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<document id="9ED33838E5E12834079F6F648B49D461" ID-CLB-Dataset="24795" ID-DOI="10.11646/zootaxa.4701.1.3" ID-GBIF-Dataset="1fb77063-018a-4222-8733-9450729ac222" ID-ISSN="1175-5326" ID-Zenodo-Dep="3557803" ID-ZooBank="668205B0-B6E9-48F3-9E21-87C534CDC837" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1575043522256" checkinUser="plazi" docAuthor="Huang, Sunbin, Tú, Výõng Tân, Phú, Ph ạm Văn, Tian, Mingyi &amp; Faille, Arnaud" docDate="2019" docId="0399DA2B7F38A810FF33F8CFFD33F284" docLanguage="en" docName="zootaxa.4701.1.3.pdf" docOrigin="Zootaxa 4701 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tonkinaphaenops jingxicus Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="48" masterDocId="FFA0A2537F3FA81DFFA4FF9CFFD8F02F" masterDocTitle="Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae)" masterLastPageNumber="53" masterPageNumber="35" pageNumber="42" updateTime="1698769588363" updateUser="ExternalLinkService">
<mods:mods id="764D5A276DD9B662AFCB100BB137FBFC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="21B04B46B1495EB955E2ABE36FC38495">
<mods:title id="2A33B2FBB52731B3B01F133239A99E5D">Contributions to the knowledge of the genus Tonkinaphaenops Deuve, 2013 from China and Vietnam (Coleoptera: Carabidae: Trechinae)</mods:title>
</mods:titleInfo>
<mods:name id="49F2A51FC43DC131A5BDB728E7BBAD4F" type="personal">
<mods:role id="E472406D6AD010F32A1F2A7A76CD9967">
<mods:roleTerm id="3411E590166A56EA68A58B71FF89E500">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3B4C35A0F521F39423BDE177903B1196">Huang, Sunbin</mods:namePart>
</mods:name>
<mods:name id="C9DF4F20F30F2CAC3ED50447C7CCE17A" type="personal">
<mods:role id="0F2AD0D81FDD5EE6E8367728F08E932D">
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</mods:role>
<mods:namePart id="F2CF1FFED52B71408E9F1E131482712C">Tú, Výõng Tân</mods:namePart>
</mods:name>
<mods:name id="39E83898003268D126E84CB8519A1D06" type="personal">
<mods:role id="45DE2AE1430BFA94840BBA6599BFFBDD">
<mods:roleTerm id="D3ED5D7F5D52E5518C691ED26505C6B7">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="3EB47C44722F1B612642A716E262C00D">Phú, Ph ạm Văn</mods:namePart>
</mods:name>
<mods:name id="0C2715D65D27643BF523D70178ED251A" type="personal">
<mods:role id="609A42AD934FCB11BC0D4B11E2353FE3">
<mods:roleTerm id="0B6E9715A81173638886E3CEFD6F1F00">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="1B4581F587F3E6AF5814875FEA99D662">Tian, Mingyi</mods:namePart>
</mods:name>
<mods:name id="08D5E26B9EFB7093DEDB92A813C592A1" type="personal">
<mods:role id="90AD0EA16BABA910735647548A28B688">
<mods:roleTerm id="4590BE5672B2DEC4E864007683A9B0EE">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="BE3A4BFD22C2AD504C2CBBC8D5093714">Faille, Arnaud</mods:namePart>
</mods:name>
<mods:typeOfResource id="02A68780FB32AE30178791A7FBDE160C">text</mods:typeOfResource>
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<mods:titleInfo id="8D7A31E233DDF850BB19DAD5D2F42FED">
<mods:title id="56D1FFC56E16E93B4D08795DD003CE47">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="C026132681B3F0BBB5B622DBBD3EC412">
<mods:date id="6AEC904B2B0634BFA7EC62B8ECFAB426">2019</mods:date>
<mods:detail id="92B14F4E28710BE6FF2FE7772CFDA571" type="pubDate">
<mods:number id="E2EBE5442043479C9441748ACC9118E4">2019-11-26</mods:number>
</mods:detail>
<mods:detail id="9A0F00A693B09CD09D0D0D7651BEEB77" type="volume">
<mods:number id="315E863216D635DEAADD0AC06A3DD710">4701</mods:number>
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<mods:number id="9923F01003209531FC4AA951A7BE3515">1</mods:number>
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<mods:start id="B9E6E2C845B5E41E3BD1E08FBE5CA57B">35</mods:start>
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<mods:classification id="67E8396F19AF83B317292478C0E25BCC">journal article</mods:classification>
<mods:identifier id="8A0C396FE9E0999C760EB10AC0DDC524" type="CLB-Dataset">24795</mods:identifier>
<mods:identifier id="4C1B8643756DA48807AC6011C340271D" type="DOI">10.11646/zootaxa.4701.1.3</mods:identifier>
<mods:identifier id="5C1B82E515433287E636F37CC2A24CA5" type="GBIF-Dataset">1fb77063-018a-4222-8733-9450729ac222</mods:identifier>
<mods:identifier id="0999EB6DE2CDC8302A2F4DBD7CEC4340" type="ISSN">1175-5326</mods:identifier>
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<treatment id="0399DA2B7F38A810FF33F8CFFD33F284" ID-DOI="http://doi.org/10.5281/zenodo.5929266" ID-GBIF-Taxon="160605350" ID-Zenodo-Dep="5929266" LSID="urn:lsid:plazi:treatment:0399DA2B7F38A810FF33F8CFFD33F284" httpUri="http://treatment.plazi.org/id/0399DA2B7F38A810FF33F8CFFD33F284" lastPageId="13" lastPageNumber="48" pageId="7" pageNumber="42">
<subSubSection id="C32A38B67F38A81AFF33F8CFFCD6F741" box="[151,782,1875,1902]" pageId="7" pageNumber="42" type="nomenclature">
<paragraph id="8B8F6B3D7F38A81AFF33F8CFFCD6F741" blockId="7.[151,782,1875,1938]" box="[151,782,1875,1902]" pageId="7" pageNumber="42">
<heading id="D0C7DC517F38A81AFF33F8CFFCD6F741" bold="true" box="[151,782,1875,1902]" fontSize="11" level="1" pageId="7" pageNumber="42" reason="1">
<emphasis id="B944B72F7F38A81AFF33F8CFFCD6F741" bold="true" box="[151,782,1875,1902]" pageId="7" pageNumber="42">
<taxonomicName id="4C3010BE7F38A81AFF33F8CFFD7CF742" ID-CoL="8MCJ4" authority="Huang &amp; Tian" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[151,676,1875,1902]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="42" phylum="Arthropoda" rank="species" species="jingxicus" status="sp. nov.">
<emphasis id="B944B72F7F38A81AFF33F8CFFE3DF741" bold="true" box="[151,485,1875,1902]" italics="true" pageId="7" pageNumber="42">Tonkinaphaenops jingxicus</emphasis>
Huang &amp; Tian
</taxonomicName>
,
<taxonomicNameLabel id="A2770A547F38A81AFD15F8C8FCD6F741" box="[689,782,1876,1902]" pageId="7" pageNumber="42" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF33F8E4FEAFF7BD" box="[151,375,1912,1938]" pageId="7" pageNumber="42" type="description">
<paragraph id="8B8F6B3D7F38A81AFF33F8E4FEAFF7BD" blockId="7.[151,782,1875,1938]" box="[151,375,1912,1938]" pageId="7" pageNumber="42">
(
<figureCitation id="130B77B87F38A81AFF3BF8E4FF35F7BD" box="[159,237,1912,1938]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1826]" captionTargetBox="[159,1428,877,1779]" captionTargetId="figure@1.[151,1436,877,1781]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Tonkinaphaenops Deuve, 2013" figureDoi="http://doi.org/10.5281/zenodo.3557805" httpUri="https://zenodo.org/record/3557805/files/figure.png" pageId="7" pageNumber="42">Figs. 1</figureCitation>
,
<figureCitation id="130B77B87F38A81AFF5FF8E4FEEDF7BD" box="[251,309,1912,1938]" captionStart-0="FIGURE 9" captionStart-1="FIGURE 10" captionStartId-0="11.[151,250,1999,2024]" captionStartId-1="12.[151,250,1968,1993]" captionTargetBox-1="[164,1424,166,1949]" captionTargetId-0="figure@11.[381,1207,166,1979]" captionTargetId-1="figure@12.[164,1424,166,1949]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionText-0="FIGURE 9. Tonkinaphaenops jingxicus sp. nov., habitus of holotype, female." captionText-1="FIGURE 10. Cave Nianluo Dong, type locality of Tonkinaphaenops jingxicus sp. nov. (A. red arrow showing the location of entrance; B. entrance; C. a running individual of Tonkinaphaenops jingxicus)." figureDoi-0="http://doi.org/10.5281/zenodo.3614846" figureDoi-1="http://doi.org/10.5281/zenodo.3557817" httpUri-0="https://zenodo.org/record/3614846/files/figure.png" httpUri-1="https://zenodo.org/record/3557817/files/figure.png" pageId="7" pageNumber="42">
9–
<geoCoordinate id="EE040DFA7F38A81AFEBDF8E4FEEDF7BD" box="[281,309,1912,1938]" degrees="10, 14" direction="east" orientation="longitude" pageId="7" pageNumber="42" precision="555" value="10.14">10</geoCoordinate>
</figureCitation>
,
<figureCitation id="130B77B87F38A81AFEE5F8E4FEB7F7BD" box="[321,367,1912,1938]" captionStart="FIGURE 14" captionStartId="16.[151,250,1912,1937]" captionTargetBox="[214,1391,181,1869]" captionTargetId="figure@16.[189,1398,181,1887]" captionTargetPageId="16" captionText="FIGURE 14. Chaetotaxal pattern of Tonkinaphaenops (A. Tonkinaphaenops marinae Deuve, 2013, holotype, male from cave TH20; B, C. Tonkinaphaenops yinquanicus sp. nov.: B. male from Yinquan Dong, C. female from Yinquan Dong; D. Tonkinaphaenops impunctatus sp. nov., male from Hang Rắn cave; E. Tonkinaphaenops jingxicus sp. nov., female from Nianluo Dong; F. Tonkinaphaenops anthonyi sp. nov., male from Hang Dõi cave)." figureDoi="http://doi.org/10.5281/zenodo.3557823" httpUri="https://zenodo.org/record/3557823/files/figure.png" pageId="7" pageNumber="42">14E</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F38A81AFF33F826FB63F7D7" pageId="7" pageNumber="42" type="materials_examined">
<paragraph id="8B8F6B3D7F38A81AFF33F826FB63F7D7" blockId="7.[151,1436,1977,2041]" pageId="7" pageNumber="42">
<materialsCitation id="3B5861607F38A81AFF33F826FB63F7D7" ID-GBIF-Occurrence="2460169377" collectingDate="2018-02-01" collectionCode="SCAU" collectorName="Sunbin Huang &amp; Dianmei Wang &amp; Mengzhen Chen &amp; Ruiqi Han" country="China" county="Jingxi" elevation="927" latitude="23.168152" location="Biaoliang" longLatPrecision="1" longitude="106.27055" municipality="Ludong" pageId="7" pageNumber="42" specimenCount="1" specimenCount-female="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B944B72F7F38A81AFF33F826FED3F7FB" bold="true" box="[151,267,1978,2004]" pageId="7" pageNumber="42">
<typeStatus id="548BD59F7F38A81AFF33F826FEDEF7FB" box="[151,262,1978,2004]" pageId="7" pageNumber="42" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="9D36A0B47F38A81AFEB5F827FEFFF7FA" box="[273,295,1979,2005]" pageId="7" pageNumber="42" type="female">♀</specimenCount>
;
<collectingCountry id="F3272BAD7F38A81AFE90F826FEA3F7FB" box="[308,379,1978,2004]" name="China" pageId="7" pageNumber="42">China</collectingCountry>
:
<collectingRegion id="49F4A5DF7F38A81AFE20F826FE31F7FB" box="[388,489,1978,2004]" country="China" name="Guangxi" pageId="7" pageNumber="42">Guangxi</collectingRegion>
:
<collectingCounty id="62EE13B17F38A81AFE57F826FDE4F7FB" box="[499,572,1978,2004]" pageId="7" pageNumber="42">Jingxi</collectingCounty>
:
<collectingMunicipality id="6BEBF1477F38A81AFDE1F826FD79F7FB" box="[581,673,1978,2004]" pageId="7" pageNumber="42">Ludong</collectingMunicipality>
:
<location id="8EEF3DE67F38A81AFD0EF826FCC3F7FB" LSID="urn:lsid:plazi:treatment:0399DA2B7F38A810FF33F8CFFD33F284:8EEF3DE67F38A81AFD0EF826FCC3F7FB" box="[682,795,1978,2004]" country="China" county="Jingxi" latitude="23.168152" longLatPrecision="1" longitude="106.27055" municipality="Ludong" name="Biaoliang" pageId="7" pageNumber="42" stateProvince="Guangxi">Biaoliang</location>
:
<location id="8EEF3DE67F38A81AFC80F827FBD8F7FA" LSID="urn:lsid:plazi:treatment:0399DA2B7F38A810FF33F8CFFD33F284:8EEF3DE67F38A81AFC80F827FBD8F7FA" box="[804,1024,1978,2005]" country="China" county="Jingxi" latitude="23.168152" longLatPrecision="1" longitude="106.27055" municipality="Ludong" name="cave Nianluo Dong" pageId="7" pageNumber="42" stateProvince="Guangxi">cave Nianluo Dong</location>
;
<geoCoordinate id="EE040DFA7F38A81AFBAEF825FB71F7FB" box="[1034,1193,1977,2004]" degrees="23" direction="north" minutes="10" orientation="latitude" pageId="7" pageNumber="42" precision="1" seconds="5.35" value="23.168152">23°10’5.35”N</geoCoordinate>
<geoCoordinate id="EE040DFA7F38A81AFB0AF826FABFF7FB" box="[1198,1383,1978,2004]" degrees="106" direction="east" minutes="16" orientation="longitude" pageId="7" pageNumber="42" precision="1" seconds="13.98" value="106.27055">106°16’13.98”E</geoCoordinate>
,
<quantity id="4CC8C6D87F38A81AFAD6F826FF68F7D6" metricMagnitude="2" metricUnit="m" metricValue="9.27" pageId="7" pageNumber="42" unit="m" value="927.0">
<elevation id="001D8C0E7F38A81AFAD6F826FF68F7D6" metricMagnitude="2" metricUnit="m" metricValue="9.27" pageId="7" pageNumber="42" unit="m" value="927.0">927 m</elevation>
</quantity>
; coll.
<collectorName id="26C50EEB7F38A81AFF50F842FE4FF7D6" box="[244,407,2014,2041]" pageId="7" pageNumber="42">Sunbin Huang</collectorName>
,
<collectorName id="26C50EEB7F38A81AFE07F842FD96F7D6" box="[419,590,2014,2041]" pageId="7" pageNumber="42">Dianmei Wang</collectorName>
,
<collectorName id="26C50EEB7F38A81AFDFCF842FCCBF7D7" box="[600,787,2014,2040]" pageId="7" pageNumber="42">Mengzhen Chen</collectorName>
,
<collectorName id="26C50EEB7F38A81AFCBBF842FC4EF7D6" box="[799,918,2014,2041]" pageId="7" pageNumber="42">Ruiqi Han</collectorName>
;
<date id="FF8E4DFD7F38A81AFC05F842FB85F7D7" box="[929,1117,2014,2040]" pageId="7" pageNumber="42" value="2018-02-01">
<collectingDate id="EFCAB4157F38A81AFC05F842FB85F7D7" box="[929,1117,2014,2040]" pageId="7" pageNumber="42" value="2018-02-01">1 February 2018</collectingDate>
</date>
;
<collectionCode id="ED21F3F87F38A81AFBCDF842FB6FF7D7" box="[1129,1207,2014,2040]" pageId="7" pageNumber="42">SCAU</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<caption id="DF4F3BB57F37A815FF33F845FC62F7DD" ID-DOI="http://doi.org/10.5281/zenodo.3614842" ID-Zenodo-Dep="3614842" box="[151,954,2009,2034]" httpUri="https://zenodo.org/record/3614842/files/figure.png" pageId="8" pageNumber="43" targetBox="[401,1173,188,1975]" targetPageId="8">
<paragraph id="8B8F6B3D7F37A815FF33F845FC62F7DD" blockId="8.[151,954,2009,2034]" box="[151,954,2009,2034]" pageId="8" pageNumber="43">
<emphasis id="B944B72F7F37A815FF33F845FECBF7DD" bold="true" box="[151,275,2009,2034]" pageId="8" pageNumber="43">FIGURE 6.</emphasis>
<taxonomicName id="4C3010BE7F37A815FEBDF846FD96F7DE" authority="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille, 2019" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[281,590,2010,2034]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="43" phylum="Arthropoda" rank="species" species="impunctatus" status="sp. nov.">
<emphasis id="B944B72F7F37A815FEBDF846FD96F7DE" box="[281,590,2010,2034]" italics="true" pageId="8" pageNumber="43">Tonkinaphaenops impunctatus</emphasis>
</taxonomicName>
<emphasis id="B944B72F7F37A815FDF0F846FD7CF7DD" bold="true" box="[596,676,2010,2034]" pageId="8" pageNumber="43">
<taxonomicNameLabel id="A2770A547F37A815FDF0F846FD7CF7DD" box="[596,676,2010,2034]" pageId="8" pageNumber="43" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus of holotype, male.
</paragraph>
</caption>
<caption id="DF4F3BB57F36A814FF33FB6FFAC8F523" ID-DOI="http://doi.org/10.5281/zenodo.3557813" ID-Zenodo-Dep="3557813" box="[151,1296,1267,1292]" httpUri="https://zenodo.org/record/3557813/files/figure.png" pageId="9" pageNumber="44" startId="9.[151,250,1267,1292]" targetBox="[169,1424,191,1220]" targetPageId="9">
<paragraph id="8B8F6B3D7F36A814FF33FB6FFAC8F523" blockId="9.[151,1296,1267,1292]" box="[151,1296,1267,1292]" pageId="9" pageNumber="44">
<emphasis id="B944B72F7F36A814FF33FB6FFECBF523" bold="true" box="[151,275,1267,1292]" pageId="9" pageNumber="44">FIGURE 7.</emphasis>
Male genitalia of
<taxonomicName id="4C3010BE7F36A814FE69FB68FCD9F523" authority="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille, 2019" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[461,769,1268,1292]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="impunctatus" status="sp. nov.">
<emphasis id="B944B72F7F36A814FE69FB68FCD9F523" box="[461,769,1268,1292]" italics="true" pageId="9" pageNumber="44">Tonkinaphaenops impunctatus</emphasis>
</taxonomicName>
<emphasis id="B944B72F7F36A814FCA3FB68FC80F523" bold="true" box="[775,856,1268,1292]" pageId="9" pageNumber="44">
<taxonomicNameLabel id="A2770A547F36A814FCA3FB68FC80F523" box="[775,856,1268,1292]" pageId="9" pageNumber="44" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(median lobe and parameres, lateral view).
</paragraph>
</caption>
<subSubSection id="C32A38B67F36A810FF63FAA5FE50F1A4" lastPageId="13" lastPageNumber="48" pageId="9" pageNumber="44" type="description">
<paragraph id="8B8F6B3D7F36A814FF63FAA5FDF3F557" blockId="9.[151,1437,1337,1977]" pageId="9" pageNumber="44">
<emphasis id="B944B72F7F36A814FF63FAA5FE81F57B" bold="true" box="[199,345,1337,1364]" pageId="9" pageNumber="44">Description.</emphasis>
Length:
<quantity id="4CC8C6D87F36A814FE64FAA6FDC0F57A" box="[448,536,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="9" pageNumber="44" unit="mm" value="6.2">6.2 mm</quantity>
(from apical margin of labrum to elytral apex),
<quantity id="4CC8C6D87F36A814FB99FAA6FB4DF57A" box="[1085,1173,1338,1365]" metricMagnitude="-3" metricUnit="m" metricValue="6.6" pageId="9" pageNumber="44" unit="mm" value="6.6">6.6 mm</quantity>
(including mandibles); width:
<quantity id="4CC8C6D87F36A814FF42FAC2FEE7F556" box="[230,319,1374,1401]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="9" pageNumber="44" unit="mm" value="2.1">2.1 mm</quantity>
. Habitus as in
<figureCitation id="130B77B87F36A814FE47FAC2FDFFF557" box="[483,551,1374,1400]" captionStart="FIGURE 9" captionStartId="11.[151,250,1999,2024]" captionTargetId="figure@11.[381,1207,166,1979]" captionTargetPageId="11" captionText="FIGURE 9. Tonkinaphaenops jingxicus sp. nov., habitus of holotype, female." figureDoi="http://doi.org/10.5281/zenodo.3614846" httpUri="https://zenodo.org/record/3614846/files/figure.png" pageId="9" pageNumber="44">Fig. 9</figureCitation>
.
</paragraph>
<paragraph id="8B8F6B3D7F36A814FF63FA1EFAFFF5CB" blockId="9.[151,1437,1337,1977]" pageId="9" pageNumber="44">
Similar to
<taxonomicName id="4C3010BE7F36A814FEE4FA1FFE62F5B3" authorityName="Deuve" authorityYear="2013" box="[320,442,1411,1436]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F36A814FEE4FA1FFE62F5B3" box="[320,442,1411,1436]" italics="true" pageId="9" pageNumber="44">T. marinae</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3010BE7F36A814FE57FA1FFD4FF5B3" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[499,663,1411,1436]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="44" phylum="Arthropoda" rank="species" species="yinquanicus">
<emphasis id="B944B72F7F36A814FE57FA1FFD4FF5B3" box="[499,663,1411,1436]" italics="true" pageId="9" pageNumber="44">T. yinquanicus</emphasis>
</taxonomicName>
. Fore body long and narrow, hind body elongated obovate, strongly convex on dorsum; concolourously reddish-brown, but antennae, palps and tarsi slightly lighter at the apical part; pronotum glabrous on its dorsal surface, sparsely pubescent on the head and widely pubescent on elytra.
</paragraph>
<paragraph id="8B8F6B3D7F36A814FF63FA72FEF5F762" blockId="9.[151,1437,1337,1977]" pageId="9" pageNumber="44">
Head (without mandibles) as long as wide, HLm/HW = 1.6, HLl/HW = 1.0; frontal furrows deep and long, subparallel-sided at the anterior half, gradually curved at the posterior half, effaced near neck constriction; clypeus quadrisetose, without additional setae; right mandible bidentate; eyes absent, two pairs of supraorbital pores present; mentum bisetose, base widely concave, mental tooth very short and blunt; lateral lobe sharp and pointed at apex; labial suture indistinct; submentum 6-setose; ligula broadened at apex, multi-setose; labial palpomere 2 about 1.3 times longer than palpomere 3, bisetose on inner margin, with two additional setae on outer side at subapical and apical portions, maxillary palpomere 3 as long as 4; antennae extending to about the middle of elytra; scape with several setae; the 2
<superScript id="7C45C6757F36A814FE05F975FE19F72B" attach="none" box="[417,449,1769,1796]" fontSize="6" pageId="9" pageNumber="44">nd–</superScript>
11
<superScript id="7C45C6757F36A814FE7EF975FE3FF6D8" attach="left" box="[474,487,1769,1783]" fontSize="6" pageId="9" pageNumber="44">th</superScript>
antennomeres pubescent; the 2
<superScript id="7C45C6757F36A814FCE7F975FC8BF6D8" attach="left" box="[835,851,1769,1783]" fontSize="6" pageId="9" pageNumber="44">nd</superScript>
antennomere a little longer than scape; the 4
<superScript id="7C45C6757F36A814FAE7F975FA88F6D8" attach="left" box="[1347,1360,1769,1783]" fontSize="6" pageId="9" pageNumber="44">th</superScript>
antennomere the longest, other antennomeres gradually decreasing in length posteriorly until the 10
<superScript id="7C45C6757F36A814FB6BF891FB04F734" attach="left" box="[1231,1244,1805,1819]" fontSize="6" pageId="9" pageNumber="44">th</superScript>
which is shorter than the 11
<superScript id="7C45C6757F36A814FEB7F8ADFEF8F710" attach="left" box="[275,288,1841,1855]" fontSize="6" pageId="9" pageNumber="44">th</superScript>
.
</paragraph>
<paragraph id="8B8F6B3D7F36A814FF63F8CBFBC9F797" blockId="9.[151,1437,1337,1977]" pageId="9" pageNumber="44">Pronotum longer than wide, PW/PL = 0.8, wider than head, PW/HW = 1.2, base as wide as front, PbW/PfW = 1.0; lateral margins bordered throughout, thick and a little widened in the middle part; two pairs of marginal setae present, at the level of 1/5 from front and a little before hind angle respectively.</paragraph>
<caption id="DF4F3BB57F35A817FF33F830FE28F7C6" ID-DOI="http://doi.org/10.5281/zenodo.3557815" ID-Zenodo-Dep="3557815" httpUri="https://zenodo.org/record/3557815/files/figure.png" pageId="10" pageNumber="45" startId="10.[151,250,1964,1989]" targetBox="[168,1420,174,1945]" targetPageId="10">
<paragraph id="8B8F6B3D7F35A817FF33F830FE28F7C6" blockId="10.[151,1436,1964,2025]" pageId="10" pageNumber="45">
<emphasis id="B944B72F7F35A817FF33F830FECAF7EA" bold="true" box="[151,274,1964,1989]" pageId="10" pageNumber="45">FIGURE 8.</emphasis>
Cave Hang Rắn, type locality of
<taxonomicName id="4C3010BE7F35A817FDFAF830FC49F7EB" authority="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille, 2019" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[606,913,1964,1988]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="45" phylum="Arthropoda" rank="species" species="impunctatus" status="sp. nov.">
<emphasis id="B944B72F7F35A817FDFAF830FC49F7EB" box="[606,913,1964,1988]" italics="true" pageId="10" pageNumber="45">Tonkinaphaenops impunctatus</emphasis>
</taxonomicName>
<emphasis id="B944B72F7F35A817FC33F830FC3EF7EB" bold="true" box="[919,998,1964,1988]" pageId="10" pageNumber="45">
<taxonomicNameLabel id="A2770A547F35A817FC33F830FC3EF7EB" box="[919,998,1964,1988]" pageId="10" pageNumber="45" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A. entrance; B. the wettest part of the cave where the beetles were collected).
</paragraph>
</caption>
<caption id="DF4F3BB57F34A816FF33F853FC76F7C7" ID-DOI="http://doi.org/10.5281/zenodo.3614846" ID-Zenodo-Dep="3614846" box="[151,942,1999,2024]" httpUri="https://zenodo.org/record/3614846/files/figure.png" pageId="11" pageNumber="46" targetBox="[393,1197,181,1964]" targetPageId="11">
<paragraph id="8B8F6B3D7F34A816FF33F853FC76F7C7" blockId="11.[151,942,1999,2024]" box="[151,942,1999,2024]" pageId="11" pageNumber="46">
<emphasis id="B944B72F7F34A816FF33F853FECBF7C7" bold="true" box="[151,275,1999,2024]" pageId="11" pageNumber="46">FIGURE 9.</emphasis>
<taxonomicName id="4C3010BE7F34A816FEBDF853FDF5F7C8" authority="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille, 2019" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[281,557,1999,2023]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="46" phylum="Arthropoda" rank="species" species="jingxicus" status="sp. nov.">
<emphasis id="B944B72F7F34A816FEBDF853FDF5F7C8" box="[281,557,1999,2023]" italics="true" pageId="11" pageNumber="46">Tonkinaphaenops jingxicus</emphasis>
</taxonomicName>
<emphasis id="B944B72F7F34A816FD97F853FD5CF7C8" bold="true" box="[563,644,1999,2023]" pageId="11" pageNumber="46">
<taxonomicNameLabel id="A2770A547F34A816FD97F853FD5CF7C8" box="[563,644,1999,2023]" pageId="11" pageNumber="46" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, habitus of holotype, female.
</paragraph>
</caption>
<caption id="DF4F3BB57F33A811FF33F82CFC72F7C2" ID-DOI="http://doi.org/10.5281/zenodo.3557817" ID-Zenodo-Dep="3557817" httpUri="https://zenodo.org/record/3557817/files/figure.png" pageId="12" pageNumber="47" startId="12.[151,250,1968,1993]" targetBox="[164,1424,166,1949]" targetPageId="12">
<paragraph id="8B8F6B3D7F33A811FF33F82CFC72F7C2" blockId="12.[151,1436,1968,2029]" pageId="12" pageNumber="47">
<emphasis id="B944B72F7F33A811FF33F82CFEF9F7E6" bold="true" box="[151,289,1968,1993]" pageId="12" pageNumber="47">FIGURE 10.</emphasis>
Cave Nianluo Dong, type locality of
<taxonomicName id="4C3010BE7F33A811FD00F82DFC60F7E6" authority="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille, 2019" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[676,952,1969,1993]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="jingxicus" status="sp. nov.">
<emphasis id="B944B72F7F33A811FD00F82DFC60F7E6" box="[676,952,1969,1993]" italics="true" pageId="12" pageNumber="47">Tonkinaphaenops jingxicus</emphasis>
</taxonomicName>
<emphasis id="B944B72F7F33A811FC64F82DFBC9F7E6" bold="true" box="[960,1041,1969,1993]" pageId="12" pageNumber="47">
<taxonomicNameLabel id="A2770A547F33A811FC64F82DFBC9F7E6" box="[960,1041,1969,1993]" pageId="12" pageNumber="47" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(A. red arrow showing the location of entrance; B. entrance; C. a running individual of
<taxonomicName id="4C3010BE7F33A811FD2CF849FC43F7C2" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[648,923,2005,2029]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="47" phylum="Arthropoda" rank="species" species="jingxicus">
<emphasis id="B944B72F7F33A811FD2CF849FC43F7C2" box="[648,923,2005,2029]" italics="true" pageId="12" pageNumber="47">Tonkinaphaenops jingxicus</emphasis>
</taxonomicName>
).
</paragraph>
</caption>
<paragraph id="8B8F6B3D7F32A810FF63FF05FD64F148" blockId="13.[151,1437,153,683]" pageId="13" pageNumber="48">
Elytra slightly more oviform-shaped, longer than wide, EL/EW = 1.7, widest at about the level of 1/3 from base, much longer than pronotum, EL/PL = 2.5, wider than pronotum, EW/PW = 1.9; chaetotaxy as in
<figureCitation id="130B77B87F32A810FB54FF21FA8EF0F8" box="[1264,1366,189,215]" captionStart="FIGURE 14" captionStartId="16.[151,250,1912,1937]" captionTargetBox="[214,1391,181,1869]" captionTargetId="figure@16.[189,1398,181,1887]" captionTargetPageId="16" captionText="FIGURE 14. Chaetotaxal pattern of Tonkinaphaenops (A. Tonkinaphaenops marinae Deuve, 2013, holotype, male from cave TH20; B, C. Tonkinaphaenops yinquanicus sp. nov.: B. male from Yinquan Dong, C. female from Yinquan Dong; D. Tonkinaphaenops impunctatus sp. nov., male from Hang Rắn cave; E. Tonkinaphaenops jingxicus sp. nov., female from Nianluo Dong; F. Tonkinaphaenops anthonyi sp. nov., male from Hang Dõi cave)." figureDoi="http://doi.org/10.5281/zenodo.3557823" httpUri="https://zenodo.org/record/3557823/files/figure.png" pageId="13" pageNumber="48">Fig. 14E</figureCitation>
: basal pores present at the beginning of the 2
<superScript id="7C45C6757F32A810FDE2FF7CFD8EF0C1" attach="left" box="[582,598,224,238]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
stria; 4 dorsal setiferous pores present in 3
<superScript id="7C45C6757F32A810FB9EFF7CFB90F0C1" attach="left" box="[1082,1096,224,238]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
stria; the first umbilicate pore shifted to level of the 7
<superScript id="7C45C6757F32A810FE3FFE98FE70F13D" attach="left" box="[411,424,260,274]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
stria and a little before the 2
<superScript id="7C45C6757F32A810FD4EFE98FD22F13D" attach="left" box="[746,762,260,274]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
pore; the 3
<superScript id="7C45C6757F32A810FCDDFE98FC5FF13D" attach="left" box="[889,903,260,274]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
pore of marginal umbilicate series much closer to 2
<superScript id="7C45C6757F32A810FF65FEB4FF09F119" attach="left" box="[193,209,296,310]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
than to 4
<superScript id="7C45C6757F32A810FE9DFEB4FE9EF119" attach="none" box="[313,326,296,310]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
, making distance between 3
<superScript id="7C45C6757F32A810FD25FEB4FD57F119" attach="left" box="[641,655,296,310]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
and 4
<superScript id="7C45C6757F32A810FD76FEB4FD07F119" attach="left" box="[722,735,296,310]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
twice as that between 2
<superScript id="7C45C6757F32A810FC43FEB4FC2FF119" attach="left" box="[999,1015,296,310]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
and 3
<superScript id="7C45C6757F32A810FB9FFEB4FB91F119" attach="none" box="[1083,1097,296,310]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
; the middle group at about the 3/5 of elytra from base; two apical pores present.
</paragraph>
<paragraph id="8B8F6B3D7F32A810FF63FEEDFE50F1A4" blockId="13.[151,1437,153,683]" box="[199,392,369,395]" pageId="13" pageNumber="48">
<emphasis id="B944B72F7F32A810FF63FEEDFED3F1A4" bold="true" box="[199,267,369,395]" pageId="13" pageNumber="48">Male.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F32A810FF63FE09FBC1F180" box="[199,1049,405,431]" pageId="13" pageNumber="48" type="etymology">
<paragraph id="8B8F6B3D7F32A810FF63FE09FBC1F180" blockId="13.[151,1437,153,683]" box="[199,1049,405,431]" pageId="13" pageNumber="48">
<emphasis id="B944B72F7F32A810FF63FE09FE96F180" bold="true" box="[199,334,405,431]" pageId="13" pageNumber="48">Etymology.</emphasis>
The species epithet is an adjective referring to the
<typeStatus id="548BD59F7F32A810FC2CFE09FC60F180" box="[904,952,405,431]" pageId="13" pageNumber="48">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F32A810FF63FE25FC77F1D8" pageId="13" pageNumber="48" type="discussion">
<paragraph id="8B8F6B3D7F32A810FF63FE25FC77F1D8" blockId="13.[151,1437,153,683]" pageId="13" pageNumber="48">
<emphasis id="B944B72F7F32A810FF63FE25FEE3F1FC" bold="true" box="[199,315,441,467]" pageId="13" pageNumber="48">Remarks.</emphasis>
This new species is close to
<taxonomicName id="4C3010BE7F32A810FD27FE26FCFEF1FC" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[643,806,442,467]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="yinquanicus">
<emphasis id="B944B72F7F32A810FD27FE26FCFEF1FC" box="[643,806,442,467]" italics="true" pageId="13" pageNumber="48">T. yinquanicus</emphasis>
</taxonomicName>
, but differs in elytra with shoulders less developed, and the 3
<superScript id="7C45C6757F32A810FF6AFE40FF04F1C5" attach="left" box="[206,220,476,490]" fontSize="6" pageId="13" pageNumber="48">rd</superScript>
pore of marginal umbilicate series much closer to 2
<superScript id="7C45C6757F32A810FC86FE40FCEAF1C5" attach="left" box="[802,818,476,490]" fontSize="6" pageId="13" pageNumber="48">nd</superScript>
than to 4
<superScript id="7C45C6757F32A810FC3FFE40FC70F1C5" attach="none" box="[923,936,476,490]" fontSize="6" pageId="13" pageNumber="48">th</superScript>
.
</paragraph>
</subSubSection>
<subSubSection id="C32A38B67F32A810FF63FD9CFD33F284" pageId="13" pageNumber="48" type="distribution">
<paragraph id="8B8F6B3D7F32A810FF63FD9CFD01F234" blockId="13.[151,1437,153,683]" box="[199,729,512,540]" pageId="13" pageNumber="48">
<emphasis id="B944B72F7F32A810FF63FD9CFEB9F234" bold="true" box="[199,353,512,539]" pageId="13" pageNumber="48">Distribution.</emphasis>
<collectingRegion id="49F4A5DF7F32A810FEC3FD9DFE1FF234" box="[359,455,513,539]" country="China" name="Guangxi" pageId="13" pageNumber="48">Guangxi</collectingRegion>
(Jingxi),
<collectingCountry id="F3272BAD7F32A810FD90FD9DFDAFF234" box="[564,631,513,539]" name="China" pageId="13" pageNumber="48">China</collectingCountry>
(
<figureCitation id="130B77B87F32A810FD22FD9DFD10F234" box="[646,712,513,539]" captionStart="FIGURE 1" captionStartId="1.[151,250,1801,1826]" captionTargetBox="[159,1428,877,1779]" captionTargetId="figure@1.[151,1436,877,1781]" captionTargetPageId="1" captionText="FIGURE 1. Distribution map of Tonkinaphaenops Deuve, 2013" figureDoi="http://doi.org/10.5281/zenodo.3557805" httpUri="https://zenodo.org/record/3557805/files/figure.png" pageId="13" pageNumber="48">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8B8F6B3D7F32A810FF63FDB9FD33F284" blockId="13.[151,1437,153,683]" pageId="13" pageNumber="48">
Known so far only from a limestone cave called Nianluo Dong, at Biaoliang Cun, Ludong Zhen. Nianluo Dong is
<quantity id="4CC8C6D87F32A810FF15FDD5FED6F24C" box="[177,270,585,611]" metricMagnitude="4" metricUnit="m" metricValue="3.1399999999999997" pageId="13" pageNumber="48" unit="km" value="31.4">31.4 km</quantity>
and
<quantity id="4CC8C6D87F32A810FEE1FDD4FE7AF24C" box="[325,418,584,611]" metricMagnitude="4" metricUnit="m" metricValue="4.59" pageId="13" pageNumber="48" unit="km" value="45.9">45.9 km</quantity>
from the
<typeStatus id="548BD59F7F32A810FDB5FDD5FD99F24C" box="[529,577,585,611]" pageId="13" pageNumber="48">type</typeStatus>
localities of
<taxonomicName id="4C3010BE7F32A810FD70FDD6FCAFF24C" authorityName="Huang &amp; Tú &amp; Phú &amp; Tian &amp; Faille" authorityYear="2019" box="[724,887,586,611]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="yinquanicus">
<emphasis id="B944B72F7F32A810FD70FDD6FCAFF24C" box="[724,887,586,611]" italics="true" pageId="13" pageNumber="48">T. yinquanicus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3010BE7F32A810FC0BFDD6FBF0F24C" authorityName="Deuve" authorityYear="2013" box="[943,1064,586,611]" class="Insecta" family="Carabidae" genus="Tonkinaphaenops" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="48" phylum="Arthropoda" rank="species" species="marinae">
<emphasis id="B944B72F7F32A810FC0BFDD6FBF0F24C" box="[943,1064,586,611]" italics="true" pageId="13" pageNumber="48">T. marinae</emphasis>
</taxonomicName>
respectively. The cave is located at the middle of a hill, with a small and unnoticeable entrance. The trechine beetles were found and collected in a large chamber which was wet and muddy (
<figureCitation id="130B77B87F32A810FDD3FD0DFD06F284" box="[631,734,657,683]" captionStart="FIGURE 10" captionStartId="12.[151,250,1968,1993]" captionTargetBox="[164,1424,166,1949]" captionTargetId="figure@12.[164,1424,166,1949]" captionTargetPageId="12" captionText="FIGURE 10. Cave Nianluo Dong, type locality of Tonkinaphaenops jingxicus sp. nov. (A. red arrow showing the location of entrance; B. entrance; C. a running individual of Tonkinaphaenops jingxicus)." figureDoi="http://doi.org/10.5281/zenodo.3557817" httpUri="https://zenodo.org/record/3557817/files/figure.png" pageId="13" pageNumber="48">Fig. 10C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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