4CC7894D" authorityName="A. FeRUSSAC, Tableaux" authorityYear="1821" box="[584,702,1086,1110]" class="Gastropoda" family="Helicidae" genus="Helix" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="barbata">
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<emphasis id="B951750CEF5EFFD45B5BFBEC4CC7894D" box="[584,702,1086,1110]" italics="true" pageId="2" pageNumber="3">L. barbata</emphasis>
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</taxonomicName>
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, where the umbilicus is almost completely covered); aperture narrow, oblique and curved (in lateral view); peristome narrow to broad, reflexed.
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</paragraph>
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<paragraph id="8B9AA91EEF5EFFD459C8FB7C4C9B88D7" blockId="2.[182,1072,825,1596]" pageId="2" pageNumber="3">
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Genital organs (
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<figureCitation id="131EB59BEF5EFFD4588AFB7C4FA789DD" box="[409,478,1198,1222]" captionStart="Figs" captionStartId="3.[144,183,1240,1258]" captionTargetBox="[156,1028,117,1185]" captionTargetId="figure-6@3.[144,1028,113,1185]" captionTargetPageId="3" captionText="Figs. 1–2. Genital organs of Helicodontidae. Fig. 1: Lindholmiola barbata; Greece, Kríti, Nomos Haniá, Bay of Soúda, 100–150 m from the mouth of the river S of the English cemetery. Fig. 2: Helicodonta obvoluta (O. F. MüLLER, 1774); Belgium, valley of the Meuse, 500 m from Annevoie in direction to Dinant, 2a) situs of genital organs, 2b) interior structure of penis. — Abbreviations used for the description of the genital organs: A = atrium, Al = albumen gland, E = Epiphallus, G = distal glandular part of epiphallus, Gl = Glandula, Hd = hermaphroditic duct, Hg = hermaphroditic gland, P = Penis, R = retractor muscle, Sp = Spermoviduct, V = vesicle, Vag = Vagina, Vd = Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5842344" httpUri="https://zenodo.org/record/5842344/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
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) simple; atrium short; penis a club-shaped tube, epiphallus distally strongly contracted (often, this area is covered by a glandular tissue); penial retractor muscle attaching at the central part of epiphallus, which connects to vas deferens without any visible distinction; vagina short (completely reduced in
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<taxonomicName id="4C25D29DEF5EFFD45B07FA964CC88847" authorityName="FALKNER, Heldia" authorityYear="1995" box="[532,689,1348,1372]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="2" pageNumber="3" phylum="Mollusca" rank="species" species="reischuetzi">
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<emphasis id="B951750CEF5EFFD45B07FA964CC88847" box="[532,689,1348,1372]" italics="true" pageId="2" pageNumber="3">L. reischuetzi</emphasis>
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</taxonomicName>
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); glandula a single tube with a narrow stem and a broader hose often sculptured by crossing furrows; bursa copulatrix with a narrow pedunculus and a more broadened vesicle (often not clearly visible) (Schileyko 2006: 1919, Fig. 2439).
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</paragraph>
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<paragraph id="8B9AA91EEF5EFFD259CFFA0B4DD58DE7" blockId="2.[182,1072,825,1596]" lastBlockId="4.[182,1071,116,1036]" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3">
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Interior of the terminal genital organs with a long fold starting at the genital pore and ending at the distal vagina; atrial part of the fold often subdivided by small crossing furrows (an additional semicircular crested fold in the atrium can be found in
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<taxonomicName id="4C25D29DEF58FFD2587DFFA64C798D97" baseAuthorityName="A. J. WAGNER" baseAuthorityYear="1927" box="[366,512,116,140]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="regisborisi">
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<emphasis id="B951750CEF58FFD2587DFFA64C798D97" box="[366,512,116,140]" italics="true" pageId="4" pageNumber="5">L. regisborisi</emphasis>
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</taxonomicName>
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); characters of the penial lumen differ according to the species, usually with 2–3 raised longitudinal pilasters filling at least the distal half of the penial lumen (exception: In
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<taxonomicName id="4C25D29DEF58FFD25BD6FF6C4D1B8DCD" baseAuthorityName="ROSSMASSLER" baseAuthorityYear="1838" box="[709,866,190,214]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="corcyrensis">
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<emphasis id="B951750CEF58FFD25BD6FF6C4D1B8DCD" box="[709,866,190,214]" italics="true" pageId="4" pageNumber="5">L. corcyrensis</emphasis>
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</taxonomicName>
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, the longitudinal pilasters are replaced by v-shaped, obliquely arranged, short folds).
