299 lines
37 KiB
XML
299 lines
37 KiB
XML
<document id="7B8100731FE7735A3E34CA465D102A0D" ID-CLB-Dataset="217329" ID-DOI="10.37520/aemnp.2022.017" ID-GBIF-Dataset="795cf556-a4a9-477b-b55d-cb80ed8a0f2d" ID-ISSN="1804-6487" ID-Zenodo-Dep="7399702" ID-ZooBank="28D55112-98B1-49A5-B382-58B1B068570B" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1670240514228" checkinUser="felipe" docAuthor="Hansen, Aslak Kappel, Brunke, Adam, Simonsen, Thomas & Solodovnikov, Alexey" docDate="2022" docId="038987A0FFC74B27B137A5DF14ADF78A" docLanguage="en" docName="ActaEntMusNatPra.62.1.225-299.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae (Acta. Ent. Mus. Natl. Pragae) 62 (1)" docSource="http://dx.doi.org/10.37520/aemnp.2022.017" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Quedius strenuus Casey 1915" docType="treatment" docVersion="7" lastPageNumber="287" masterDocId="FFB0FFD8FFFA4B19B016A005151AFF9F" masterDocTitle="Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)" masterLastPageNumber="299" masterPageNumber="225" pageNumber="286" updateTime="1699412962807" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="1E55D7546A2102EEB362FFBC908D9088">Revision of Quedius sensu stricto (Coleoptera: Staphylinidae)</mods:title>
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<mods:namePart id="84A3168A6C79F833F497B04DBB913C61">Hansen, Aslak Kappel</mods:namePart>
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<mods:affiliation id="F4DD1881033370F53A18534E3945FD64">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark & Department of Bioscience, University of Aarhus, Ny Munkegade 116, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:namePart id="F5A7C92DB20448E45CCE7FD2F42A4557">Brunke, Adam</mods:namePart>
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<mods:affiliation id="938CEC3ACFE78DED228AD7B3DCDE2DD6">Agriculture and Agri-Food Canada, 960 Carling Avenue, Ottawa, Ontario, K</mods:affiliation>
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<mods:namePart id="7FDD293D49E47A6D4E4897DFB2602235">Simonsen, Thomas</mods:namePart>
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<mods:affiliation id="B6F2F1D10F245B11A16D2430EE54F883">Natural History Museum Aarhus, Wilhelm Meyers Allé 10, 8000 Aarhus, Denmark</mods:affiliation>
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<mods:namePart id="FD1BACFEAC7C80E7368FA52D90CF898F">Solodovnikov, Alexey</mods:namePart>
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<mods:affiliation id="396DEEF50700742F8C5DAFA2398E62F4">Natural History Museum of Denmark, Zoological Museum, Universitetsparken 15, 2100 Copenhagen, Denmark; e-mails: akhansen @ snm. ku. dk; asolodovnikov @ snm. ku. dk & Zoological Institute, Russian Academy of Science, Universitetskaja nab.</mods:affiliation>
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<treatment id="038987A0FFC74B27B137A5DF14ADF78A" ID-DOI="http://doi.org/10.5281/zenodo.7503816" ID-GBIF-Taxon="208533256" ID-Zenodo-Dep="7503816" LSID="urn:lsid:plazi:treatment:038987A0FFC74B27B137A5DF14ADF78A" httpUri="http://treatment.plazi.org/id/038987A0FFC74B27B137A5DF14ADF78A" lastPageId="62" lastPageNumber="287" pageId="61" pageNumber="286">
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<paragraph id="8B9F36B6FFC74B24B137A5DF1774FA6D" blockId="61.[289,622,1498,1522]" box="[289,622,1498,1522]" pageId="61" pageNumber="286">
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<heading id="D0D781DAFFC74B24B137A5DF1774FA6D" box="[289,622,1498,1522]" centered="true" fontSize="10" level="2" pageId="61" pageNumber="286" reason="3">
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<taxonomicName id="4C204D35FFC74B24B137A5DF1774FA6D" ID-CoL="4R2Z5" authority="Casey, 1915" authorityName="Casey" authorityYear="1915" box="[289,622,1498,1522]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="286" phylum="Arthropoda" rank="species" species="strenuus">
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<emphasis id="B954EAA4FFC74B24B137A5DF1774FA6D" bold="true" box="[289,622,1498,1522]" pageId="61" pageNumber="286">
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<emphasis id="B954EAA4FFC74B24B137A5DF14F8FA6D" bold="true" box="[289,482,1498,1522]" italics="true" pageId="61" pageNumber="286">Quedius strenuus</emphasis>
