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<mods:title id="F13DA8DAB258DA269724A3253E8C1767">The tadpoles of nine Cameroonian Leptodactylodon species (Amphibia, Anura, Arthroleptidae)</mods:title>
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<mods:namePart id="D7779320223A1B242BC5D22A03823126">Loader, Simon P.</mods:namePart>
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<mods:namePart id="795AAD2DC1A4FD9E1020C60F5AAB502E">Gonwouo, L. Nono</mods:namePart>
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<treatment id="038987F9FF8CFFE1FAC25314FD87FC66" ID-DOI="http://doi.org/10.5281/zenodo.5626137" ID-GBIF-Taxon="127667039" ID-Zenodo-Dep="5626137" LSID="urn:lsid:plazi:treatment:038987F9FF8CFFE1FAC25314FD87FC66" httpUri="http://treatment.plazi.org/id/038987F9FF8CFFE1FAC25314FD87FC66" lastPageId="14" lastPageNumber="43" pageId="13" pageNumber="42">
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<paragraph id="8B9F36EFFF8CFFE2FAC25314FCA8F9A6" blockId="13.[151,847,1575,1600]" box="[151,847,1575,1600]" pageId="13" pageNumber="42">
<heading id="D0D78183FF8CFFE2FAC25314FCA8F9A6" bold="true" box="[151,847,1575,1600]" fontSize="10" level="2" pageId="13" pageNumber="42" reason="2">
<taxonomicName id="4C204D6CFF8CFFE2FAC25314FCA8F9A6" ID-CoL="5JB5F" authority="Amiet, 1971" authorityName="Amiet" authorityYear="1971" box="[151,847,1575,1600]" class="Amphibia" family="Arthroleptidae" genus="Leptodactylodon" kingdom="Animalia" order="Anura" pageId="13" pageNumber="42" phylum="Chordata" rank="subSpecies" species="polyacanthus" subSpecies="punctiventris">
<emphasis id="B954EAFDFF8CFFE2FAC25314FCA8F9A6" bold="true" box="[151,847,1575,1600]" pageId="13" pageNumber="42">
<emphasis id="B954EAFDFF8CFFE2FAC25314FEB1F9A6" bold="true" box="[151,342,1575,1600]" italics="true" pageId="13" pageNumber="42">Leptodactylodon</emphasis>
cf.
<emphasis id="B954EAFDFF8CFFE2FBD45314FD5EF9A6" bold="true" box="[385,697,1575,1600]" italics="true" pageId="13" pageNumber="42">polyacanthus punctiventris</emphasis>
Amiet, 1971
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<paragraph id="8B9F36EFFF8CFFE2FAC2535CFD27F915" blockId="13.[151,1436,1647,2032]" pageId="13" pageNumber="42">
Three tadpoles from Fotabong were tentatively assigned to
<taxonomicName id="4C204D6CFF8CFFE2F9385343FB2FF961" box="[877,1224,1647,1671]" class="Amphibia" family="Arthroleptidae" genus="Leptodactylodon" kingdom="Animalia" order="Anura" pageId="13" pageNumber="42" phylum="Chordata" rank="subSpecies" species="polyacanthus" subSpecies="punctiventris">
<emphasis id="B954EAFDFF8CFFE2F9385343FB2FF961" box="[877,1224,1647,1671]" italics="true" pageId="13" pageNumber="42">L. polyacanthus punctiventris</emphasis>
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based on known amphibian assemblages from that area (e.g.
<bibRefCitation id="EFB14B1EFF8CFFE2F8F053A0FCD4F94A" author="Amiet" box="[677,819,1683,1708]" pageId="13" pageNumber="42" refString="Amiet, J. - L. (1980) Revision du genre Leptodactylodon Andersson (Amphibia, Anura, Astylosterninae). Annales de la Faculte des Sciences de Yaounde, 27, 69 - 224." type="journal article" year="1980">Amiet 1980</bibRefCitation>
). One specimen was genotyped: ZMB 78551 (three tadpoles, Gosner stage 25, Fotabong,
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,
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,
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,
<date id="FF9E102FFF8CFFE2F9B1538BFB51F936" box="[996,1206,1719,1744]" pageId="13" pageNumber="42" value="2011-11-01">1 November 2011</date>
). The tadpoles were collected from shallow, sandy streams at
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.
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<paragraph id="8B9F36EFFF8CFFE2FA9253CCFB28F8DA" blockId="13.[151,1436,1647,2032]" pageId="13" pageNumber="42">The description is based on the genotyped individual (Gosner stage 25). Body length/total length ratio and description of tail tips was based on non-genotyped specimens of the same developmental stage.</paragraph>
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<emphasis id="B954EAFDFF8CFFE2FA925274FE86F886" bold="true" box="[199,353,1863,1888]" pageId="13" pageNumber="42">Morphology.</emphasis>
Long and slender tadpole with long, muscular tail (non-genotyped tadpoles not as slender as drawn in
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a, b); body elliptical in dorsal and depressed in lateral view; body length 27.529.3% (N= 2) of total length; body height 37.2% of body length, body width 46.3% of body length, maximum body width on level of spiracle insertion; snout nearly rounded in dorsal view; nostrils oval and situated laterally; nostrils equidistant from eye and snout tip; eye diameter 9.9% of body length; inter-orbital distance equals inter-nostril distance; tail fins narrow, dorsal fin originating
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anterior to tail base, reaching deepest point in first third of tail, slightly deeper than ventral fin; ventral fin originates at tail base, runs parallel to tail axis; tail tip rounded to slightly pointed; tail axis broad, continuously converging towards tail tip; maximum height of tail axis 57.060.5% (N= 2) of total tail height; vent tube dextral; lateral sacs present, extending from spiracle to end of body, covering lower two thirds of flanks; very short spiracle, sinistral, translucent, opening lateral, not visible in dorsal view, originating at mid-body; mouth opening frontal; labial tooth row formula 0/0; both jaw sheaths completely keratinized, strongly serrated, each with lateral pair of caniniform projections (fangs); median part of lower jaw with six to seven needle-like cusps (
<figureCitation id="131B2A6AFF8FFFE1FBFB54A7FDE6FE4B" box="[430,513,404,429]" captionStart="FIGURE 12" captionStartId="14.[151,250,1679,1701]" captionTargetBox="[168,1402,944,1656]" captionTargetId="figure@14.[163,1408,940,1662]" captionTargetPageId="14" captionText="FIGURE 12. Dorsal (a) and lateral view (b) of Leptodactylodon cf. polyacanthus punctiventris with sketches of mouthpart in frontal view (c) and jaw sheaths with caniniform projections (d); scale bars: 1 mm." httpUri="https://zenodo.org/record/285572/files/figure.png" pageId="14" pageNumber="43">Fig. 12</figureCitation>
d); three distinct serrations abaxial to fangs; upper jaw narrow and long, lower jaw shorter; large, broad bilobate posterior lip covered with 21 large papillae (
<figureCitation id="131B2A6AFF8FFFE1FE135484FB44FE36" box="[1094,1187,439,464]" captionStart="FIGURE 12" captionStartId="14.[151,250,1679,1701]" captionTargetBox="[168,1402,944,1656]" captionTargetId="figure@14.[163,1408,940,1662]" captionTargetPageId="14" captionText="FIGURE 12. Dorsal (a) and lateral view (b) of Leptodactylodon cf. polyacanthus punctiventris with sketches of mouthpart in frontal view (c) and jaw sheaths with caniniform projections (d); scale bars: 1 mm." httpUri="https://zenodo.org/record/285572/files/figure.png" pageId="14" pageNumber="43">Fig. 12</figureCitation>
c); papillae arranged symmetrically along in vertical body axis; papillae arranged in two semicircular rows; almost straight skin fold on lower lip, just posterior to lower jaw sheath; oral disc width 39.7% of body length; mouth width 25.0% of oral disc width.
</paragraph>
<paragraph id="8B9F36EFFF8FFFE1FA925774FF38FD4E" blockId="14.[151,1436,151,896]" pageId="14" pageNumber="43">
The two individuals with intact tails (both Gosner stage 25) had total lengths of 39.6 and
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, respectively (body length: 11.6 &amp; 12.0 mm; tail length: 28.0 &amp;
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). Metamorphs measure about
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SVL (
<bibRefCitation id="EFB14B1EFF8FFFE1FF01575FFF34FD4E" author="Amiet" pageId="14" pageNumber="43" refString="Amiet, J. - L. (1980) Revision du genre Leptodactylodon Andersson (Amphibia, Anura, Astylosterninae). Annales de la Faculte des Sciences de Yaounde, 27, 69 - 224." type="journal article" year="1980">Amiet 1980</bibRefCitation>
).
</paragraph>
<paragraph id="8B9F36EFFF8FFFE1FA925787FC30FCF3" blockId="14.[151,1436,151,896]" pageId="14" pageNumber="43">
<emphasis id="B954EAFDFF8FFFE1FA925787FDF2FD2B" bold="true" box="[199,533,692,717]" pageId="14" pageNumber="43">Coloration in preservation.</emphasis>
Body and tail light brown, mottled with bigger and smaller dark spots and speckles on dorsal part of body and tail; anterior half of tail axis with median, longitudinal dark line (not visible in life); venter lighter with fewer dark patterns; fins translucent cream-white.
</paragraph>
<paragraph id="8B9F36EFFF8FFFE1FA92562CFD87FC66" blockId="14.[151,1436,151,896]" pageId="14" pageNumber="43">
<emphasis id="B954EAFDFF8FFFE1FA92562CFE7FFCDE" bold="true" box="[199,408,799,824]" pageId="14" pageNumber="43">Coloration in life.</emphasis>
Light brown with light to dark brown speckles; dark line from snout tip to anterior corner of eye; anterior part of tail axis with white spots, posterior part with black dots; fins light with dark spots, posterior part almost white; venter lighter (
<figureCitation id="131B2A6AFF8FFFE1F8585654FDB5FC66" box="[525,594,871,896]" captionStart="FIGURE 6" captionStartId="7.[151,250,1271,1293]" captionTargetBox="[151,1436,411,1250]" captionTargetId="figure@7.[151,1436,411,1250]" captionTargetPageId="7" captionText="FIGURE 6. Life coloration of Leptodactylodon tadpoles, from top to bottom: L. cf. polyacanthus punctiventris (ZMB 78551, Fotabong), L. boulengeri (ZMB 78471, Fotabong), L. ventrimarmoratus (ZMB 78554, Mt. Kala); pictures not to scale." httpUri="https://zenodo.org/record/285566/files/figure.png" pageId="14" pageNumber="43">Fig. 6</figureCitation>
).
</paragraph>
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