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</paragraph>
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<caption id="DF5AF996EF5FFFD55983FB0A4DBB8891" ID-DOI="http://doi.org/10.5281/zenodo.5842344" ID-Zenodo-Dep="5842344" httpUri="https://zenodo.org/record/5842344/files/figure.png" pageId="3" pageNumber="4" startId="3.[144,183,1240,1258]" targetBox="[156,1028,117,1185]" targetPageId="3">
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<paragraph id="8B9AA91EEF5FFFD55983FB0A4DBB8891" blockId="3.[144,1029,1240,1418]" pageId="3" pageNumber="4">
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Figs. 1–2. Genital organs of
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<taxonomicName id="4C25D29DEF5FFFD55890FB0A4C7389F1" authorityName="Kobelt" authorityYear="1904" box="[387,522,1240,1258]" class="Gastropoda" family="Helicodontidae" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="family">Helicodontidae</taxonomicName>
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. Fig. 1:
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<taxonomicName id="4C25D29DEF5FFFD55B5EFB0A4D7189F1" baseAuthorityName="A. FeRUSSAC" baseAuthorityYear="1821" box="[589,776,1240,1258]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="barbata">
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<emphasis id="B951750CEF5FFFD55B5EFB0A4D7189F1" bold="true" box="[589,776,1240,1258]" italics="true" pageId="3" pageNumber="4">Lindholmiola barbata</emphasis>
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</taxonomicName>
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; Greece, Kríti, Nomos Haniá, Bay of Soúda, 100–150 m from the mouth of the river S of the English cemetery. Fig. 2:
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<taxonomicName id="4C25D29DEF5FFFD55AB5FB214FCF8804" authority="(O. F. MuLLER, 1774)" baseAuthorityName="O. F. MuLLER" baseAuthorityYear="1774" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="3" pageNumber="4" phylum="Mollusca" rank="species" species="obvoluta">
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<emphasis id="B951750CEF5FFFD55AB5FB214E8D8804" bold="true" italics="true" pageId="3" pageNumber="4">Helicodonta obvoluta</emphasis>
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(O. F. MüLLER, 1774)
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</taxonomicName>
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; Belgium, valley of the Meuse, 500 m from Annevoie in direction to Dinant, 2a) situs of genital organs, 2b) interior structure of penis. — Abbreviations used for the description of the genital organs: A = atrium, Al = albumen gland, E = Epiphallus, G = distal glandular part of epiphallus, Gl = Glandula, Hd = hermaphroditic duct, Hg = hermaphroditic gland, P = Penis, R = retractor muscle, Sp = Spermoviduct, V = vesicle, Vag = Vagina, Vd = Vas deferens.
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</paragraph>
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</caption>
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<paragraph id="8B9AA91EEF58FFD259C8FEDB4C478C77" blockId="4.[182,1071,116,1036]" pageId="4" pageNumber="5">It has to be stressed that the characters of the terminal genital organs as described above can only be seen in sexually mature specimens. In juveniles, the folds are missing or very weak.</paragraph>
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</subSubSection>
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<subSubSection id="C33FFA95EF58FFD259C8FEAB4D7D8917" pageId="4" pageNumber="5" type="discussion">
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<paragraph id="8B9AA91EEF58FFD259C8FEAB4D5E8F8D" blockId="4.[182,1071,116,1036]" pageId="4" pageNumber="5">
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Differentiating remarks: The shells of
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<taxonomicName id="4C25D29DEF58FFD25BB9FEAB4D408C8A" authorityName="A. Ferussac" authorityYear="1821" box="[682,825,377,401]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis id="B951750CEF58FFD25BB9FEAB4D408C8A" box="[682,825,377,401]" italics="true" pageId="4" pageNumber="5">Helicodonta</emphasis>
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</taxonomicName>
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are also depressed but never lentiform and usually concave. In species of this genus, the whorls increase faster in width, particularly the ultimate and the penultimate whorl, which are always broader than those in
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<taxonomicName id="4C25D29DEF58FFD25B9FFE3B4D5B8F1A" authorityName="HESSE, Zoologica" authorityYear="1931" box="[652,802,489,513]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
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<emphasis id="B951750CEF58FFD25B9FFE3B4D5B8F1A" box="[652,802,489,513]" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
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</taxonomicName>
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. Additionally, they are laterally rounded and only slightly shouldered. The umbilicus of
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<taxonomicName id="4C25D29DEF58FFD25AB3FDDC4E9E8F57" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B951750CEF58FFD25AB3FDDC4A508F3D" box="[928,1065,526,550]" italics="true" pageId="4" pageNumber="5">Helicodonta</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
is broader and more perspective if compared to
|
||
<taxonomicName id="4C25D29DEF58FFD25A0DFDE64DCA8F57" authorityName="HESSE, Zoologica" authorityYear="1931" box="[798,947,564,588]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951750CEF58FFD25A0DFDE64DCA8F57" box="[798,947,564,588]" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
|
||
</taxonomicName>
|
||
, and their apertures have often three indentations which are even thickened to form teeth. The peristome is usually broadened to form a lip.