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<bibRefCitation id="EFB14B47FFC74B24B1FFA5DE1774FA6D" author="CASEY T. L." box="[489,622,1499,1522]" pageId="61" pageNumber="286" pagination="395 - 450" refId="ref81365" refString="CASEY T. L. 1915: Studies in some staphylinid genera of North America. Memoirs on the Coleoptera 6: 395 - 450." type="journal article" year="1915">Casey, 1915</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A653DFFC74B24B170A5FC1732F993" box="[358,552,1529,1548]" pageId="61" pageNumber="286" type="description">
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<paragraph id="8B9F36B6FFC74B24B170A5FC1732F993" blockId="61.[151,644,1529,1582]" box="[358,552,1529,1548]" pageId="61" pageNumber="286">
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(
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<figureCitation id="131B2A33FFC74B24B17AA5FC14B9F993" box="[364,419,1529,1548]" captionStart="Fig" captionStartId="1.[152,183,2006,2025]" captionTargetBox="[222,1381,1063,1978]" captionTargetId="figure-474@1.[207,1384,1060,1988]" captionTargetPageId="1" captionText="Fig. 1. Phylogenetic tree of Quedius s. str. based on preliminary target capture molecular dataset of Bඋඎඇĸൾ et al. (2021) and Hൺඇඌൾඇ et al. (in prep). Species groups shown in color. Q. altanai Hansen & Brunke, sp. nov. of the Q. molochinoides-group was not included in the dataset. Habitus sketch is of Q. fuliginosus (Gravenhorst, 1802). Lettering denotes species distribution within their respective groups." figureDoi="http://doi.org/10.5281/zenodo.7399704" httpUri="https://zenodo.org/record/7399704/files/figure.png" pageId="61" pageNumber="286">Figs 1</figureCitation>
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,
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<figureCitation id="131B2A33FFC74B24B1BBA5FC14A3F993" box="[429,441,1529,1548]" captionStart="Fig" captionStartId="12.[151,182,2008,2027]" captionTargetBox="[182,1406,199,1981]" captionTargetId="figure-13@12.[182,1407,189,1989]" captionTargetPageId="12" captionText="Fig. 4. Phylogeny of Quedius s. str. species based on Bayesian Inference (BI) and Maximum Likelihood (ML) COI barcodes (part 2). BI posterior probabilities (PP), ML ultrafast bootstrap support (UFB) and ML SH-aLRT test are shown. Nodes coloured as: green, PP ≥ 0.90, and either UFB ≥ 95 or SH-aLRT ≥ 80; yellow, one of PP ≥ 0.90, UFB ≥ 95 or SH-aLRT ≥ 80; red, PP <0.90, UFB <95 and SH-aLRT <80." figureDoi="http://doi.org/10.5281/zenodo.7399710" httpUri="https://zenodo.org/record/7399710/files/figure.png" pageId="61" pageNumber="286">4</figureCitation>
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,
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<figureCitation id="131B2A33FFC74B24B1D4A5FC14C0F993" box="[450,474,1529,1548]" captionStart="Fig" captionStartId="19.[151,182,2028,2047]" captionTargetBox="[203,1360,198,1958]" captionTargetId="figure-13@19.[183,1405,186,1971]" captionTargetPageId="19" captionText="Fig. 9. Habitus of Quedius s. str. A – Q. hispanicus Bernhauer, 1898; B – Q. hispanicus (pale sutures variant); C – Q. pallipes Lucas, 1846; D – Q. laticollis (Gravenhorst, 1802) (black elytra variant); E – Q. laticollis (red elytra variant); F – Q. strenuus Casey, 1915." figureDoi="http://doi.org/10.5281/zenodo.7399724" httpUri="https://zenodo.org/record/7399724/files/figure.png" pageId="61" pageNumber="286">9F</figureCitation>
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,
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<figureCitation id="131B2A33FFC74B24B1F4A5FC1719F993" box="[482,515,1529,1548]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="61" pageNumber="286">14F</figureCitation>
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,
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<figureCitation id="131B2A33FFC74B24B21AA5FC1738F993" box="[524,546,1529,1548]" captionStart="Fig" captionStartId="61.[151,182,1421,1440]" captionTargetBox="[189,1399,189,1406]" captionTargetId="figure-13@61.[189,1400,189,1408]" captionTargetPageId="61" captionText="Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399754" httpUri="https://zenodo.org/record/7399754/files/figure.png" pageId="61" pageNumber="286">22</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A653DFFC74B24B081A61E176FF9EC" pageId="61" pageNumber="286" type="reference_group">
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<paragraph id="8B9F36B6FFC74B24B081A61E179EF9B1" blockId="61.[151,644,1529,1582]" box="[151,644,1563,1582]" pageId="61" pageNumber="286">
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<treatmentCitationGroup id="AB301198FFC74B24B081A61E179EF9B1" box="[151,644,1563,1582]" pageId="61" pageNumber="286">
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<treatmentCitation id="0A8110A7FFC74B24B081A61E14DEF9B1" author="CASEY T. L." box="[151,452,1563,1582]" page="401" pageId="61" pageNumber="286" year="1915">
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<taxonomicName id="4C204D35FFC74B24B081A61E14DEF9B1" ID-CoL="4R2Z5" authority="Casey, 1915: 401" authorityName="Casey" authorityPageNumber="401" authorityYear="1915" box="[151,452,1563,1582]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="286" phylum="Arthropoda" rank="species" species="strenuus">
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<emphasis id="B954EAA4FFC74B24B081A61E1436F9B1" box="[151,300,1563,1582]" italics="true" pageId="61" pageNumber="286">Quedius strenuus</emphasis>
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<bibRefCitation id="EFB14B47FFC74B24B126A61E14DEF9B1" author="CASEY T. L." box="[304,452,1563,1582]" pageId="61" pageNumber="286" pagination="395 - 450" refId="ref81365" refString="CASEY T. L. 1915: Studies in some staphylinid genera of North America. Memoirs on the Coleoptera 6: 395 - 450." type="journal article" year="1915">Casey, 1915: 401</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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[
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<typeStatus id="549B8814FFC74B24B1D8A61E14E1F9B1" box="[462,507,1563,1582]" pageId="61" pageNumber="286">Type</typeStatus>
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locality:
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<collectingRegion id="49E4F854FFC74B24B25DA61E1765F9B1" box="[587,639,1563,1582]" country="United States of America" name="Texas" pageId="61" pageNumber="286">Texas</collectingRegion>
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]
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8B9F36B6FFC74B24B081A640176FF9EC" blockId="61.[151,761,1605,1933]" pageId="61" pageNumber="286">
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<emphasis id="B954EAA4FFC74B24B081A640141AF9C7" bold="true" box="[151,256,1605,1624]" pageId="61" pageNumber="286">References.</emphasis>
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Sආൾඍൺඇൺ (1971a): 132 (
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<typeStatus id="549B8814FFC74B24B1CCA6401730F9C7" box="[474,554,1605,1624]" pageId="61" pageNumber="286" type="lectotype">lectotype</typeStatus>
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designation, characters and distribution); (1981): 638, (1990): 98 (distribution).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33A653DFFC74B24B081A68F1443F812" pageId="61" pageNumber="286" type="materials_examined">
|
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<paragraph id="8B9F36B6FFC74B24B081A68F1484F94D" blockId="61.[151,761,1605,1933]" pageId="61" pageNumber="286">
|
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<emphasis id="B954EAA4FFC74B24B081A68F146AF902" bold="true" box="[151,368,1674,1693]" pageId="61" pageNumber="286">
|
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<typeStatus id="549B8814FFC74B24B081A68F15D9F902" box="[151,195,1674,1693]" pageId="61" pageNumber="286">Type</typeStatus>
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material examined.
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</emphasis>
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Lൾർඍඈඍඒඉൾ:
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<specimenCount id="9D26FD3FFFC74B24B1CCA68614F0F903" box="[474,490,1667,1692]" pageId="61" pageNumber="286" type="female">♀</specimenCount>
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, designated by Sආൾඍൺඇൺ (1971a) (
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<collectionCode id="ED31AE73FFC74B24B08BA6A115C7F928" box="[157,221,1700,1719]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="61" pageNumber="286" type="Museum">USNM</collectionCode>
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) [not examined], labelled: “Tex / Casey bequest 1925/
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<typeStatus id="549B8814FFC74B24B2DBA6A117EDF928" box="[717,759,1700,1719]" pageId="61" pageNumber="286">Type</typeStatus>
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||
USNM 48279 /
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<taxonomicName id="4C204D35FFC74B24B136A6BA1488F94D" authority="Csy" authorityName="Casey" authorityYear="1915" box="[288,402,1727,1746]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="61" pageNumber="286" phylum="Arthropoda" rank="species" species="strenuus">strenuus Csy</taxonomicName>
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”.
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</paragraph>
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<paragraph id="8B9F36B6FFC74B24B081A6DF1443F812" blockId="61.[151,761,1605,1933]" pageId="61" pageNumber="286">
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||
<emphasis id="B954EAA4FFC74B24B081A6DF1732F972" bold="true" box="[151,552,1754,1773]" pageId="61" pageNumber="286">
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Additional material examined.
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<collectingCountry id="F3377626FFC74B24B1B1A6DF14CEF972" box="[423,468,1754,1773]" name="United States of America" pageId="61" pageNumber="286">USA</collectingCountry>
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: Kൺඇඌൺඌ:
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</emphasis>
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Lawrence, below bridge,
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<collectingRegion id="49E4F854FFC74B24B081A6F115C9F898" box="[151,211,1780,1799]" country="United States of America" name="Kansas" pageId="61" pageNumber="286">Kansas</collectingRegion>
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River,[38.96, -95.26],under chunk of mud flat,
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<date id="FF9E1076FFC74B24B24BA6F117ADF898" box="[605,695,1780,1799]" pageId="61" pageNumber="286" value="1936-10-16">16.X.1936</date>
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, leg. M. W. Sanderson (1
|
||
<collectionCode id="ED31AE73FFC74B24B13FA70A1443F8BD" box="[297,345,1807,1826]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="61" pageNumber="286">CNC</collectionCode>
|
||
).
|
||
<emphasis id="B954EAA4FFC74B24B170A70A14BFF8BD" bold="true" box="[358,421,1807,1826]" pageId="61" pageNumber="286">Tൾඑൺඌ:</emphasis>
|
||
Braunfels, [29.69, -98.11],
|
||
<date id="FF9E1076FFC74B24B287A70A17EEF8BD" box="[657,756,1807,1826]" pageId="61" pageNumber="286" value="1927-06-13">13.VI.1927</date>
|
||
, leg. Darlington (1
|
||
<collectionCode id="ED31AE73FFC74B24B12EA72F1470F8A2" box="[312,362,1834,1853]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33791" lsid="urn:lsid:biocol.org:col:33791" name="Museum of Comparative Zoology" pageId="61" pageNumber="286" type="Museum">MCZ</collectionCode>
|
||
); Del Rio, [29.36, -100.9],
|
||
<date id="FF9E1076FFC74B24B243A72F17AEF8A2" box="[597,692,1834,1853]" pageId="61" pageNumber="286" value="1938-07-08">8.VII.1938</date>
|
||
, leg. R. L. Sailer (3
|
||
<collectionCode id="ED31AE73FFC74B24B115A7411429F8C8" box="[259,307,1860,1879]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="61" pageNumber="286">CNC</collectionCode>
|
||
); Kerrville, [30.02, -99.13],
|
||
<date id="FF9E1076FFC74B24B23AA7411796F8C8" box="[556,652,1860,1879]" pageId="61" pageNumber="286" value="1998-04-11">11.IV.1998</date>
|
||
, leg. Chamberlain (3
|
||
<collectionCode id="ED31AE73FFC74B24B0E4A75A1429F8ED" box="[242,307,1887,1906]" pageId="61" pageNumber="286">TAMU</collectionCode>
|
||
); Uvalde County, [29.30, -99.70],
|
||
<date id="FF9E1076FFC74B24B276A75A17A0F8ED" box="[608,698,1887,1906]" pageId="61" pageNumber="286" value="1938-05-26">26.V.1938</date>
|
||
, leg. J. H. Robinson (1
|
||
<collectionCode id="ED31AE73FFC74B24B109A77F1455F812" box="[287,335,1914,1933]" country="Canada" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="61" pageNumber="286">CNC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFC74B27B081A7A01433FD0A" lastPageId="62" lastPageNumber="287" pageId="61" pageNumber="286" type="description">
|
||
<paragraph id="8B9F36B6FFC74B24B081A7A0169FF965" blockId="61.[151,760,1952,2076]" lastBlockId="61.[828,1437,1507,2074]" pageId="61" pageNumber="286">
|
||
<emphasis id="B954EAA4FFC74B24B081A7A0145AF823" bold="true" box="[151,320,1957,1980]" pageId="61" pageNumber="286">Redescription.</emphasis>
|
||
Measurements JJ (n = 3): HW = 1.60– 1.82 (1.73); HL = 1.33–1.51 (1.44); HL/HW 0.80–0.85 (0.83); PW = 2.20–2.47 (2.37); PL = 2.02–2.22 (2.14); PL/PW 0.89–0.92 (0.90); EW = 2.33–2.60 (2.50); EL = 2.00–2.24 (2.16); EL/EW 0.85–0.88 (0.86); EL/PL 0.99–1.03 (1.01); PW/HW 1.63–1.67 (1.65); forebody length 5.36–5.96 (5.73).
|
||
<specimenCount id="9D26FD3FFFC74B24B444A6191162F9A4" box="[1106,1144,1564,1595]" pageId="61" pageNumber="286" type="female">♀♀</specimenCount>
|
||
(n = 2): HW = 1.69–1.76 (1.72); HL = 1.44–1.47 (1.46); HL/HW 0.82–0.87 (0.85); PW = 2.33–2.44 (2.39); PL = 2.07–2.09 (2.08); PL/PW 0.85–0.89 (0.87); EW = 2.62–2.64 (2.63); EL = 2.18–2.29 (2.23); EL/EW 0.83–0.87 (0.85); EL/PL 1.05–1.10 (1.07); PW/HW 1.59–1.69 (1.64); forebody length 5.71–5.82 (5.77).
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFC74B24B34CA706109CF885" blockId="61.[828,1437,1507,2074]" box="[858,1414,1795,1818]" pageId="61" pageNumber="286">Medium sized, robust species; body black (Fig, 9F).</paragraph>
|
||
<paragraph id="8B9F36B6FFC74B24B34CA726117AF825" blockId="61.[828,1437,1507,2074]" pageId="61" pageNumber="286">
|
||
Head black, distinctly transverse, with eyes medium sized (EyL/TL = 1.76–2.00 (1.88)); microsculpture of transverse waves; no additional punctures between anterior frontal punctures (
|
||
<figureCitation id="131B2A33FFC74B24B41FA7861145F805" box="[1033,1119,1923,1946]" captionStart="Fig" captionStartId="16.[151,181,1101,1120]" captionTargetBox="[171,736,194,1069]" captionTargetId="figure-533@16.[151,760,186,1075]" captionTargetPageId="16" captionText="Fig.6. Head and pronotal chaetotaxy of Quedius s. str.A – general scheme; B – Q. curtipennis Bernhauer, 1908, C – Q. fuliginosus (Gravenhorst, 1802); D – Q. levicollis Brullé, 1832; E – Q. afrofuliginosus Gusarov, 1991; F – Q. molochinus- and Q. laticollis-groups; G – Q. molochinoides-group. Abbreviations as follows: anterior frontal puncture (AFP); basal punctures (BP); interocular punctures (IOP); lateral puncture (LP); lateral row (LR); posterior frontal puncture (PFP); parocular punctures (POP); sublateral row (SLR)." figureDoi="http://doi.org/10.5281/zenodo.7399716" httpUri="https://zenodo.org/record/7399716/files/figure.png" pageId="61" pageNumber="286">Fig. 6F</figureCitation>
|
||
); all antennomeres elongate, antennae and palpi reddish.
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFC74B27B34CA7C61722FECA" blockId="61.[828,1437,1507,2074]" lastBlockId="62.[151,760,190,2069]" lastPageId="62" lastPageNumber="287" pageId="61" pageNumber="286">Thorax: pronotum black, slightly wider than long, wider than head, with microsculpture of transverse waves; three punctures in dorsal row and one to two in sublateral row with its posteriormost puncture reaching just beyond first puncture of dorsal row; scutellum punctured and pubescent; elytra black, uniformly pubescent, slightly wider than long, roughly as long as pronotum; legs reddish brown with inner face of tibia and femur darkened.</paragraph>
|
||
<paragraph id="8B9F36B6FFC44B27B0A3A15B1443FE0A" blockId="62.[151,760,190,2069]" pageId="62" pageNumber="287">Abdomen black, tergites uniformly punctured, with slight iridescence.</paragraph>
|
||
<paragraph id="8B9F36B6FFC44B27B0A3A1981433FD0A" blockId="62.[151,760,190,2069]" pageId="62" pageNumber="287">
|
||
<emphasis id="B954EAA4FFC44B27B0A3A19815ECFE2A" bold="true" box="[181,246,413,437]" italics="true" pageId="62" pageNumber="287">Male.</emphasis>
|
||
Aedeagus (
|
||
<figureCitation id="131B2A33FFC44B27B169A19B14FCFE2A" box="[383,486,414,437]" captionStart="Fig" captionStartId="38.[151,182,1981,2000]" captionTargetBox="[199,1380,186,1956]" captionTargetId="figure-13@38.[183,1399,178,1966]" captionTargetPageId="38" captionText="Fig. 14. Aedeagus of Quedius s. str. A – Q. unicolor Kiesenwetter, 1847; B – Q. levasseuri Coiffait, 1964 (syn. of Q. pallipes Lucas, 1846); C – Q. hispanicus Bernhauer, 1898; D – Q. pallipes Lucas, 1846; E – Q. laticollis (Gravenhorst, 1802); F – Q. strenuus Casey, 1915. I – Parameral view of whole aedeagus, II – apices of paramere in antiparameral view with peg setae, III – lateral view of whole aedeagus, IV – apices of median lobe at an angle between parameral and lateral view." figureDoi="http://doi.org/10.5281/zenodo.7399734" httpUri="https://zenodo.org/record/7399734/files/figure.png" pageId="62" pageNumber="287">Fig. 14F</figureCitation>
|
||
): paramere rather broad lanceolate, slightly protruding beyond apex of median lobe, with sensory peg setae forming two distinct short and wide rows of multiple punctures; median lobe broad with gentle constriction towards pointed apex, on parameral side with two small hooked teeth protruding slightly basad, positioned at level near lower level of peg setae patch of the paramere.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFC44B27B081A29B1465F9AA" pageId="62" pageNumber="287" type="diagnosis">
|
||
<paragraph id="8B9F36B6FFC44B27B081A29B145CFACA" blockId="62.[151,760,190,2069]" pageId="62" pageNumber="287">
|
||
<emphasis id="B954EAA4FFC44B27B081A29B1489FD2A" bold="true" box="[151,403,670,693]" pageId="62" pageNumber="287">Differential diagnosis.</emphasis>
|
||
<taxonomicName id="4C204D35FFC44B27B18FA29B174EFD2A" authorityName="Casey" authorityYear="1915" box="[409,596,670,693]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B18FA29B174EFD2A" box="[409,596,670,693]" italics="true" pageId="62" pageNumber="287">Quedius strenuus</emphasis>
|
||
</taxonomicName>
|
||
is similar to
|
||
<taxonomicName id="4C204D35FFC44B27B2CBA29B15EFFD4A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="B954EAA4FFC44B27B2CBA29B15EFFD4A" italics="true" pageId="62" pageNumber="287">Q. laticollis</emphasis>
|
||
</taxonomicName>
|
||
from which it normally differs by the completely dark body with pale appendages. For confident delimitation of both species from each other the position of the peg setae of the paramere can be used, which form two distinct rows in
|
||
<taxonomicName id="4C204D35FFC44B27B151A33B14DEFCCA" authorityName="Casey" authorityYear="1915" box="[327,452,830,853]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B151A33B14DEFCCA" box="[327,452,830,853]" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
and a triangular patch in
|
||
<taxonomicName id="4C204D35FFC44B27B2CBA33B15EDFCEA" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="B954EAA4FFC44B27B2CBA33B15EDFCEA" italics="true" pageId="62" pageNumber="287">Q. laticollis</emphasis>
|
||
</taxonomicName>
|
||
. Co-occurrence of the two species is unlikely since
|
||
<taxonomicName id="4C204D35FFC44B27B0CFA37B1443FC0A" authorityName="Casey" authorityYear="1915" box="[217,345,894,917]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B0CFA37B1443FC0A" box="[217,345,894,917]" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
replaces
|
||
<taxonomicName id="4C204D35FFC44B27B1D1A37B1751FC0A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[455,587,894,917]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="B954EAA4FFC44B27B1D1A37B1751FC0A" box="[455,587,894,917]" italics="true" pageId="62" pageNumber="287">Q. laticollis</emphasis>
|
||
</taxonomicName>
|
||
to the west, in the forested areas of Great Plains of central-eastern and lowland areas of the southwestern
|
||
<collectingCountry id="F3377626FFC44B27B212A3BB178CFC4A" box="[516,662,958,981]" name="United States of America" pageId="62" pageNumber="287">United States</collectingCountry>
|
||
.
