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<document id="9BFBF844446B196F2A5A487A1CF321AE" ID-CLB-Dataset="8228" ID-DOI="10.11646/zootaxa.4858.4.6" ID-GBIF-Dataset="8260832b-9a33-48e0-b1ab-a0cb90656b9a" ID-ISSN="1175-5326" ID-Zenodo-Dep="4412724" ID-ZooBank="4B56C5A4-3B35-40A6-B6F2-EEB3B4C3F885" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609691798849" checkinUser="plazi" docAuthor="Ferreira, Luciane Augusto De Azevedo, Santana-Moreno, L. Daniel &amp; Anker, Arthur" docDate="2020" docId="038987C34E36FFCED7CA043A2FE6FD3A" docLanguage="en" docName="zootaxa.4858.4.6.pdf" docOrigin="Zootaxa 4858 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Petrolisthes lazarus Ferreira &amp; Santana-Moreno &amp; Anker 2020, sp. nov." docType="treatment" docVersion="9" lastPageNumber="587" masterDocId="FFB0FFBB4E37FFC5D75D00582E24FFBF" masterDocTitle="A new species of the porcelain crab genus Petrolisthes from the Pacific coast of Panama, with taxonomic notes on closely related species (Decapoda: Anomura Porcellanidae)" masterLastPageNumber="588" masterPageNumber="576" pageNumber="577" updateTime="1698851651748" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="0AB057C3CE9F81895BFD59F81173FD5B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="5B24CC1C6BA2AEACDE7618B77E2D0F34">
<mods:title id="78A9E8BCFCB18ED36F60CD096E213BCB">A new species of the porcelain crab genus Petrolisthes from the Pacific coast of Panama, with taxonomic notes on closely related species (Decapoda: Anomura Porcellanidae)</mods:title>
</mods:titleInfo>
<mods:name id="87F269E718474C8F767DD2EECB7AF44C" type="personal">
<mods:role id="527A3167784B315815C781C71918EC3A">
<mods:roleTerm id="4EF57435EE9E16B3809DB279164298EF">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="44E8C9AA6BE1D7A2099E29529CDED4C7">Ferreira, Luciane Augusto De Azevedo</mods:namePart>
<mods:nameIdentifier id="DD356B4588563E224B82E0DB0D95CCE1" type="email">lucianeaaf@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2099123C5FE7D0E034B1F4EDD2B98CB2" type="personal">
<mods:role id="3C71865FDBB49563487AB7B41FC8EED4">
<mods:roleTerm id="E54A2017B4C853491C0F4C8318E076C1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5608357294C52563F0B6FE90FA8E9B74">Santana-Moreno, L. Daniel</mods:namePart>
<mods:affiliation id="39B0A88FBAA72DC84FE16C0CAFBC5721">danielsantana 1386 @ gmail. com; http: // orcid. org / 0000 - 0002 - 0638 - 8590</mods:affiliation>
<mods:nameIdentifier id="C51893E172F51226B8FB882380A86469" type="email">danielsantana1386@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="E5135F0C83FB7DF0C43B68DC2B3C6DD3" type="personal">
<mods:role id="ACD15D6A67EC483698C7CFE6C21D9864">
<mods:roleTerm id="1F977565CFFA033037DD5FCC25453E99">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="07F9363C74C9ABDEE9F37917C415652F">Anker, Arthur</mods:namePart>
<mods:affiliation id="CE3F2FE7A9D8E8470DE7CD559E256F68">Universidade Federal de Goiás, Campus Samambaia, Instituto de Ciências Biológicas-ICB- 5. Av. Esperança, s / n., 74690 - 900, Goiânia, Goiás, Brazil. arthuranker 7 @ gmail. com; http: // orcid. org / 0000 - 0002 - 5350 - 4267</mods:affiliation>
<mods:nameIdentifier id="26A8572682AAC321A5BCD01F253BDCC2" type="email">lucianeaaf@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="FFA1BDF699E11C073F81507EB831665A">text</mods:typeOfResource>
<mods:relatedItem id="4F37EEF1B38C137A444BAF59EBBFA658" type="host">
<mods:titleInfo id="3E4CC85B855B92514A7B59BFBB62902E">
<mods:title id="B760C9DDE87DAC5EC3A8684309DF648B">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="8DE866319A4B065D41DE86BB8E12783F">
<mods:date id="4A9710407A2397849B27920C660F1976">2020</mods:date>
<mods:detail id="28036976E3FD173E656FABE60FC91B0A" type="pubDate">
<mods:number id="F2922D49609A27ECA04ADE8973B311D4">2020-10-02</mods:number>
</mods:detail>
<mods:detail id="4D9834406570D7B3E52BCD0803231C30" type="volume">
<mods:number id="570A2EEA479D60B422105690A964202C">4858</mods:number>
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<mods:number id="49A82FE54C0CECA2BFC09EB361D630FC">4</mods:number>
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<mods:start id="53349CF65FC45F83C0212563F44780E5">576</mods:start>
<mods:end id="A3C91B92FAFCE430CC1AEC45D8A1DA46">588</mods:end>
</mods:extent>
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<mods:classification id="16E6663090C9BA40EE2C5B72118DFDE2">journal article</mods:classification>
<mods:identifier id="25E639A839463B8CDB9F664FF40663D1" type="CLB-Dataset">8228</mods:identifier>
<mods:identifier id="80FA6D6EB0C26ABE402F5801B2AA9C7F" type="DOI">10.11646/zootaxa.4858.4.6</mods:identifier>
<mods:identifier id="8BFC9C79BB5762BFBEC2802B5B598FFA" type="GBIF-Dataset">8260832b-9a33-48e0-b1ab-a0cb90656b9a</mods:identifier>
<mods:identifier id="F572C2F319CF4081CE6088C92604AA15" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="D0C6860588F4732D231EFC9FC57C4E9C" type="Zenodo-Dep">4412724</mods:identifier>
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<treatment id="038987C34E36FFCED7CA043A2FE6FD3A" ID-DOI="http://doi.org/10.5281/zenodo.4538551" ID-GBIF-Taxon="177324755" ID-Zenodo-Dep="4538551" LSID="urn:lsid:plazi:treatment:038987C34E36FFCED7CA043A2FE6FD3A" httpUri="http://treatment.plazi.org/id/038987C34E36FFCED7CA043A2FE6FD3A" lastPageId="11" lastPageNumber="587" pageId="1" pageNumber="577">
<subSubSection id="C33A655E4E36FFC4D7CA043A2FCDFBC2" box="[151,489,1122,1149]" pageId="1" pageNumber="577" type="nomenclature">
<paragraph id="8B9F36D54E36FFC4D7CA043A2FCDFBC2" blockId="1.[151,489,1122,1185]" box="[151,489,1122,1149]" pageId="1" pageNumber="577">
<heading id="D0D781B94E36FFC4D7CA043A2FCDFBC2" bold="true" box="[151,489,1122,1149]" fontSize="11" level="1" pageId="1" pageNumber="577" reason="1">
<emphasis id="B954EAC74E36FFC4D7CA043A2FCDFBC2" bold="true" box="[151,489,1122,1149]" pageId="1" pageNumber="577">
<taxonomicName id="4C204D564E36FFC4D7CA043A2FA0FBC2" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[151,388,1122,1149]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="577" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E36FFC4D7CA043A2FA0FBC2" bold="true" box="[151,388,1122,1149]" italics="true" pageId="1" pageNumber="577">Petrolisthes lazarus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26757BC4E36FFC4D6D1043B2FCDFBC2" box="[396,489,1123,1149]" pageId="1" pageNumber="577" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C33A655E4E36FFC4D7CA04DF2F36FB1E" box="[151,274,1159,1185]" pageId="1" pageNumber="577" type="description">
<paragraph id="8B9F36D54E36FFC4D7CA04DF2F36FB1E" blockId="1.[151,489,1122,1185]" box="[151,274,1159,1185]" pageId="1" pageNumber="577">
(
<figureCitation id="131B2A504E36FFC4D7C204DF2F2DFB1E" box="[159,265,1159,1185]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,1860,1884]" captionStartId-1="3.[151,250,1962,1986]" captionStartId-2="4.[151,250,985,1009]" captionTargetBox-0="[273,1313,188,1826]" captionTargetBox-1="[182,1406,181,1938]" captionTargetBox-2="[151,1435,181,961]" captionTargetId-0="figure-18@2.[264,1323,181,1836]" captionTargetId-1="figure-23@3.[182,1406,181,1938]" captionTargetId-2="figure-18@4.[151,1435,181,961]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) carapace, dorsal view; (B) fronto-orbital region of carapace, anterior view; (C) carapace, lateral view of branchiostegite; (D) right antennule, ventral view; (E) right antenna, dorsal view (flagellum omitted); (F) left Mxp 3, ventral (exterior) view (flagellum of exopod omitted); (G) left P1, dorsal view; (H) right P1, dorsal view; (I) right P2, lateral view; (J) telson, external view; (K) thoracic sternum, ventral view. Setae omitted for clarity. Scale bars: 1 mm." captionText-1="FIGURE 2. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker." captionText-2="FIGURE 3. Petrolisthes lazarus sp. nov., female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker." figureDoi-0="http://doi.org/10.5281/zenodo.4412726" figureDoi-1="http://doi.org/10.5281/zenodo.4412728" figureDoi-2="http://doi.org/10.5281/zenodo.4412730" httpUri-0="https://zenodo.org/record/4412726/files/figure.png" httpUri-1="https://zenodo.org/record/4412728/files/figure.png" httpUri-2="https://zenodo.org/record/4412730/files/figure.png" pageId="1" pageNumber="577">Figs. 1–3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C33A655E4E36FFC4D7CA04972C03FA22" pageId="1" pageNumber="577" type="materials_examined">
<paragraph id="8B9F36D54E36FFC4D7CA04972C03FA22" blockId="1.[151,1437,1230,1977]" pageId="1" pageNumber="577">
<emphasis id="B954EAC74E36FFC4D7CA04972F64FB57" bold="true" box="[151,320,1230,1257]" pageId="1" pageNumber="577">Type material</emphasis>
.
<materialsCitation id="3B483C884E36FFC4D60D04972F3DFAEA" ID-GBIF-Occurrence="3034646301" collectingDate="2015-04-22" collectionCode="R, T" collectorName="A. Anker &amp; Anker, J &amp; Luque, A. &amp; Palmer &amp; Kaji" country="Panama" county="Saboga" elevation="8" latitude="8.644305" location="Pacific" longLatPrecision="1" longitude="-79.06209" municipality="Las Perlas Islands" pageId="1" pageNumber="577" specimenCode="MZUSP 33807" specimenCount="1" typeStatus="holotype">
<typeStatus id="549B88774E36FFC4D60D04972F99FB56" box="[336,445,1231,1257]" pageId="1" pageNumber="577" type="holotype">Holotype</typeStatus>
: female, cl
<quantity id="4CD89B304E36FFC4D51F04972C8EFB56" box="[578,682,1231,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="1" pageNumber="577" unit="mm" value="10.8">10.8 mm</quantity>
, cw
<quantity id="4CD89B304E36FFC4D5BC04962D63FB56" box="[737,839,1230,1257]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="577" unit="mm" value="12.5">12.5 mm</quantity>
(
<specimenCode id="DB869EAE4E36FFC4D40A04972A2CFB56" box="[855,1032,1231,1257]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="577">MZUSP 33807</specimenCode>
),
<collectingCountry id="F33776454E36FFC4D34304972A5EFB56" box="[1054,1146,1231,1257]" name="Panama" pageId="1" pageNumber="577">Panama</collectingCountry>
,
<location id="8EFF600E4E36FFC4D3DB04972AF0FB56" LSID="urn:lsid:plazi:treatment:038987C34E36FFCED7CA043A2FE6FD3A:8EFF600E4E36FFC4D3DB04972AF0FB56" box="[1158,1236,1231,1257]" country="Panama" county="Saboga" latitude="8.644305" longLatPrecision="1" longitude="-79.06209" municipality="Las Perlas Islands" name="Pacific" pageId="1" pageNumber="577">Pacific</location>
coast,
<collectingMunicipality id="6BFBACAF4E36FFC4D27B04972ECDFAB2" pageId="1" pageNumber="577">Las Perlas Islands</collectingMunicipality>
, small rocky islet off north-eastern coast of Isla
<collectingCounty id="62FE4E594E36FFC4D44604AB2D54FAB2" box="[795,880,1267,1293]" pageId="1" pageNumber="577">Saboga</collectingCounty>
,
<geoCoordinate id="EE1450124E36FFC4D42304AA2A3EFAB2" box="[894,1050,1266,1293]" degrees="8" direction="north" minutes="38" orientation="latitude" pageId="1" pageNumber="577" precision="1" seconds="39.5" value="8.644305">8°38’39.5”N–</geoCoordinate>
<geoCoordinate id="EE1450124E36FFC4D34704AA2AE4FAB2" box="[1050,1216,1266,1293]" degrees="79" direction="west" minutes="03" orientation="longitude" pageId="1" pageNumber="577" precision="1" seconds="43.5" value="-79.06209">79°03’43.5”W</geoCoordinate>
, rock-sand bottom,
<quantity id="4CD89B304E36FFC4D7CA054E2ED7FA8E" box="[151,243,1302,1329]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="15.0" metricValueMin="0.0" pageId="1" pageNumber="577" unit="m" value="0.75" valueMax="1.5" valueMin="0.0">
<elevation id="000DD1E64E36FFC4D7CA054E2ED7FA8E" box="[151,243,1302,1329]" metricMagnitude="-1" metricUnit="m" metricValue="7.5" metricValueMax="15.0" metricValueMin="0.0" pageId="1" pageNumber="577" unit="m" value="0.75" valueMax="1.5" valueMin="0.0">0–1.5 m</elevation>
</quantity>
at low tide, strong current, under rocks and in dead corals, leg.
<collectorName id="26D553034E36FFC4D4E7054F2A06FA8E" box="[954,1058,1303,1329]" pageId="1" pageNumber="577">A. Anker</collectorName>
, J.
<collectorName id="26D553034E36FFC4D31B054F2A91FA8E" box="[1094,1205,1303,1329]" pageId="1" pageNumber="577">Luque, A.</collectorName>
<collectionCode id="ED31AE104E36FFC4D3E8054F2AEFFA8E" box="[1205,1227,1303,1329]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="577">R</collectionCode>
.
<collectorName id="26D553034E36FFC4D38B054F2B01FA8E" box="[1238,1317,1303,1329]" pageId="1" pageNumber="577">Palmer</collectorName>
&amp;
<collectionCode id="ED31AE104E36FFC4D215054F2B7EFA8E" box="[1352,1370,1303,1329]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="577">T</collectionCode>
.
<collectorName id="26D553034E36FFC4D238054F2BBCFA8E" box="[1381,1432,1303,1329]" pageId="1" pageNumber="577">Kaji</collectorName>
,
<date id="FF9E10154E36FFC4D7CA05622F3DFAEA" box="[151,281,1338,1365]" pageId="1" pageNumber="577" value="2015-04-22">
<collectingDate id="EFDAE9FD4E36FFC4D7CA05622F3DFAEA" box="[151,281,1338,1365]" pageId="1" pageNumber="577" value="2015-04-22">22.04.2015</collectingDate>
</date>
</materialsCitation>
.
<materialsCitation id="3B483C884E36FFC4D67E05632C07FA22" ID-GBIF-Occurrence="3034646302" collectingDate="2015-03-20" collectionCode="T" collectorName="A. Anker &amp; Anker, J. F &amp; Kaji" country="Panama" latitude="8.455333" location="Pacific" longLatPrecision="1" longitude="-79.96928" municipality="Rio Mar" pageId="1" pageNumber="577" specimenCode="MZUSP 33686" specimenCount="1" typeStatus="paratype">
<typeStatus id="549B88774E36FFC4D67E05632FADFAEA" box="[291,393,1339,1365]" pageId="1" pageNumber="577" type="paratype">Paratype</typeStatus>
: ovigerous female, cl
<quantity id="4CD89B304E36FFC4D5DD05622CFDFAEA" box="[640,729,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="1" pageNumber="577" unit="mm" value="8.5">8.5 mm</quantity>
, cw
<quantity id="4CD89B304E36FFC4D45105632D47FAEA" box="[780,867,1339,1365]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="1" pageNumber="577" unit="mm" value="9.9">9.9 mm</quantity>
(
<specimenCode id="DB869EAE4E36FFC4D42C05632A05FAE9" box="[881,1057,1339,1366]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="577">MZUSP 33686</specimenCode>
),
<collectingCountry id="F33776454E36FFC4D36905632AB4FAEA" box="[1076,1168,1339,1365]" name="Panama" pageId="1" pageNumber="577">Panama</collectingCountry>
,
<location id="8EFF600E4E36FFC4D3C605632ACDFAEA" LSID="urn:lsid:plazi:treatment:038987C34E36FFCED7CA043A2FE6FD3A:8EFF600E4E36FFC4D3C605632ACDFAEA" box="[1179,1257,1339,1365]" country="Panama" latitude="8.455333" longLatPrecision="1" longitude="-79.96928" municipality="Rio Mar" name="Pacific" pageId="1" pageNumber="577">Pacific</location>
coast,
<collectingMunicipality id="6BFBACAF4E36FFC4D26505632BBCFAEA" box="[1336,1432,1339,1365]" pageId="1" pageNumber="577">Río Mar</collectingMunicipality>
,
<geoCoordinate id="EE1450124E36FFC4D7CA05062F17FAC6" box="[151,307,1374,1401]" degrees="8" direction="north" minutes="27" orientation="latitude" pageId="1" pageNumber="577" precision="1" seconds="19.2" value="8.455333">8°27’19.2”N–</geoCoordinate>
<geoCoordinate id="EE1450124E36FFC4D66E05062FFEFAC6" box="[307,474,1374,1401]" degrees="79" direction="west" minutes="58" orientation="longitude" pageId="1" pageNumber="577" precision="1" seconds="09.4" value="-79.96928">79°58’09.4”W</geoCoordinate>
, rocky intertidal, low tide, under rocks and in rock crevices, leg.
<collectorName id="26D553034E36FFC4D3E405072B07FAC6" box="[1209,1315,1375,1401]" pageId="1" pageNumber="577">A. Anker</collectorName>
, J.F. Lazarus-Agudelo &amp;
<collectionCode id="ED31AE104E36FFC4D61B05DB2F7CFA22" box="[326,344,1411,1437]" name="Tavera, Department of Geology and Geophysics" pageId="1" pageNumber="577">T</collectionCode>
.
<collectorName id="26D553034E36FFC4D63E05DB2FB2FA22" box="[355,406,1411,1437]" pageId="1" pageNumber="577">Kaji</collectorName>
,
<date id="FF9E10154E36FFC4D6FF05DA2C07FA22" box="[418,547,1410,1437]" pageId="1" pageNumber="577" value="2015-03-20">
<collectingDate id="EFDAE9FD4E36FFC4D6FF05DA2C07FA22" box="[418,547,1410,1437]" pageId="1" pageNumber="577" value="2015-03-20">20.03.2015</collectingDate>
</date>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C33A655E4E36FFC0D79A05FE2DFCFEDA" lastPageId="5" lastPageNumber="581" pageId="1" pageNumber="577" type="description">
<paragraph id="8B9F36D54E36FFC4D79A05FE2EF0F896" blockId="1.[151,1437,1230,1977]" pageId="1" pageNumber="577">
<emphasis id="B954EAC74E36FFC4D79A05FE2F75FA7F" bold="true" box="[199,337,1446,1472]" pageId="1" pageNumber="577">Description</emphasis>
. Carapace subcircular, distinctly wider than long; dorsal surface rugose and sparsely granular. Frontal and orbital regions depressed, rugose. Front about 1/3 width of maximal carapace width; margins granulate, trilobate in dorsal and frontal views, devoid of setae; median lobe broadly subtriangular, distally rounded, slightly overreaching lateral lobes, lateral lobes rounded, somewhat protruding anteriorly. Outer orbital angle well defined, rounded, with granular margin. Orbit with slightly granulate margin. Lateral carapace margin poorly defined by longitudinal row of granules extending posteriorly from epibranchial angle to mesobranchial margin. Protogastric region swollen; epibranchial and hepatic regions swollen, delimited by shallow grooves; epibranchial regions with conspicuous granules, without spines. Epibranchial angle and cervical groove well defined. Posterolateral carapace surface rugose; posterior region depressed; posterior margin concave, devoid of setae. Branchiostegite with longitudinal, subaparallel wrinkles, fringed with long plumose setae; anterodorsal region with slightly projecting, granular crest.
</paragraph>
<paragraph id="8B9F36D54E36FFC4D79A076B2DFBF82A" blockId="1.[151,1437,1230,1977]" pageId="1" pageNumber="577">Thoracic sternite 3 with median lobe broad, subtriangular, unarmed, not overreaching lateral lobes; lateral lobes narrow, subtriangular. Thoracic sternites 4–7 broad, sutures between sternites incomplete, distinct only laterally. Thoracic sternite 8 with anterior margin convex, posterior margin concave.</paragraph>
<paragraph id="8B9F36D54E36FFC4D79A07C72AEBF806" blockId="1.[151,1437,1230,1977]" box="[199,1231,1951,1977]" pageId="1" pageNumber="577">Ocular peduncle subcylindrical dorsally, ventral surface flattened. Cornea small, not dilated.</paragraph>
<caption id="DF5F665D4E35FFC7D7CA071C2D93F854" ID-DOI="http://doi.org/10.5281/zenodo.4412726" ID-Zenodo-Dep="4412726" httpUri="https://zenodo.org/record/4412726/files/figure.png" pageId="2" pageNumber="578" startId="2.[151,250,1860,1884]" targetBox="[273,1313,188,1826]" targetPageId="2">
<paragraph id="8B9F36D54E35FFC7D7CA071C2D93F854" blockId="2.[151,1437,1860,2028]" pageId="2" pageNumber="578">
<emphasis id="B954EAC74E35FFC7D7CA071C2F29F8E3" bold="true" box="[151,269,1860,1884]" pageId="2" pageNumber="578">FIGURE 1</emphasis>
.
<taxonomicName id="4C204D564E35FFC7D644071C2FC6F8E3" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[281,482,1860,1884]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="578" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E35FFC7D644071C2FC6F8E3" box="[281,482,1860,1884]" italics="true" pageId="2" pageNumber="578">Petrolisthes lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E35FFC7D6B5071C2C1CF8E3" bold="true" box="[488,568,1860,1884]" pageId="2" pageNumber="578">
<taxonomicNameLabel id="A26757BC4E35FFC7D6B5071C2C1CF8E3" box="[488,568,1860,1884]" pageId="2" pageNumber="578" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) carapace, dorsal view; (B) fronto-orbital region of carapace, anterior view; (C) carapace, lateral view of branchiostegite; (D) right antennule, ventral view; (E) right antenna, dorsal view (flagellum omitted); (F) left Mxp 3, ventral (exterior) view (flagellum of exopod omitted); (G) left P1, dorsal view; (H) right P1, dorsal view; (I) right P2, lateral view; (J) telson, external view; (K) thoracic sternum, ventral view. Setae omitted for clarity. Scale bars: 1 mm.
</paragraph>
</caption>
<caption id="DF5F665D4E34FFC6D7CA07F22DCCF858" ID-DOI="http://doi.org/10.5281/zenodo.4412728" ID-Zenodo-Dep="4412728" httpUri="https://zenodo.org/record/4412728/files/figure.png" pageId="3" pageNumber="579" startId="3.[151,250,1962,1986]" targetBox="[182,1406,181,1938]" targetPageId="3">
<paragraph id="8B9F36D54E34FFC6D7CA07F22DCCF858" blockId="3.[151,1436,1962,2023]" pageId="3" pageNumber="579">
<emphasis id="B954EAC74E34FFC6D7CA07F22F29F87C" bold="true" box="[151,269,1962,1987]" pageId="3" pageNumber="579">FIGURE 2</emphasis>
.
<taxonomicName id="4C204D564E34FFC6D64407F32FC6F87C" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[281,482,1963,1987]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="579" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E34FFC6D64407F32FC6F87C" box="[281,482,1963,1987]" italics="true" pageId="3" pageNumber="579">Petrolisthes lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E34FFC6D6B507F32C1CF87C" bold="true" box="[488,568,1963,1987]" pageId="3" pageNumber="579">
<taxonomicNameLabel id="A26757BC4E34FFC6D6B507F32C1CF87C" box="[488,568,1963,1987]" pageId="3" pageNumber="579" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker.
</paragraph>
</caption>
<caption id="DF5F665D4E33FFC1D7CA03812F3CFBAA" ID-DOI="http://doi.org/10.5281/zenodo.4412730" ID-Zenodo-Dep="4412730" httpUri="https://zenodo.org/record/4412730/files/figure.png" pageId="4" pageNumber="580" startId="4.[151,250,985,1009]" targetBox="[151,1435,181,961]" targetPageId="4">
<paragraph id="8B9F36D54E33FFC1D7CA03812F3CFBAA" blockId="4.[151,1436,985,1045]" pageId="4" pageNumber="580">
<emphasis id="B954EAC74E33FFC1D7CA03812F2AFC4E" bold="true" box="[151,270,985,1009]" pageId="4" pageNumber="580">FIGURE 3</emphasis>
.
<taxonomicName id="4C204D564E33FFC1D64103812FC2FC4E" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[284,486,985,1009]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="580" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E33FFC1D64103812FC2FC4E" box="[284,486,985,1009]" italics="true" pageId="4" pageNumber="580">Petrolisthes lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E33FFC1D6B003812C1BFC4E" bold="true" box="[493,575,985,1009]" pageId="4" pageNumber="580">
<taxonomicNameLabel id="A26757BC4E33FFC1D6B003812C1BFC4E" box="[493,575,985,1009]" pageId="4" pageNumber="580" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker.
</paragraph>
</caption>
<paragraph id="8B9F36D54E33FFC1D79A041B2D18FB3E" blockId="4.[151,1437,1091,2018]" pageId="4" pageNumber="580">Antennular basal article unarmed; anterior lateral part slightly concave. Articles 2 and 3 unarmed, subcylindrical, equal in length to each other; article 3 widening distally.</paragraph>
<paragraph id="8B9F36D54E33FFC1D79A04D32C17FB51" blockId="4.[151,1437,1091,2018]" pageId="4" pageNumber="580">Antennal articles unarmed, slightly granulate. Article 1 acutely pointed anteriorly, with granular margins. Article 2 about half-length of article 3. Article 4 about 1/3 of total length of article 3. Flagellum long, composed of short units with inconspicuous setae.</paragraph>
<paragraph id="8B9F36D54E33FFC1D79A04A02A47FAE6" blockId="4.[151,1437,1091,2018]" pageId="4" pageNumber="580">Mxp 3 with ventral surface rugose. Ischium with lateral lobe narrow, distinct, separated by deep V-shaped incision. Merus as long as ischium; flexor lobe broad, with rounded tip. Exopod lanceolate, fringed with long plumose setae on lateral margin, reaching about half length of merus; flagellum well developed.</paragraph>
<paragraph id="8B9F36D54E33FFC1D79A053C2AF8F98D" blockId="4.[151,1437,1091,2018]" pageId="4" pageNumber="580">P1 equal in size from right to left, subsymmetrical in shape, robust, flattened, granulate dorsally; slightly rugose ventrally. Merus about 1/4 length of carpus; dorsomesial lobe well developed, rounded, up-turned, granular; ventral distomesial spine well developed. Carpus as long as palm, with 4 or 5 blunt, marginally granulate teeth on proximal half of anterodorsal margin; posterodorsal margin somewhat uneven, unarmed. Both palm and carpus with dense field of moderately long plumose setae on lateral (outer) surface. Fixed finger about 1/3 total length of entire chela, equal in length to dactylus. Fingertips curved; cutting edges of fingers granulate, with short setae.</paragraph>
<paragraph id="8B9F36D54E33FFC1D79A06632BA6F95A" blockId="4.[151,1437,1091,2018]" pageId="4" pageNumber="580">Walking legs decreasing in size posteriorly from P2 to P4, slightly rugose laterally and mesially, densely fringed with long plumose setae dorsaly and ventrally; margins of P2–P4 meri and carpi unarmed; P2–P3 carpi about halflength of P2–P3 meri; P4 carpus about 2/3 length of P4 merus; P2–P4 carpi nearly equal in length to P2–P4 propodi; P2–P4 propodi with 6 or 7 movable corneous spines ventrally, 4 or 5 medially aligned, 2 on distal margin near articulation with dactylus; P2–P4 dactyli about half-length of P2–P4 propodi, ventral margin with row of 3 spines.</paragraph>
<paragraph id="8B9F36D54E33FFC1D79A06B72B62F8C9" blockId="4.[151,1437,1091,2018]" pageId="4" pageNumber="580">Pleon subrectangular, with dorsal surface smooth; lateral margins with dense row of long plumose setae. Configuration of male pleopods unknown. Female pleopods well developed on somites 4 and 5. Uropods broad, reaching distal margin of telson; lateral margins with densely inserted, long plumose setae. Telson smooth on external surface, wider than long, subdivided into 7 plates; lateral margins densely fringed with long plumose setae.</paragraph>
<paragraph id="8B9F36D54E33FFC0D79A07272DFCFEDA" blockId="4.[151,1437,1091,2018]" lastBlockId="5.[151,1436,151,357]" lastPageId="5" lastPageNumber="581" pageId="4" pageNumber="580">
<emphasis id="B954EAC74E33FFC1D79A07272F5FF825" bold="true" box="[199,379,1919,1946]" pageId="4" pageNumber="580">Colour pattern</emphasis>
. Carapace mostly brown or pale brown, symmetrically marbled with brown-greenish and pale orange areas, with whitish or pale orange striation; frontal and hepatic areas darker red-brown; cardiac region pale orange with two 2 whitish patches; median posterior region with more or less marked white patches; branchiostegite pale yellow or whitish. Antennule pale brownish, with brown or grey area subdistally. Antenna pale orange-brown; flagellum red-brown. Mxp 3 brown-greenish to greyish dorsally (on interior surface), yellowish ventrally (on exterior surface). P1 dorsal surface pale brown with pinkish or orange tinge dorsally, with some brown spots on merus, carpus and palm, distolateral area of palm and fingers darker brown (more in
<typeStatus id="549B88774E32FFC0D345015B2A5AFEA1" box="[1048,1150,259,286]" pageId="5" pageNumber="581" type="holotype">holotype</typeStatus>
), except for whitish tips; ventral surface pale orange or pale pink. P2–5 whitish with pale yellow or brownish tinge or pale brown, partly concealed brown by setae. Pleon pale pink on external surface (
<figureCitation id="131B2A504E32FFC0D40201132D8AFEDA" box="[863,942,331,357]" captionStart="FIGURE 2" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[182,1406,181,1938]" captionTargetId="figure-23@3.[182,1406,181,1938]" captionTargetPageId="3" captionText="FIGURE 2. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412728" httpUri="https://zenodo.org/record/4412728/files/figure.png" pageId="5" pageNumber="581">Figs. 2</figureCitation>
,
<figureCitation id="131B2A504E32FFC0D4E701132DEEFEDA" box="[954,970,331,357]" captionStart="FIGURE 3" captionStartId="4.[151,250,985,1009]" captionTargetBox="[151,1435,181,961]" captionTargetId="figure-18@4.[151,1435,181,961]" captionTargetPageId="4" captionText="FIGURE 3. Petrolisthes lazarus sp. nov., female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412730" httpUri="https://zenodo.org/record/4412730/files/figure.png" pageId="5" pageNumber="581">3</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF5F665D4E32FFC0D7CA07F82A9EF862" ID-DOI="http://doi.org/10.5281/zenodo.4412732" ID-Zenodo-Dep="4412732" httpUri="https://zenodo.org/record/4412732/files/figure.png" pageId="5" pageNumber="581" startId="5.[151,250,1952,1976]" targetBox="[151,1435,396,1929]" targetPageId="5">
<paragraph id="8B9F36D54E32FFC0D7CA07F82A9EF862" blockId="5.[151,1437,1952,2013]" pageId="5" pageNumber="581">
<emphasis id="B954EAC74E32FFC0D7CA07F82F29F807" bold="true" box="[151,269,1952,1976]" pageId="5" pageNumber="581">FIGURE 4</emphasis>
. Extensive rocky intertidal at Río Mar, Pacific coast of Panama, collection locality of paratype of
<taxonomicName id="4C204D564E32FFC0D3B607F92E93F863" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="581" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E32FFC0D3B607F92E93F863" italics="true" pageId="5" pageNumber="581">Petrolisthes lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E32FFC0D7E3079D2F2AF862" bold="true" box="[190,270,1989,2013]" pageId="5" pageNumber="581">
<taxonomicNameLabel id="A26757BC4E32FFC0D7E3079D2F2AF862" box="[190,270,1989,2013]" pageId="5" pageNumber="581" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
: (A) beach view, showing coastal development; (B) ocean view. Photographs by I.N. Marin.
</paragraph>
</caption>
<caption id="DF5F665D4E31FFC3D7CA07FF2AB1F85B" ID-DOI="http://doi.org/10.5281/zenodo.4412734" ID-Zenodo-Dep="4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="6" pageNumber="582" startId="6.[151,250,1959,1983]" targetBox="[175,1412,181,1935]" targetPageId="6">
<paragraph id="8B9F36D54E31FFC3D7CA07FF2AB1F85B" blockId="6.[151,1437,1959,2020]" pageId="6" pageNumber="582">
<emphasis id="B954EAC74E31FFC3D7CA07FF2F2AF87F" bold="true" box="[151,270,1959,1984]" pageId="6" pageNumber="582">FIGURE 5</emphasis>
.
<taxonomicName id="4C204D564E31FFC3D64107FF2CE1F87F" authority="Lockington, 1878" authorityName="Lockington" authorityYear="1878" box="[284,709,1959,1984]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="582" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E31FFC3D64107FF2C22F800" box="[284,518,1959,1983]" italics="true" pageId="6" pageNumber="582">Petrolisthes crenulatus</emphasis>
<bibRefCitation id="EFB14B244E31FFC3D55007FF2CE1F87F" author="Lockington, W. N." box="[525,709,1959,1984]" pageId="6" pageNumber="582" pagination="394 - 406" refId="ref5372" refString="Lockington, W. N. (1878) Remarks upon the Porcellanidea of the West Coast of North America. Annals of the Magazine of Natural History, Series 5, 2, 394 - 406. https: // doi. org / 10.1080 / 00222937808682444" type="journal article" year="1878">Lockington, 1878</bibRefCitation>
</taxonomicName>
, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno.
</paragraph>
</caption>
<caption id="DF5F665D4E30FFC2D7CA05952ECBF992" ID-DOI="http://doi.org/10.5281/zenodo.4412736" ID-Zenodo-Dep="4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="7" pageNumber="583" startId="7.[151,250,1485,1509]" targetBox="[151,1435,181,1461]" targetPageId="7">
<paragraph id="8B9F36D54E30FFC2D7CA05952ECBF992" blockId="7.[151,1436,1485,1581]" pageId="7" pageNumber="583">
<emphasis id="B954EAC74E30FFC2D7CA05952F2AFA5A" bold="true" box="[151,270,1485,1509]" pageId="7" pageNumber="583">FIGURE 6</emphasis>
.
<taxonomicName id="4C204D564E30FFC2D64105952CE2FA5A" authority="Lockington, 1878" authorityName="Lockington" authorityYear="1878" box="[284,710,1485,1509]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E30FFC2D64105952C22FA5A" box="[284,518,1485,1509]" italics="true" pageId="7" pageNumber="583">Petrolisthes crenulatus</emphasis>
<bibRefCitation id="EFB14B244E30FFC2D55005952CE2FA5A" author="Lockington, W. N." box="[525,710,1485,1509]" pageId="7" pageNumber="583" pagination="394 - 406" refId="ref5372" refString="Lockington, W. N. (1878) Remarks upon the Porcellanidea of the West Coast of North America. Annals of the Magazine of Natural History, Series 5, 2, 394 - 406. https: // doi. org / 10.1080 / 00222937808682444" type="journal article" year="1878">Lockington, 1878</bibRefCitation>
</taxonomicName>
: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno.
</paragraph>
</caption>
<subSubSection id="C33A655E4E30FFC2D79A06032BB9F907" pageId="7" pageNumber="583" type="etymology">
<paragraph id="8B9F36D54E30FFC2D79A06032BB9F907" blockId="7.[151,1437,1627,2009]" pageId="7" pageNumber="583">
<emphasis id="B954EAC74E30FFC2D79A06032F6CF9CA" bold="true" box="[199,328,1627,1653]" pageId="7" pageNumber="583">Etymology</emphasis>
. The new species is named after our colleague and friend, Dr. Juan Felipe Lazarus-Agudelo (Universidad del Valle, Cali,
<collectingCountry id="F33776454E30FFC2D6DB06252FD2F927" box="[390,502,1661,1688]" name="Colombia" pageId="7" pageNumber="583">Colombia</collectingCountry>
) for his invaluable help in the field and for generously sharing his excellent knowledge of the ecology and taxonomy of porcelain crabs, one of his favourite study groups. Used as a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C33A655E4E30FFC2D79A069A2FE2F8BE" pageId="7" pageNumber="583" type="distribution">
<paragraph id="8B9F36D54E30FFC2D79A069A2FE2F8BE" blockId="7.[151,1437,1627,2009]" pageId="7" pageNumber="583">
<emphasis id="B954EAC74E30FFC2D79A069A2F7DF963" bold="true" box="[199,345,1730,1756]" pageId="7" pageNumber="583">Distribution</emphasis>
. Currently known only from two localities on the Pacific coast of
<collectingCountry id="F33776454E30FFC2D361069B2ABCF962" box="[1084,1176,1731,1757]" name="Panama" pageId="7" pageNumber="583">Panama</collectingCountry>
, viz. Las Perlas Islands (
<collectingCountry id="F33776454E30FFC2D7C206BE2EEDF8BF" box="[159,201,1766,1792]" name="Iceland" pageId="7" pageNumber="583">Isla</collectingCountry>
Saboga) and Río Mar.
</paragraph>
</subSubSection>
<subSubSection id="C33A655E4E30FFC2D79A07532F89F8D2" pageId="7" pageNumber="583" type="biology_ecology">
<paragraph id="8B9F36D54E30FFC2D79A07532F89F8D2" blockId="7.[151,1437,1627,2009]" pageId="7" pageNumber="583">
<emphasis id="B954EAC74E30FFC2D79A07532F02F89A" bold="true" box="[199,294,1803,1829]" pageId="7" pageNumber="583">Ecology</emphasis>
. Both
<typeStatus id="549B88774E30FFC2D62C07532F85F89A" box="[369,417,1803,1829]" pageId="7" pageNumber="583">type</typeStatus>
specimens of
<taxonomicName id="4C204D564E30FFC2D51C07532C8BF89B" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[577,687,1802,1828]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E30FFC2D51C07532C8BF89B" box="[577,687,1802,1828]" italics="true" pageId="7" pageNumber="583">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E30FFC2D5EB07532D2BF89A" bold="true" box="[694,783,1803,1829]" pageId="7" pageNumber="583">
<taxonomicNameLabel id="A26757BC4E30FFC2D5EB07532D2BF89A" box="[694,783,1803,1829]" pageId="7" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
were collected under large rocks on sand-rock bottoms, in the lower rocky intertidal area (Río Mar,
<figureCitation id="131B2A504E30FFC2D53807762C83F8F6" box="[613,679,1838,1865]" captionStart="FIGURE 4" captionStartId="5.[151,250,1952,1976]" captionTargetBox="[151,1435,396,1929]" captionTargetId="figure-124@5.[151,1435,381,1929]" captionTargetPageId="5" captionText="FIGURE 4. Extensive rocky intertidal at Río Mar, Pacific coast of Panama, collection locality of paratype of Petrolisthes lazarus sp. nov.: (A) beach view, showing coastal development; (B) ocean view. Photographs by I.N. Marin." figureDoi="http://doi.org/10.5281/zenodo.4412732" httpUri="https://zenodo.org/record/4412732/files/figure.png" pageId="7" pageNumber="583">Fig. 4</figureCitation>
) or in the shallow sandy-rocky subtidal area, less than
<quantity id="4CD89B304E30FFC2D24D07762B74F8F6" box="[1296,1360,1838,1865]" metricMagnitude="0" metricUnit="m" metricValue="1.5" pageId="7" pageNumber="583" unit="m" value="1.5">1.5 m</quantity>
at low tide (Las Perlas Islands).
</paragraph>
</subSubSection>
<subSubSection id="C33A655E4E30FFCED79A072F2FE6FD3A" lastPageId="11" lastPageNumber="587" pageId="7" pageNumber="583" type="discussion">
<paragraph id="8B9F36D54E30FFCDD79A072F2F5EFE36" blockId="7.[151,1437,1627,2009]" lastBlockId="8.[151,1436,151,393]" lastPageId="8" lastPageNumber="584" pageId="7" pageNumber="583">
<emphasis id="B954EAC74E30FFC2D79A072F2F10F82E" bold="true" box="[199,308,1911,1937]" pageId="7" pageNumber="583">Remarks</emphasis>
.
<taxonomicName id="4C204D564E30FFC2D61E072E2C05F82F" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[323,545,1910,1936]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E30FFC2D61E072E2C05F82F" box="[323,545,1910,1936]" italics="true" pageId="7" pageNumber="583">Petrolisthes lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E30FFC2D574072F2CA7F82E" bold="true" box="[553,643,1911,1937]" pageId="7" pageNumber="583">
<taxonomicNameLabel id="A26757BC4E30FFC2D574072F2CA7F82E" box="[553,643,1911,1937]" pageId="7" pageNumber="583" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is morphologically closest to
<taxonomicName id="4C204D564E30FFC2D48A072F2A4CF82F" authorityName="Lockington" authorityYear="1878" box="[983,1128,1910,1936]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E30FFC2D48A072F2A4CF82F" box="[983,1128,1910,1936]" italics="true" pageId="7" pageNumber="583">P. crenulatus</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A504E30FFC2D325072E2AECF82E" box="[1144,1224,1910,1937]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="7" pageNumber="583">Figs. 5</figureCitation>
,
<figureCitation id="131B2A504E30FFC2D388072E2AC1F82E" box="[1237,1253,1910,1937]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="7" pageNumber="583">6</figureCitation>
), and to a lesser degree also to
<taxonomicName id="4C204D564E30FFC2D61F07C32F85F80B" baseAuthorityName="Glassell" baseAuthorityYear="1936" box="[322,417,1946,1972]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="B954EAC74E30FFC2D61F07C32F85F80B" box="[322,417,1946,1972]" italics="true" pageId="7" pageNumber="583">P. lewisi</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A504E30FFC2D6EE07C22C14F80A" box="[435,560,1946,1973]" captionStart="FIGURE 8" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1920]" captionTargetId="figure-23@9.[164,1423,181,1920]" captionTargetPageId="9" captionText="FIGURE 8. Petrolisthes lewisi (Glassell, 1936) [A, B] and Petrolisthes ortmanni Nobili, 1901 [C]: (A) male from Los Negritos, off Bahía Málaga, Pacific coast of Colombia (CERBMcr UV 2019-103), dorsal view; (B) ovigerous female from the same locality (CERBMcr UV 2019-102); male from the same locality (CERBMcr UV 2019-102). Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412740" httpUri="https://zenodo.org/record/4412740/files/figure.png" pageId="7" pageNumber="583">Fig. 8A, B</figureCitation>
) and
<taxonomicName id="4C204D564E30FFC2D52807C32CDEF80B" authorityName="Nobili" authorityYear="1901" box="[629,762,1947,1972]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E30FFC2D52807C32CDEF80B" box="[629,762,1947,1972]" italics="true" pageId="7" pageNumber="583">P. ortmanni</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A504E30FFC2D45607C22D43F80A" box="[779,871,1946,1973]" captionStart="FIGURE 8" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1920]" captionTargetId="figure-23@9.[164,1423,181,1920]" captionTargetPageId="9" captionText="FIGURE 8. Petrolisthes lewisi (Glassell, 1936) [A, B] and Petrolisthes ortmanni Nobili, 1901 [C]: (A) male from Los Negritos, off Bahía Málaga, Pacific coast of Colombia (CERBMcr UV 2019-103), dorsal view; (B) ovigerous female from the same locality (CERBMcr UV 2019-102); male from the same locality (CERBMcr UV 2019-102). Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412740" httpUri="https://zenodo.org/record/4412740/files/figure.png" pageId="7" pageNumber="583">Fig. 8C</figureCitation>
). As correctly pointed out by
<bibRefCitation id="EFB14B244E30FFC2D39F07C22F81F866" author="Garcia-Madrigal, M. S. &amp; Andreu-Sanchez, L. I." pageId="7" pageNumber="583" pagination="23 - 54" refId="ref4779" refString="Garcia-Madrigal, M. S. &amp; Andreu-Sanchez, L. I. (2009) Los cangrejos porcelanidos (Decapoda: Anomura) del Pacifico sur de Mexico, incluyendo una lista y clave de identificacion para todas las especies del Pacifico oriental tropical. Ciencia y Mar, 13 (39), 23 - 54." type="journal article" year="2009">García-Madrigal &amp; Andréu-Sanchez (2009)</bibRefCitation>
,
<taxonomicName id="4C204D564E30FFC2D6ED07E72C64F867" authorityName="Lockington" authorityYear="1878" box="[432,576,1982,2008]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E30FFC2D6ED07E72C64F867" box="[432,576,1982,2008]" italics="true" pageId="7" pageNumber="583">P. crenulatus</emphasis>
</taxonomicName>
can be most easily separated from
<taxonomicName id="4C204D564E30FFC2D49A07E72A6DF867" authorityName="Nobili" authorityYear="1901" box="[967,1097,1983,2008]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="583" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E30FFC2D49A07E72A6DF867" box="[967,1097,1983,2008]" italics="true" pageId="7" pageNumber="583">P. ortmanni</emphasis>
</taxonomicName>
by the presence of dense seta- tion on the P1 carpus, in addition to the very clear differences in the proportions of this article, as mentioned and illustrated by
<bibRefCitation id="EFB14B244E3FFFCDD66F00E32F99FF69" author="Haig, J." box="[306,445,187,214]" pageId="8" pageNumber="584" pagination="1 - 440" refId="ref5082" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960)</bibRefCitation>
. In other words, in
<emphasis id="B954EAC74E3FFFCDD5CA00E32D0BFF6A" box="[663,815,187,213]" italics="true" pageId="8" pageNumber="584">
<taxonomicName id="4C204D564E3FFFCDD5CA00E32D0FFF6A" authorityName="Lockington" authorityYear="1878" box="[663,811,187,213]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">P. crenulatus</taxonomicName>
,
</emphasis>
the carpus is relatively slender and has a dense field of setae along its lateral (outer) surface (
<figureCitation id="131B2A504E3FFFCDD51900872CADFF46" box="[580,649,223,249]" captionStart="FIGURE 4" captionStartId="5.[151,250,1952,1976]" captionTargetBox="[151,1435,396,1929]" captionTargetId="figure-124@5.[151,1435,381,1929]" captionTargetPageId="5" captionText="FIGURE 4. Extensive rocky intertidal at Río Mar, Pacific coast of Panama, collection locality of paratype of Petrolisthes lazarus sp. nov.: (A) beach view, showing coastal development; (B) ocean view. Photographs by I.N. Marin." figureDoi="http://doi.org/10.5281/zenodo.4412732" httpUri="https://zenodo.org/record/4412732/files/figure.png" pageId="8" pageNumber="584">Fig. 4</figureCitation>
,
<figureCitation id="131B2A504E3FFFCDD5CB00862C81FF46" box="[662,677,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="8" pageNumber="584">5</figureCitation>
), whereas in
<taxonomicName id="4C204D564E3FFFCDD46200872DE6FF47" authorityName="Nobili" authorityYear="1901" box="[831,962,223,248]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E3FFFCDD46200872DE6FF47" box="[831,962,223,248]" italics="true" pageId="8" pageNumber="584">P. ortmanni</emphasis>
</taxonomicName>
, the carpus is noticeably more robust and without dense setation laterally (
<figureCitation id="131B2A504E3FFFCDD55B015B2C7BFEA2" box="[518,607,259,285]" captionStart="FIGURE 8" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1920]" captionTargetId="figure-23@9.[164,1423,181,1920]" captionTargetPageId="9" captionText="FIGURE 8. Petrolisthes lewisi (Glassell, 1936) [A, B] and Petrolisthes ortmanni Nobili, 1901 [C]: (A) male from Los Negritos, off Bahía Málaga, Pacific coast of Colombia (CERBMcr UV 2019-103), dorsal view; (B) ovigerous female from the same locality (CERBMcr UV 2019-102); male from the same locality (CERBMcr UV 2019-102). Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412740" httpUri="https://zenodo.org/record/4412740/files/figure.png" pageId="8" pageNumber="584">Fig. 8C</figureCitation>
). The same is true for
<taxonomicName id="4C204D564E3FFFCDD401015B2D9DFEA2" baseAuthorityName="Glassell" baseAuthorityYear="1936" box="[860,953,259,285]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="B954EAC74E3FFFCDD401015B2D9DFEA2" box="[860,953,259,285]" italics="true" pageId="8" pageNumber="584">P. lewisi</emphasis>
</taxonomicName>
, in which the P1 carpus is similarly robust and lacking setation, thereby clearly differing from the P1 carpi of
<taxonomicName id="4C204D564E3FFFCDD4DB017F2DD7FEFE" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[902,1011,295,321]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3FFFCDD4DB017F2DD7FEFE" box="[902,1011,295,321]" italics="true" pageId="8" pageNumber="584">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3FFFCDD4A6017F2A71FEFE" bold="true" box="[1019,1109,295,321]" pageId="8" pageNumber="584">
<taxonomicNameLabel id="A26757BC4E3FFFCDD4A6017F2A71FEFE" box="[1019,1109,295,321]" pageId="8" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C204D564E3FFFCDD3D6017F2B38FEFE" authorityName="Lockington" authorityYear="1878" box="[1163,1308,295,321]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD3D6017F2B38FEFE" box="[1163,1308,295,321]" italics="true" pageId="8" pageNumber="584">P. crenulatus</emphasis>
</taxonomicName>
. These and several other morphological characters that might be useful in the separation of these four eastern Pacific species are listed in
<tableCitation id="C6A2036E4E3FFFCDD67D01372F50FE36" box="[288,372,367,393]" captionStart="TABLE 1" captionStartId="10.[151,239,517,543]" captionTargetPageId="10" captionText="TABLE 1. Characters (morphology and colour pattern) that may be used in the separation of four putatively related species of Petrolisthes from the eastern Pacific, including the" httpUri="http://table.plazi.org/id/DF5F665D4E3DFFCFD7CA025D29D3FDA0" pageId="8" pageNumber="584" tableUuid="DF5F665D4E3DFFCFD7CA025D29D3FDA0">Table 1</tableCitation>
.
</paragraph>
<caption id="DF5F665D4E3FFFCDD7CA06782A77F93F" ID-DOI="http://doi.org/10.5281/zenodo.4412738" ID-Zenodo-Dep="4412738" httpUri="https://zenodo.org/record/4412738/files/figure.png" pageId="8" pageNumber="584" startId="8.[151,250,1568,1592]" targetBox="[151,1435,417,1545]" targetPageId="8">
<paragraph id="8B9F36D54E3FFFCDD7CA06782A77F93F" blockId="8.[151,1437,1568,1664]" pageId="8" pageNumber="584">
<emphasis id="B954EAC74E3FFFCDD7CA06782F34F987" bold="true" box="[151,272,1568,1592]" pageId="8" pageNumber="584">FIGURE 7</emphasis>
. Mixed sand-rock intertidal at La Cruz de Huanacaxtle, Nayarit, Pacific coast of Mexico, collection locality of
<taxonomicName id="4C204D564E3FFFCDD7CA061C2C74F9E3" authority="(Lockington, 1878)" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[151,592,1604,1628]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD7CA061C2FA4F9E3" box="[151,384,1604,1628]" italics="true" pageId="8" pageNumber="584">Petrolisthes crenulatus</emphasis>
(
<bibRefCitation id="EFB14B244E3FFFCDD6D0061C2C6CF9E3" author="Lockington, W. N." box="[397,584,1604,1628]" pageId="8" pageNumber="584" pagination="394 - 406" refId="ref5372" refString="Lockington, W. N. (1878) Remarks upon the Porcellanidea of the West Coast of North America. Annals of the Magazine of Natural History, Series 5, 2, 394 - 406. https: // doi. org / 10.1080 / 00222937808682444" type="journal article" year="1878">Lockington, 1878</bibRefCitation>
)
</taxonomicName>
: (A) general ocean view; (B) detail of rocks and sand; (C) one of the collected porcelain crabs
<emphasis id="B954EAC74E3FFFCDD64B06312F71F93F" box="[278,341,1641,1664]" italics="true" pageId="8" pageNumber="584">in situ</emphasis>
, exposed after flipping a large rock. Photographs by L.D. Santana-Moreno.
</paragraph>
</caption>
<paragraph id="8B9F36D54E3FFFCFD79A06F62FF1FDFC" blockId="8.[151,1437,1710,2025]" lastBlockId="10.[151,2039,517,579]" lastPageId="10" lastPageNumber="586" pageId="8" pageNumber="584">
<taxonomicName id="4C204D564E3FFFCDD79A06F62F83F977" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[199,423,1710,1736]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3FFFCDD79A06F62F83F977" box="[199,423,1710,1736]" italics="true" pageId="8" pageNumber="584">Petrolisthes lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3FFFCDD6EC06F62C29F977" bold="true" box="[433,525,1710,1736]" pageId="8" pageNumber="584">
<taxonomicNameLabel id="A26757BC4E3FFFCDD6EC06F62C29F977" box="[433,525,1710,1736]" pageId="8" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is distinguishable from
<taxonomicName id="4C204D564E3FFFCDD47906F72D93F977" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[804,951,1710,1736]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD47906F72D93F977" box="[804,951,1710,1736]" italics="true" pageId="8" pageNumber="584">P. crenulatus</emphasis>
</taxonomicName>
by the relatively wider carapace, with the frontal width being about 1/3 of the maximal carapace width (
<emphasis id="B954EAC74E3FFFCDD434068B2DA3F953" box="[873,903,1747,1772]" italics="true" pageId="8" pageNumber="584">vs.</emphasis>
approximately 1/
<quantity id="4CD89B304E3FFFCDD30D068A2A5BF953" box="[1104,1151,1746,1772]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="584" unit="in" value="2.0">2 in</quantity>
<emphasis id="B954EAC74E3FFFCDD3D4068B2B01F953" box="[1161,1317,1746,1772]" italics="true" pageId="8" pageNumber="584">
<taxonomicName id="4C204D564E3FFFCDD3D4068B2B39F953" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[1161,1309,1746,1772]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">P. crenulatus</taxonomicName>
)
</emphasis>
; the more strongly protruding frontal lateral lobes; and both the palm and the carpus of P1 fringed with dense plumose setae laterally (
<emphasis id="B954EAC74E3FFFCDD65907432F3FF88B" box="[260,283,1819,1844]" italics="true" pageId="8" pageNumber="584">vs</emphasis>
. only P1 carpus fringed with plumose setae laterally in
<taxonomicName id="4C204D564E3FFFCDD4D307432A3BF88B" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[910,1055,1818,1844]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD4D307432A3BF88B" box="[910,1055,1818,1844]" italics="true" pageId="8" pageNumber="584">P. crenulatus</emphasis>
</taxonomicName>
). This last character, i.e. the presence of dense field of plumose setae on the lateral (outer) surface of the palm (as well as of the carpus) is the most obvious one, separating
<taxonomicName id="4C204D564E3FFFCDD6F6073B2C3DF8C3" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[427,537,1890,1916]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3FFFCDD6F6073B2C3DF8C3" box="[427,537,1890,1916]" italics="true" pageId="8" pageNumber="584">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3FFFCDD57F073A2C59F8C3" bold="true" box="[546,637,1890,1916]" pageId="8" pageNumber="584">
<taxonomicNameLabel id="A26757BC4E3FFFCDD57F073A2C59F8C3" box="[546,637,1890,1916]" pageId="8" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
at once from
<taxonomicName id="4C204D564E3FFFCDD441073B2D8AF8C3" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[796,942,1890,1916]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD441073B2D8AF8C3" box="[796,942,1890,1916]" italics="true" pageId="8" pageNumber="584">P. crenulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D564E3FFFCDD4E0073B2A65F8C3" authorityName="Nobili" authorityYear="1901" box="[957,1089,1891,1916]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E3FFFCDD4E0073B2A65F8C3" box="[957,1089,1891,1916]" italics="true" pageId="8" pageNumber="584">P. ortmanni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C204D564E3FFFCDD324073B2AF3F8C3" baseAuthorityName="Glassell" baseAuthorityYear="1936" box="[1145,1239,1890,1916]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="B954EAC74E3FFFCDD324073B2AF3F8C3" box="[1145,1239,1890,1916]" italics="true" pageId="8" pageNumber="584">P. lewisi</emphasis>
</taxonomicName>
(cf.
<figureCitation id="131B2A504E3FFFCDD253073A2B79F8C3" box="[1294,1373,1890,1916]" captionStart="FIGURE 2" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[182,1406,181,1938]" captionTargetId="figure-23@3.[182,1406,181,1938]" captionTargetPageId="3" captionText="FIGURE 2. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412728" httpUri="https://zenodo.org/record/4412728/files/figure.png" pageId="8" pageNumber="584">Figs. 2</figureCitation>
,
<figureCitation id="131B2A504E3FFFCDD237073A2B5DF8C3" box="[1386,1401,1890,1916]" captionStart="FIGURE 3" captionStartId="4.[151,250,985,1009]" captionTargetBox="[151,1435,181,961]" captionTargetId="figure-18@4.[151,1435,181,961]" captionTargetPageId="4" captionText="FIGURE 3. Petrolisthes lazarus sp. nov., female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412730" httpUri="https://zenodo.org/record/4412730/files/figure.png" pageId="8" pageNumber="584">3</figureCitation>
,
<figureCitation id="131B2A504E3FFFCDD2DA073A2BB3F8C2" box="[1415,1431,1890,1917]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="8" pageNumber="584">5</figureCitation>
,
<figureCitation id="131B2A504E3FFFCDD7CA07DE2E83F81F" box="[151,167,1926,1952]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="8" pageNumber="584">6</figureCitation>
,
<figureCitation id="131B2A504E3FFFCDD7E907DE2EE0F81F" box="[180,196,1926,1952]" captionStart="FIGURE 8" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1920]" captionTargetId="figure-23@9.[164,1423,181,1920]" captionTargetPageId="9" captionText="FIGURE 8. Petrolisthes lewisi (Glassell, 1936) [A, B] and Petrolisthes ortmanni Nobili, 1901 [C]: (A) male from Los Negritos, off Bahía Málaga, Pacific coast of Colombia (CERBMcr UV 2019-103), dorsal view; (B) ovigerous female from the same locality (CERBMcr UV 2019-102); male from the same locality (CERBMcr UV 2019-102). Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412740" httpUri="https://zenodo.org/record/4412740/files/figure.png" pageId="8" pageNumber="584">8</figureCitation>
). In addition,
<taxonomicName id="4C204D564E3FFFCDD63907DF2FF6F81F" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[356,466,1926,1952]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3FFFCDD63907DF2FF6F81F" box="[356,466,1926,1952]" italics="true" pageId="8" pageNumber="584">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3FFFCDD68607DE2C11F81F" bold="true" box="[475,565,1926,1952]" pageId="8" pageNumber="584">
<taxonomicNameLabel id="A26757BC4E3FFFCDD68607DE2C11F81F" box="[475,565,1926,1952]" pageId="8" pageNumber="584" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be separated from
<taxonomicName id="4C204D564E3FFFCDD46307DF2DEBF81F" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[830,975,1926,1952]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD46307DF2DEBF81F" box="[830,975,1926,1952]" italics="true" pageId="8" pageNumber="584">P. crenulatus</emphasis>
</taxonomicName>
by the much less contrasting colour pattern. Most importantly,
<taxonomicName id="4C204D564E3FFFCDD6FA07F32C1EF87B" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[423,570,1962,1988]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="584" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3FFFCDD6FA07F32C1EF87B" box="[423,570,1962,1988]" italics="true" pageId="8" pageNumber="584">P. crenulatus</emphasis>
</taxonomicName>
has a very conspicuous, large, bright white area on the post-cardiac region of carapace, contrasting with the brown-orange colour of the remaining carapace (
<figureCitation id="131B2A504E3FFFCDD36A07962AA3F856" box="[1079,1159,1998,2025]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="8" pageNumber="584">Figs. 5</figureCitation>
,
<figureCitation id="131B2A504E3FFFCDD3CE07962A87F856" box="[1171,1187,1998,2025]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="8" pageNumber="584">6</figureCitation>
). Such a white area is herein described new species.
</paragraph>
<caption id="DF5F665D4E3EFFCCD7CA07CF2B3CF847" ID-DOI="http://doi.org/10.5281/zenodo.4412740" ID-Zenodo-Dep="4412740" httpUri="https://zenodo.org/record/4412740/files/figure.png" pageId="9" pageNumber="585" startId="9.[151,250,1943,1967]" targetBox="[164,1423,181,1920]" targetPageId="9">
<paragraph id="8B9F36D54E3EFFCCD7CA07CF2B3CF847" blockId="9.[151,1437,1943,2040]" pageId="9" pageNumber="585">
<emphasis id="B954EAC74E3EFFCCD7CA07CF2F28F810" bold="true" box="[151,268,1943,1967]" pageId="9" pageNumber="585">FIGURE 8</emphasis>
.
<taxonomicName id="4C204D564E3EFFCCD64A07C02C52F80F" authority="(Glassell, 1936)" baseAuthorityName="Glassell" baseAuthorityYear="1936" box="[279,630,1944,1968]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="585" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="B954EAC74E3EFFCCD64A07C02FF4F80F" box="[279,464,1944,1968]" italics="true" pageId="9" pageNumber="585">Petrolisthes lewisi</emphasis>
(
<bibRefCitation id="EFB14B244E3EFFCCD68007C02C4BF80F" author="Glassell, S. A." box="[477,623,1944,1968]" pageId="9" pageNumber="585" pagination="227 - 304" refId="ref4838" refString="Glassell, S. A. (1936) New porcellanids and pinnotherids from tropical North American waters. Transactions of the San Diego Society of Natural History, 8, 227 - 304. https: // doi. org / 10.5962 / bhl. part. 14900" type="journal article" year="1936">Glassell, 1936</bibRefCitation>
)
</taxonomicName>
[A, B] and
<taxonomicName id="4C204D564E3EFFCCD5B607C02A6EF80F" authority="Nobili, 1901" authorityName="Nobili" authorityYear="1901" box="[747,1098,1944,1968]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="585" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E3EFFCCD5B607C02DE1F810" box="[747,965,1944,1968]" italics="true" pageId="9" pageNumber="585">Petrolisthes ortmanni</emphasis>
<bibRefCitation id="EFB14B244E3EFFCCD49607C02A6EF80F" author="Nobili, G." box="[971,1098,1944,1968]" pageId="9" pageNumber="585" pagination="58" refId="ref5544" refString="Nobili, G. (1901) Viaggio del Dr. Enrico Festa nella Repubblica dell'Ecuador e regioni vicine, 23: Decapodi e Stomatopodi. Bollettino dei Musei di Zoologia ed Anatomia Comparata della R. Universita di Torino, 16 (415), l- 58." type="journal article" year="1901">Nobili, 1901</bibRefCitation>
</taxonomicName>
[C]: (A) male from Los Negritos, off Bahía Málaga, Pacific coast of Colombia (CERBMcr UV 2019-103), dorsal view; (B) ovigerous female from the same locality (CERBMcr UV 2019-102); male from the same locality (CERBMcr UV 2019-102). Photographs by A. Anker.
</paragraph>
</caption>
<caption id="DF5F665D4E3DFFCFD7CA025D29D3FDA0" ID-Table-UUID="DF5F665D4E3DFFCFD7CA025D29D3FDA0" box="[151,2039,517,543]" httpUri="http://table.plazi.org/id/DF5F665D4E3DFFCFD7CA025D29D3FDA0" pageId="10" pageNumber="586" targetBox="[166,2024,594,1673]" targetIsTable="true" targetPageId="10">
<paragraph id="8B9F36D54E3DFFCFD7CA025D29D3FDA0" blockId="10.[151,2039,517,579]" box="[151,2039,517,543]" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7CA025D2F2EFDA0" bold="true" box="[151,266,517,543]" pageId="10" pageNumber="586">TABLE 1.</emphasis>
Characters (morphology and colour pattern) that may be used in the separation of four putatively related species of
<taxonomicName id="4C204D564E3DFFCFD286025D287DFDA0" authorityName="Stimpson" authorityYear="1858" box="[1499,1625,517,543]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="586" phylum="Arthropoda" rank="genus">
<emphasis id="B954EAC74E3DFFCFD286025D287DFDA0" box="[1499,1625,517,543]" italics="true" pageId="10" pageNumber="586">Petrolisthes</emphasis>
</taxonomicName>
from the eastern Pacific, including the
</paragraph>
</caption>
<paragraph id="8B9F36D54E3DFFCFD7FB020A29B9F9DA" pageId="10" pageNumber="586">
<table id="F920C4754E3D003AD7FB020A29CCF936" box="[166,2024,594,1673]" gridcols="5" gridrows="12" pageId="10" pageNumber="586">
<tr id="351034974E3D003AD7FB020A29CCFD30" box="[166,2024,594,655]" gridrow="0" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB020A2C24FD30" box="[166,512,594,655]" gridcol="0" gridrow="0" pageId="10" pageNumber="586">
Species
<emphasis id="B954EAC74E3DFFCFD7FB022E2F3BFD30" bold="true" box="[166,287,630,655]" pageId="10" pageNumber="586">Characters</emphasis>
</th>
<th id="76C15DEB4E3D003AD569020A2DA1FD30" box="[564,901,594,655]" gridcol="1" gridrow="0" pageId="10" pageNumber="586">
<taxonomicName id="4C204D564E3DFFCFD569020A2CBCFDD4" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[564,664,594,619]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="586" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3DFFCFD569020A2CBCFDD4" box="[564,664,594,619]" italics="true" pageId="10" pageNumber="586">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3DFFCFD5C3020A2CCAFDD4" bold="true" box="[670,750,594,619]" pageId="10" pageNumber="586">
<taxonomicNameLabel id="A26757BC4E3DFFCFD5C3020A2CCAFDD4" box="[670,750,594,619]" pageId="10" pageNumber="586" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</th>
<th id="76C15DEB4E3D003AD4F5020A2ADEFD30" box="[936,1274,594,655]" gridcol="2" gridrow="0" pageId="10" pageNumber="586">
<taxonomicName id="4C204D564E3DFFCFD4F5020A2A0FFDD4" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[936,1067,594,619]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="586" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3DFFCFD4F5020A2A0FFDD4" box="[936,1067,594,619]" italics="true" pageId="10" pageNumber="586">P. crenulatus</emphasis>
</taxonomicName>
</th>
<th id="76C15DEB4E3D003AD240020A2850FD30" box="[1309,1652,594,655]" gridcol="3" gridrow="0" pageId="10" pageNumber="586">
<taxonomicName id="4C204D564E3DFFCFD240020A2BB7FDD4" authorityName="Nobili" authorityYear="1901" box="[1309,1427,594,619]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="586" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E3DFFCFD240020A2BB7FDD4" box="[1309,1427,594,619]" italics="true" pageId="10" pageNumber="586">P. ortmanni</emphasis>
</taxonomicName>
</th>
<th id="76C15DEB4E3D003AD1CF020A29CCFD30" box="[1682,2024,594,655]" gridcol="4" gridrow="0" pageId="10" pageNumber="586">
<taxonomicName id="4C204D564E3DFFCFD1CF020A28C1FDD4" baseAuthorityName="Glassell" baseAuthorityYear="1936" box="[1682,1765,594,619]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="586" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="B954EAC74E3DFFCFD1CF020A28C1FDD4" box="[1682,1765,594,619]" italics="true" pageId="10" pageNumber="586">P. lewisi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="351034974E3D003AD7FB02C629CCFD40" box="[166,2024,670,767]" gridrow="1" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB02C62C24FD40" box="[166,512,670,767]" gridcol="0" gridrow="1" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7FB02C62FE0FD08" bold="true" box="[166,452,670,695]" pageId="10" pageNumber="586">General shape of carapace</emphasis>
</th>
<td id="76C15DEB4E3D003AD56902C62DA1FD40" box="[564,901,670,767]" gridcol="1" gridrow="1" pageId="10" pageNumber="586">Subcircular, wider than long (lateral median region strongly convex)</td>
<td id="76C15DEB4E3D003AD4F502C62ADEFD40" box="[936,1274,670,767]" gridcol="2" gridrow="1" pageId="10" pageNumber="586">Subcircular, wider than long</td>
<td id="76C15DEB4E3D003AD24002C62850FD40" box="[1309,1652,670,767]" gridcol="3" gridrow="1" pageId="10" pageNumber="586">Subcircular, as long as wide</td>
<td id="76C15DEB4E3D003AD1CF02C629CCFD40" box="[1682,2024,670,767]" gridcol="4" gridrow="1" pageId="10" pageNumber="586">Subquadrate, as long as wide</td>
</tr>
<tr id="351034974E3D003AD7FB035629CCFCF4" box="[166,2024,782,843]" gridrow="2" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB03562C24FCF4" box="[166,512,782,843]" gridcol="0" gridrow="2" pageId="10" pageNumber="586">Frontal width / maximal carapace width (cw)</th>
<td id="76C15DEB4E3D003AD56903562DA1FCF4" box="[564,901,782,843]" gridcol="1" gridrow="2" pageId="10" pageNumber="586">Frontal width about 1/3 of maximal cw</td>
<td id="76C15DEB4E3D003AD4F503562ADEFCF4" box="[936,1274,782,843]" gridcol="2" gridrow="2" pageId="10" pageNumber="586">Frontal width close to 1/2 of maximal cw</td>
<td id="76C15DEB4E3D003AD24003562850FCF4" box="[1309,1652,782,843]" gridcol="3" gridrow="2" pageId="10" pageNumber="586">Frontal width about 1/2 of maximal cw</td>
<td id="76C15DEB4E3D003AD1CF035629CCFCF4" box="[1682,2024,782,843]" gridcol="4" gridrow="2" pageId="10" pageNumber="586">Frontal width about 1/2 of maximal cw</td>
</tr>
<tr id="351034974E3D003AD7FB030229CCFCCC" box="[166,2024,858,883]" gridrow="3" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB03022C24FCCC" box="[166,512,858,883]" gridcol="0" gridrow="3" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7FB03022FA5FCCC" bold="true" box="[166,385,858,883]" pageId="10" pageNumber="586">Frontal lateral lobes</emphasis>
</th>
<td id="76C15DEB4E3D003AD56903022DA1FCCC" box="[564,901,858,883]" gridcol="1" gridrow="3" pageId="10" pageNumber="586">Distinctly protruding</td>
<td id="76C15DEB4E3D003AD4F503022ADEFCCC" box="[936,1274,858,883]" gridcol="2" gridrow="3" pageId="10" pageNumber="586">Moderately protruding</td>
<td id="76C15DEB4E3D003AD24003022850FCCC" box="[1309,1652,858,883]" gridcol="3" gridrow="3" pageId="10" pageNumber="586">Feebly protruding</td>
<td id="76C15DEB4E3D003AD1CF030229CCFCCC" box="[1682,2024,858,883]" gridcol="4" gridrow="3" pageId="10" pageNumber="586">Feebly protruding</td>
</tr>
<tr id="351034974E3D003AD7FB03DA29CCFC23" box="[166,2024,898,924]" gridrow="4" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB03DA2C24FC23" box="[166,512,898,924]" gridcol="0" gridrow="4" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7FB03DA2FDCFC24" bold="true" box="[166,504,898,923]" pageId="10" pageNumber="586">Protogastric region of carapace</emphasis>
</th>
<td id="76C15DEB4E3D003AD56903DA2DA1FC23" box="[564,901,898,924]" gridcol="1" gridrow="4" pageId="10" pageNumber="586">Elevated</td>
<td id="76C15DEB4E3D003AD4F503DA2ADEFC23" box="[936,1274,898,924]" gridcol="2" gridrow="4" pageId="10" pageNumber="586">Elevated</td>
<td id="76C15DEB4E3D003AD24003DA2850FC23" box="[1309,1652,898,924]" gridcol="3" gridrow="4" pageId="10" pageNumber="586">Elevated</td>
<td id="76C15DEB4E3D003AD1CF03DA29CCFC23" box="[1682,2024,898,924]" gridcol="4" gridrow="4" pageId="10" pageNumber="586">Not elevated</td>
</tr>
<tr id="351034974E3D003AD7FB039629CCFBB3" box="[166,2024,974,1036]" gridrow="5" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB03962C24FBB3" box="[166,512,974,1036]" gridcol="0" gridrow="5" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7FB03962F51FC58" bold="true" box="[166,373,974,999]" pageId="10" pageNumber="586">Antennal flagellum</emphasis>
</th>
<td id="76C15DEB4E3D003AD56903962DA1FBB3" box="[564,901,974,1036]" gridcol="1" gridrow="5" pageId="10" pageNumber="586">With inconspicuous setae between units</td>
<td id="76C15DEB4E3D003AD4F503962ADEFBB3" box="[936,1274,974,1036]" gridcol="2" gridrow="5" pageId="10" pageNumber="586">With inconspicuous setae between units</td>
<td id="76C15DEB4E3D003AD24003962850FBB3" box="[1309,1652,974,1036]" gridcol="3" gridrow="5" pageId="10" pageNumber="586">With long setae between units</td>
<td id="76C15DEB4E3D003AD1CF039629CCFBB3" box="[1682,2024,974,1036]" gridcol="4" gridrow="5" pageId="10" pageNumber="586">With long setae between units</td>
</tr>
<tr id="351034974E3D003AD7FB044329CCFBE7" box="[166,2024,1051,1112]" gridrow="6" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB04432C24FBE7" box="[166,512,1051,1112]" gridcol="0" gridrow="6" pageId="10" pageNumber="586">P1 carpus</th>
<td id="76C15DEB4E3D003AD56904432DA1FBE7" box="[564,901,1051,1112]" gridcol="1" gridrow="6" pageId="10" pageNumber="586">Elongate, about 1.5 times as long as wide</td>
<td id="76C15DEB4E3D003AD4F504432ADEFBE7" box="[936,1274,1051,1112]" gridcol="2" gridrow="6" pageId="10" pageNumber="586">Elongate, about 1.5 times as long as wide</td>
<td id="76C15DEB4E3D003AD24004432850FBE7" box="[1309,1652,1051,1112]" gridcol="3" gridrow="6" pageId="10" pageNumber="586">Stout, much less than 1.5 times as long as wide</td>
<td id="76C15DEB4E3D003AD1CF044329CCFBE7" box="[1682,2024,1051,1112]" gridcol="4" gridrow="6" pageId="10" pageNumber="586">Stout, less than 1.5 times as long as wide</td>
</tr>
<tr id="351034974E3D003AD7FB043F29CCFB77" box="[166,2024,1127,1224]" gridrow="7" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB043F2C24FB77" box="[166,512,1127,1224]" gridcol="0" gridrow="7" pageId="10" pageNumber="586">Setation on P1</th>
<td id="76C15DEB4E3D003AD569043F2DA1FB77" box="[564,901,1127,1224]" gridcol="1" gridrow="7" pageId="10" pageNumber="586">Palm and carpus fringed with dense plumose setae on lateral (outer) surface</td>
<td id="76C15DEB4E3D003AD4F5043F2ADEFB77" box="[936,1274,1127,1224]" gridcol="2" gridrow="7" pageId="10" pageNumber="586">Carpus only fringed with dense plumose setae on lateral (outer) surface</td>
<td id="76C15DEB4E3D003AD240043F2850FB77" box="[1309,1652,1127,1224]" gridcol="3" gridrow="7" pageId="10" pageNumber="586">Absent</td>
<td id="76C15DEB4E3D003AD1CF043F29CCFB77" box="[1682,2024,1127,1224]" gridcol="4" gridrow="7" pageId="10" pageNumber="586">Absent</td>
</tr>
<tr id="351034974E3D003AD7FB048F29CCFB4E" box="[166,2024,1239,1265]" gridrow="8" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB048F2C24FB4E" box="[166,512,1239,1265]" gridcol="0" gridrow="8" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7FB048F2ECEFB4F" bold="true" box="[166,234,1239,1264]" pageId="10" pageNumber="586">P2–P4</emphasis>
</th>
<td id="76C15DEB4E3D003AD569048F2DA1FB4E" box="[564,901,1239,1265]" gridcol="1" gridrow="8" pageId="10" pageNumber="586">Slender</td>
<td id="76C15DEB4E3D003AD4F5048F2ADEFB4E" box="[936,1274,1239,1265]" gridcol="2" gridrow="8" pageId="10" pageNumber="586">Slender</td>
<td id="76C15DEB4E3D003AD240048F2850FB4E" box="[1309,1652,1239,1265]" gridcol="3" gridrow="8" pageId="10" pageNumber="586">Robust</td>
<td id="76C15DEB4E3D003AD1CF048F29CCFB4E" box="[1682,2024,1239,1265]" gridcol="4" gridrow="8" pageId="10" pageNumber="586">Robust</td>
</tr>
<tr id="351034974E3D003AD7FB055829CCFA3A" box="[166,2024,1280,1413]" gridrow="9" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB05582C24FA3A" box="[166,512,1280,1413]" gridcol="0" gridrow="9" pageId="10" pageNumber="586">Colouration of carapace</th>
<td id="76C15DEB4E3D003AD56905582DA1FA3A" box="[564,901,1280,1413]" gridcol="1" gridrow="9" pageId="10" pageNumber="586">Brown to pale brown, with some darker or paler orange areas; post-cardiac region with inconspicuous whitish areas</td>
<td id="76C15DEB4E3D003AD4F505582ADEFA3A" box="[936,1274,1280,1413]" gridcol="2" gridrow="9" pageId="10" pageNumber="586">Red-brown to orange brown with paler areas; post-cardiac region with large white area</td>
<td id="76C15DEB4E3D003AD24005582850FA3A" box="[1309,1652,1280,1413]" gridcol="3" gridrow="9" pageId="10" pageNumber="586">
Variable from pale yellow to greenish white, sometimes with reddish blotches (
<bibRefCitation id="EFB14B244E3DFFCFD28D05102819FADE" author="Haig, J." box="[1488,1597,1352,1377]" pageId="10" pageNumber="586" pagination="1 - 440" refId="ref5082" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig 1960</bibRefCitation>
)
</td>
<td id="76C15DEB4E3D003AD1CF055829CCFA3A" box="[1682,2024,1280,1413]" gridcol="4" gridrow="9" pageId="10" pageNumber="586">Almost uniform dark brown, with slightly paler striation; posterior region dark orange</td>
</tr>
<tr id="351034974E3D003AD7FB05CC29CCF9A6" box="[166,2024,1428,1561]" gridrow="10" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB05CC2C24F9A6" box="[166,512,1428,1561]" gridcol="0" gridrow="10" pageId="10" pageNumber="586">
<emphasis id="B954EAC74E3DFFCFD7FB05CC2C24FA12" bold="true" box="[166,512,1428,1453]" pageId="10" pageNumber="586">Colouration of P1 chela (dorsal)</emphasis>
</th>
<td id="76C15DEB4E3D003AD56905CC2DA1F9A6" box="[564,901,1428,1561]" gridcol="1" gridrow="10" pageId="10" pageNumber="586">Pale brown, somewhat darker on distodorsal surface of palm and fingers</td>
<td id="76C15DEB4E3D003AD4F505CC2ADEF9A6" box="[936,1274,1428,1561]" gridcol="2" gridrow="10" pageId="10" pageNumber="586">Orange brown to pale greenish, contrasting with purple-red- brown distodorsal surface of palm and fingers</td>
<td id="76C15DEB4E3D003AD24005CC2850F9A6" box="[1309,1652,1428,1561]" gridcol="3" gridrow="10" pageId="10" pageNumber="586">Typically with dark green-brown or brown spots</td>
<td id="76C15DEB4E3D003AD1CF05CC29CCF9A6" box="[1682,2024,1428,1561]" gridcol="4" gridrow="10" pageId="10" pageNumber="586">Uniform dark brown</td>
</tr>
<tr id="351034974E3D003AD7FB067029CCF936" box="[166,2024,1576,1673]" gridrow="11" pageId="10" pageNumber="586">
<th id="76C15DEB4E3D003AD7FB06702C24F936" box="[166,512,1576,1673]" gridcol="0" gridrow="11" pageId="10" pageNumber="586">Colouration of P2–P4</th>
<td id="76C15DEB4E3D003AD56906702DA1F936" box="[564,901,1576,1673]" gridcol="1" gridrow="11" pageId="10" pageNumber="586">Pale yellow to pale brown</td>
<td id="76C15DEB4E3D003AD4F506702ADEF936" box="[936,1274,1576,1673]" gridcol="2" gridrow="11" pageId="10" pageNumber="586">White with straw-yellow</td>
<td id="76C15DEB4E3D003AD24006702850F936" box="[1309,1652,1576,1673]" gridcol="3" gridrow="11" pageId="10" pageNumber="586">Dark brown or reddish brown (contrasting with usually much paler carapace)</td>
<td id="76C15DEB4E3D003AD1CF067029CCF936" box="[1682,2024,1576,1673]" gridcol="4" gridrow="11" pageId="10" pageNumber="586">Pale to dark brown (not contrasting with carapace)</td>
</tr>
</table>
</paragraph>
<paragraph id="8B9F36D54E3CFFCED7CA00CF2FE6FD3A" blockId="11.[151,1437,151,645]" pageId="11" pageNumber="587">
at most indicated in
<taxonomicName id="4C204D564E3CFFCED6DC00CF2FD4FF0E" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[385,496,151,177]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3CFFCED6DC00CF2FD4FF0E" box="[385,496,151,177]" italics="true" pageId="11" pageNumber="587">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3CFFCED6A400CF2C70FF0E" bold="true" box="[505,596,151,177]" pageId="11" pageNumber="587">
<taxonomicNameLabel id="A26757BC4E3CFFCED6A400CF2C70FF0E" box="[505,596,151,177]" pageId="11" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A504E3CFFCED53800CF2C92FF0E" box="[613,694,151,177]" captionStart="FIGURE 2" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[182,1406,181,1938]" captionTargetId="figure-23@3.[182,1406,181,1938]" captionTargetPageId="3" captionText="FIGURE 2. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412728" httpUri="https://zenodo.org/record/4412728/files/figure.png" pageId="11" pageNumber="587">Figs. 2</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED59900CF2CF7FF0E" box="[708,723,151,177]" captionStart="FIGURE 3" captionStartId="4.[151,250,985,1009]" captionTargetBox="[151,1435,181,961]" captionTargetId="figure-18@4.[151,1435,181,961]" captionTargetPageId="4" captionText="FIGURE 3. Petrolisthes lazarus sp. nov., female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412730" httpUri="https://zenodo.org/record/4412730/files/figure.png" pageId="11" pageNumber="587">3</figureCitation>
), being not nearly as conspicuous and large as in the Mexican species. In addition, the distolateral area of the palm and fingers of P1 of
<taxonomicName id="4C204D564E3CFFCED48F00E32A47FF6A" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[978,1123,187,213]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3CFFCED48F00E32A47FF6A" box="[978,1123,187,213]" italics="true" pageId="11" pageNumber="587">P. crenulatus</emphasis>
</taxonomicName>
are dark red-purple (except for whitish fingertips), contrasting with a paler, orange-brown or greenish palm (
<figureCitation id="131B2A504E3CFFCED37900872A50FF46" box="[1060,1140,222,249]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="11" pageNumber="587">Figs. 5</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED3DC00872AB5FF45" box="[1153,1169,223,250]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="11" pageNumber="587">6</figureCitation>
). In
<taxonomicName id="4C204D564E3CFFCED39800872B17FF46" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[1221,1331,223,249]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3CFFCED39800872B17FF46" box="[1221,1331,223,249]" italics="true" pageId="11" pageNumber="587">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3CFFCED26600872BB1FF46" bold="true" box="[1339,1429,223,249]" pageId="11" pageNumber="587">
<taxonomicNameLabel id="A26757BC4E3CFFCED26600872BB1FF46" box="[1339,1429,223,249]" pageId="11" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, these areas are also darker (
<figureCitation id="131B2A504E3CFFCED68F015B2C33FEA2" box="[466,535,259,285]" captionStart="FIGURE 2" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[182,1406,181,1938]" captionTargetId="figure-23@3.[182,1406,181,1938]" captionTargetPageId="3" captionText="FIGURE 2. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412728" httpUri="https://zenodo.org/record/4412728/files/figure.png" pageId="11" pageNumber="587">Fig. 2</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED578015B2C10FEA2" box="[549,564,259,285]" captionStart="FIGURE 3" captionStartId="4.[151,250,985,1009]" captionTargetBox="[151,1435,181,961]" captionTargetId="figure-18@4.[151,1435,181,961]" captionTargetPageId="4" captionText="FIGURE 3. Petrolisthes lazarus sp. nov., female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412730" httpUri="https://zenodo.org/record/4412730/files/figure.png" pageId="11" pageNumber="587">3</figureCitation>
), but not nearly as contrasting as in
<taxonomicName id="4C204D564E3CFFCED48F015B2A40FEA2" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[978,1124,259,285]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3CFFCED48F015B2A40FEA2" box="[978,1124,259,285]" italics="true" pageId="11" pageNumber="587">P. crenulatus</emphasis>
</taxonomicName>
(
<figureCitation id="131B2A504E3CFFCED328015B2AE1FEA2" box="[1141,1221,258,285]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="11" pageNumber="587">Figs. 5</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED38F015B2AC6FEA1" box="[1234,1250,259,286]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="11" pageNumber="587">6</figureCitation>
), although more specimens are needed to document the variability of the colour pattern in the new species. It must be noted that the detailed description of the colour pattern of
<taxonomicName id="4C204D564E3CFFCED5D401132D3EFEDA" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[649,794,331,357]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3CFFCED5D401132D3EFEDA" box="[649,794,331,357]" italics="true" pageId="11" pageNumber="587">P. crenulatus</emphasis>
</taxonomicName>
made by A. Petersen and published by
<bibRefCitation id="EFB14B244E3CFFCED38101132BB8FEDA" author="Haig, J." box="[1244,1436,331,357]" pageId="11" pageNumber="587" pagination="1 - 440" refId="ref5082" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960: 111)</bibRefCitation>
was possibly based on a relatively young or an unusually pale specimen. Nevertheless,
<bibRefCitation id="EFB14B244E3CFFCED33001372AD1FE35" author="Haig, J." box="[1133,1269,367,394]" pageId="11" pageNumber="587" pagination="1 - 440" refId="ref5082" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960)</bibRefCitation>
mentioned the P1 fingers being “light russet with reddish purple tint”. All our specimens from a single locality in
<collectingRegion id="49E4F8374E3CFFCED3B001CB2B65FE12" box="[1261,1345,403,429]" country="Mexico" name="Nayarit" pageId="11" pageNumber="587">Nayarit</collectingRegion>
(
<figureCitation id="131B2A504E3CFFCED20D01CB2BB0FE12" box="[1360,1428,403,429]" captionStart="FIGURE 7" captionStartId="8.[151,250,1568,1592]" captionTargetBox="[151,1435,417,1545]" captionTargetId="figure-146@8.[151,1435,417,1545]" captionTargetPageId="8" captionText="FIGURE 7. Mixed sand-rock intertidal at La Cruz de Huanacaxtle, Nayarit, Pacific coast of Mexico, collection locality of Petrolisthes crenulatus (Lockington, 1878): (A) general ocean view; (B) detail of rocks and sand; (C) one of the collected porcelain crabs in situ, exposed after flipping a large rock. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412738" httpUri="https://zenodo.org/record/4412738/files/figure.png" pageId="11" pageNumber="587">Fig. 7</figureCitation>
) identified as
<taxonomicName id="4C204D564E3CFFCED67501EF2F9DFE6E" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[296,441,439,465]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3CFFCED67501EF2F9DFE6E" box="[296,441,439,465]" italics="true" pageId="11" pageNumber="587">P. crenulatus</emphasis>
</taxonomicName>
have a very similar colour pattern (
<figureCitation id="131B2A504E3CFFCED41601EF2DBEFE6E" box="[843,922,438,465]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="11" pageNumber="587">Figs. 5</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED4FB01EF2D91FE6D" box="[934,949,439,466]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="11" pageNumber="587">6</figureCitation>
) and are darker than the material described by
<bibRefCitation id="EFB14B244E3CFFCED7E101832F6DFE49" author="Haig, J." box="[188,329,475,502]" pageId="11" pageNumber="587" pagination="1 - 440" refId="ref5082" refString="Haig, J. (1960) The Porcellanidae (Crustacea: Anomura) of the eastern Pacific. Allan Hancock Pacific Expedition, 24, 1 - 440." type="journal article" year="1960">Haig (1960)</bibRefCitation>
. Also noteworthy is that the low-resolution colour photographs of
<taxonomicName id="4C204D564E3CFFCED31901832AF3FE4A" baseAuthorityName="Lockington" baseAuthorityYear="1878" box="[1092,1239,475,501]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="crenulatus">
<emphasis id="B954EAC74E3CFFCED31901832AF3FE4A" box="[1092,1239,475,501]" italics="true" pageId="11" pageNumber="587">P. crenulatus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C204D564E3CFFCED3BA01832B4FFE4B" authorityName="Nobili" authorityYear="1901" box="[1255,1387,475,500]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="ortmanni">
<emphasis id="B954EAC74E3CFFCED3BA01832B4FFE4B" box="[1255,1387,475,500]" italics="true" pageId="11" pageNumber="587">P. ortmanni</emphasis>
</taxonomicName>
and
<taxonomicName id="4C204D564E3CFFCED7CA01A72ED2FDA6" baseAuthorityName="Glassell" baseAuthorityYear="1936" box="[151,246,511,537]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="lewisi">
<emphasis id="B954EAC74E3CFFCED7CA01A72ED2FDA6" box="[151,246,511,537]" italics="true" pageId="11" pageNumber="587">P. lewisi</emphasis>
</taxonomicName>
published by García-Madrigal &amp; Abréu-Sanchez (2009) show dead and largely discoloured specimens. Therefore, the colour photographs of these three species provided herein (
<figureCitation id="131B2A504E3CFFCED488027B2A00FD82" box="[981,1060,546,573]" captionStart="FIGURE 5" captionStartId="6.[151,250,1959,1983]" captionTargetBox="[175,1412,181,1935]" captionTargetId="figure-18@6.[175,1412,181,1935]" captionTargetPageId="6" captionText="FIGURE 5. Petrolisthes crenulatus Lockington, 1878, male from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35708): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by L.D. Santana-Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412734" httpUri="https://zenodo.org/record/4412734/files/figure.png" pageId="11" pageNumber="587">Figs. 5</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED36D027B2A64FD82" box="[1072,1088,547,573]" captionStart="FIGURE 6" captionStartId="7.[151,250,1485,1509]" captionTargetBox="[151,1435,181,1461]" captionTargetId="figure-23@7.[151,1435,181,1461]" captionTargetPageId="7" captionText="FIGURE 6. Petrolisthes crenulatus Lockington, 1878: (A) ovigerous female from La Cruz de Huanacaxtle, Nayarit, Mexico (CNCR 35709), dorsal view; (B) male from the same locality (CNCR 35710), dorsal view. Photographs by L.D. Santana- Moreno." figureDoi="http://doi.org/10.5281/zenodo.4412736" httpUri="https://zenodo.org/record/4412736/files/figure.png" pageId="11" pageNumber="587">6</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED310027B2A78FD82" box="[1101,1116,547,573]" captionStart="FIGURE 8" captionStartId="9.[151,250,1943,1967]" captionTargetBox="[164,1423,181,1920]" captionTargetId="figure-23@9.[164,1423,181,1920]" captionTargetPageId="9" captionText="FIGURE 8. Petrolisthes lewisi (Glassell, 1936) [A, B] and Petrolisthes ortmanni Nobili, 1901 [C]: (A) male from Los Negritos, off Bahía Málaga, Pacific coast of Colombia (CERBMcr UV 2019-103), dorsal view; (B) ovigerous female from the same locality (CERBMcr UV 2019-102); male from the same locality (CERBMcr UV 2019-102). Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412740" httpUri="https://zenodo.org/record/4412740/files/figure.png" pageId="11" pageNumber="587">8</figureCitation>
), as well as of
<taxonomicName id="4C204D564E3CFFCED258027B2B57FD82" authority="Ferreira &amp; Santana-Moreno &amp; Anker, 2020" authorityName="Ferreira &amp; Santana-Moreno &amp; Anker" authorityYear="2020" box="[1285,1395,547,573]" class="Malacostraca" family="Porcellanidae" genus="Petrolisthes" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="587" phylum="Arthropoda" rank="species" species="lazarus" status="sp. nov.">
<emphasis id="B954EAC74E3CFFCED258027B2B57FD82" box="[1285,1395,547,573]" italics="true" pageId="11" pageNumber="587">P. lazarus</emphasis>
</taxonomicName>
<emphasis id="B954EAC74E3CFFCED226027B2EECFDDE" bold="true" pageId="11" pageNumber="587">
<taxonomicNameLabel id="A26757BC4E3CFFCED226027B2EECFDDE" pageId="11" pageNumber="587" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="131B2A504E3CFFCED78A021F2F02FDDE" box="[215,294,583,609]" captionStart="FIGURE 2" captionStartId="3.[151,250,1962,1986]" captionTargetBox="[182,1406,181,1938]" captionTargetId="figure-23@3.[182,1406,181,1938]" captionTargetPageId="3" captionText="FIGURE 2. Petrolisthes lazarus sp. nov., female holotype from Isla Saboga, Las Perlas Islands, Panama (MZUSP 33807): (A) dorsal view; (B) ventral view; (C) anterior (frontal) view. Photographs by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412728" httpUri="https://zenodo.org/record/4412728/files/figure.png" pageId="11" pageNumber="587">Figs. 2</figureCitation>
,
<figureCitation id="131B2A504E3CFFCED66C021F2F64FDDE" box="[305,320,583,609]" captionStart="FIGURE 3" captionStartId="4.[151,250,985,1009]" captionTargetBox="[151,1435,181,961]" captionTargetId="figure-18@4.[151,1435,181,961]" captionTargetPageId="4" captionText="FIGURE 3. Petrolisthes lazarus sp. nov., female paratype from Río Mar, Panama (MZUSP 33686), dorsal view. Photograph by A. Anker." figureDoi="http://doi.org/10.5281/zenodo.4412730" httpUri="https://zenodo.org/record/4412730/files/figure.png" pageId="11" pageNumber="587">3</figureCitation>
), should be used as a quick visual reference for future field studies and easy discrimination of living or freshly dead specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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