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<document id="E708DA8DF75E1105C7FB37B83561467C" ID-DOI="10.11646/phytotaxa.73.1.7" ID-ISSN="1179-3163" ID-Zenodo-Dep="5066503" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625292153320" checkinUser="felipe" docAuthor="Rua, Gabriel H. &amp; Valls, José Francisco M." docDate="2012" docId="03898781DF7D110EFF15CBEEFA8E1DD4" docLanguage="en" docName="Phytotaxa.73.60-66.pdf" docOrigin="Phytotaxa 73 (1)" docSource="http://biotaxa.org/Phytotaxa/article/view/phytotaxa.73.1.7" docStyle="DocumentStyle:13F189B31CB4CCCE9197361A4E100104.3:Phytotaxa.2011-2013.journal_article" docStyleId="13F189B31CB4CCCE9197361A4E100104" docStyleName="Phytotaxa.2011-2013.journal_article" docStyleVersion="3" docTitle="Paspalum repandum G. H. Rua &amp; Valls 2012, comb. nov." docType="treatment" docVersion="2" lastPageNumber="65" masterDocId="FFB0FFF9DF79110BFF82CB79FFB91F55" masterDocTitle="On the taxonomic status of the genus Thrasyopsis (Poaceae, Panicoideae, Paspaleae): new combinations in Paspalum" masterLastPageNumber="66" masterPageNumber="60" pageNumber="64" updateTime="1698987767244" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="39B48BE067C260C9C864E6951A69F2EA">On the taxonomic status of the genus Thrasyopsis (Poaceae, Panicoideae, Paspaleae): new combinations in Paspalum</mods:title>
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<mods:namePart id="EAE96D21027094BDFE3907BF9650B312">Rua, Gabriel H.</mods:namePart>
<mods:affiliation id="C8748673F1E20E80DACB4AFF5049EC7E">Cátedra de Botánica Agrícola, Facultad de Agronomía, Universidad de Buenos Aires, Av. San Martín 4453, C 1417 DSE Buenos Aires, Argentina; E-mail: ruagabri @ agro. uba. ar</mods:affiliation>
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<mods:namePart id="7FB3FB686A2B420D1725A495515C1E03">Valls, José Francisco M.</mods:namePart>
<mods:affiliation id="1316B79C5AC2E5EB51D50F0BBFE2B30C">EMBRAPA Recursos Genéticos e Biotecnologia, Parque Estação Biológica, Final W- 5 Norte, Caixa Postal 02372, CEP 70770 - 900 Brasília, DF, Brazil; E-mail: valls @ cenargen. embrapa. br</mods:affiliation>
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<mods:title id="33CD15532ED36AA3DCD0C06F86D6B2A3">Phytotaxa</mods:title>
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<heading id="D0D781FBDF7D110FFF15CBEEFBA31FE1" bold="true" box="[151,1050,151,180]" fontSize="12" level="1" pageId="4" pageNumber="64" reason="1">
<treatmentCitationGroup id="AB3011B9DF7D110FFF15CBEEFBA31FE1" box="[151,1050,151,180]" pageId="4" pageNumber="64">
<emphasis id="B954EA85DF7D110FFF15CBEEFBA31FE1" bold="true" box="[151,1050,151,180]" pageId="4" pageNumber="64">
<treatmentCitation id="0A811086DF7D110FFF15CBEEFCC61FE1" author="Nees von Esenbeck, C. G." box="[151,895,151,180]" page="98" pageId="4" pageNumber="64" year="1829">
<taxonomicName id="4C204D14DF7D110FFF15CBEEFCC61FE1" ID-CoL="4DWP2" authority="(Nees 1829: 98) G. H. Rua &amp; Valls" authorityName="G. H. Rua &amp; Valls" authorityYear="2012" baseAuthorityName="Nees" baseAuthorityPageNumber="98" baseAuthorityYear="1829" box="[151,895,151,180]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum" status="comb. nov.">
<emphasis id="B954EA85DF7D110FFF15CBEEFE101FE6" bold="true" box="[151,425,151,179]" italics="true" pageId="4" pageNumber="64">Paspalum repandum</emphasis>
(
<bibRefCitation id="EFB14B66DF7D110FFE38CBEEFDC01FE1" author="Nees von Esenbeck, C. G." box="[442,633,151,180]" pageId="4" pageNumber="64" refId="ref4357" refString="Nees von Esenbeck, C. G. (1829) Agrostologia Brasiliensis, seu, Descriptio graminum in imperio Brasiliensi huc usque detectorum. In: Martius, C. F. P. (ed.) Flora Brasiliensis seu Enumeratio Plantarum 2 (1). J. G. Cotta, Stuttgart and Tubingen, 608 pp." type="book" year="1829">Nees 1829: 98</bibRefCitation>
) G. H. Rua &amp; Valls
</taxonomicName>
</treatmentCitation>
,
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<emphasis id="B954EA85DF7D110FFC0FCBEEFBAB1FE6" bold="true" box="[909,1042,151,179]" italics="true" pageId="4" pageNumber="64">comb. nov</emphasis>
.
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</paragraph>
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Basionym:—
<taxonomicName id="4C204D14DF7D110FFEA1CB99FD091FA2" ID-CoL="4CF2P" authority="Nees (1829: 98)" authorityName="Nees" authorityPageNumber="98" authorityYear="1829" box="[291,688,224,247]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFEA1CB99FE4D1FA2" box="[291,500,224,247]" italics="true" pageId="4" pageNumber="64">Panicum repandum</emphasis>
<bibRefCitation id="EFB14B66DF7D110FFE7FCB99FD091FA2" author="Nees von Esenbeck, C. G." box="[509,688,224,247]" pageId="4" pageNumber="64" refId="ref4357" refString="Nees von Esenbeck, C. G. (1829) Agrostologia Brasiliensis, seu, Descriptio graminum in imperio Brasiliensi huc usque detectorum. In: Martius, C. F. P. (ed.) Flora Brasiliensis seu Enumeratio Plantarum 2 (1). J. G. Cotta, Stuttgart and Tubingen, 608 pp." type="book" year="1829">Nees (1829: 98)</bibRefCitation>
</taxonomicName>
.
<taxonomicName id="4C204D14DF7D110FFD3FCB99FB7F1FA2" ID-CoL="56LJM" authority="(Nees) Parodi (1946: 297)" authorityName="Parodi" authorityPageNumber="297" authorityYear="1946" baseAuthorityName="Nees" box="[701,1222,224,247]" class="Liliopsida" family="Poaceae" genus="Thrasyopsis" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repanda">
<emphasis id="B954EA85DF7D110FFD3FCB99FC251FA2" box="[701,924,224,247]" italics="true" pageId="4" pageNumber="64">Thrasyopsis repanda</emphasis>
(Nees)
<bibRefCitation id="EFB14B66DF7D110FFC74CB99FB7F1FA2" author="Parodi, L. R." box="[1014,1222,224,247]" pageId="4" pageNumber="64" pagination="293 - 297" refId="ref4419" refString="Parodi, L. R. (1946) Nuevo genero de gramineas del Brasil. Boletin de la Sociedad Argentina de Botanica 1: 293 - 297." type="journal article" year="1946">Parodi (1946: 297)</bibRefCitation>
</taxonomicName>
. TYPE:—
<collectingCountry id="F3377607DF7D110FFAB4CB99FA211FA2" box="[1334,1432,224,247]" name="Brazil" pageId="4" pageNumber="64">BRAZIL</collectingCountry>
. “Brasilia australiore”, Barra do
<collectingRegion id="49E4F875DF7D110FFD90CA79FDF81E42" box="[530,577,256,279]" country="Brazil" name="Para" pageId="4" pageNumber="64">Pará</collectingRegion>
(as “Bra. do Paxa”), date unknown,
<emphasis id="B954EA85DF7D110FFC3CCA79FB1B1E42" box="[958,1186,256,279]" italics="true" pageId="4" pageNumber="64">Sellow B.1246. C.194</emphasis>
(
<typeStatus id="549B8835DF7D110FFB2DCA79FAAF1E42" box="[1199,1302,256,279]" pageId="4" pageNumber="64" type="holotype">Holotype</typeStatus>
: B [barcode] B 10 0249432 digital image!; isotypes: BAA [barcode BAA00000622] fragment ex B!,
<collectingCountry id="F3377607DF7D110FFB03CA59FB1A1E62" box="[1153,1187,288,311]" name="United States of America" pageId="4" pageNumber="64">US</collectingCountry>
[barcode
<collectingCountry id="F3377607DF7D110FFA93CA59FA881E62" box="[1297,1329,288,311]" name="United States of America" pageId="4" pageNumber="64">US</collectingCountry>
00139930 p.p.] fragment ex B, digital image!; probable isotypes: G [barcodes G00168437, G00168438 and G00168439] digital images!, K [barcodes K000643320 and K000643322] digital images!, LE, P [P02328829 and P02328833] digital images!,
<collectingCountry id="F3377607DF7D110FFEA5CAF9FEF01EC2" box="[295,329,384,407]" name="United States of America" pageId="4" pageNumber="64">US</collectingCountry>
[barcode
<collectingCountry id="F3377607DF7D110FFE33CAF9FE681EC2" box="[433,465,384,407]" name="United States of America" pageId="4" pageNumber="64">US</collectingCountry>
00139930 p.p.] fragment ex LE, digital image!, W [W0030638] digital image!).
</paragraph>
<paragraph id="8B9F3697DF7D110FFF15CABFFD541D2E" blockId="4.[151,1436,454,1988]" pageId="4" pageNumber="64">
Cespitose perennials with erect culms. Leaf sheaths hirsute, fibrous and persistent after senescence, leaf blades flat, flexuous, pale, silvery-hirsute. Racemes paired or solitary,
<quantity id="4CD89B72DF7D110FFC65CA94FBEF1D52" box="[999,1110,493,519]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="4.0" metricValueMin="1.5" pageId="4" pageNumber="64" unit="cm" value="2.75" valueMax="4.0" valueMin="1.5">1.54 cm</quantity>
long, with rachis narrowly winged, not covering the spikelets. Spikelets paired, subglabrous, lower glumes 57-nerved, up to ¾ the length of the spikelets; upper glumes obovate, cucullate, 11-nerved; lower floret usually male, the lower lemma 9-nerved, glume-like; upper floret bisexual.
</paragraph>
</subSubSection>
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<paragraph id="8B9F3697DF7D110FFF44C9FEFD791CDF" blockId="4.[151,1436,454,1988]" pageId="4" pageNumber="64">
<emphasis id="B954EA85DF7D110FFF44C9FEFCA81DF4" bold="true" box="[198,785,647,673]" pageId="4" pageNumber="64">Distribution, habitat, and conservation status:</emphasis>
―Endemic from the Brazilian states of
<collectingRegion id="49E4F875DF7D110FFB77C9F1FA2E1DF7" box="[1269,1431,648,674]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="64">Minas Gerais</collectingRegion>
,
<collectingRegion id="49E4F875DF7D110FFF15C9D7FEB71D9D" box="[151,270,686,712]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="64">São Paulo</collectingRegion>
(most ancient collections known from both states), and
<collectingRegion id="49E4F875DF7D110FFC2AC9D7FC451D9D" box="[936,1020,686,712]" country="Brazil" name="Parana" pageId="4" pageNumber="64">Paraná</collectingRegion>
. This species has been consistently collected near Ponta Grossa (state of
<collectingRegion id="49E4F875DF7D110FFDD6C9ACFD1F1DBA" box="[596,678,725,751]" country="Brazil" name="Parana" pageId="4" pageNumber="64">Paraná</collectingRegion>
) in the vicinity of Vila Velha (
<bibRefCitation id="EFB14B66DF7D110FFB91C9ACFB051DBA" author="Burman, A. G." box="[1043,1212,725,751]" pageId="4" pageNumber="64" pagination="101 - 116" refId="ref3666" refString="Burman, A. G. (1983) The genus Thrasyopsis Parodi (Gramineae). Phyton (Austria) 23: 101 - 116." type="journal article" year="1983">Burman 1983</bibRefCitation>
). Nevertheless, we have recently conducted intensive field collections in this area and only found a single tuft. Although the current conservation status of this species has not yet been established, it is very likely to be a threatened species, because of its extreme rarity. Flowering of
<taxonomicName id="4C204D14DF7D110FFD77C830FC511C37" authorityName="G. H. Rua &amp; Valls" authorityYear="2012" baseAuthorityName="Nees" baseAuthorityYear="1829" box="[757,1000,841,866]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFD77C830FC511C37" box="[757,1000,841,866]" italics="true" pageId="4" pageNumber="64">Paspalum repandum</emphasis>
</taxonomicName>
seems to be fire-dependent, since all known herbarium specimens are burned plants.
</paragraph>
<paragraph id="8B9F3697DF7D110FFF44C8EFFEC51B71" blockId="4.[151,1436,454,1988]" pageId="4" pageNumber="64">
<emphasis id="B954EA85DF7D110FFF44C8EFFDD11CE5" bold="true" box="[198,616,918,944]" pageId="4" pageNumber="64">Additional specimens examined:</emphasis>
<taxonomicName id="4C204D14DF7D110FFD07C8EEFCC41CE5" authorityName="G. H. Rua &amp; Valls" authorityYear="2012" baseAuthorityName="Nees" baseAuthorityYear="1829" box="[645,893,919,944]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFD07C8EEFCC41CE5" box="[645,893,919,944]" italics="true" pageId="4" pageNumber="64">Paspalum repandum</emphasis>
</taxonomicName>
.
<materialsCitation id="3B483CCADF7D110FFC10C8EFFE1A1CAB" collectingDate="2006-02-12" collectionCode="CEN" country="Brazil" elevation="915" latitude="-25.265556" location="Ponta Grossa" longLatPrecision="21" longitude="-49.980556" pageId="4" pageNumber="64" specimenCount="1" stateProvince="Parana">
<collectingCountry id="F3377607DF7D110FFC10C8EFFBB91CE5" box="[914,1024,918,944]" name="Brazil" pageId="4" pageNumber="64">BRAZIL</collectingCountry>
.
<collectingRegion id="49E4F875DF7D110FFB93C8EFFBDC1CE5" box="[1041,1125,918,944]" country="Brazil" name="Parana" pageId="4" pageNumber="64">Paraná</collectingRegion>
:
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, Faz. Vila Velha, BR-367, km 520, borda de barranco com arenito, elev.
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<elevation id="000DD1A4DF7D110FFCFAC8C4FC7F1C82" box="[888,966,957,983]" metricMagnitude="2" metricUnit="m" metricValue="9.15" pageId="4" pageNumber="64" unit="m" value="915.0">915 m</elevation>
</quantity>
,
<geoCoordinate id="EE145050DF7D110FFC51C8C4FBE71C82" box="[979,1118,957,983]" degrees="25" direction="south" minutes="15" orientation="latitude" pageId="4" pageNumber="64" precision="15" seconds="56" value="-25.265556">25º1556”S</geoCoordinate>
,
<geoCoordinate id="EE145050DF7D110FFBE9C8C4FAB91C82" box="[1131,1280,957,983]" degrees="49" direction="west" minutes="58" orientation="longitude" pageId="4" pageNumber="64" precision="15" seconds="50" value="-49.980556">49º5850”W</geoCoordinate>
,
<date id="FF9E1057DF7D110FFA8EC8C4FF6F1CAB" pageId="4" pageNumber="64" value="2006-02-12">
<collectingDate id="EFDAE9BFDF7D110FFA8EC8C4FF6F1CAB" pageId="4" pageNumber="64" value="2006-02-12">12 February 2006</collectingDate>
</date>
,
<emphasis id="B954EA85DF7D110FFF61C89DFEFE1CA8" box="[227,327,996,1021]" italics="true" pageId="4" pageNumber="64">Rua s.n.</emphasis>
(
<collectionCode id="ED31AE52DF7D110FFEDAC89DFE231CAB" box="[344,410,996,1022]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15624" lsid="urn:lsid:biocol.org:col:15624" name="EMBRAPA Recursos Geneticos e Biotecnologia - CENARGEN" pageId="4" pageNumber="64" type="Herbarium">CEN</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CCADF7D110FFE32C89DFBB51CAB" box="[432,1036,996,1022]" collectingDate="2010-11-13" collectionCode="HUPG" country="Brazil" pageId="4" pageNumber="64" specimenCount="1" stateProvince="Parana">
id.,
<date id="FF9E1057DF7D110FFE62C89DFD741CAB" box="[480,717,996,1022]" pageId="4" pageNumber="64" value="2010-11-13">
<collectingDate id="EFDAE9BFDF7D110FFE62C89DFD741CAB" box="[480,717,996,1022]" pageId="4" pageNumber="64" value="2010-11-13">13 November 2010</collectingDate>
</date>
,
<emphasis id="B954EA85DF7D110FFD59C89DFCCD1CA8" box="[731,884,996,1021]" italics="true" pageId="4" pageNumber="64">Carmo et al.</emphasis>
22 (
<collectionCode id="ED31AE52DF7D110FFC2EC89DFBBC1CAB" box="[940,1029,996,1022]" country="Brazil" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15109" lsid="urn:lsid:biocol.org:col:15109" name="State University of Ponta Grossa" pageId="4" pageNumber="64" type="Herbarium">HUPG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B483CCADF7D110FFB98C89DFEC11B71" collectingDate="1952-11-26" collectionCode="BAA" country="Brazil" location="Vila Velha" pageId="4" pageNumber="64" specimenCount="1" stateProvince="Parana">
<location id="8EFF604CDF7D110FFB98C89DFB241CAB" LSID="urn:lsid:plazi:treatment:03898781DF7D110EFF15CBEEFA8E1DD4:8EFF604CDF7D110FFB98C89DFB241CAB" box="[1050,1181,996,1022]" country="Brazil" name="Vila Velha" pageId="4" pageNumber="64" stateProvince="Parana">Vila Velha</location>
,
<date id="FF9E1057DF7D110FFB28C89DFA2E1CAB" box="[1194,1431,996,1022]" pageId="4" pageNumber="64" value="1952-11-26">
<collectingDate id="EFDAE9BFDF7D110FFB28C89DFA2E1CAB" box="[1194,1431,996,1022]" pageId="4" pageNumber="64" value="1952-11-26">26 November 1952</collectingDate>
</date>
,
<emphasis id="B954EA85DF7D110FFF15CF73FEA21B76" box="[151,283,1034,1059]" italics="true" pageId="4" pageNumber="64">Araujo 174</emphasis>
(
<collectionCode id="ED31AE52DF7D110FFEA9CF73FED61B71" box="[299,367,1034,1060]" country="Argentina" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15008" lsid="urn:lsid:biocol.org:col:15008" name="Universidad de Buenos Aires" pageId="4" pageNumber="64" type="Herbarium">BAA</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B9F3697DF7D110FFF44CF49FAE819DA" blockId="4.[151,1436,454,1988]" pageId="4" pageNumber="64">
<emphasis id="B954EA85DF7D110FFF44CF49FE641B1F" bold="true" box="[198,477,1072,1098]" pageId="4" pageNumber="64">Nomenclatural notes:</emphasis>
―There are two descriptions for this species published in 1829:
<taxonomicName id="4C204D14DF7D110FFAB6CF48FEE31B27" authority="Nees" authorityName="Nees" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFAB6CF48FEAA1B24" italics="true" pageId="4" pageNumber="64">Panicum repandum</emphasis>
Nees
</taxonomicName>
in Martius,
<emphasis id="B954EA85DF7D110FFE78CF21FBC21B24" box="[506,1147,1112,1137]" italics="true" pageId="4" pageNumber="64">Flora Brasiliensis seu Enumeratio Plantarum 2(1)</emphasis>
: 9899, and
<taxonomicName id="4C204D14DF7D110FFAACCF21FE011BCD" authority="Nees ex Trin." authorityName="Trin." class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFAACCF21FEB41BCD" italics="true" pageId="4" pageNumber="64">Panicum repandum</emphasis>
Nees ex Trin.
</taxonomicName>
,
<emphasis id="B954EA85DF7D110FFE46CF06FD551BCD" box="[452,748,1151,1176]" italics="true" pageId="4" pageNumber="64">Species Graminum 2(13)</emphasis>
: t. 150. The first work was published between
<date id="FF9E1057DF7D110FFA9ECF07FEA81BEA" pageId="4" pageNumber="64" value="1829-03" valueMax="1829-06-31" valueMin="1829-03-00">March and June 1829</date>
, whereas the exact date of publication of the part 13 of
<emphasis id="B954EA85DF7D110FFC1BCFDCFBCE1BEB" box="[921,1143,1189,1214]" italics="true" pageId="4" pageNumber="64">Species Graminum</emphasis>
is still unknown (
<bibRefCitation id="EFB14B66DF7D110FFACACFDCFEF51BB3" author="Stafleu, F. A. &amp; Cowan, R. S." pageId="4" pageNumber="64" refId="ref4625" refString="Stafleu, F. A. &amp; Cowan, R. S. (1986) Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types (TL 2) 6. Regnum vegetabile. Scheltema &amp; Holkema, Utrecht &amp; Bohn, 926 pp." type="book" year="1986">Stafleu &amp; Cowan 1986</bibRefCitation>
), so that it is impossible to know which of both publications is earlier. The original material on which the species was described was collected by Sellow in southern
<collectingCountry id="F3377607DF7D110FFC66CF8BFB941A59" box="[996,1069,1266,1292]" name="Brazil" pageId="4" pageNumber="64">Brazil</collectingCountry>
and originally deposited at B, where it was studied by Nees [
<emphasis id="B954EA85DF7D110FFD81CE60FA951A67" box="[515,1324,1305,1330]" italics="true" pageId="4" pageNumber="64">“Habitat in Brasilia australiori. (Sellow. Vidi in Herb. Reg. Berol.)”</emphasis>
, cf.
<bibRefCitation id="EFB14B66DF7D110FFAE3CE60FEBB1A0F" author="Nees von Esenbeck, C. G." pageId="4" pageNumber="64" refId="ref4357" refString="Nees von Esenbeck, C. G. (1829) Agrostologia Brasiliensis, seu, Descriptio graminum in imperio Brasiliensi huc usque detectorum. In: Martius, C. F. P. (ed.) Flora Brasiliensis seu Enumeratio Plantarum 2 (1). J. G. Cotta, Stuttgart and Tubingen, 608 pp." type="book" year="1829">Nees 1829: 99</bibRefCitation>
]. The specimen still conserved at B [barcode B 10 0249432] has a label in Sellows handwriting with the inscription “Bra. do Paxa, B 1246, c.194”, without indication of date. There is no place in
<collectingCountry id="F3377607DF7D110FFB6FCE1FFA8D1AD5" box="[1261,1332,1382,1408]" name="Brazil" pageId="4" pageNumber="64">Brazil</collectingCountry>
with the name “Paxa”, and probably there is a handwriting mistake in Sellow´s label. Between 1814 and 1831, Sellow made six collection trips in
<collectingCountry id="F3377607DF7D110FFE6BCECDFD8F1A9B" box="[489,566,1460,1486]" name="Brazil" pageId="4" pageNumber="64">Brazil</collectingCountry>
. In the third trip across
<collectingRegion id="49E4F875DF7D110FFCDACECDFC441A9B" box="[856,1021,1460,1486]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="64">Minas Gerais</collectingRegion>
,
<collectingRegion id="49E4F875DF7D110FFB88CECDFB3D1A9B" box="[1034,1156,1460,1486]" country="Brazil" name="Sao Paulo" pageId="4" pageNumber="64">São Paulo</collectingRegion>
and
<collectingRegion id="49E4F875DF7D110FFB42CECDFACB1A9B" box="[1216,1394,1460,1486]" country="Brazil" name="Rio de Janeiro" pageId="4" pageNumber="64">Rio de Janeiro</collectingRegion>
, he double-numbered the specimens from about 2,500 locations under the letters B and c (
<bibRefCitation id="EFB14B66DF7D110FFB15CEA3FA931AA1" author="Urban, I." box="[1175,1322,1498,1524]" pageId="4" pageNumber="64" pagination="177 - 198" refId="ref4755" refString="Urban, I. (1893) Biographische Skizzen. Botanische Jahrbucher 17: 177 - 198." type="journal article" year="1893">Urban 1893</bibRefCitation>
), thus the
<typeStatus id="549B8835DF7D110FFF15CD78FF72194E" box="[151,203,1537,1563]" pageId="4" pageNumber="64">type</typeStatus>
material of
<taxonomicName id="4C204D14DF7D110FFEE0CD78FE43194F" box="[354,506,1537,1562]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFEE0CD78FE43194F" box="[354,506,1537,1562]" italics="true" pageId="4" pageNumber="64">P. repandum</emphasis>
</taxonomicName>
belongs to this collection. The itinerary of Sellows trips detailed by
<bibRefCitation id="EFB14B66DF7D110FFAD2CD78FEE01917" author="Urban, I." pageId="4" pageNumber="64" pagination="1 - 154" refId="ref4773" refString="Urban, I. (1906) Vitae itineraque collectorum botanicorum. In: Urban, I. (ed.), Flora Brasiliensis 1 (1). Oldenburg, Munchen, pp. 1 - 154." type="book chapter" year="1906">Urban (1906: 105-112)</bibRefCitation>
casts light on the actual locality and date of collection: Sellow was in
<date id="FF9E1057DF7D110FFB28CD51FADE1917" box="[1194,1383,1576,1602]" pageId="4" pageNumber="64" value="1818-12">
December
<quantity id="4CD89B72DF7D110FFAAECD51FADE1917" box="[1324,1383,1576,1602]" metricMagnitude="1" metricUnit="m" metricValue="4.61772" pageId="4" pageNumber="64" unit="in" value="1818.0">1818</quantity>
</date>
in a place named “Barra do
<collectingRegion id="49E4F875DF7D110FFE29CD37FE5D193D" box="[427,484,1614,1640]" country="Brazil" name="Para" pageId="4" pageNumber="64">Pará</collectingRegion>
”, i.e. the mouth of the
<collectingRegion id="49E4F875DF7D110FFD6FCD37FC98193D" box="[749,801,1614,1640]" country="Brazil" name="Para" pageId="4" pageNumber="64">Pará</collectingRegion>
river in the São Francisco river, near the present town of Pompeu, in the state of
<collectingRegion id="49E4F875DF7D110FFE49CD0CFDD219DA" box="[459,619,1653,1679]" country="Brazil" name="Minas Gerais" pageId="4" pageNumber="64">Minas Gerais</collectingRegion>
, and this is probably the locality where the plant was collected.
</paragraph>
<paragraph id="8B9F3697DF7D110FFF44CDE5FC111891" blockId="4.[151,1436,454,1988]" pageId="4" pageNumber="64">
Trinius, who explicitly attributed the species to Nees, based his description and illustration on a sheet deposited at LE (LE-TRIN-0922.01), and labelled “
<taxonomicName id="4C204D14DF7D110FFC91CDBAFBF81989" authority="Nees" authorityName="Nees" box="[787,1089,1730,1756]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFC91CDBAFC441989" box="[787,1021,1731,1756]" italics="true" pageId="4" pageNumber="64">Panicum repandum</emphasis>
Nees
</taxonomicName>
ab Ess., Sellow, Brasilia, de Schlechtendal 27” (R. Soreng, pers. comm.). This specimen (not seen) is presumably a duplicate of the original Sellows material sent to Trinius by Schlechtendal. The hypothesis of Soreng that this specimen belongs to a collection of cultivated material from seeds of the Sellow plant (pers. comm., see also the entry for
<taxonomicName id="4C204D14DF7D110FFF40CC24FEEE1823" box="[194,343,1885,1910]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="4" pageNumber="64" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7D110FFF40CC24FEEE1823" box="[194,343,1885,1910]" italics="true" pageId="4" pageNumber="64">P. repandum</emphasis>
</taxonomicName>
in Tropicos.org (
<bibRefCitation id="EFB14B66DF7D110FFDA5CC24FC0F1822" author="Missouri Botanical Garden" box="[551,950,1885,1911]" pageId="4" pageNumber="64" refId="ref4320" refString="Missouri Botanical Garden (2012) Tropicos. org, Missouri Botanical Garden. Available from: http: // www. tropicos. org / Name / 25522634 (Accessed: 21 June 2012)." type="url" year="2012">Missouri Botanical Garden 2012</bibRefCitation>
): “
<emphasis id="B954EA85DF7D110FFC5ACC24FDB818C8" italics="true" pageId="4" pageNumber="64">
F. Sellow; cult. coll. de Schlechtendal 27; Brasilia, cult. in
<collectingCountry id="F3377607DF7D110FFE11CCFDFDB818C8" box="[403,513,1924,1949]" name="Germany" pageId="4" pageNumber="64">Germany</collectingCountry>
</emphasis>
”) seems improbable, because it is a plant difficult to bring into flower under cultivation since flowering seems to be fire-dependent (see above).
</paragraph>
<paragraph id="8B9F3697DF7C110EFF44CBE1FBB21FAA" blockId="5.[151,1436,152,642]" pageId="5" pageNumber="65">
As exact publication dates of Trinius work are uncertain (
<bibRefCitation id="EFB14B66DF7C110EFC08CBE1FB511FE7" author="Stafleu, F. A. &amp; Cowan, R. S." box="[906,1256,152,178]" pageId="5" pageNumber="65" refId="ref4625" refString="Stafleu, F. A. &amp; Cowan, R. S. (1986) Taxonomic literature: a selective guide to botanical publications and collections with dates, commentaries and types (TL 2) 6. Regnum vegetabile. Scheltema &amp; Holkema, Utrecht &amp; Bohn, 926 pp." type="book" year="1986">Stafleu &amp; Cowan 1986: 495</bibRefCitation>
), we prefer, in agreement with Soreng (comm. pers.) to take the Nees description (and the corresponding
<typeStatus id="549B8835DF7C110EFB52CBC7FA8E1F8D" box="[1232,1335,190,216]" pageId="5" pageNumber="65" type="holotype">holotype</typeStatus>
at B) as the valid one, and consider
<taxonomicName id="4C204D14DF7C110EFE5BCB9CFCDD1FAA" authority="Nees ex Trin." authorityName="Trin." box="[473,868,229,255]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="65" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7C110EFE5BCB9CFD071FAB" box="[473,702,229,254]" italics="true" pageId="5" pageNumber="65">Panicum repandum</emphasis>
Nees ex Trin.
</taxonomicName>
as an isonym.
</paragraph>
<paragraph id="8B9F3697DF7C110EFF44CA75FA8E1DD4" blockId="5.[151,1436,152,642]" pageId="5" pageNumber="65">
Further duplicates of the Sellows collection were apparently distributed to several herbaria (G, K, P, W) by Schlechtendal, each specimen identified as
<taxonomicName id="4C204D14DF7C110EFD48CA4BFC4E1E1E" authority="Nees" authorityName="Nees" box="[714,1015,306,331]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="65" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7C110EFD48CA4BFC4E1E1E" box="[714,1015,306,331]" italics="true" pageId="5" pageNumber="65">Panicum repandum Nees</emphasis>
</taxonomicName>
in Schlechtendal handwriting on a label printed with the inscription “
<emphasis id="B954EA85DF7C110EFDC1CA20FB2B1E27" box="[579,1170,345,370]" italics="true" pageId="5" pageNumber="65">Herb. Reg. Berolinense. Brasilia. Sellow legit.”</emphasis>
, with no reference to numbering nor locality. On the other hand, fragments at BAA and
<collectingCountry id="F3377607DF7C110EFC26CAF9FC701ECF" box="[932,969,384,410]" name="United States of America" pageId="5" pageNumber="65">US</collectingCountry>
were taken directly from B by Lorenzo R. Parodi and M. Agnes Chase respectively, who transcribed the original Sellows label. Further, a specimen at HAL that was indicated erroneously as the
<typeStatus id="549B8835DF7C110EFD4ECAB4FCB91EB2" box="[716,768,461,487]" pageId="5" pageNumber="65">type</typeStatus>
of
<taxonomicName id="4C204D14DF7C110EFCA9CAB4FBA01EB3" box="[811,1049,461,486]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="65" phylum="Tracheophyta" rank="species" species="repandum">
<emphasis id="B954EA85DF7C110EFCA9CAB4FBA01EB3" box="[811,1049,461,486]" italics="true" pageId="5" pageNumber="65">Panicum repandum</emphasis>
</taxonomicName>
(barcode HAL0063595, digital image!) corresponds to the species
<taxonomicName id="4C204D14DF7C110EFDB4CA8DFB381D5B" authority="Nees ex Trinius (1826: 121)" authorityName="Trinius" authorityPageNumber="121" authorityYear="1826" box="[566,1153,500,526]" class="Liliopsida" family="Poaceae" genus="Paspalum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="65" phylum="Tracheophyta" rank="species" species="erianthum">
<emphasis id="B954EA85DF7C110EFDB4CA8DFC921D58" box="[566,811,500,525]" italics="true" pageId="5" pageNumber="65">Paspalum erianthum</emphasis>
Nees ex
<bibRefCitation id="EFB14B66DF7C110EFC1BCA8DFB381D5B" author="Trinius, C. B." box="[921,1153,500,526]" pageId="5" pageNumber="65" refId="ref4678" refString="Trinius, C. B. (1826) De Graminibus Paniceis. Dissertatio botanica altera. Impensis Academiae Imperialis Scientiarum, Saint Petersburg, 289 pp." type="book" year="1826">Trinius (1826: 121)</bibRefCitation>
</taxonomicName>
. This specimen has two labels in Schlechtendal handwriting, probably dated after Schlechtendal moved to Halle in 1833: one of the labels includes the proper identification followed by a latin description beginning with the expression
<emphasis id="B954EA85DF7C110EFF15C911FE491DD4" box="[151,496,616,641]" italics="true" pageId="5" pageNumber="65">
<taxonomicName id="4C204D14DF7C110EFF15C911FE601DD4" box="[151,473,616,641]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="65" phylum="Tracheophyta" rank="subSpecies" species="repandum" subSpecies="spicam">Panicum repandum spicam</taxonomicName>
</emphasis>
, the other label reads “
<emphasis id="B954EA85DF7C110EFC82C911FA8E1DD4" box="[768,1335,616,641]" italics="true" pageId="5" pageNumber="65">
<taxonomicName id="4C204D14DF7C110EFC82C911FC5F1DD4" box="[768,998,616,641]" class="Liliopsida" family="Poaceae" genus="Panicum" kingdom="Plantae" order="Poales" pageId="5" pageNumber="65" phylum="Tracheophyta" rank="species" species="repandum">Panicum repandum</taxonomicName>
N. ab E. Sellow. Brasilia…”
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>