|
||
<taxonomicName id="4C204D35FFC44B27B288A3BB15E8FC6A" authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B288A3BB15E8FC6A" italics="true" pageId="62" pageNumber="287">Quedius strenuus</emphasis>
|
||
</taxonomicName>
|
||
is also very similar to the introduced species
|
||
<taxonomicName id="4C204D35FFC44B27B081A3FB1427FB8A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[151,317,1022,1045]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="molochinus">
|
||
<emphasis id="B954EAA4FFC44B27B081A3FB1427FB8A" box="[151,317,1022,1045]" italics="true" pageId="62" pageNumber="287">Q. molochinus</emphasis>
|
||
</taxonomicName>
|
||
, which has not yet been found within the distribution range of
|
||
<taxonomicName id="4C204D35FFC44B27B1BAA41B1730FBAA" authorityName="Casey" authorityYear="1915" box="[428,554,1054,1077]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B1BAA41B1730FBAA" box="[428,554,1054,1077]" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
. Other differences include that, in
|
||
<taxonomicName id="4C204D35FFC44B27B157A43B14C5FBCA" baseAuthorityName="Gravenhorst" baseAuthorityYear="1806" box="[321,479,1086,1109]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="molochinus">
|
||
<emphasis id="B954EAA4FFC44B27B157A43B14C5FBCA" box="[321,479,1086,1109]" italics="true" pageId="62" pageNumber="287">Q. molochinus</emphasis>
|
||
</taxonomicName>
|
||
, the antennae are slightly stouter, less elongate, with darker middle antennomeres; as well as elytra are finer and more densely punctate, and either brick red or dark red (always dark in
|
||
<taxonomicName id="4C204D35FFC44B27B2B3A49B15D0FB4A" authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B2B3A49B15D0FB4A" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<taxonomicName id="4C204D35FFC44B27B0F5A4BB14BBFB4A" authorityName="Casey" authorityYear="1915" box="[227,417,1214,1237]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B0F5A4BB14BBFB4A" box="[227,417,1214,1237]" italics="true" pageId="62" pageNumber="287">Quedius strenuus</emphasis>
|
||
</taxonomicName>
|
||
cannot be confused with other North American
|
||
<taxonomicName id="4C204D35FFC44B27B158A4DB14B2FB6A" authorityName="Stephens" authorityYear="1829" box="[334,424,1246,1269]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B954EAA4FFC44B27B158A4DB14B2FB6A" box="[334,424,1246,1269]" italics="true" pageId="62" pageNumber="287">Quedius</emphasis>
|
||
</taxonomicName>
|
||
as they either have interocular punctures on frons or (those from other subgenera) have clear medial incision of labrum and abdomen notably tapering apicad.
|
||
</paragraph>
|
||
<paragraph id="8B9F36B6FFC44B27B081A55B1465F9AA" blockId="62.[151,760,190,2069]" pageId="62" pageNumber="287">
|
||
<emphasis id="B954EAA4FFC44B27B081A55B140DFAEA" bold="true" box="[151,279,1374,1397]" pageId="62" pageNumber="287">Bionomics.</emphasis>
|
||
Little is known about the bionomics of
|
||
<taxonomicName id="4C204D35FFC44B27B2CBA55B15E8FA0A" authorityName="Casey" authorityYear="1915" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B2CBA55B15E8FA0A" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
. Based on current records,
|
||
<taxonomicName id="4C204D35FFC44B27B235A57B17BAFA0A" authorityName="Casey" authorityYear="1915" box="[547,672,1406,1429]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B235A57B17BAFA0A" box="[547,672,1406,1429]" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
is most likely restricted to lowland sites. As the region covered by its distribution is rather arid and most specimens were found near creeks, rivers, and lakes, it is likely to be strictly confined to moist microhabitats like wet flood or water-edge debris.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A653DFFC44B27B081A63B14ADF78A" pageId="62" pageNumber="287" type="distribution">
|
||
<paragraph id="8B9F36B6FFC44B27B081A63B14ADF78A" blockId="62.[151,760,190,2069]" pageId="62" pageNumber="287">
|
||
<emphasis id="B954EAA4FFC44B27B081A63B1430F9CA" bold="true" box="[151,298,1598,1621]" pageId="62" pageNumber="287">Distribution.</emphasis>
|
||
<taxonomicName id="4C204D35FFC44B27B126A63B14F2F9CA" authorityName="Casey" authorityYear="1915" box="[304,488,1598,1621]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B126A63B14F2F9CA" box="[304,488,1598,1621]" italics="true" pageId="62" pageNumber="287">Quedius strenuus</emphasis>
|
||
</taxonomicName>
|
||
is found across the Great Plains (ecoregion 9.4) and potentially a large part of lowland southwest
|
||
<collectingCountry id="F3377626FFC44B27B17EA67B14E3F90A" box="[360,505,1662,1685]" name="United States of America" pageId="62" pageNumber="287">United States</collectingCountry>
|
||
(parts of ecoregion 10,
|
||
<figureCitation id="131B2A33FFC44B27B081A69B15FDF92A" box="[151,231,1694,1717]" captionStart="Fig" captionStartId="61.[151,182,1421,1440]" captionTargetBox="[189,1399,189,1406]" captionTargetId="figure-13@61.[189,1400,189,1408]" captionTargetPageId="61" captionText="Fig. 22. Distribution of Quedius molochinoides Smetana, 1965 (top), and Q. laticollis (Gravenhorst, 1802) and Q. strenuus Casey, 1915 (bottom)." figureDoi="http://doi.org/10.5281/zenodo.7399754" httpUri="https://zenodo.org/record/7399754/files/figure.png" pageId="62" pageNumber="287">Fig. 22</figureCitation>
|
||
). Only a few specimens, which we were not able to examine, were imprecisely recorded from lowland Southwest United State, from
|
||
<collectingRegion id="49E4F854FFC44B27B1ECA6DB174EF96A" box="[506,596,1758,1781]" country="United States of America" name="Arizona" pageId="62" pageNumber="287">Arizona</collectingRegion>
|
||
and Southern
|
||
<collectingRegion id="49E4F854FFC44B27B081A6FB141DF88A" box="[151,263,1790,1813]" country="United States of America" name="California" pageId="62" pageNumber="287">California</collectingRegion>
|
||
. Of these, only a record from Lyman Lake in Apache County,
|
||
<collectingRegion id="49E4F854FFC44B27B147A71B14B2F8AA" box="[337,424,1822,1845]" country="United States of America" name="Arizona" pageId="62" pageNumber="287">Arizona</collectingRegion>
|
||
(Sආൾඍൺඇൺ 1981) is exact and fully reliable. Earlier Sආൾඍൺඇൺ (1971a) mentioned subtle differences between these lowland specimens and ones from the Great Plains, but not clear enough to describe an additional species. The known range of
|
||
<taxonomicName id="4C204D35FFC44B27B23BA79B17B9F82A" authorityName="Casey" authorityYear="1915" box="[557,675,1950,1973]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B23BA79B17B9F82A" box="[557,675,1950,1973]" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
extends southwards almost to the border between
|
||
<collectingCountry id="F3377626FFC44B27B229A7BB1768F84A" box="[575,626,1982,2005]" name="United States of America" pageId="62" pageNumber="287">USA</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3377626FFC44B27B2B7A7BB17EEF84A" box="[673,756,1982,2005]" name="Mexico" pageId="62" pageNumber="287">Mexico</collectingCountry>
|
||
, northwards to Lawrence in Kansas. To the east
|
||
<taxonomicName id="4C204D35FFC44B27B296A7DB17E2F86A" authorityName="Casey" authorityYear="1915" box="[640,760,2014,2037]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="strenuus">
|
||
<emphasis id="B954EAA4FFC44B27B296A7DB17E2F86A" box="[640,760,2014,2037]" italics="true" pageId="62" pageNumber="287">Q. strenuus</emphasis>
|
||
</taxonomicName>
|
||
is replaced by
|
||
<taxonomicName id="4C204D35FFC44B27B124A7FB14AAF78A" baseAuthorityName="Gravenhorst" baseAuthorityYear="1802" box="[306,432,2046,2069]" class="Insecta" family="Staphylinidae" genus="Quedius" kingdom="Animalia" order="Coleoptera" pageId="62" pageNumber="287" phylum="Arthropoda" rank="species" species="laticollis">
|
||
<emphasis id="B954EAA4FFC44B27B124A7FB14AAF78A" box="[306,432,2046,2069]" italics="true" pageId="62" pageNumber="287">Q. laticollis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |