229 lines
27 KiB
XML
229 lines
27 KiB
XML
<document id="57B606741D38033A30E29F5D789680F6" ID-CLB-Dataset="5815" ID-DOI="10.11646/zootaxa.3779.3.2" ID-GBIF-Dataset="2055f347-d478-4d50-aad0-7e78c4b3b9c5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4910403" ID-ZooBank="23584B50-EFAC-4BCE-A20E-9B09C22529B0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623118476769" checkinUser="felipe" docAuthor="Deein, Gridsada, Tangjitjaroen, Weerapongse & Page, Lawrence M." docDate="2014" docId="038987E7FFCCDB2BC0EF5EAA5E85A7F0" docLanguage="en" docName="zt03779p352.pdf" docOrigin="Zootaxa 3779 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lepidocephalus pahangensis" docType="treatment" docVersion="8" lastPageNumber="348" masterDocId="FFB0FF9FFFCBDB2CC0785C1C5D08A042" masterDocTitle="A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae)" masterLastPageNumber="352" masterPageNumber="341" pageNumber="348" updateTime="1699003940845" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods id="09306793F50ADCEA668D4BFAA3C7027F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="6E435F53376248DC6B1ADE15F26789A1">
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<mods:title id="B347732A7972765ACAB7727D02CC4047">A revision of the spirit loaches, genus Lepidocephalus (Cypriniformes, Cobitidae)</mods:title>
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</mods:titleInfo>
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<mods:name id="BD5211E559C5144B56CCC39B14373C07" type="personal">
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<mods:role id="893AFC9C95DB3625ACB74FE0CC94DE7A">
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<mods:roleTerm id="67C3AB374405E32CD5351E006ABFA335">Author</mods:roleTerm>
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<mods:namePart id="A8B23B82E14C0BC2A0CE5C0CF8C4C25E">Deein, Gridsada</mods:namePart>
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<mods:affiliation id="F5E3980E449188ECC8F9D8DF46C33B08">Inland Fisheries Research and Development Bureau, Department of Fisheries, NIFI Building, Kasetklang, Chatuchak, Bangkok 10900 Thailand. E-mail: dgridsada @ hotmail. com</mods:affiliation>
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</mods:name>
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<mods:name id="F6972A67D1267A4B68E4216D14924CD6" type="personal">
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<mods:roleTerm id="D3F9D264145E0769ABCEE4C2E4DC8360">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="B555BF5311C1416019655C31DEA075C4">Tangjitjaroen, Weerapongse</mods:namePart>
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<mods:affiliation id="222E07F0730C7A033AFA2C5FFDD0C125">Faculty of Veterinary Medicine, Chiang Mai University, Chiang Mai, 50100, Thailand. E-mail: weerapongse @ yahoo. com</mods:affiliation>
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<mods:name id="8D97472112DC7CDAA824F7A91A338E57" type="personal">
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<mods:roleTerm id="C5492F4D54359DE573F4A11E4273993B">Author</mods:roleTerm>
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<mods:namePart id="42F8B2EE24879FEC4846388D541FD176">Page, Lawrence M.</mods:namePart>
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<mods:affiliation id="EEBD50B9F8E5A34980F5186A5797A544">Florida Museum of Natural History, University of Florida, Gainesville, FL 32611 - 7800 USA. E-mail: lpage 1 @ ufl. edu</mods:affiliation>
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<mods:typeOfResource id="B784B10B21D4EC189FFA9E064D952408">text</mods:typeOfResource>
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<mods:title id="1814712C3BDFDB5893F9749D1F70A7A7">Zootaxa</mods:title>
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<mods:part id="B986E9699AB7B45AA33134E9CB66D2D5">
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<mods:date id="6C711FCBD52122F3EAFA95684EAA3A26">2014</mods:date>
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<mods:number id="796E3C5B4BF40B93713F35327B566492">2014-03-17</mods:number>
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<mods:number id="C4FB40127F2055C661ECBE2E825CE1B7">3779</mods:number>
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<treatment id="038987E7FFCCDB2BC0EF5EAA5E85A7F0" ID-DOI="http://doi.org/10.5281/zenodo.5061025" ID-GBIF-Taxon="183237225" ID-Zenodo-Dep="5061025" LSID="urn:lsid:plazi:treatment:038987E7FFCCDB2BC0EF5EAA5E85A7F0" httpUri="http://treatment.plazi.org/id/038987E7FFCCDB2BC0EF5EAA5E85A7F0" lastPageNumber="348" pageId="7" pageNumber="348">
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<subSubSection id="C33A657AFFCCDB2BC0EF5EAA5C73A2B0" pageId="7" pageNumber="348" type="nomenclature">
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<paragraph id="8B9F36F1FFCCDB2BC0EF5EAA5FE3A292" blockId="7.[151,747,694,790]" box="[151,747,694,720]" pageId="7" pageNumber="348">
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<heading id="D0D7819DFFCCDB2BC0EF5EAA5FE3A292" bold="true" box="[151,747,694,720]" fontSize="11" level="1" pageId="7" pageNumber="348" reason="1">
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<taxonomicName id="4C204D72FFCCDB2BC0EF5EAA5FE3A292" authority="(de Beaufort, 1933)" baseAuthorityName="de Beaufort" baseAuthorityYear="1933" box="[151,747,694,720]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pahangensis">
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<emphasis id="B954EAE3FFCCDB2BC0EF5EAA5FE3A292" bold="true" box="[151,747,694,720]" pageId="7" pageNumber="348">
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<emphasis id="B954EAE3FFCCDB2BC0EF5EAA5CFBA292" bold="true" box="[151,499,694,720]" italics="true" pageId="7" pageNumber="348">Lepidocephalus pahangensis</emphasis>
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(
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<bibRefCitation id="EFB14B00FFCCDB2BC27C5EAA5FEBA292" author="de Beaufort, L. F." box="[516,739,694,720]" pageId="7" pageNumber="348" pagination="31 - 36" refId="ref7518" refString="de Beaufort, L. F. (1933) On some new or rare species of Ostariophysi from the Malay Peninsula and a new species of Betta from Borneo. Bulletin of the Raffles Museum, 8, 31 - 36." type="journal article" year="1933">de Beaufort, 1933</bibRefCitation>
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||
)
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||
</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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||
<paragraph id="8B9F36F1FFCCDB2BC0EF5EC55C73A2B0" blockId="7.[151,747,694,790]" box="[151,379,729,754]" pageId="7" pageNumber="348">
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<collectingRegion id="49E4F813FFCCDB2BC0EF5EC55DE2A2B0" box="[151,234,729,754]" country="Malaysia" name="Pahang" pageId="7" pageNumber="348">Pahang</collectingRegion>
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Spirit Loach
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A657AFFCCDB2BC0EF5EE15DF7A354" box="[151,255,765,790]" pageId="7" pageNumber="348" type="description">
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<paragraph id="8B9F36F1FFCCDB2BC0EF5EE15DF7A354" blockId="7.[151,747,694,790]" box="[151,255,765,790]" pageId="7" pageNumber="348">
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(
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<figureCitation id="131B2A74FFCCDB2BC0E75EE15DFEA354" box="[159,246,765,790]" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="7" pageNumber="348">Fig. 1D</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A657AFFCCDB2BC0EF5F595E6CA338" pageId="7" pageNumber="348" type="reference_group">
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<paragraph id="8B9F36F1FFCCDB2BC0EF5F595E6CA338" blockId="7.[151,1437,837,890]" pageId="7" pageNumber="348">
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<treatmentCitationGroup id="AB3011DFFFCCDB2BC0EF5F595E6CA338" pageId="7" pageNumber="348">
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<treatmentCitation id="0A8110E0FFCCDB2BC0EF5F595FC8A319" author="de Beaufort, L. F." box="[151,704,837,859]" page="31" pageId="7" pageNumber="348" year="1933">
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<taxonomicName id="4C204D72FFCCDB2BC0EF5F595FC8A319" ID-CoL="8Z8S" authority="de Beaufort, 1933: 31" authorityName=", de Beaufort" authorityPageNumber="31" authorityYear="1933" box="[151,704,837,859]" family="Cobitidae" genus="Acanthophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pahangensis">
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<emphasis id="B954EAE3FFCCDB2BC0EF5F595CD3A319" box="[151,475,837,859]" italics="true" pageId="7" pageNumber="348">Acanthophthalmus pahangensis</emphasis>
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<bibRefCitation id="EFB14B00FFCCDB2BC19B5F595FC8A319" author="de Beaufort, L. F." box="[483,704,837,859]" pageId="7" pageNumber="348" pagination="31 - 36" refId="ref7518" refString="de Beaufort, L. F. (1933) On some new or rare species of Ostariophysi from the Malay Peninsula and a new species of Betta from Borneo. Bulletin of the Raffles Museum, 8, 31 - 36." type="journal article" year="1933">de Beaufort, 1933:31</bibRefCitation>
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</taxonomicName>
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||
</treatmentCitation>
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||
. Type locality: off Mentakab,
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<collectingRegion id="49E4F813FFCCDB2BC3815F59594EA319" box="[1017,1094,837,859]" country="Malaysia" name="Pahang" pageId="7" pageNumber="348">Pahang</collectingRegion>
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,
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<collectingRegion id="49E4F813FFCCDB2BC42A5F595995A319" box="[1106,1181,837,859]" country="Malaysia" name="Pahang" pageId="7" pageNumber="348">Pahang</collectingRegion>
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River, Malay Peninsula.
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<typeStatus id="549B8853FFCCDB2BC0BF5F785C2DA338" box="[199,293,868,890]" pageId="7" pageNumber="348" type="holotype">Holotype</typeStatus>
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||
ZRC 490; see
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||
<treatmentCitation id="0A8110E0FFCCDB2BC1C55F795F77A338" author="Hora, S. L." box="[445,639,868,890]" page="55" pageId="7" pageNumber="348" year="1941">
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||
<bibRefCitation id="EFB14B00FFCCDB2BC1C55F795F77A338" author="Hora, S. L." box="[445,639,868,890]" pageId="7" pageNumber="348" pagination="44 - 64" refId="ref7983" refString="Hora, S. L. (1941) Notes on Malayan fishes in the collection of the Raffles Museum, Singapore. Parts 2 and 3. Bulletin of the Raffles Museum, 17, 44 - 64." type="journal article" year="1941">Hora (1941:55–56)</bibRefCitation>
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</treatmentCitation>
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and
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<treatmentCitation id="0A8110E0FFCCDB2BC2C85F785E68A338" author="Alfred, E. R." box="[688,864,868,890]" page="70" pageId="7" pageNumber="348" year="1970">
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||
<bibRefCitation id="EFB14B00FFCCDB2BC2C85F785E68A338" author="Alfred, E. R." box="[688,864,868,890]" pageId="7" pageNumber="348" pagination="68 - 73" refId="ref7451" refString="Alfred, E. R. (1970) Type specimens of fishes in the National Museum, Singapore. Journal of the Singapore National Academy of Sciences, 2 (2), 68 - 73." type="journal article" year="1970">Alfred (1970:70)</bibRefCitation>
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</treatmentCitation>
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||
.
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||
</treatmentCitationGroup>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A657AFFCCDB2BC0EF5FBA5C49A4F9" pageId="7" pageNumber="348" type="diagnosis">
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<paragraph id="8B9F36F1FFCCDB2BC0EF5FBA5C49A4F9" blockId="7.[151,1437,934,1970]" pageId="7" pageNumber="348">
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<emphasis id="B954EAE3FFCCDB2BC0EF5FBA5C19A3FC" box="[151,273,934,959]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">
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||
<emphasis id="B954EAE3FFCCDB2BC0EF5FBA5C02A3FD" bold="true" box="[151,266,934,959]" pageId="7" pageNumber="348">Diagnosis</emphasis>
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.
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</emphasis>
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A species of
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<taxonomicName id="4C204D72FFCCDB2BC1C85FBB5F69A3FD" authorityName="Bleeker" authorityYear="1858" box="[432,609,935,959]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="genus">
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<emphasis id="B954EAE3FFCCDB2BC1C85FBB5F69A3FD" box="[432,609,935,959]" italics="true" pageId="7" pageNumber="348">Lepidocephalus</emphasis>
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</taxonomicName>
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that is readily distinguishable (
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<figureCitation id="131B2A74FFCCDB2BC3B25FBA5906A3FD" box="[970,1038,934,959]" captionStart="FIGURE 1" captionStartId="2.[151,249,1690,1712]" captionTargetBox="[153,1434,151,1673]" captionTargetId="figure-188@2.[153,1438,141,1673]" captionTargetPageId="2" captionText="FIGURE 1. Species of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. macrochir, syntype, BMNH 1866.5.2.55, 77.7 mm SL, Sumatra or Java; (C) L. pallens, holotype, RMNH 7783, 41.9 mm SL; (D) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (E) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos A, C, D and E by Z. Randall; B by M. Sabaj Perez." figureDoi="http://doi.org/10.5281/zenodo.4910405" httpUri="https://zenodo.org/record/4910405/files/figure.png" pageId="7" pageNumber="348">Fig. 1</figureCitation>
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,
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<tableCitation id="C6A2034AFFCCDB2BC4665FBA5979A3FD" box="[1054,1137,934,959]" captionStart="TABLE 1" captionStartId="9.[151,241,476,499]" captionTargetPageId="9" captionText="TABLE 1. Characteristics that vary among species of Lepidocephalus. Measurements are in mm or percentages of standard length (SL). Range is followed by average in parentheses." httpUri="http://table.plazi.org/id/DF5F6679FFC2DB25C0EF5DC05E6AA251" pageId="7" pageNumber="348" tableUuid="DF5F6679FFC2DB25C0EF5DC05E6AA251">Table 1</tableCitation>
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) from all other species of
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<taxonomicName id="4C204D72FFCCDB2BC0EF5FD75C40A3A1" authorityName="Bleeker" authorityYear="1858" box="[151,328,971,995]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="genus">
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<emphasis id="B954EAE3FFCCDB2BC0EF5FD75C40A3A1" box="[151,328,971,995]" italics="true" pageId="7" pageNumber="348">Lepidocephalus</emphasis>
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</taxonomicName>
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by the absence of scales on top of the head. It is further distinguished from
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<taxonomicName id="4C204D72FFCCDB2BC4BC5FD75843A3A0" authorityName="Roberts" authorityYear="1989" box="[1220,1355,971,994]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="spectrum">
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||
<emphasis id="B954EAE3FFCCDB2BC4BC5FD75843A3A0" box="[1220,1355,971,994]" italics="true" pageId="7" pageNumber="348">L. spectrum</emphasis>
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</taxonomicName>
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by the presence of eyes and dark pigment on the body, and absence of tubules along the lateral line; from
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<taxonomicName id="4C204D72FFCCDB2BC57F5FEC587FA445" baseAuthorityName="Vaillant" baseAuthorityYear="1902" box="[1287,1399,1007,1031]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pallens">
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<emphasis id="B954EAE3FFCCDB2BC57F5FEC587FA445" box="[1287,1399,1007,1031]" italics="true" pageId="7" pageNumber="348">L. pallens</emphasis>
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</taxonomicName>
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by having the dorsal-fin origin well behind (vs. over) the pelvic-fin origin (predorsal length 68.2 vs. 59.9% SL); from
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<taxonomicName id="4C204D72FFCCDB2BC0EF58245C1EA40D" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[151,278,1080,1103]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="nanensis">
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<emphasis id="B954EAE3FFCCDB2BC0EF58245C1EA40D" box="[151,278,1080,1103]" italics="true" pageId="7" pageNumber="348">L. nanensis</emphasis>
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</taxonomicName>
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by having large papillae on the lips (
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<figureCitation id="131B2A74FFCCDB2BC2CD582A5E04A40D" box="[693,780,1078,1103]" captionStart="FIGURE 3" captionStartId="5.[151,249,945,967]" captionTargetBox="[165,1424,354,917]" captionTargetId="figure-182@5.[159,1428,350,920]" captionTargetPageId="5" captionText="FIGURE 3. Mouths of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (C) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos by Z. Randall." figureDoi="http://doi.org/10.5281/zenodo.4910409" httpUri="https://zenodo.org/record/4910409/files/figure.png" pageId="7" pageNumber="348">Fig. 3B</figureCitation>
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); and from
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<taxonomicName id="4C204D72FFCCDB2BC3F758245928A40D" baseAuthorityName="Bleeker" baseAuthorityYear="1854" box="[911,1056,1079,1103]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="macrochir">
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<emphasis id="B954EAE3FFCCDB2BC3F758245928A40D" box="[911,1056,1079,1103]" italics="true" pageId="7" pageNumber="348">L. macrochir</emphasis>
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</taxonomicName>
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by having a longer head (21.8 vs. 16.6–18.8% SL). Although counts for
|
||
<taxonomicName id="4C204D72FFCCDB2BC23458475FFFA431" authorityName=", de Beaufort" authorityYear="1933" box="[588,759,1115,1139]" family="Cobitidae" genus="Acanthophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pahangensis">
|
||
<emphasis id="B954EAE3FFCCDB2BC23458475FFFA431" box="[588,759,1115,1139]" italics="true" pageId="7" pageNumber="348">L. pahangensis</emphasis>
|
||
</taxonomicName>
|
||
are available only for the
|
||
<typeStatus id="549B8853FFCCDB2BC45D58465982A431" box="[1061,1162,1114,1139]" pageId="7" pageNumber="348" type="holotype">holotype</typeStatus>
|
||
, it differs from all other species in having 4 (vs. 5–6) branched anal-fin rays, and from all except
|
||
<taxonomicName id="4C204D72FFCCDB2BC3A9589C595DA4D5" authorityName="Roberts" authorityYear="1989" box="[977,1109,1152,1175]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="spectrum">
|
||
<emphasis id="B954EAE3FFCCDB2BC3A9589C595DA4D5" box="[977,1109,1152,1175]" italics="true" pageId="7" pageNumber="348">L. spectrum</emphasis>
|
||
</taxonomicName>
|
||
in having 9 (vs. 8) branched dorsal-fin rays.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFCCDB2BC0BF58DA5C5AA6D0" pageId="7" pageNumber="348" type="description">
|
||
<paragraph id="8B9F36F1FFCCDB2BC0BF58DA5919A565" blockId="7.[151,1437,934,1970]" pageId="7" pageNumber="348">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF58DA5C50A49C" box="[199,344,1222,1247]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF58DA5C59A49D" bold="true" box="[199,337,1222,1247]" pageId="7" pageNumber="348">Description</emphasis>
|
||
.
|
||
</emphasis>
|
||
Body deep, slab-sided; greatest depth 17.0% SL. Head narrow, length 21.8% SL. Eye small, in dorsal half of head in shallow depression above bifid suborbital spine. Origin of dorsal fin behind origin of pelvic fin. Small axillary lobe on pelvic fin. Large fleshy lobe at origin of pectoral fin.
|
||
</paragraph>
|
||
<paragraph id="8B9F36F1FFCCDB2BC0BF592F59ABA608" blockId="7.[151,1437,934,1970]" pageId="7" pageNumber="348">
|
||
No scales on top of head; scales present on cheek and opercle; body completely covered with minute, partially embedded scales. Lateral line complete, 174 pores along lateral line, 6 pores on caudal fin; no tubules on lateral line. Mouth horseshoe-shaped; lips with large papillae on outer surfaces; upper lip without indentation; lower lip with median indentation on lower edge (
|
||
<figureCitation id="131B2A74FFCCDB2BC21559825FC0A5F5" box="[621,712,1438,1463]" captionStart="FIGURE 3" captionStartId="5.[151,249,945,967]" captionTargetBox="[165,1424,354,917]" captionTargetId="figure-182@5.[159,1428,350,920]" captionTargetPageId="5" captionText="FIGURE 3. Mouths of Lepidocephalus. (A) L. nanensis, holotype, NIFI 4388, 72.0 mm SL, Nan River, Thailand; (B) L. pahangensis, holotype, ZRC 490, 35.8 mm SL; (C) L. spectrum, paratype, USNM 230267, 49.2 mm SL, Kalimantan. Photos by Z. Randall." figureDoi="http://doi.org/10.5281/zenodo.4910409" httpUri="https://zenodo.org/record/4910409/files/figure.png" pageId="7" pageNumber="348">Fig. 3B</figureCitation>
|
||
). Two pairs of rostral barbels; inner pair reaching to corner of mouth, outer pair reaching past corner of mouth; one pair of maxillary barbels, reaching approximately to vertical at posterior end of groove containing suborbital spine. Large flap on anterior nostril. Dorsal rays iii,9; pectoral rays 11; pelvic rays 7; anal rays ii,4; upper branched caudal rays 7; lower branched caudal rays 7. Pectoral fin falcate with 2
|
||
<superScript id="7C559BB9FFCCDB2BC0A55A315DE4A679" attach="left" box="[221,236,1581,1595]" fontSize="6" pageId="7" pageNumber="348">nd</superScript>
|
||
ray longer than other rays (15.6% SL), but not thickened. Pelvic fin small, 6.4% SL.
|
||
</paragraph>
|
||
<paragraph id="8B9F36F1FFCCDB2BC0BF5A495C5AA6D0" blockId="7.[151,1437,934,1970]" pageId="7" pageNumber="348">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF5A495C47A62F" box="[199,335,1621,1646]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF5A495C40A62C" bold="true" box="[199,328,1621,1646]" pageId="7" pageNumber="348">Coloration</emphasis>
|
||
.
|
||
</emphasis>
|
||
Described as being uniformly colored (presumably pinkish gray-brown as in
|
||
<taxonomicName id="4C204D72FFCCDB2BC4B15A4A5843A62F" authorityName="Deein & Tangjitjaroen & Page" authorityYear="2014" box="[1225,1355,1622,1645]" class="Actinopterygii" family="Cobitidae" genus="Lepidocephalus" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="nanensis">
|
||
<emphasis id="B954EAE3FFCCDB2BC4B15A4A5843A62F" box="[1225,1355,1622,1645]" italics="true" pageId="7" pageNumber="348">L. nanensis</emphasis>
|
||
</taxonomicName>
|
||
) by
|
||
<bibRefCitation id="EFB14B00FFCCDB2BC5FA5A495C46A6D0" author="de Beaufort, L. F." pageId="7" pageNumber="348" pagination="31 - 36" refId="ref7518" refString="de Beaufort, L. F. (1933) On some new or rare species of Ostariophysi from the Malay Peninsula and a new species of Betta from Borneo. Bulletin of the Raffles Museum, 8, 31 - 36." type="journal article" year="1933">de Beaufort (1933)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFCCDB2BC0BF5A815894A6F4" box="[199,1436,1693,1718]" pageId="7" pageNumber="348" type="distribution">
|
||
<paragraph id="8B9F36F1FFCCDB2BC0BF5A815894A6F4" blockId="7.[151,1437,934,1970]" box="[199,1436,1693,1718]" pageId="7" pageNumber="348">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF5A815C68A6F7" box="[199,352,1693,1718]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF5A815C51A6F4" bold="true" box="[199,345,1693,1718]" pageId="7" pageNumber="348">Distribution</emphasis>
|
||
.
|
||
</emphasis>
|
||
Known only from the
|
||
<typeStatus id="549B8853FFCCDB2BC2275A825F87A6F4" box="[607,655,1694,1718]" pageId="7" pageNumber="348">type</typeStatus>
|
||
locality, the
|
||
<collectingRegion id="49E4F813FFCCDB2BC3675A815E79A6F4" box="[799,881,1693,1718]" country="Malaysia" name="Pahang" pageId="7" pageNumber="348">Pahang</collectingRegion>
|
||
River at Mentakab, Peninsular
|
||
<collectingCountry id="F3377661FFCCDB2BC4AC5A815833A6F4" box="[1236,1339,1693,1718]" name="Malaysia" pageId="7" pageNumber="348">Malaysia</collectingCountry>
|
||
(
|
||
<figureCitation id="131B2A74FFCCDB2BC5315A815886A6F4" box="[1353,1422,1693,1718]" captionStart="FIGURE 2" captionStartId="4.[151,249,1699,1721]" captionTargetBox="[153,1429,349,1673]" captionTargetId="figure-196@4.[151,1436,339,1677]" captionTargetPageId="4" captionText="FIGURE 2. Distribution of specimens of Lepidocephalus examined." figureDoi="http://doi.org/10.5281/zenodo.4910407" httpUri="https://zenodo.org/record/4910407/files/figure.png" pageId="7" pageNumber="348">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A657AFFCCDB2BC0BF5ADD5E85A7F0" pageId="7" pageNumber="348" type="discussion">
|
||
<paragraph id="8B9F36F1FFCCDB2BC0BF5ADD5E85A7F0" blockId="7.[151,1437,934,1970]" pageId="7" pageNumber="348">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF5ADD5C33A69B" box="[199,315,1729,1754]" inLineHeading="true" pageId="7" pageNumber="348" reason="1">
|
||
<emphasis id="B954EAE3FFCCDB2BC0BF5ADD5C3CA698" bold="true" box="[199,308,1729,1754]" pageId="7" pageNumber="348">Remarks</emphasis>
|
||
.
|
||
</emphasis>
|
||
In the original description of
|
||
<taxonomicName id="4C204D72FFCCDB2BC2D95ADD59F0A698" authority=", de Beaufort (1933)" authorityName=", de Beaufort" authorityYear="1933" box="[673,1272,1729,1754]" family="Cobitidae" genus="Acanthophthalmus" higherTaxonomySource="CoL" kingdom="Animalia" order="Cypriniformes" pageId="7" pageNumber="348" phylum="Chordata" rank="species" species="pahangensis">
|
||
<emphasis id="B954EAE3FFCCDB2BC2D95ADD5902A69B" box="[673,1034,1729,1753]" italics="true" pageId="7" pageNumber="348">Acanthophthalmus pahangensis</emphasis>
|
||
,
|
||
<bibRefCitation id="EFB14B00FFCCDB2BC4645ADD59F0A698" author="de Beaufort, L. F." box="[1052,1272,1729,1754]" pageId="7" pageNumber="348" pagination="31 - 36" refId="ref7518" refString="de Beaufort, L. F. (1933) On some new or rare species of Ostariophysi from the Malay Peninsula and a new species of Betta from Borneo. Bulletin of the Raffles Museum, 8, 31 - 36." type="journal article" year="1933">de Beaufort (1933)</bibRefCitation>
|
||
</taxonomicName>
|
||
described the head as scaleless. The
|
||
<typeStatus id="549B8853FFCCDB2BC1EA5AF95CFCA6BC" box="[402,500,1765,1790]" pageId="7" pageNumber="348" type="holotype">holotype</typeStatus>
|
||
has no scales on top of the head, but has scales on the cheeks and opercles. It seems likely that de Beaufort was referring only to the top of the head in his description, as scales are readily apparent on the cheek and opercle.
|
||
<bibRefCitation id="EFB14B00FFCCDB2BC1E35B325F2DA704" author="Hora, S. L." box="[411,549,1837,1862]" pageId="7" pageNumber="348" pagination="44 - 64" refId="ref7983" refString="Hora, S. L. (1941) Notes on Malayan fishes in the collection of the Raffles Museum, Singapore. Parts 2 and 3. Bulletin of the Raffles Museum, 17, 44 - 64." type="journal article" year="1941">Hora (1941)</bibRefCitation>
|
||
examined the
|
||
<typeStatus id="549B8853FFCCDB2BC2B45B315E26A704" box="[716,814,1837,1862]" pageId="7" pageNumber="348" type="holotype">holotype</typeStatus>
|
||
and found scales “on the head in the region below and behind the eyes.” Oddly, Hora included a drawing of the head and anterior body of the
|
||
<typeStatus id="549B8853FFCCDB2BC4005B4D59D2A728" box="[1144,1242,1873,1898]" pageId="7" pageNumber="348" type="holotype">holotype</typeStatus>
|
||
that differs from his verbal description in showing scales on top of the head as well as on the cheek and opercle. De Beaufort was correct in originally noting the absence of scales on top of the head.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |