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<document id="11D71579E6D96AEEF0183EE91F25BD71" ID-CLB-Dataset="21868" ID-DOI="10.5852/ejt.2016.237" ID-GBIF-Dataset="9e616425-484c-4252-ac56-38de6af08d4e" ID-ISSN="2118-9773" ID-Zenodo-Dep="10076742" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1590437334157" checkinUser="valdenar" docAuthor="Ačanski, Jelena, Vujić, Ante, Djan, Mihajla, Vidaković, Dragana Obreht, Ståhls, Gunilla &amp; Radenković, Snežana" docDate="2016" docId="03851D0F3215FFAC7E8D301EB23FFE2E" docLanguage="en" docName="ejt-2016-237.pdf.imf" docOrigin="European Journal of Taxonomy 237" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Merodon megavidus Vujic &amp; Radenkovic 2016, sp. nov." docType="treatment" docUuid="B2016D80-A7ED-4958-AE91-EE5E6A8C7C51" docUuidSource="ZooBank" docVersion="9" lastPageNumber="12" masterDocId="FFBC65773210FFA17F7F323AB37FFFD7" masterDocTitle="Defining species boundaries in the Merodon avidus complex (Diptera, Syrphidae) using integrative taxonomy, with the description of a new species" masterLastPageNumber="25" masterPageNumber="1" pageNumber="6" updateTime="1699764339099" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2B9150579FAB78D6D63898F9E019FFFC">Defining species boundaries in the Merodon avidus complex (Diptera, Syrphidae) using integrative taxonomy, with the description of a new species</mods:title>
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<mods:namePart id="DEA37296E0EB7DB8094A96CBAA600581">Ačanski, Jelena</mods:namePart>
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<mods:affiliation id="219BDB2DF0418B4418EDA8F000B99569">University of Novi Sad, BioSense Institute - Research Institute for Information Technologies in Biosystems, Trg Dr Zorana Đinđića 1, 21000 Novi Sad, Serbia. &amp; urn: lsid: zoobank. org: author: F 02841 B 0 - AC 6 B- 4 FC 4 - A 64 A-D 13 A 3355 B 079 &amp; Corresponding author: acanskijelena @ gmail. com</mods:affiliation>
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<mods:affiliation id="B6F14B91FC822188DB4CE884C1B9EE53">Department of Biology and Ecology, Faculty of Sciences, University of Novi Sad, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia. &amp; Email: ante. vujic @ dbe. uns. ac. rs &amp; urn: lsid: zoobank. org: author: A 20 D 5863 - CF 18 - 4 BF 7 - BB 68 - 0 DA 75 D 34 B 7 A 8</mods:affiliation>
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<mods:namePart id="862F618D50A671FF8E63ABF8FD3E98AB">Djan, Mihajla</mods:namePart>
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<mods:namePart id="B1437F8C91FB77F9BDA51A69DBFD2564">Vidaković, Dragana Obreht</mods:namePart>
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<mods:affiliation id="9882FE0CC66690DD58705DEC422F34C7">Email: dragana. obreht @ dbe. uns. ac. rs &amp; urn: lsid: zoobank. org: author: 75 E 52 F 0 E-EEDC- 4 EF 1 - 907 B- 9202 DBDEAC 9 F &amp; Department of Biology and Ecology, Faculty of Sciences, University of Novi Sad, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<mods:namePart id="691EC840FBBC9DB3F3F1422FF02A11A7">Ståhls, Gunilla</mods:namePart>
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<mods:affiliation id="8D192631544A227473CAAAAD4FD41CD2">Zoology Unit, Finnish Museum of Natural History, PO Box 17, 00014 University of Helsinki, Finland. &amp; Email: gunilla. stahls @ helsinki. fi &amp; urn: lsid: zoobank. org: author: D 19 B 1262 - 503 A- 48 E 2 - 91 AE-F 99 D 0 F 2 EA 3 A 3</mods:affiliation>
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<mods:namePart id="7DD88A93BB706E204780DABEF66914C6">Radenković, Snežana</mods:namePart>
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<mods:affiliation id="14295CAB1BF8DDF9B4FA3194BCAAF6A0">Email: snezana. radenkovic @ dbe. uns. ac. rs &amp; urn: lsid: zoobank. org: author: 26 DF 35 D 9 - 55 FA- 4485 - 8 E 8 C-C 1 F 90 EFE 1036 &amp; Department of Biology and Ecology, Faculty of Sciences, University of Novi Sad, Trg Dositeja Obradovića 2, 21000 Novi Sad, Serbia.</mods:affiliation>
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<taxonomicName id="4C2CD79A3215FFA47E8D301EB09FFDE9" ID-CoL="9HJJ2" authority="Vujic &amp; Radenkovic" authorityName="Vujic &amp; Radenkovic" authorityYear="2016" box="[498,992,548,575]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3215FFA47E8D301EB196FDE8" bold="true" box="[498,745,548,575]" italics="true" pageId="5" pageNumber="6">Merodon megavidus</emphasis>
Vujić &amp; Radenković
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<paragraph id="8B93AC193215FFA47DC33051B008FD51" blockId="5.[361,1227,548,646]" box="[700,887,619,646]" pageId="5" pageNumber="6">
<figureCitation id="1317B09C3215FFA47DC33051B07BFD52" box="[700,772,619,645]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="5" pageNumber="6">Figs 2</figureCitation>
C–D, 3–5
</paragraph>
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<subSubSection id="C336FF923215FFA47FC23096B71AFBD8" pageId="5" pageNumber="6" type="diagnosis">
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<emphasis id="B958700B3215FFA47FC23096B249FD11" bold="true" box="[189,310,684,710]" pageId="5" pageNumber="6">Diagnosis</emphasis>
</heading>
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<paragraph id="8B93AC193215FFA47FC230E1B71AFBD8" blockId="5.[189,1399,731,1039]" pageId="5" pageNumber="6">
Medium- to large-sized species(
<quantity id="4CD401FC3215FFA47D5230E1B1D6FD22" box="[557,681,731,757]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="1.8" metricValueMin="1.3" pageId="5" pageNumber="6" unit="mm" value="15.5" valueMax="18.0" valueMin="13.0">13–18mm</quantity>
);black mesoscutum with four white microtrichose longitudinal stripes; tapering orange and black abdomen with white, transverse, microtrichose bands on tergites 2–4 (exceptionally without bands on tergite 2); tarsi reddish-orange dorsally; hind femur medium wide and slightly curved (
<figureCitation id="1317B09C3215FFA47EC8317EB281FC88" box="[439,510,836,863]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[208,1380,727,1927]" captionTargetId="figure@6.[189,1399,707,1948]" captionTargetPageId="6" captionText="Fig. 3. Hind leg, lateral view. A–B. Merodon avidus (Rossi, 1790). A. Ƌ. B. ♀. — C–D. M. megavidus Vujić &amp; Radenković sp. nov. C. Ƌ. D. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115163" httpUri="https://zenodo.org/record/10115163/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
C–D), with very short pile posteroventrally. 
<taxonomicName id="4C2CD79A3215FFA47B66317FB236FC55" authority="Vujic &amp; Radenkovic" authorityName="Vujic &amp; Radenkovic" authorityYear="2016" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3215FFA47B66317FB674FC88" box="[1049,1291,837,863]" italics="true" pageId="5" pageNumber="6">Merodon megavidus</emphasis>
Vujić &amp; Radenković
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<taxonomicNameLabel id="A26BCD703215FFA47E2E3152B2D2FC55" box="[337,429,872,898]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
belongs to the 
<taxonomicName id="4C2CD79A3215FFA47D1C3152B1CFFC55" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[611,688,872,898]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3215FFA47D1C3152B1CFFC55" box="[611,688,872,898]" italics="true" pageId="5" pageNumber="6">avidus</emphasis>
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complex (male genitalia in all species identical in shape, as on 
<figureCitation id="1317B09C3215FFA47F9B31B1B253FC72" box="[228,300,907,933]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="5" pageNumber="6">Fig. 2</figureCitation>
). 
<taxonomicName id="4C2CD79A3215FFA47E3B31B1B14AFC72" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[324,565,907,933]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3215FFA47E3B31B1B14AFC72" box="[324,565,907,933]" italics="true" pageId="5" pageNumber="6">Merodon megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703215FFA47D4031B1B1E4FC72" box="[575,667,907,933]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
can be separated from the other members of the complex by larger size, golden body pile, bright orange colour of the pale parts of legs and extremely short pile on hind femur (
<figureCitation id="1317B09C3215FFA47E3231EBB2F0FC3C" box="[333,399,977,1003]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[208,1380,727,1927]" captionTargetId="figure@6.[189,1399,707,1948]" captionTargetPageId="6" captionText="Fig. 3. Hind leg, lateral view. A–B. Merodon avidus (Rossi, 1790). A. Ƌ. B. ♀. — C–D. M. megavidus Vujić &amp; Radenković sp. nov. C. Ƌ. D. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115163" httpUri="https://zenodo.org/record/10115163/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
C–D). These characteristics contrast with other species from the complex, which have yellow to grayish pale body pile and longer pile on the hind femur (
<figureCitation id="1317B09C3215FFA47CA931CEB765FBD8" box="[982,1050,1012,1039]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[208,1380,727,1927]" captionTargetId="figure@6.[189,1399,707,1948]" captionTargetPageId="6" captionText="Fig. 3. Hind leg, lateral view. A–B. Merodon avidus (Rossi, 1790). A. Ƌ. B. ♀. — C–D. M. megavidus Vujić &amp; Radenković sp. nov. C. Ƌ. D. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115163" httpUri="https://zenodo.org/record/10115163/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
A–B).
</paragraph>
</subSubSection>
<subSubSection id="C336FF923215FFA47FC2360CB2FDFB75" pageId="5" pageNumber="6" type="etymology">
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<emphasis id="B958700B3215FFA47FC2360CB23BFB87" bold="true" box="[189,324,1078,1104]" pageId="5" pageNumber="6">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC193215FFA47FC2365EB2FDFB75" blockId="5.[189,1398,1124,1186]" pageId="5" pageNumber="6">
The name 
<taxonomicName id="4C2CD79A3215FFA47E49365FB2CCFBA8" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[310,435,1125,1151]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="megavidus">
<emphasis id="B958700B3215FFA47E49365FB2CCFBA8" box="[310,435,1125,1151]" italics="true" pageId="5" pageNumber="6">megavidus</emphasis>
</taxonomicName>
refers to the large size (Greek word 
<emphasis id="B958700B3215FFA47C29365FB0DEFBA9" box="[854,929,1125,1150]" italics="true" pageId="5" pageNumber="6">megas</emphasis>
means “large”) and great similarity with 
<taxonomicName id="4C2CD79A3215FFA47FC236B2B203FB75" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[189,380,1160,1186]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3215FFA47FC236B2B203FB75" box="[189,380,1160,1186]" italics="true" pageId="5" pageNumber="6">Merodon avidus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C336FF923215FFA77FC236F3B774FD51" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6" type="materials_examined">
<paragraph id="8B93AC193215FFA47FC236F3B213FB34" blockId="5.[189,364,1225,1251]" box="[189,364,1225,1251]" pageId="5" pageNumber="6">
<heading id="D0DB1B753215FFA47FC236F3B213FB34" bold="true" box="[189,364,1225,1251]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
<emphasis id="B958700B3215FFA47FC236F3B213FB34" bold="true" box="[189,364,1225,1251]" pageId="5" pageNumber="6">Type material</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC193215FFA47F9536C2B223FAC5" blockId="5.[189,1321,1272,1334]" box="[234,348,1272,1298]" pageId="5" pageNumber="6">
<emphasis id="B958700B3215FFA47F9536C2B223FAC5" bold="true" box="[234,348,1272,1298]" pageId="5" pageNumber="6">
<typeStatus id="549712BB3215FFA47F9536C2B223FAC5" box="[234,348,1272,1298]" pageId="5" pageNumber="6" type="holotype">Holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="8B93AC193215FFA77FC23721B218FEF2" blockId="5.[189,1321,1272,1334]" lastBlockId="6.[189,1399,267,646]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<materialsCitation id="3B44A6443215FFA77FC23721B218FEF2" ID-GBIF-Occurrence="2625466301" collectingDate="2015-07-26" collectionCode="FSUNS" collectorName="A. Vujic &amp; S. Radenkovic" country="Greece" lastPageId="6" lastPageNumber="7" location="Lesvos" municipality="Petrounta" pageId="5" pageNumber="6" specimenCode="FSUNS 10132" specimenCount="1" typeStatus="paratype">
<collectingCountry id="F33BEC893215FFA47FC23721B24BFAE2" box="[189,308,1307,1333]" name="Greece" pageId="5" pageNumber="6">GREECE</collectingCountry>
: Ƌ, 
<location id="8EF3FAC23215FFA47E1C3721B2C6FAE2" LSID="urn:lsid:plazi:treatment:03851D0F3215FFAC7E8D301EB23FFE2E:8EF3FAC23215FFA47E1C3721B2C6FAE2" box="[355,441,1307,1333]" country="Greece" municipality="Petrounta" name="Lesvos" pageId="5" pageNumber="6">Lesvos</location>
, 
<collectingMunicipality id="6BF736633215FFA47EBA3721B146FAE2" box="[453,569,1307,1333]" pageId="5" pageNumber="6">Petrounta</collectingMunicipality>
, 
<date id="FF928AD93215FFA47D3B3721B1A7FAE1" box="[580,728,1307,1334]" pageId="5" pageNumber="6" value="2015-07-26">
<collectingDate id="EFD673313215FFA47D3B3721B1A7FAE1" box="[580,728,1307,1334]" pageId="5" pageNumber="6" value="2015-07-26">
<date id="FF928AD93215FFA47D3B3721B1ECFAE2" box="[580,659,1307,1333]" day="26" month="07" monthName="Jul" pageId="5" pageNumber="6" year="1926">
26 
<date id="FF928AD93215FFA47D163721B1ECFAE2" box="[617,659,1307,1333]" month="07" monthName="Jul" pageId="5" pageNumber="6" year="2015">Jul.</date>
</date>
2015
</collectingDate>
</date>
, leg. 
<collectorName id="26D9C9CF3215FFA47C6B3721B009FAE2" box="[788,886,1307,1333]" pageId="5" pageNumber="6">A. Vujić</collectorName>
and 
<collectorName id="26D9C9CF3215FFA47CCF3721B724FAE2" box="[944,1115,1307,1333]" pageId="5" pageNumber="6">S. Radenković</collectorName>
(
<specimenCode id="DB8A04623215FFA47B153721B663FAE2" box="[1130,1308,1307,1333]" collectionCode="FSUNS" country="0" httpUri="http://grbio.org/cool/zzce-zzpb" name="Faculty of Science, The University of Novi Sad" pageId="5" pageNumber="6">FSUNS 10132</specimenCode>
). 
<emphasis id="B958700B3216FFA77F953331B218FEF2" bold="true" box="[234,359,267,293]" pageId="6" pageNumber="7">
<typeStatus id="549712BB3216FFA77F953331B218FEF2" box="[234,359,267,293]" pageId="6" pageNumber="7" type="paratype">Paratypes</typeStatus>
</emphasis>
</materialsCitation>
</paragraph>
<caption id="DF53FC913215FFA47FC335B0B62AF83C" ID-DOI="http://doi.org/10.5281/zenodo.10115161" ID-Zenodo-Dep="10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="5" pageNumber="6" startId="5.[188,230,1930,1956]" targetBox="[221,1353,1380,1891]" targetPageId="5">
<paragraph id="8B93AC193215FFA47FC335B0B62AF83C" blockId="5.[188,1399,1930,2028]" pageId="5" pageNumber="6">
<emphasis id="B958700B3215FFA47FC335B0B274F873" bold="true" box="[188,267,1930,1956]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
<taxonomicName id="4C2CD79A3215FFA47E6935B0B07EF873" authority="Vujic &amp; Radenkovic" authorityName="Vujic &amp; Radenkovic" authorityYear="2016" box="[278,769,1930,1957]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3215FFA47E6935B0B17BF873" box="[278,516,1930,1956]" italics="true" pageId="5" pageNumber="6">Merodon megavidus</emphasis>
Vujić &amp; Radenković
</taxonomicName>
<taxonomicNameLabel id="A26BCD703215FFA47C7435B1B015F872" box="[779,874,1931,1957]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, male genitalia. 
<emphasis id="B958700B3215FFA47B5035B0B73BF873" bold="true" box="[1071,1092,1930,1956]" pageId="5" pageNumber="6">A</emphasis>
. Epandrium, lateral view. 
<emphasis id="B958700B3215FFA47FC33594B3B0F81F" bold="true" box="[188,207,1966,1992]" pageId="5" pageNumber="6">B</emphasis>
. Left surstylus, anterior view. 
<emphasis id="B958700B3215FFA47D583597B143F810" bold="true" box="[551,572,1965,1991]" pageId="5" pageNumber="6">C</emphasis>
. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm.
</paragraph>
</caption>
<paragraph id="8B93AC193216FFA77FC23314B774FD51" blockId="6.[189,1399,267,646]" pageId="6" pageNumber="7">
<collectingCountry id="F33BEC893216FFA77FC23314B24BFE9F" box="[189,308,302,328]" name="Greece" pageId="6" pageNumber="7">GREECE</collectingCountry>
: Lesvos: 
<specimenCount id="9D2A67903216FFA77EDD3314B2B0FE9E" box="[418,463,302,329]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
, (
<collectionCode id="ED3D34DC3216FFA77E993314B150FE9F" box="[486,559,302,328]" pageId="6" pageNumber="7">WML</collectionCode>
Misc 38). Agiassos: 1 Ƌ, 
<date id="FF928AD93216FFA77C1B3315B095FE9F" box="[868,1002,302,329]" pageId="6" pageNumber="7" value="2007-07-07">7 Jul. 2007</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77BE83314B79FFE9F" box="[1175,1248,302,328]" pageId="6" pageNumber="7">WML</collectionCode>
H723); 
<specimenCount id="9D2A67903216FFA77A3D3314B610FE9E" box="[1346,1391,302,329]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
, 
<date id="FF928AD93216FFA77FC2336BB238FEBC" box="[189,327,337,364]" pageId="6" pageNumber="7" value="2003-06-08">8 Jun. 2003</date>
, leg. G. Ståhls (
<collectionCode id="ED3D34DC3216FFA77E80336BB123FEBC" box="[511,604,337,363]" country="0" httpUri="http://grbio.org/cool/zzce-zzpb" name="Faculty of Science, The University of Novi Sad" pageId="6" pageNumber="7">FSUNS</collectionCode>
04385); 
<specimenCount id="9D2A67903216FFA77DB9336BB18DFEBB" box="[710,754,337,364]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
, 
<date id="FF928AD93216FFA77C7F336BB0F5FEBC" box="[768,906,337,364]" pageId="6" pageNumber="7" value="2003-06-08">8 Jun. 2003</date>
, leg. G. Ståhls (
<collectionCode id="ED3D34DC3216FFA77B3C336BB7F5FEBC" box="[1091,1162,337,363]" country="Finland" httpUri="http://grbio.org/cool/2vmj-fp0v" name="Finnish Museum of Natural History" pageId="6" pageNumber="7" type="Museum">MZH</collectionCode>
); 1 Ƌ, 
<date id="FF928AD93216FFA77BA6336BB60DFEBC" box="[1241,1394,337,364]" pageId="6" pageNumber="7" value="2004-06-16">16 Jun. 2004</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77E19334FB2D0FE58" box="[358,431,373,399]" pageId="6" pageNumber="7">WML</collectionCode>
Hu-93); 3 ƋƋ, 
<date id="FF928AD93216FFA77D05334FB065FE58" box="[634,794,373,399]" pageId="6" pageNumber="7" value="2003-06-19">19 Jun. 2003</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77CAC334FB763FE58" box="[979,1052,373,399]" pageId="6" pageNumber="7">WML</collectionCode>
Hu70, Hu-78, Hu-81); 1 Ƌ, 
<date id="FF928AD93216FFA77FC233A2B227FE65" box="[189,344,408,434]" pageId="6" pageNumber="7" value="2004-04-20">20 Apr. 2004</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77D7D33A2B134FE65" box="[514,587,408,434]" pageId="6" pageNumber="7">WML</collectionCode>
Hu-92); 4 ƋƋ, 
<date id="FF928AD93216FFA77C7933A2B0DFFE65" box="[774,928,408,434]" pageId="6" pageNumber="7" value="2004-06-20">20 Jun. 2004</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77B3633A2B7EDFE65" box="[1097,1170,408,434]" pageId="6" pageNumber="7">WML</collectionCode>
Hu-77, Hu-94-96); 5 ƋƋ, 
<date id="FF928AD93216FFA77E733381B2DBFE02" box="[268,420,443,469]" pageId="6" pageNumber="7" value="2004-06-22">22 Jun. 2004</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77D353381B1ECFE02" box="[586,659,443,469]" pageId="6" pageNumber="7">WML</collectionCode>
Hu-76, Hu-97-100); 2 ƋƋ, 
<date id="FF928AD93216FFA77CA53381B70CFE02" box="[986,1139,443,469]" pageId="6" pageNumber="7" value="2004-06-22">22 Jun. 2004</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77A663381B609FE02" box="[1305,1398,443,469]" pageId="6" pageNumber="7">FSUSN</collectionCode>
04390, 04391); 2 ƋƋ, 
<date id="FF928AD93216FFA77EB033E4B113FE2F" box="[463,620,478,505]" pageId="6" pageNumber="7" value="2009-06-22">22 Jun. 2009</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77C6333E4B01AFE2F" box="[796,869,478,504]" pageId="6" pageNumber="7">WML</collectionCode>
H1432, H1433); 
<specimenCount id="9D2A67903216FFA77B4633E4B719FE2E" box="[1081,1126,478,505]" pageId="6" pageNumber="7" type="female">1 ♀</specimenCount>
, 
<date id="FF928AD93216FFA77B0833E4B66CFE2F" box="[1143,1299,478,505]" pageId="6" pageNumber="7" value="1999-06-23">23 Jun. 1999</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77E7D303BB220FDCC" box="[258,351,513,539]" country="0" httpUri="http://grbio.org/cool/zzce-zzpb" name="Faculty of Science, The University of Novi Sad" pageId="6" pageNumber="7">FSUNS</collectionCode>
04392); 3 ƋƋ, 
<date id="FF928AD93216FFA77D5D303BB1C0FDCC" box="[546,703,513,540]" pageId="6" pageNumber="7" value="2003-06-23">23 Jun. 2003</date>
, leg. M. Hull (
<collectionCode id="ED3D34DC3216FFA77C0E303BB0C5FDCC" box="[881,954,513,539]" pageId="6" pageNumber="7">WML</collectionCode>
Hu-69, Hu-79, Hu-80). Vatoussa: 16 ƋƋ, 
<specimenCount id="9D2A67903216FFA77F89301FB239FDE8" box="[246,326,549,575]" pageId="6" pageNumber="7" type="female">10 ♀♀</specimenCount>
, 
<date id="FF928AD93216FFA77E2C301EB29CFDE8" box="[339,483,548,575]" pageId="6" pageNumber="7" value="2015-07-26">26 Jul. 2015</date>
, leg. A. Vujić and S. Radenković (
<collectionCode id="ED3D34DC3216FFA77C13301FB0B6FDE8" box="[876,969,549,575]" country="0" httpUri="http://grbio.org/cool/zzce-zzpb" name="Faculty of Science, The University of Novi Sad" pageId="6" pageNumber="7">FSUNS</collectionCode>
10133-10151, 10159, 10160, 10162, 10163, 10243, 10253, 10256); 1 Ƌ, 
<date id="FF928AD93216FFA77D133072B067FDB5" box="[620,792,584,610]" pageId="6" pageNumber="7" value="2012-06-01" valueMax="2012-06-04" valueMin="2012-06-01">1–4 Jun. 2012</date>
, leg. Nakas (
<collectionCode id="ED3D34DC3216FFA77CCC3072B76FFDB5" box="[947,1040,584,610]" country="0" httpUri="http://grbio.org/cool/zzce-zzpb" name="Faculty of Science, The University of Novi Sad" pageId="6" pageNumber="7">FSUNS</collectionCode>
Ć94). Plomari: 2 ƋƋ, 
<date id="FF928AD93216FFA77A583072B384FD52" pageId="6" pageNumber="7" value="2004-07-14">14 Jul. 2004</date>
, (
<collectionCode id="ED3D34DC3216FFA77E713051B214FD52" box="[270,363,619,645]" country="0" httpUri="http://grbio.org/cool/zzce-zzpb" name="Faculty of Science, The University of Novi Sad" pageId="6" pageNumber="7">FSUNS</collectionCode>
02325, 03966); 1 Ƌ, 
<date id="FF928AD93216FFA77D173051B07CFD52" box="[616,771,619,645]" pageId="6" pageNumber="7" value="2004-07-14">14. Jul. 2004</date>
, leg. H. Dahm (S532).
</paragraph>
</subSubSection>
<caption id="DF53FC913216FFA77FC23597B000F83C" ID-DOI="http://doi.org/10.5281/zenodo.10115163" ID-Zenodo-Dep="10115163" httpUri="https://zenodo.org/record/10115163/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1965,1991]" targetBox="[208,1380,727,1927]" targetPageId="6">
<paragraph id="8B93AC193216FFA77FC23597B000F83C" blockId="6.[189,1398,1965,2027]" pageId="6" pageNumber="7">
<emphasis id="B958700B3216FFA77FC23597B275F81F" bold="true" box="[189,266,1965,1992]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
Hind leg, lateral view. 
<emphasis id="B958700B3216FFA77D623597B12BF810" bold="true" box="[541,596,1965,1991]" pageId="6" pageNumber="7">A–B</emphasis>
. 
<taxonomicName id="4C2CD79A3216FFA77D1C3594B0B4F81F" authority="(Rossi, 1790)" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[611,971,1966,1992]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3216FFA77D1C3594B05DF81F" box="[611,802,1966,1992]" italics="true" pageId="6" pageNumber="7">Merodon avidus</emphasis>
(Rossi, 1790)
</taxonomicName>
. 
<emphasis id="B958700B3216FFA77CA93597B094F810" bold="true" box="[982,1003,1965,1991]" pageId="6" pageNumber="7">A</emphasis>
. Ƌ. 
<emphasis id="B958700B3216FFA77B613594B74DF81F" bold="true" box="[1054,1074,1966,1992]" pageId="6" pageNumber="7">B</emphasis>
. ♀. — 
<emphasis id="B958700B3216FFA77BF53597B7BCF810" bold="true" box="[1162,1219,1965,1991]" pageId="6" pageNumber="7">C–D</emphasis>
. 
<taxonomicName id="4C2CD79A3216FFA77BAD3594B614F81F" box="[1234,1387,1966,1992]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="megavidu">
<emphasis id="B958700B3216FFA77BAD3594B614F81F" box="[1234,1387,1966,1992]" italics="true" pageId="6" pageNumber="7">M. megavidu</emphasis>
</taxonomicName>
s Vujić &amp; Radenković sp. nov. 
<emphasis id="B958700B3216FFA77D6835EBB153F83C" bold="true" box="[535,556,2001,2027]" pageId="6" pageNumber="7">C</emphasis>
. Ƌ. 
<emphasis id="B958700B3216FFA77D2035EBB10BF83C" bold="true" box="[607,628,2001,2027]" pageId="6" pageNumber="7">D</emphasis>
. ♀. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C336FF923217FFA97FC23331B048FD10" lastPageId="8" lastPageNumber="9" pageId="7" pageNumber="8" type="description">
<paragraph id="8B93AC193217FFA67FC23331B231FEF2" blockId="7.[189,334,267,293]" box="[189,334,267,293]" pageId="7" pageNumber="8">
<heading id="D0DB1B753217FFA67FC23331B231FEF2" bold="true" box="[189,334,267,293]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="B958700B3217FFA67FC23331B231FEF2" bold="true" box="[189,334,267,293]" pageId="7" pageNumber="8">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC193217FFA67F953300B15BFE83" blockId="7.[189,1399,313,691]" box="[234,548,313,340]" pageId="7" pageNumber="8">
<emphasis id="B958700B3217FFA67F953300B255FE83" bold="true" box="[234,298,314,340]" pageId="7" pageNumber="8">Male</emphasis>
(
<figureCitation id="1317B09C3217FFA67E463303B2E6FE83" box="[313,409,313,340]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="7" pageNumber="8">Figs 2C</figureCitation>
, 
<figureCitation id="1317B09C3217FFA67ED93300B2C9FE83" box="[422,438,314,340]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[208,1380,727,1927]" captionTargetId="figure@6.[189,1399,707,1948]" captionTargetPageId="6" captionText="Fig. 3. Hind leg, lateral view. A–B. Merodon avidus (Rossi, 1790). A. Ƌ. B. ♀. — C–D. M. megavidus Vujić &amp; Radenković sp. nov. C. Ƌ. D. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115163" httpUri="https://zenodo.org/record/10115163/files/figure.png" pageId="7" pageNumber="8">3</figureCitation>
, 
<figureCitation id="1317B09C3217FFA67EBC3300B296FE83" box="[451,489,314,340]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[203,1383,1369,1932]" captionTargetId="figure@7.[189,1399,1353,1945]" captionTargetPageId="7" captionText="Fig. 4. Merodon megavidus Vujić &amp; Radenković sp. nov., head, antero-lateral view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115165" httpUri="https://zenodo.org/record/10115165/files/figure.png" pageId="7" pageNumber="8">4A</figureCitation>
, 
<figureCitation id="1317B09C3217FFA67E893303B165FE83" box="[502,538,313,340]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="7" pageNumber="8">5A</figureCitation>
)
</paragraph>
<paragraph id="8B93AC193217FFA67FC23367B05CFD64" blockId="7.[189,1399,313,691]" pageId="7" pageNumber="8">
HEAD (
<figureCitation id="1317B09C3217FFA67E703366B211FEA0" box="[271,366,348,375]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[203,1383,1369,1932]" captionTargetId="figure@7.[189,1399,1353,1945]" captionTargetPageId="7" captionText="Fig. 4. Merodon megavidus Vujić &amp; Radenković sp. nov., head, antero-lateral view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115165" httpUri="https://zenodo.org/record/10115165/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
). Antenna (
<figureCitation id="1317B09C3217FFA67E883366B12BFEA0" box="[503,596,348,375]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[203,1383,1369,1932]" captionTargetId="figure@7.[189,1399,1353,1945]" captionTargetPageId="7" captionText="Fig. 4. Merodon megavidus Vujić &amp; Radenković sp. nov., head, antero-lateral view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115165" httpUri="https://zenodo.org/record/10115165/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
) orange, first flagellomere 1.8–2.0 times as long as wide, 2.0 times longer than pedicel, concave, apex acute; arista: second, third and basal part of fourth flagellomeres pale, fourth flagellomere dark brown in apical ½ and thickened basally, 1.4 times longer than first flagellomere; with short, dense microtrichia. Face and frons black, covered with long golden pile and silver, dense microtrichia. Oral margin shiny black, except for the lateral microtrichose areas (
<figureCitation id="1317B09C3217FFA67A7433D3B615FDD3" box="[1291,1386,489,516]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[203,1383,1369,1932]" captionTargetId="figure@7.[189,1399,1353,1945]" captionTargetPageId="7" captionText="Fig. 4. Merodon megavidus Vujić &amp; Radenković sp. nov., head, antero-lateral view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115165" httpUri="https://zenodo.org/record/10115165/files/figure.png" pageId="7" pageNumber="8">Fig. 4A</figureCitation>
). Vertical triangle isosceles, shiny black except in front of the anterior ocellus that has pale microtrichia, covered with long orange pile except for black pile on the ocellar triangle. Ocellar triangle equilateral. Eye contiguity about 12 ommatidia long. Vertical triangle: eye contiguity: ocellar triangle = 1.5: 0.7: 1. Eye pile dense, white. Occiput with orange pile, along the eye margin with dense white microtrichia and posteriorly with metallic, bluish-greenish lustre.
</paragraph>
<paragraph id="8B93AC193217FFA67FC230E1B2F7FBD9" blockId="7.[189,1399,731,1039]" pageId="7" pageNumber="8">
THORAX. Mesoscutum and scutellum black with bronze lustre, covered with relatively long, dense, erect golden pile. Side of mesoscutum above wing-base with a patch of black pile. Mesoscutum with two lateral and two submedian, longitudinal, white microtrichose stripes. Proepimeron, posterior anepisternum, anteroventral and posterodorsal part of katepisternum, anepimeron, metasternum and katatergite with long golden pile and grey-green microtrichia. Wing hyaline, with dense microtrichia; veins dark brown except for light brown C, Sc and R1. Calypter pale yellow. Haltere with light brown pedicel and yellow capitulum. Legs orange, except for the black basal ¾ of the front- and mid-femora. Pile on legs golden. Hind femur (
<figureCitation id="1317B09C3217FFA67E2731EBB2CCFC3C" box="[344,435,977,1003]" captionStart="Fig" captionStartId="6.[189,232,1965,1991]" captionTargetBox="[208,1380,727,1927]" captionTargetId="figure@6.[189,1399,707,1948]" captionTargetPageId="6" captionText="Fig. 3. Hind leg, lateral view. A–B. Merodon avidus (Rossi, 1790). A. Ƌ. B. ♀. — C–D. M. megavidus Vujić &amp; Radenković sp. nov. C. Ƌ. D. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115163" httpUri="https://zenodo.org/record/10115163/files/figure.png" pageId="7" pageNumber="8">Fig. 3C</figureCitation>
) moderately thickened and curved, about 3.6 times as long as deep. Pile on hind femur very short.
</paragraph>
<paragraph id="8B93AC193217FFA67FC2360CB78AFAD7" blockId="7.[189,1399,1078,1280]" pageId="7" pageNumber="8">
ABDOMEN (
<figureCitation id="1317B09C3217FFA67E42360CB2E3FB86" box="[317,412,1078,1105]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="7" pageNumber="8">Fig. 5A</figureCitation>
). Dark with white microtrichose bands, tapering, 1.4 times longer than mesonotum (including scutellum). Tergites orange and reddish except for black tergite 1 and central parts of tergites 2–3 (and 4) (
<figureCitation id="1317B09C3217FFA67E1E3646B2BCFB40" box="[353,451,1148,1175]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="7" pageNumber="8">Fig. 5A</figureCitation>
); orange-reddish parts of variable size on tergite 3 and 4, laterally and along microtrichose bands. Tergites 2–4 each with a pair of white microtrichose marks (exceptionally absent only on tergite 2); tergites 3-4 with wide, oblique bands (
<figureCitation id="1317B09C3217FFA67C0936F9B0A7FB0A" box="[886,984,1218,1245]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="7" pageNumber="8">Fig. 5A</figureCitation>
). Pile on tergites golden. Sternites translucent, orange to brown towards the tip of the abdomen, covered with long yellow pile.
</paragraph>
<caption id="DF53FC913217FFA67FC23597B232F83C" ID-DOI="http://doi.org/10.5281/zenodo.10115165" ID-Zenodo-Dep="10115165" httpUri="https://zenodo.org/record/10115165/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1965,1991]" targetBox="[203,1383,1369,1932]" targetPageId="7">
<paragraph id="8B93AC193217FFA67FC23597B232F83C" blockId="7.[189,1399,1965,2027]" pageId="7" pageNumber="8">
<emphasis id="B958700B3217FFA67FC23597B274F810" bold="true" box="[189,267,1965,1991]" pageId="7" pageNumber="8">Fig. 4.</emphasis>
<taxonomicName id="4C2CD79A3217FFA67E6B3597B287F810" box="[276,504,1965,1991]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="megavidu">
<emphasis id="B958700B3217FFA67E6B3597B287F810" box="[276,504,1965,1991]" italics="true" pageId="7" pageNumber="8">Merodon megavidu</emphasis>
</taxonomicName>
s Vujić &amp; Radenković sp. nov., head, antero-lateral view. 
<emphasis id="B958700B3217FFA67BDB3597B7C6F810" bold="true" box="[1188,1209,1965,1991]" pageId="7" pageNumber="8">A</emphasis>
. Ƌ. 
<emphasis id="B958700B3217FFA67B913597B67DF810" bold="true" box="[1262,1282,1965,1991]" pageId="7" pageNumber="8">B</emphasis>
. ♀. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8B93AC193218FFA97FC23331B1B6FE58" blockId="8.[189,1399,267,399]" pageId="8" pageNumber="9">
Male genitalia (
<figureCitation id="1317B09C3218FFA97E0A3331B2C3FEF2" box="[373,444,267,293]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
). Similar to all species of the 
<taxonomicName id="4C2CD79A3218FFA97C693336B0F5FEF2" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[790,906,267,293]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3218FFA97C693336B0F5FEF2" box="[790,906,267,293]" italics="true" pageId="8" pageNumber="9">M. avidus</emphasis>
</taxonomicName>
complex. Anterior lobe of surstylus broad and hairy (
<figureCitation id="1317B09C3218FFA97E3D3314B2F3FE9E" box="[322,396,302,329]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
A–B); posterior lobe of surstylus ellipsoidal at ventral margin (
<figureCitation id="1317B09C3218FFA97BF13314B7A7FE9E" box="[1166,1240,302,329]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Fig. 2</figureCitation>
A–B); cercus rectangular, without prominences (
<figureCitation id="1317B09C3218FFA97D223368B1C1FEBB" box="[605,702,338,364]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Fig. 2A</figureCitation>
). Hypandrium elongate and sickle–shaped, without lateral projections (
<figureCitation id="1317B09C3218FFA97E30334FB2D3FE58" box="[335,428,373,399]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
); lingula long (
<figureCitation id="1317B09C3218FFA97D20334FB1C3FE58" box="[607,700,373,399]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Fig. 2C</figureCitation>
).
</paragraph>
<paragraph id="8B93AC193218FFA97F95338DB15FFE06" blockId="8.[189,1399,438,711]" box="[234,544,438,465]" pageId="8" pageNumber="9">
<emphasis id="B958700B3218FFA97F95338DB23AFE06" bold="true" box="[234,325,439,465]" pageId="8" pageNumber="9">Female</emphasis>
(
<figureCitation id="1317B09C3218FFA97E2A338CB2C8FE06" box="[341,439,438,465]" captionStart="Fig" captionStartId="5.[188,230,1930,1956]" captionTargetBox="[221,1353,1380,1891]" captionTargetId="figure@5.[200,1385,1371,1898]" captionTargetPageId="5" captionText="Fig. 2. Merodon megavidus Vujić &amp; Radenković sp. nov., male genitalia. A. Epandrium, lateral view. B. Left surstylus, anterior view. C. Hypandrium, lateral view. Abbreviations: psl = posterior surstylus lobe; asl = anterior surstylus lobe; c = cercus; ae = aedeagus; ea = ejaculatory apodeme. Scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.10115161" httpUri="https://zenodo.org/record/10115161/files/figure.png" pageId="8" pageNumber="9">Figs 2D</figureCitation>
, 
<figureCitation id="1317B09C3218FFA97EBC338DB298FE06" box="[451,487,439,465]" captionStart="Fig" captionStartId="7.[189,232,1965,1991]" captionTargetBox="[203,1383,1369,1932]" captionTargetId="figure@7.[189,1399,1353,1945]" captionTargetPageId="7" captionText="Fig. 4. Merodon megavidus Vujić &amp; Radenković sp. nov., head, antero-lateral view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115165" httpUri="https://zenodo.org/record/10115165/files/figure.png" pageId="8" pageNumber="9">4B</figureCitation>
, 
<figureCitation id="1317B09C3218FFA97E8B338CB169FE06" box="[500,534,438,465]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="8" pageNumber="9">5B</figureCitation>
)
</paragraph>
<paragraph id="8B93AC193218FFA97FC233E0B048FD10" blockId="8.[189,1399,438,711]" pageId="8" pageNumber="9">
Similar to the male except for typical sexual dimorphism and for the following characteristics: first flagellomere broader and longer; frons with two wide (about 0.34 width of frons) lateral silver microtrichose longitudinal stripes; frons in the widest part about 0.25 width of head; white microtrichose longitudinal stripes on mesoscutum more visible; broad stripe of black pile between wing bases; tergites predominately red except for tergite 1 and darkened parts of tergites 
<quantity id="4CD401FC3218FFA97C97305DB74CFD57" box="[1000,1075,614,641]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" metricValueMax="10.16" metricValueMin="5.08" pageId="8" pageNumber="9" unit="in" value="3.0" valueMax="4.0" valueMin="2.0">2–4 in</quantity>
some specimens (
<figureCitation id="1317B09C3218FFA97A74305CB617FD56" box="[1291,1384,614,641]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="8" pageNumber="9">Fig. 5B</figureCitation>
); white, microtrichose, transverse bands on tergites 3–4 (
<figureCitation id="1317B09C3218FFA97C2B30B0B0CDFD73" box="[852,946,649,676]" captionStart="Fig" captionStartId="8.[189,232,1964,1990]" captionTargetBox="[208,1383,956,1930]" captionTargetId="figure@8.[189,1399,936,1947]" captionTargetPageId="8" captionText="Fig. 5. Merodon megavidus Vujić &amp; Radenković sp. nov., abdomen, dorsal view. A. Ƌ. B. ♀. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="8" pageNumber="9">Fig. 5B</figureCitation>
); tergites 2–3 with black pile on dark parts; white microtrichose bands solely with pale pile.
</paragraph>
</subSubSection>
<subSubSection id="C336FF923218FFA87FC230D4B033FEBB" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="discussion">
<paragraph id="8B93AC193218FFA97FC230D4B250FCDF" blockId="8.[189,303,750,776]" box="[189,303,750,776]" pageId="8" pageNumber="9">
<heading id="D0DB1B753218FFA97FC230D4B250FCDF" bold="true" box="[189,303,750,776]" fontSize="11" level="3" pageId="8" pageNumber="9" reason="3">
<emphasis id="B958700B3218FFA97FC230D4B250FCDF" bold="true" box="[189,303,750,776]" pageId="8" pageNumber="9">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC193218FFA87FC23127B033FEBB" blockId="8.[189,1399,796,859]" lastBlockId="9.[189,1398,267,364]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
This species was mentioned as 
<taxonomicName id="4C2CD79A3218FFA97D463127B1DCFCE0" authorityName="Meigen" authorityYear="1803" box="[569,675,797,823]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="genus">
<emphasis id="B958700B3218FFA97D463127B1DCFCE0" box="[569,675,797,823]" italics="true" pageId="8" pageNumber="9">Merodon</emphasis>
</taxonomicName>
sp. nova 
<quantity id="4CD401FC3218FFA97C633127B033FCE0" box="[796,844,797,823]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="8" pageNumber="9" unit="in" value="2.0">2 in</quantity>
<bibRefCitation id="EFBDD1E83218FFA97C293126B742FCE0" author="Stahls G. &amp; Vujic A. &amp; Perez-Banon C. &amp; Radenkovic S. &amp; Rojo S. &amp; Petanidou T." box="[854,1085,796,823]" pageId="8" pageNumber="9" pagination="1431 - 1438" refId="ref10841" refString="Stahls G., Vujic A., Perez-Banon C., Radenkovic S., Rojo S. &amp; Petanidou T. 2009. COI barcodes for identification of Merodon hoverflies (Diptera, Syrphidae) of Lesvos Island, Greece. Molecular Ecology Resources 9 (6): 1431 - 1438. http: // dx. doi. org / 10.1111 / j. 1755 - 0998.2009.02592. x" type="journal article" year="2009">
Ståhls 
<emphasis id="B958700B3218FFA97CD83124B09AFCE0" box="[935,997,797,823]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2009)
</bibRefCitation>
and Ricarte 
<emphasis id="B958700B3218FFA97BA53124B66FFCE0" box="[1242,1296,797,823]" italics="true" pageId="8" pageNumber="9">et al</emphasis>
. (2012). 
<bibRefCitation id="EFBDD1E83218FFA97FC2317AB2DCFC8D" author="Stahls G. &amp; Vujic A. &amp; Perez-Banon C. &amp; Radenkovic S. &amp; Rojo S. &amp; Petanidou T." box="[189,419,832,859]" pageId="8" pageNumber="9" pagination="1431 - 1438" refId="ref10841" refString="Stahls G., Vujic A., Perez-Banon C., Radenkovic S., Rojo S. &amp; Petanidou T. 2009. COI barcodes for identification of Merodon hoverflies (Diptera, Syrphidae) of Lesvos Island, Greece. Molecular Ecology Resources 9 (6): 1431 - 1438. http: // dx. doi. org / 10.1111 / j. 1755 - 0998.2009.02592. x" type="journal article" year="2009">
Ståhls 
<emphasis id="B958700B3218FFA97E71317BB234FC8D" box="[270,331,832,858]" italics="true" pageId="8" pageNumber="9">et al.</emphasis>
(2009)
</bibRefCitation>
assumed it to be a distinct species based on the different COI barcode sequences obtained from two specimens morphologically similar to 
<taxonomicName id="4C2CD79A3219FFA87C283336B0B5FEF2" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[855,970,267,293]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87C283336B0B5FEF2" box="[855,970,267,293]" italics="true" pageId="9" pageNumber="10">M. avidus</emphasis>
</taxonomicName>
taken from Lesvos Island. In the key prepared as supporting material for 
<bibRefCitation id="EFBDD1E83219FFA87D1A3314B036FE9E" author="Stahls G. &amp; Vujic A. &amp; Perez-Banon C. &amp; Radenkovic S. &amp; Rojo S. &amp; Petanidou T." box="[613,841,302,329]" pageId="9" pageNumber="10" pagination="1431 - 1438" refId="ref10841" refString="Stahls G., Vujic A., Perez-Banon C., Radenkovic S., Rojo S. &amp; Petanidou T. 2009. COI barcodes for identification of Merodon hoverflies (Diptera, Syrphidae) of Lesvos Island, Greece. Molecular Ecology Resources 9 (6): 1431 - 1438. http: // dx. doi. org / 10.1111 / j. 1755 - 0998.2009.02592. x" type="journal article" year="2009">
Ståhls 
<emphasis id="B958700B3219FFA87DCA3315B18EFE9E" box="[693,753,303,329]" italics="true" pageId="9" pageNumber="10">et al.</emphasis>
(2009)
</bibRefCitation>
, 
<taxonomicName id="4C2CD79A3219FFA87C283315B0B3FE9E" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[855,972,303,329]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87C283315B0B3FE9E" box="[855,972,303,329]" italics="true" pageId="9" pageNumber="10">M. avidus</emphasis>
</taxonomicName>
and 
<emphasis id="B958700B3219FFA87B793315B759FE9F" box="[1030,1062,303,328]" italics="true" pageId="9" pageNumber="10">M.</emphasis>
sp. nova 2 key out together, without any morphological differences being described.
</paragraph>
</subSubSection>
<caption id="DF53FC913218FFA97FC23596B277F83D" ID-DOI="http://doi.org/10.5281/zenodo.10115167" ID-Zenodo-Dep="10115167" httpUri="https://zenodo.org/record/10115167/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1964,1990]" targetBox="[208,1383,956,1930]" targetPageId="8">
<paragraph id="8B93AC193218FFA97FC23596B277F83D" blockId="8.[189,1399,1964,2026]" pageId="8" pageNumber="9">
<emphasis id="B958700B3218FFA97FC23596B276F810" bold="true" box="[189,265,1964,1991]" pageId="8" pageNumber="9">Fig. 5.</emphasis>
<taxonomicName id="4C2CD79A3218FFA97E703596B28DF811" box="[271,498,1964,1990]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="megavidu">
<emphasis id="B958700B3218FFA97E703596B28DF811" box="[271,498,1964,1990]" italics="true" pageId="8" pageNumber="9">Merodon megavidu</emphasis>
</taxonomicName>
s Vujić &amp; Radenković sp. nov., abdomen, dorsal view. 
<emphasis id="B958700B3218FFA97B153596B700F811" bold="true" box="[1130,1151,1964,1990]" pageId="8" pageNumber="9">A</emphasis>
. Ƌ. 
<emphasis id="B958700B3218FFA97BCF3597B7BBF810" bold="true" box="[1200,1220,1965,1991]" pageId="8" pageNumber="9">B</emphasis>
. ♀. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="C336FF923219FFAC7FC233A9B23FFE2E" lastPageId="13" lastPageNumber="14" pageId="9" pageNumber="10" type="distribution">
<paragraph id="8B93AC193219FFA87FC233A9B153FE7A" blockId="9.[189,556,403,429]" box="[189,556,403,429]" pageId="9" pageNumber="10">
<heading id="D0DB1B753219FFA87FC233A9B153FE7A" bold="true" box="[189,556,403,429]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B958700B3219FFA87FC233A9B153FE7A" bold="true" box="[189,556,403,429]" pageId="9" pageNumber="10">Distribution and habitat data</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC193219FFA87FC233F8B1C8FE0B" blockId="9.[189,695,450,476]" box="[189,695,450,476]" pageId="9" pageNumber="10">
Lesvos Island (
<collectingCountry id="F33BEC893219FFA87E1033F8B2B6FE0B" box="[367,457,450,476]" name="Greece" pageId="9" pageNumber="10">Greece</collectingCountry>
). Maquis shrubland.
</paragraph>
<paragraph id="8B93AC193219FFA87FC23032B2BAFDF3" blockId="9.[189,453,519,548]" box="[189,453,519,548]" pageId="9" pageNumber="10">
<emphasis id="B958700B3219FFA87FC23032B2BAFDF3" bold="true" box="[189,453,519,548]" italics="true" pageId="9" pageNumber="10">Species delimitation</emphasis>
</paragraph>
<paragraph id="8B93AC193219FFA87FC23006B204FD81" blockId="9.[189,379,572,598]" box="[189,379,572,598]" pageId="9" pageNumber="10">
<heading id="D0DB1B753219FFA87FC23006B204FD81" bold="true" box="[189,379,572,598]" fontSize="11" level="3" pageId="9" pageNumber="10" reason="3">
<emphasis id="B958700B3219FFA87FC23006B204FD81" bold="true" box="[189,379,572,598]" pageId="9" pageNumber="10">Molecular data</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC193219FFA87FC23050B262FC49" blockId="9.[189,1399,618,926]" pageId="9" pageNumber="10">
<taxonomicName id="4C2CD79A3219FFA87FC23050B2D1FD53" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[189,430,618,644]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3219FFA87FC23050B2D1FD53" box="[189,430,618,644]" italics="true" pageId="9" pageNumber="10">Merodon megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703219FFA87EC73051B16AFD52" box="[440,533,619,645]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
clearly differs from other members of 
<taxonomicName id="4C2CD79A3219FFA87C923051B71BFD53" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1005,1124,618,644]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87C923051B71BFD53" box="[1005,1124,618,644]" italics="true" pageId="9" pageNumber="10">M. avidus</emphasis>
</taxonomicName>
complex based on our barcoding fragment of COI. All conducted phylogenetic analyses resulted in similar tree topologies (
<figureCitation id="1317B09C3219FFA87FBA308BB24EFD1C" box="[197,305,689,715]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="9.[189,232,1928,1954]" captionStartId-1="10.[189,232,1930,1956]" captionTargetBox-0="[191,1396,1266,1871]" captionTargetBox-1="[191,1391,961,1889]" captionTargetId-0="figure@9.[189,1399,1253,1888]" captionTargetId-1="figure@10.[189,1399,943,1903]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Fig. 6. Maximum parsimony strict consensus tree based on DNA barcode COI sequences. Length 136 steps, Consistency Index (CI) = 93, Retention Index (RI) = 95. Filled circles denote unique changes, open circles non-unique." captionText-1="Fig. 7. Maximum Likelihood tree based on a 5’ fragment of COI mtDNA sequences from the Merodon avidus complex. Bootstrap values (1000 replicates) are shown next to the branches. The tree is drawn to scale, with branch lengths proportional to the number of substitutions per site." figureDoi-0="http://doi.org/10.5281/zenodo.10115169" figureDoi-1="http://doi.org/10.5281/zenodo.10115171" httpUri-0="https://zenodo.org/record/10115169/files/figure.png" httpUri-1="https://zenodo.org/record/10115171/files/figure.png" pageId="9" pageNumber="10">Figs 6–7</figureCitation>
; Appendix 3). Sequences from the 
<taxonomicName id="4C2CD79A3219FFA87DAA308BB0EAFD1C" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[725,917,689,715]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87DAA308BB0EAFD1C" box="[725,917,689,715]" italics="true" pageId="9" pageNumber="10">Merodon avidus</emphasis>
</taxonomicName>
complex formed three separate clusters: one cluster represented 
<taxonomicName id="4C2CD79A3219FFA87E9F30EFB115FD39" authorityName="Vujic" authorityYear="2015" box="[480,618,724,750]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B3219FFA87E9F30EFB115FD39" box="[480,618,724,750]" italics="true" pageId="9" pageNumber="10">M. ibericus</emphasis>
</taxonomicName>
, a second comprised all 
<taxonomicName id="4C2CD79A3219FFA87CDE30EFB736FD39" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[929,1097,724,750]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3219FFA87CDE30EFB736FD39" box="[929,1097,724,750]" italics="true" pageId="9" pageNumber="10">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703219FFA87B2930EFB7CAFD38" box="[1110,1205,725,751]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
sequences with highly-significant bootstrap values (100), the third grouped sequences of 
<taxonomicName id="4C2CD79A3219FFA87B5230C2B7DBFCC6" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1069,1188,759,785]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87B5230C2B7DBFCC6" box="[1069,1188,759,785]" italics="true" pageId="9" pageNumber="10">M. avidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A3219FFA87B9E30C2B608FCC6" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[1249,1399,760,785]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B3219FFA87B9E30C2B608FCC6" box="[1249,1399,760,785]" italics="true" pageId="9" pageNumber="10">M. moenium</emphasis>
</taxonomicName>
together. All four 
<taxonomicName id="4C2CD79A3219FFA87EF23121B14EFCE3" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[397,561,794,820]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3219FFA87EF23121B14EFCE3" box="[397,561,794,820]" italics="true" pageId="9" pageNumber="10">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703219FFA87D473121B1EDFCE2" box="[568,658,795,821]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
sequences were identical, defining one haplotype unique to the species. The number of mutational steps between 
<taxonomicName id="4C2CD79A3219FFA87C703104B0CBFC8F" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[783,948,830,856]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3219FFA87C703104B0CBFC8F" box="[783,948,830,856]" italics="true" pageId="9" pageNumber="10">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703219FFA87CC13104B765FC8F" box="[958,1050,830,856]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C2CD79A3219FFA87B2A3104B7B5FC8F" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1109,1226,830,856]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87B2A3104B7B5FC8F" box="[1109,1226,830,856]" italics="true" pageId="9" pageNumber="10">M. avidus</emphasis>
</taxonomicName>
is 14, with 13 and 22 mutational steps between 
<taxonomicName id="4C2CD79A3219FFA87D3A3158B195FCAC" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[581,746,865,891]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3219FFA87D3A3158B195FCAC" box="[581,746,865,891]" italics="true" pageId="9" pageNumber="10">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703219FFA87D8D315BB032FCAC" box="[754,845,865,891]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C2CD79A3219FFA87CF83158B765FCAC" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[903,1050,866,891]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B3219FFA87CF83158B765FCAC" box="[903,1050,866,891]" italics="true" pageId="9" pageNumber="10">M. moenium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A3219FFA87B2B3158B7A5FCAC" authorityName="Vujic" authorityYear="2015" box="[1108,1242,865,891]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B3219FFA87B2B3158B7A5FCAC" box="[1108,1242,865,891]" italics="true" pageId="9" pageNumber="10">M. ibericus</emphasis>
</taxonomicName>
, respectively (
<figureCitation id="1317B09C3219FFA87FB931BEB273FC49" box="[198,268,900,926]" captionStart="Fig" captionStartId="11.[189,232,1613,1639]" captionTargetBox="[501,1092,1277,1563]" captionTargetId="figure@11.[495,1100,1268,1573]" captionTargetPageId="11" captionText="Fig. 8. Median-joining network of the mtDNA 5’-end of the COI gene. Circle sizes are proportional to haplotype frequencies. Each branch represents one mutational step; if more than one mutational step is present, it is denoted by the given number." figureDoi="http://doi.org/10.5281/zenodo.10115173" httpUri="https://zenodo.org/record/10115173/files/figure.png" pageId="9" pageNumber="10">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8B93AC193219FFA87FC231FCB069FBF1" blockId="9.[189,1398,965,1062]" pageId="9" pageNumber="10">
Our UPGMA tree based on genetic distances among species revealed genetic relationships among four taxa of the 
<taxonomicName id="4C2CD79A3219FFA87E3231D3B16FFBD4" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[333,528,1001,1027]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87E3231D3B16FFBD4" box="[333,528,1001,1027]" italics="true" pageId="9" pageNumber="10">Merodon avidus</emphasis>
</taxonomicName>
complex that support and strengthen the branch positions in wing and surstylus phenograms described below (see 
<figureCitation id="1317B09C3219FFA87DBE3636B078FBF1" box="[705,775,1036,1062]" captionStart="Fig" captionStartId="11.[189,232,1964,1990]" captionTargetBox="[498,1072,1802,1914]" captionTargetId="figure@11.[491,1096,1782,1921]" captionTargetPageId="11" captionText="Fig. 9. UPGMA tree based on pairwise genetic distances for four species from the Merodon avidus complex." figureDoi="http://doi.org/10.5281/zenodo.10115175" httpUri="https://zenodo.org/record/10115175/files/figure.png" pageId="9" pageNumber="10">Fig. 9</figureCitation>
).
</paragraph>
<paragraph id="8B93AC193219FFAB7FC23677B040FD03" blockId="9.[189,1399,1101,1199]" lastBlockId="10.[189,1399,592,724]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
Significant pairwise genetic divergence (ϕst value) was detected in each pairwise comparison between 
<taxonomicName id="4C2CD79A3219FFA87FC2364BB220FB5C" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[189,351,1137,1163]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B3219FFA87FC2364BB220FB5C" box="[189,351,1137,1163]" italics="true" pageId="9" pageNumber="10">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD703219FFA87E1A364BB2C2FB5C" box="[357,445,1137,1163]" pageId="9" pageNumber="10" rank="species">sp. nov.</taxonomicNameLabel>
and each of the other species 
<taxonomicName id="4C2CD79A3219FFA87C70364BB0ECFB5C" authorityName="Vujic" authorityYear="2015" box="[783,915,1137,1163]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B3219FFA87C70364BB0ECFB5C" box="[783,915,1137,1163]" italics="true" pageId="9" pageNumber="10">M. ibericus</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C2CD79A3219FFA87CE0364BB76DFB5C" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[927,1042,1137,1163]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B3219FFA87CE0364BB76DFB5C" box="[927,1042,1137,1163]" italics="true" pageId="9" pageNumber="10">M. avidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A3219FFA87B39364BB7A8FB5D" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[1094,1239,1137,1162]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B3219FFA87B39364BB7A8FB5D" box="[1094,1239,1137,1162]" italics="true" pageId="9" pageNumber="10">M. moenium</emphasis>
</taxonomicName>
(
<geoCoordinate id="EE18CADE3219FFA87B9B364BB655FB5C" box="[1252,1322,1137,1163]" degrees="0.420" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="55" value="0.42">0.420</geoCoordinate>
, 
<geoCoordinate id="EE18CADE3219FFA87A4B364BB609FB5C" box="[1332,1398,1137,1163]" degrees="0.492" direction="east" orientation="longitude" pageId="9" pageNumber="10" precision="55" value="0.492">0.492</geoCoordinate>
and 0.529, respectively). Sequence divergence (uncorrected p distance) of the COI gene was used to assess relative divergence times between the four 
<taxonomicName id="4C2CD79A321AFFAB7D84306AB01AFDBD" authorityName="Meigen" authorityYear="1803" box="[763,869,592,618]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="genus">
<emphasis id="B958700B321AFFAB7D84306AB01AFDBD" box="[763,869,592,618]" italics="true" pageId="10" pageNumber="11">Merodon</emphasis>
</taxonomicName>
taxa (
<tableCitation id="C6AE99A2321AFFAB7CD6306AB080FDBD" box="[937,1023,592,618]" captionStart="Table 1" captionStartId="10.[189,258,268,294]" captionText="Table 1. Below diagonal - pairwise p distances; above diagonal - estimated divergence times in years." pageId="10" pageNumber="11">Table 1</tableCitation>
), indicating an initial separation of 
<taxonomicName id="4C2CD79A321AFFAB7F9E304EB215FD59" authorityName="Vujic" authorityYear="2015" box="[225,362,628,654]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321AFFAB7F9E304EB215FD59" box="[225,362,628,654]" italics="true" pageId="10" pageNumber="11">M. ibericus</emphasis>
</taxonomicName>
from the rest of the complex around 800 ky BP. Divergence between 
<taxonomicName id="4C2CD79A321AFFAB7BB1304EB609FD59" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[1230,1398,628,654]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321AFFAB7BB1304EB609FD59" box="[1230,1398,628,654]" italics="true" pageId="10" pageNumber="11">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321AFFAB7FC230ADB266FD66" box="[189,281,663,689]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
and 
<emphasis id="B958700B321AFFAB7E2C30A2B11BFD66" box="[339,612,663,689]" italics="true" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7E2C30A2B2BAFD66" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[339,453,663,689]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="avidus">M. avidus</taxonomicName>
/ 
<taxonomicName id="4C2CD79A321AFFAB7EAD30ADB11BFD66" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[466,612,663,689]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="moenium">M. moenium</taxonomicName>
</emphasis>
occurred around 500 ky BP. The most recent separation happened between 
<taxonomicName id="4C2CD79A321AFFAB7E593081B2E5FD03" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[294,410,698,724]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321AFFAB7E593081B2E5FD03" box="[294,410,698,724]" italics="true" pageId="10" pageNumber="11">M. avidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321AFFAB7EAC3081B119FD03" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[467,614,699,724]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321AFFAB7EAC3081B119FD03" box="[467,614,699,724]" italics="true" pageId="10" pageNumber="11">M. moenium</emphasis>
</taxonomicName>
, around 87 ky BP.
</paragraph>
<caption id="DF53FC913219FFA87FC235B2B2A0F83E" ID-DOI="http://doi.org/10.5281/zenodo.10115169" ID-Zenodo-Dep="10115169" httpUri="https://zenodo.org/record/10115169/files/figure.png" pageId="9" pageNumber="10" startId="9.[189,232,1928,1954]" targetBox="[191,1396,1266,1871]" targetPageId="9">
<paragraph id="8B93AC193219FFA87FC235B2B2A0F83E" blockId="9.[189,1398,1928,2025]" pageId="9" pageNumber="10">
<emphasis id="B958700B3219FFA87FC235B2B274F875" bold="true" box="[189,267,1928,1954]" pageId="9" pageNumber="10">Fig. 6.</emphasis>
Maximum parsimony strict consensus tree based on DNA barcode COI sequences. Length 136 steps, Consistency Index (CI) = 93, Retention Index (RI) = 95. Filled circles denote unique changes, open circles non-unique.
</paragraph>
</caption>
<caption id="DF53FC91321AFFAB7FC23336B617FEF1" box="[189,1384,267,294]" pageId="10" pageNumber="11">
<paragraph id="8B93AC19321AFFAB7FC23336B617FEF1" blockId="10.[189,1384,267,294]" box="[189,1384,267,294]" pageId="10" pageNumber="11">
<emphasis id="B958700B321AFFAB7FC23336B25FFEF2" bold="true" box="[189,288,267,294]" pageId="10" pageNumber="11">Table 1.</emphasis>
Below diagonal - pairwise p distances; above diagonal - estimated divergence times in years.
</paragraph>
</caption>
<paragraph id="8B93AC19321AFFAB7D663376B664FDF5" pageId="10" pageNumber="11">
<table id="F92C5EB9321A005E7FB53371B62BFDF5" box="[202,1364,331,546]" gridcols="5" gridrows="5" pageId="10" pageNumber="11">
<tr id="351CAE5B321A005E7FB53371B62BFEB2" box="[202,1364,331,357]" gridrow="0" pageId="10" pageNumber="11" rowspan-0="1">
<th id="76CDC727321A005E7D663371B060FEB2" box="[537,799,331,357]" gridcol="1" gridrow="0" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7D663376B1C2FEB2" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[537,701,331,357]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321AFFAB7D663376B1C2FEB2" box="[537,701,331,357]" italics="true" pageId="10" pageNumber="11">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321AFFAB7DBA3371B060FEB2" box="[709,799,331,357]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<th id="76CDC727321A005E7C253371B09FFEB2" box="[858,992,331,357]" gridcol="2" gridrow="0" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7C253376B09FFEB2" authorityName="Vujic" authorityYear="2015" box="[858,992,331,357]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321AFFAB7C253376B09FFEB2" box="[858,992,331,357]" italics="true" pageId="10" pageNumber="11">M. ibericus</emphasis>
</taxonomicName>
</th>
<th id="76CDC727321A005E7B683371B7F1FEB2" box="[1047,1166,331,357]" gridcol="3" gridrow="0" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7B683376B7F4FEB2" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1047,1163,331,357]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321AFFAB7B683376B7F4FEB2" box="[1047,1163,331,357]" italics="true" pageId="10" pageNumber="11">M. avidus</emphasis>
</taxonomicName>
</th>
<th id="76CDC727321A005E7BBE3371B62BFEB2" box="[1217,1364,331,357]" gridcol="4" gridrow="0" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7BBE3376B62BFEB2" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[1217,1364,332,357]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321AFFAB7BBE3376B62BFEB2" box="[1217,1364,332,357]" italics="true" pageId="10" pageNumber="11">M. moenium</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="351CAE5B321A005E7FB53340B62BFE42" box="[202,1364,378,405]" gridrow="1" pageId="10" pageNumber="11">
<th id="76CDC727321A005E7FB53340B2AEFE42" box="[202,465,378,405]" gridcol="0" gridrow="1" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7FB53341B211FE43" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[202,366,378,404]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321AFFAB7FB53341B211FE43" box="[202,366,378,404]" italics="true" pageId="10" pageNumber="11">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321AFFAB7E093340B2AEFE43" box="[374,465,378,404]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
</th>
<td id="76CDC727321A005E7D663340B060FE42" box="[537,799,378,405]" gridcol="1" gridrow="1" pageId="10" pageNumber="11">–</td>
<td id="76CDC727321A005E7C253340B09FFE42" box="[858,992,378,405]" gridcol="2" gridrow="1" pageId="10" pageNumber="11">826.000</td>
<td id="76CDC727321A005E7B683340B7F1FE42" box="[1047,1166,378,405]" gridcol="3" gridrow="1" pageId="10" pageNumber="11">521.000</td>
<td id="76CDC727321A005E7BBE3340B62BFE42" box="[1217,1364,378,405]" gridcol="4" gridrow="1" pageId="10" pageNumber="11">478.000</td>
</tr>
<tr id="351CAE5B321A005E7FB53393B62BFE14" box="[202,1364,425,451]" gridrow="2" pageId="10" pageNumber="11">
<th id="76CDC727321A005E7FB53393B2AEFE14" box="[202,465,425,451]" gridcol="0" gridrow="2" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7FB53390B22FFE14" authorityName="Vujic" authorityYear="2015" box="[202,336,425,451]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321AFFAB7FB53390B22FFE14" box="[202,336,425,451]" italics="true" pageId="10" pageNumber="11">M. ibericus</emphasis>
</taxonomicName>
</th>
<td id="76CDC727321A005E7D663393B060FE14" box="[537,799,425,451]" gridcol="1" gridrow="2" pageId="10" pageNumber="11">0.038</td>
<td id="76CDC727321A005E7C253393B09FFE14" box="[858,992,425,451]" gridcol="2" gridrow="2" pageId="10" pageNumber="11">–</td>
<td id="76CDC727321A005E7B683393B7F1FE14" box="[1047,1166,425,451]" gridcol="3" gridrow="2" pageId="10" pageNumber="11">800.000</td>
<td id="76CDC727321A005E7BBE3393B62BFE14" box="[1217,1364,425,451]" gridcol="4" gridrow="2" pageId="10" pageNumber="11">760.000</td>
</tr>
<tr id="351CAE5B321A005E7FB533E2B62BFE25" box="[202,1364,472,498]" gridrow="3" pageId="10" pageNumber="11">
<th id="76CDC727321A005E7FB533E2B2AEFE25" box="[202,465,472,498]" gridcol="0" gridrow="3" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7FB533E3B241FE25" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[202,318,472,498]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321AFFAB7FB533E3B241FE25" box="[202,318,472,498]" italics="true" pageId="10" pageNumber="11">M. avidus</emphasis>
</taxonomicName>
</th>
<td id="76CDC727321A005E7D6633E2B060FE25" box="[537,799,472,498]" gridcol="1" gridrow="3" pageId="10" pageNumber="11">0.024</td>
<td id="76CDC727321A005E7C2533E2B09FFE25" box="[858,992,472,498]" gridcol="2" gridrow="3" pageId="10" pageNumber="11">0.037</td>
<td id="76CDC727321A005E7B6833E2B7F1FE25" box="[1047,1166,472,498]" gridcol="3" gridrow="3" pageId="10" pageNumber="11">–</td>
<td id="76CDC727321A005E7BBE33E2B62BFE25" box="[1217,1364,472,498]" gridcol="4" gridrow="3" pageId="10" pageNumber="11">87.000</td>
</tr>
<tr id="351CAE5B321A005E7FB5303DB62BFDF5" box="[202,1364,519,546]" gridrow="4" pageId="10" pageNumber="11">
<th id="76CDC727321A005E7FB5303DB2AEFDF5" box="[202,465,519,546]" gridcol="0" gridrow="4" pageId="10" pageNumber="11">
<taxonomicName id="4C2CD79A321AFFAB7FB53032B222FDF6" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[202,349,520,545]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321AFFAB7FB53032B222FDF6" box="[202,349,520,545]" italics="true" pageId="10" pageNumber="11">M. moenium</emphasis>
</taxonomicName>
</th>
<td id="76CDC727321A005E7D66303DB060FDF5" box="[537,799,519,546]" gridcol="1" gridrow="4" pageId="10" pageNumber="11">0.022</td>
<td id="76CDC727321A005E7C25303DB09FFDF5" box="[858,992,519,546]" gridcol="2" gridrow="4" pageId="10" pageNumber="11">0.0035</td>
<td id="76CDC727321A005E7B68303DB7F1FDF5" box="[1047,1166,519,546]" gridcol="3" gridrow="4" pageId="10" pageNumber="11">0.004</td>
<td id="76CDC727321A005E7BBE303DB62BFDF5" box="[1217,1364,519,546]" gridcol="4" gridrow="4" pageId="10" pageNumber="11">–</td>
</tr>
</table>
</paragraph>
<paragraph id="8B93AC19321AFFAB7FC230C1B068FCC2" blockId="10.[189,791,763,790]" box="[189,791,763,790]" pageId="10" pageNumber="11">
<heading id="D0DB1B75321AFFAB7FC230C1B068FCC2" bold="true" box="[189,791,763,790]" fontSize="11" level="3" pageId="10" pageNumber="11" reason="3">
<emphasis id="B958700B321AFFAB7FC230C1B068FCC2" bold="true" box="[189,791,763,790]" pageId="10" pageNumber="11">Geometric morphometrics - wing shape analysis</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC19321AFFAA7FC23110B0D6FD38" blockId="10.[189,1399,810,907]" lastBlockId="11.[189,1399,267,751]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
Principal component analysis conducted on 
<specimenCount id="9D2A6790321AFFAB7DA43110B0F2FC93" box="[731,909,810,836]" pageId="10" pageNumber="11" type="generic">444 specimens</specimenCount>
of the 
<taxonomicName id="4C2CD79A321AFFAB7C913111B718FC93" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1006,1127,810,836]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321AFFAB7C913111B718FC93" box="[1006,1127,810,836]" italics="true" pageId="10" pageNumber="11">M. avidus</emphasis>
</taxonomicName>
complex revealed six principal components (PC) that together explained 63.9% of total wing shape variability. A MANOVA with Fisher LSD post-hoc test showed that variability reflected shape changes among the investigated species in all six PCs (MANOVA: F 
<subScript id="17A8AE5C321BFFAA7D0A3327B1D3FEFC" attach="left" box="[629,684,285,299]" fontSize="6" pageId="11" pageNumber="12">18,123 0</subScript>
= 38.56; p &lt;0.00000). Further, DA showed that all species differ highly significantly in wing shape (p &lt;0.00000), and correctly classified species with an overall classification success of 90%. All specimens of 
<taxonomicName id="4C2CD79A321BFFAA7D8A3369B0E5FEBB" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[757,922,338,364]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321BFFAA7D8A3369B0E5FEBB" box="[757,922,338,364]" italics="true" pageId="11" pageNumber="12">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321BFFAA7CDC3368B081FEBB" box="[931,1022,338,364]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
were correctly classified, while the lowest classification success was for 
<taxonomicName id="4C2CD79A321BFFAA7DD6334CB04EFE58" authorityName="Vujic" authorityYear="2015" box="[681,817,373,399]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7DD6334CB04EFE58" box="[681,817,373,399]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
(78%). Specimens belonging to 
<taxonomicName id="4C2CD79A321BFFAA7BC1334CB64BFE58" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1214,1332,373,399]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321BFFAA7BC1334CB64BFE58" box="[1214,1332,373,399]" italics="true" pageId="11" pageNumber="12">M. avidus</emphasis>
</taxonomicName>
were correctly classified with 89% and 
<taxonomicName id="4C2CD79A321BFFAA7D1133A3B077FE65" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[622,776,409,434]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7D1133A3B077FE65" box="[622,776,409,434]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
with 93% certainty. Canonical analysis produced three canonical axes (CV) related to wing shape differences (
<figureCitation id="1317B09C321BFFAA7CE83381B08EFE02" box="[919,1009,443,469]" captionStart="Fig" captionStartId="12.[189,232,980,1006]" captionTargetBox="[196,1394,413,935]" captionTargetId="figure@12.[189,1399,406,939]" captionTargetPageId="12" captionText="Fig. 10. Differences in wing shape among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115177" httpUri="https://zenodo.org/record/10115177/files/figure.png" pageId="11" pageNumber="12">Fig. 10</figureCitation>
). CV1 separated 
<taxonomicName id="4C2CD79A321BFFAA7BC03386B64AFE02" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1215,1333,443,469]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321BFFAA7BC03386B64AFE02" box="[1215,1333,443,469]" italics="true" pageId="11" pageNumber="12">M. avidus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C2CD79A321BFFAA7FC233E5B231FE2F" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[189,334,479,504]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7FC233E5B231FE2F" box="[189,334,479,504]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
with 58% of total variation (CV1: Wilks’ Lambda = 0.145440; χ2 = 832.8937; p &lt;0.00000), while CV2 separated 
<taxonomicName id="4C2CD79A321BFFAA7EC63039B153FDCB" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[441,556,514,540]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321BFFAA7EC63039B153FDCB" box="[441,556,514,540]" italics="true" pageId="11" pageNumber="12">M. avidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7D1B3039B188FDCB" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[612,759,515,540]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7D1B3039B188FDCB" box="[612,759,515,540]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C2CD79A321BFFAA7C423039B09EFDCB" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[829,993,514,540]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321BFFAA7C423039B09EFDCB" box="[829,993,514,540]" italics="true" pageId="11" pageNumber="12">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321BFFAA7C973038B73DFDCB" box="[1000,1090,514,540]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
and 
<taxonomicName id="4C2CD79A321BFFAA7B053039B780FDCB" authorityName="Vujic" authorityYear="2015" box="[1146,1279,514,540]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7B053039B780FDCB" box="[1146,1279,514,540]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
with 27% of total variation (CV2: Wilks’ Lambda = 0.388320; χ2 = 408.6395; p &lt;0.00000) (
<figureCitation id="1317B09C321BFFAA7BF5301FB788FDE8" box="[1162,1271,549,575]" captionStart="Fig" captionStartId="12.[189,232,980,1006]" captionTargetBox="[196,1394,413,935]" captionTargetId="figure@12.[189,1399,406,939]" captionTargetPageId="12" captionText="Fig. 10. Differences in wing shape among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115177" httpUri="https://zenodo.org/record/10115177/files/figure.png" pageId="11" pageNumber="12">Fig. 10A</figureCitation>
). 
<taxonomicName id="4C2CD79A321BFFAA7A73301FB256FDB5" authorityName="Wiedemann in Meigen" authorityYear="1822" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7A73301FB256FDB5" italics="true" pageId="11" pageNumber="12">Merodon moenium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7E1C3073B295FDB5" authorityName="Vujic" authorityYear="2015" box="[355,490,584,610]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7E1C3073B295FDB5" box="[355,490,584,610]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
were clearly separated according to CV3 (CV3: Wilks’ Lambda = 0.698600; χ2 = 154.9484; p &lt;0.00000) (
<figureCitation id="1317B09C321BFFAA7D543051B1E6FD52" box="[555,665,619,645]" captionStart="Fig" captionStartId="12.[189,232,980,1006]" captionTargetBox="[196,1394,413,935]" captionTargetId="figure@12.[189,1399,406,939]" captionTargetPageId="12" captionText="Fig. 10. Differences in wing shape among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115177" httpUri="https://zenodo.org/record/10115177/files/figure.png" pageId="11" pageNumber="12">Fig. 10B</figureCitation>
). The phenogram constructed based on squared Mahalanobis distances clearly depicts that 
<taxonomicName id="4C2CD79A321BFFAA7D7030B5B1FDFD7E" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[527,642,655,681]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321BFFAA7D7030B5B1FDFD7E" box="[527,642,655,681]" italics="true" pageId="11" pageNumber="12">M. avidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7DC830B5B036FD7F" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[695,841,655,680]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7DC830B5B036FD7F" box="[695,841,655,680]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
have more similar wing shapes than 
<taxonomicName id="4C2CD79A321BFFAA7B8D30B5B608FD7E" authorityName="Vujic" authorityYear="2015" box="[1266,1399,655,681]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7B8D30B5B608FD7E" box="[1266,1399,655,681]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7F923089B2EFFD1B" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[237,400,690,716]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321BFFAA7F923089B2EFFD1B" box="[237,400,690,716]" italics="true" pageId="11" pageNumber="12">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321BFFAA7EE83088B28EFD1B" box="[407,497,690,716]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="1317B09C321BFFAA7E803088B12BFD1B" box="[511,596,690,716]" captionStart="Fig" captionStartId="12.[189,232,1964,1990]" captionTargetBox="[201,1387,1137,1913]" captionTargetId="figure@12.[189,1399,1118,1920]" captionTargetPageId="12" captionText="Fig. 11. UPGMA phenogram constructed using the squared Mahalanobis distances of wing shape for species of the M. avidus complex." figureDoi="http://doi.org/10.5281/zenodo.10115179" httpUri="https://zenodo.org/record/10115179/files/figure.png" pageId="11" pageNumber="12">Fig. 11</figureCitation>
). Differences in wing shape among species are depicted in 
<figureCitation id="1317B09C321BFFAA7B803088B60DFD1B" box="[1279,1394,690,716]" captionStart="Fig" captionStartId="13.[189,232,1964,1990]" captionTargetBox="[212,1370,976,1908]" captionTargetId="figure@13.[189,1398,943,1935]" captionTargetPageId="13" captionText="Fig. 12. Thin-plate spline deformation grids showing wing shape differences between analysed species. Differences between the species have been exaggerated five-fold to make them more visible." figureDoi="http://doi.org/10.5281/zenodo.10115181" httpUri="https://zenodo.org/record/10115181/files/figure.png" pageId="11" pageNumber="12">Figure 12</figureCitation>
, but have been exaggerated five-fold to make them more visible.
</paragraph>
<caption id="DF53FC91321AFFAB7FC235B0B72FF83C" ID-DOI="http://doi.org/10.5281/zenodo.10115171" ID-Zenodo-Dep="10115171" httpUri="https://zenodo.org/record/10115171/files/figure.png" pageId="10" pageNumber="11" startId="10.[189,232,1930,1956]" targetBox="[191,1391,961,1889]" targetPageId="10">
<paragraph id="8B93AC19321AFFAB7FC235B0B72FF83C" blockId="10.[189,1399,1930,2027]" pageId="10" pageNumber="11">
<emphasis id="B958700B321AFFAB7FC235B0B274F872" bold="true" box="[189,267,1930,1957]" pageId="10" pageNumber="11">Fig. 7.</emphasis>
Maximum Likelihood tree based on a 5’ fragment of COI mtDNA sequences from the 
<taxonomicName id="4C2CD79A321AFFAB7A7335B0B275F81F" baseAuthorityName="Rossi" baseAuthorityYear="1790" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321AFFAB7A7335B0B275F81F" italics="true" pageId="10" pageNumber="11">Merodon avidus</emphasis>
</taxonomicName>
complex. Bootstrap values (1000 replicates) are shown next to the branches. The tree is drawn to scale, with branch lengths proportional to the number of substitutions per site.
</paragraph>
</caption>
<paragraph id="8B93AC19321BFFAA7FC2312CB77DFC83" blockId="11.[189,1399,790,852]" pageId="11" pageNumber="12">
Additionally, the phenogram constructed based on squared Mahalanobis distances of wing shape showed that all 26 analysed populations grouped according to species (
<figureCitation id="1317B09C321BFFAA7CE23103B08BFC83" box="[925,1012,825,852]" captionStart="Fig" captionStartId="14.[189,232,1966,1992]" captionTargetBox="[199,1387,1219,1931]" captionTargetId="figure@14.[189,1398,1202,1937]" captionTargetPageId="14" captionText="Fig. 13. UPGMA phenogram constructed using squared Mahalanobis distances of wing shape for populations of species of the M. avidus complex." figureDoi="http://doi.org/10.5281/zenodo.10115183" httpUri="https://zenodo.org/record/10115183/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8B93AC19321BFFAA7FC23141B19EFC42" blockId="11.[189,737,891,917]" box="[189,737,891,917]" pageId="11" pageNumber="12">
<heading id="D0DB1B75321BFFAA7FC23141B19EFC42" bold="true" box="[189,737,891,917]" fontSize="11" level="3" pageId="11" pageNumber="12" reason="3">
<emphasis id="B958700B321BFFAA7FC23141B19EFC42" bold="true" box="[189,737,891,917]" pageId="11" pageNumber="12">Geometric morphometrics - surstylus shape</emphasis>
</heading>
</paragraph>
<paragraph id="8B93AC19321BFFAC7FC23190B23FFE2E" blockId="11.[189,1399,937,1211]" lastBlockId="13.[189,1399,267,505]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
PCA of surstylus shape revealed seven PC, of which the first six were connected with shape differences among species (MANOVA: F 
<subScript id="17A8AE5C321BFFAA7D5F31E5B130FC3A" attach="left" box="[544,591,991,1005]" fontSize="6" pageId="11" pageNumber="12">21,333</subScript>
= 5.280113; p &lt;0.00000). DA showed that all species differ highly significantly in surstylus shape (p &lt;0.00000).All specimens of 
<taxonomicName id="4C2CD79A321BFFAA7CEC31CBB77AFBDD" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[915,1029,1008,1034]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321BFFAA7CEC31CBB77AFBDD" box="[915,1029,1008,1034]" italics="true" pageId="11" pageNumber="12">M. avidus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7B4631CBB7A4FBDD" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[1081,1243,1008,1034]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321BFFAA7B4631CBB7A4FBDD" box="[1081,1243,1008,1034]" italics="true" pageId="11" pageNumber="12">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321BFFAA7B9F31CAB646FBDD" box="[1248,1337,1008,1034]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
were correctly classified (100%), while only two specimens of 
<taxonomicName id="4C2CD79A321BFFAA7C2E362EB0AAFBFA" authorityName="Vujic" authorityYear="2015" box="[849,981,1043,1069]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7C2E362EB0AAFBFA" box="[849,981,1043,1069]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7B75362EB7E4FBFA" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[1034,1179,1044,1069]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7B75362EB7E4FBFA" box="[1034,1179,1044,1069]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
were misclassified (97%). CVA found three CVs connected with shape change (
<figureCitation id="1317B09C321BFFAA7CF7360CB09FFB87" box="[904,992,1078,1104]" captionStart="Fig" captionStartId="15.[189,232,1964,1990]" captionTargetBox="[196,1386,1387,1913]" captionTargetId="figure@15.[189,1399,1372,1922]" captionTargetPageId="15" captionText="Fig. 14. Differences in the posterior part of the surstylus among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115185" httpUri="https://zenodo.org/record/10115185/files/figure.png" pageId="11" pageNumber="12">Fig. 14</figureCitation>
). CV1 clearly separated 
<taxonomicName id="4C2CD79A321BFFAA7A7D360DB608FB87" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[1282,1399,1078,1104]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="avidus">
<emphasis id="B958700B321BFFAA7A7D360DB608FB87" box="[1282,1399,1078,1104]" italics="true" pageId="11" pageNumber="12">M. avidus</emphasis>
</taxonomicName>
from 
<taxonomicName id="4C2CD79A321BFFAA7F833660B2E0FBA3" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[252,415,1114,1140]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321BFFAA7F833660B2E0FBA3" box="[252,415,1114,1140]" italics="true" pageId="11" pageNumber="12">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321BFFAA7ED93660B17DFBA3" box="[422,514,1114,1140]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, 
<taxonomicName id="4C2CD79A321BFFAA7D723660B1E0FBA4" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[525,671,1114,1139]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7D723660B1E0FBA4" box="[525,671,1114,1139]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7DA93660B024FBA3" authorityName="Vujic" authorityYear="2015" box="[726,859,1114,1140]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7DA93660B024FBA3" box="[726,859,1114,1140]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
and represented 53% of total shape variability (CV1: Wilks’ Lambda = 0.013392; χ2 = 405.4285; p &lt;0.00000) (
<figureCitation id="1317B09C321BFFAA7CC03647B752FB40" box="[959,1069,1149,1175]" captionStart="Fig" captionStartId="15.[189,232,1964,1990]" captionTargetBox="[196,1386,1387,1913]" captionTargetId="figure@15.[189,1399,1372,1922]" captionTargetPageId="15" captionText="Fig. 14. Differences in the posterior part of the surstylus among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115185" httpUri="https://zenodo.org/record/10115185/files/figure.png" pageId="11" pageNumber="12">Fig. 14A</figureCitation>
). The second canonical axis clearly separated 
<taxonomicName id="4C2CD79A321BFFAA7EF6369BB153FB6D" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[393,556,1184,1210]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321BFFAA7EF6369BB153FB6D" box="[393,556,1184,1210]" italics="true" pageId="11" pageNumber="12">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321BFFAA7D4C369AB1F2FB6D" box="[563,653,1184,1210]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
from 
<taxonomicName id="4C2CD79A321BFFAA7DAD369BB01AFB6D" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[722,869,1185,1210]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321BFFAA7DAD369BB01AFB6D" box="[722,869,1185,1210]" italics="true" pageId="11" pageNumber="12">M. moenium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321BFFAA7CE4369BB75EFB6D" authorityName="Vujic" authorityYear="2015" box="[923,1057,1184,1210]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321BFFAA7CE4369BB75EFB6D" box="[923,1057,1184,1210]" italics="true" pageId="11" pageNumber="12">M. ibericus</emphasis>
</taxonomicName>
and was responsible for 25% of the variability (CV2: Wilks’ Lambda = 0.086702; χ2 = 229.8565; p &lt;0.00000) (
<figureCitation id="1317B09C321DFFAC7BF53331B788FEF2" box="[1162,1271,267,293]" captionStart="Fig" captionStartId="15.[189,232,1964,1990]" captionTargetBox="[196,1386,1387,1913]" captionTargetId="figure@15.[189,1399,1372,1922]" captionTargetPageId="15" captionText="Fig. 14. Differences in the posterior part of the surstylus among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115185" httpUri="https://zenodo.org/record/10115185/files/figure.png" pageId="13" pageNumber="14">Fig. 14A</figureCitation>
). 
<taxonomicName id="4C2CD79A321DFFAC7A733331B256FE9F" authorityName="Wiedemann in Meigen" authorityYear="1822" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321DFFAC7A733331B256FE9F" italics="true" pageId="13" pageNumber="14">Merodon moenium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321DFFAC7E183315B28FFE9E" authorityName="Vujic" authorityYear="2015" box="[359,496,303,329]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321DFFAC7E183315B28FFE9E" box="[359,496,303,329]" italics="true" pageId="13" pageNumber="14">M. ibericus</emphasis>
</taxonomicName>
were separated by CV3, with 18% of total shape variability (CV3: Wilks’ Lambda = 0.346752; χ2 = 99.5597; p &lt;0.000305) (
<figureCitation id="1317B09C321DFFAC7C663368B0FBFEBB" box="[793,900,338,364]" captionStart="Fig" captionStartId="15.[189,232,1964,1990]" captionTargetBox="[196,1386,1387,1913]" captionTargetId="figure@15.[189,1399,1372,1922]" captionTargetPageId="15" captionText="Fig. 14. Differences in the posterior part of the surstylus among species of the M. avidus complex. A. Scatter plot of individual scores of CV1 vs CV2. B. Scatter plot of individual scores of CV2 vs CV3." figureDoi="http://doi.org/10.5281/zenodo.10115185" httpUri="https://zenodo.org/record/10115185/files/figure.png" pageId="13" pageNumber="14">Fig. 14B</figureCitation>
). According to the phenogram constructed based on squared Mahalanobis distances, 
<taxonomicName id="4C2CD79A321DFFAC7DCB334CB024FE58" authority="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković, 2016" authorityName="Ačanski &amp; Vujić &amp; Djan &amp; Vidaković &amp; Ståhls &amp; Radenković" authorityYear="2016" box="[692,859,373,399]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="megavidus" status="sp. nov.">
<emphasis id="B958700B321DFFAC7DCB334CB024FE58" box="[692,859,373,399]" italics="true" pageId="13" pageNumber="14">M. megavidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26BCD70321DFFAC7C1A334FB0BDFE58" box="[869,962,373,399]" pageId="13" pageNumber="14" rank="species">sp. nov.</taxonomicNameLabel>
has a more distinct surstylus shape, while the surstyli of 
<taxonomicName id="4C2CD79A321DFFAC7EC333A3B12CFE65" authorityName="Wiedemann in Meigen" authorityYear="1822" box="[444,595,409,434]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="moenium">
<emphasis id="B958700B321DFFAC7EC333A3B12CFE65" box="[444,595,409,434]" italics="true" pageId="13" pageNumber="14">M. moenium</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C2CD79A321DFFAC7DEC33A3B063FE65" authorityName="Vujic" authorityYear="2015" box="[659,796,408,434]" class="Insecta" family="Syrphidae" genus="Merodon" kingdom="Animalia" order="Diptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="ibericus">
<emphasis id="B958700B321DFFAC7DEC33A3B063FE65" box="[659,796,408,434]" italics="true" pageId="13" pageNumber="14">M. ibericus</emphasis>
</taxonomicName>
are the most similar (
<figureCitation id="1317B09C321DFFAC7B5233A2B7E1FE64" box="[1069,1182,408,435]" captionStart="Fig" captionStartId="16.[189,232,1560,1586]" captionTargetBox="[202,1377,1070,1512]" captionTargetId="figure@16.[189,1399,1060,1516]" captionTargetPageId="16" captionText="Fig. 15. Differences in the posterior part of the surstylus among species of the M. avidus complex. A. UPGMA phenogram constructed using squared Mahalanobis distances. B. Thin-plate spline deformation grids showing overall shape differences between analysed species." figureDoi="http://doi.org/10.5281/zenodo.10115187" httpUri="https://zenodo.org/record/10115187/files/figure.png" pageId="13" pageNumber="14">Fig. 15A</figureCitation>
). The main shape differences among all three species lie in the posterior margin of the posterior part of the surstylus lobe (
<figureCitation id="1317B09C321DFFAC7FB933E4B24CFE2E" box="[198,307,478,505]" captionStart="Fig" captionStartId="16.[189,232,1560,1586]" captionTargetBox="[202,1377,1070,1512]" captionTargetId="figure@16.[189,1399,1060,1516]" captionTargetPageId="16" captionText="Fig. 15. Differences in the posterior part of the surstylus among species of the M. avidus complex. A. UPGMA phenogram constructed using squared Mahalanobis distances. B. Thin-plate spline deformation grids showing overall shape differences between analysed species." figureDoi="http://doi.org/10.5281/zenodo.10115187" httpUri="https://zenodo.org/record/10115187/files/figure.png" pageId="13" pageNumber="14">Fig. 15A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>