|
||
</paragraph>
|
||
<paragraph id="8B9AA91EEF58FFD259CFFD764D7D8917" blockId="4.[182,1071,116,1036]" pageId="4" pageNumber="5">
|
||
Both genera differ in the morphology of their genital organs:
|
||
<taxonomicName id="4C25D29DEF58FFD25A60FD764E9A8FFA" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B951750CEF58FFD25A60FD764D8E8FA7" box="[883,1015,676,700]" italics="true" pageId="4" pageNumber="5">Helicodonta</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
have a considerably longer penis and vagina and a short tubular organ of unknown function that branches off distally to glandua and bursa copulatrix (
|
||
<figureCitation id="131EB59BEF58FFD259FAFCC64F518E37" box="[233,296,788,812]" captionStart="Figs" captionStartId="3.[144,183,1240,1258]" captionTargetBox="[156,1028,117,1185]" captionTargetId="figure-6@3.[144,1028,113,1185]" captionTargetPageId="3" captionText="Figs. 1–2. Genital organs of Helicodontidae. Fig. 1: Lindholmiola barbata; Greece, Kríti, Nomos Haniá, Bay of Soúda, 100–150 m from the mouth of the river S of the English cemetery. Fig. 2: Helicodonta obvoluta (O. F. MüLLER, 1774); Belgium, valley of the Meuse, 500 m from Annevoie in direction to Dinant, 2a) situs of genital organs, 2b) interior structure of penis. — Abbreviations used for the description of the genital organs: A = atrium, Al = albumen gland, E = Epiphallus, G = distal glandular part of epiphallus, Gl = Glandula, Hd = hermaphroditic duct, Hg = hermaphroditic gland, P = Penis, R = retractor muscle, Sp = Spermoviduct, V = vesicle, Vag = Vagina, Vd = Vas deferens." figureDoi="http://doi.org/10.5281/zenodo.5842344" httpUri="https://zenodo.org/record/5842344/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
) (not present in
|
||
<taxonomicName id="4C25D29DEF58FFD258F7FCC64C008E37" authorityName="HESSE, Zoologica" authorityYear="1931" box="[484,633,788,812]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951750CEF58FFD258F7FCC64C008E37" box="[484,633,788,812]" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
|
||
</taxonomicName>
|
||
). In the interior of the terminal genital organs, there are elongated narrow pilasters in the vagina (different in
|
||
<taxonomicName id="4C25D29DEF58FFD25AE5FCEB4F658E6D" authorityName="HESSE, Zoologica" authorityYear="1931" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951750CEF58FFD25AE5FCEB4F658E6D" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
|
||
</taxonomicName>
|
||
), and in the middle of the penis, the penial lumen is filled by two thickened pilasters (different in
|
||
<taxonomicName id="4C25D29DEF58FFD25B3BFC564CC48E87" authorityName="HESSE, Zoologica" authorityYear="1931" box="[552,701,900,924]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951750CEF58FFD25B3BFC564CC48E87" box="[552,701,900,924]" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
|
||
</taxonomicName>
|
||
). In the distal part of the penis, an additional pilaster covered by small punctiform granules is found (not present in
|
||
<taxonomicName id="4C25D29DEF58FFD259C6FC1C4F128EFD" authorityName="HESSE, Zoologica" authorityYear="1931" box="[213,363,974,998]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="genus">
|
||
<emphasis id="B951750CEF58FFD259C6FC1C4F128EFD" box="[213,363,974,998]" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
|
||
</taxonomicName>
|
||
). The glandular tissue at the distal end of the epiphallus of
|
||
<taxonomicName id="4C25D29DEF58FFD259A5FC264FFB8917" box="[182,386,1012,1036]" class="Gastropoda" family="Helicodontidae" genus="Lindholmiola" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B951750CEF58FFD259A5FC264F328917" box="[182,331,1012,1036]" italics="true" pageId="4" pageNumber="5">Lindholmiola</emphasis>
|
||
spp.
|
||
</taxonomicName>
|
||
is missing in
|
||
<taxonomicName id="4C25D29DEF58FFD25B04FC264D7D8917" box="[535,772,1012,1036]" class="Gastropoda" family="Helicodontidae" genus="Helicodonta" kingdom="Animalia" order="Stylommatophora" pageId="4" pageNumber="5" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B951750CEF58FFD25B04FC264CD98917" box="[535,672,1012,1036]" italics="true" pageId="4" pageNumber="5">Helicodonta</emphasis>
|
||
species.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |