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<document id="1E0A39030396A7A054A1775E8825E8D6" ID-CLB-Dataset="4131" ID-DOI="10.5852/ejt.2021.773.1509" ID-GBIF-Dataset="30ee5d5d-c02a-4239-836e-f921911db9a2" ID-ISSN="2118-9773" ID-Zenodo-Dep="5536301" ID-ZooBank="2A159A89-64D1-4352-8E0B-FA546983C99A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632919176088" checkinUser="felipe" docAuthor="Kantor, Yuri I. &amp; Puillandre, Nicolas" docDate="2021" docId="E43BBB1DFFD4FFD826017EB47558F337" docLanguage="en" docName="ejt-773_19-60.pdf" docOrigin="European Journal of Taxonomy 773" docSource="http://dx.doi.org/10.5852/ejt.2021.773.1509" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Sibogasyrinx pagodiformis Kantor &amp; Puillandre 2021, sp. nov." docType="treatment" docUuid="DAA613D6-5AB6-4F89-A0AD-0C718A60307E" docUuidSource="ZooBank" docVersion="8" lastPageNumber="54" masterDocId="1802C365FFF4FFFB240678167111F117" masterDocTitle="Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae)" masterLastPageNumber="60" masterPageNumber="19" pageNumber="51" updateTime="1699458178031" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="B66859A90F609ABB2294CE91013EF792" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="11BAAAEF44A7F46DC5B211F30C1D957F">
<mods:title id="FFF9F9D59AA3396633E6F6F35035A483">Rare, deep-water and similar: revision of Sibogasyrinx (Conoidea: Cochlespiridae)</mods:title>
</mods:titleInfo>
<mods:name id="78F965ADACA926161F840C4AB81C3AFB" type="personal">
<mods:role id="ACEE60E1A84FFC6B96929C4C4097F7FF">
<mods:roleTerm id="A1D37BC3E61B14E44ED0AC53907F2FD3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F8350EC40034970457815D7A1B1878B9">Kantor, Yuri I.</mods:namePart>
<mods:nameIdentifier id="40BAE14827144CE7AC68436D3C1ADC58" type="LSID">48F89A50-4CAC-4143-9D8B-73BA82735EC9</mods:nameIdentifier>
<mods:affiliation id="71D4B353A4A090D36F836348B0D6715D">Severtsov Institute of Ecology and Evolution, Russian Academy of Sciences, Leninski prospect 33, 119071 Moscow, Russian Federation. &amp; Institut Systématique Evolution Biodiversité (ISYEB), Muséum national d’histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="BB6941C3762786ECE18211B90E2A480C" type="email">kantor.yuri1956@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="2C72F5DCA62830878B01D4E4573C63D1" type="personal">
<mods:role id="5CD5F25296EA69B36AFCC3C2A520F321">
<mods:roleTerm id="93F69918BF368CEC691699EEE3ED162C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B2CAF6A472CA4C22BC8BDC2D05E44C2D">Puillandre, Nicolas</mods:namePart>
<mods:nameIdentifier id="7E5DB86B30594975730EF712B1A90E66" type="LSID">00565F2A-C170-48A1-AAD9-16559C536E4F</mods:nameIdentifier>
<mods:affiliation id="C62975C4971E1D61C4C7368A5CA9370E">Institut Systématique Evolution Biodiversité (ISYEB), Muséum national d’histoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, 57 rue Cuvier, 75005 Paris, France.</mods:affiliation>
<mods:nameIdentifier id="47A21B7C9661C3336D46D244F74391B7" type="email">puillandre@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="1ECC3BBCA17ED7F5D31E102CCC3B5A05">text</mods:typeOfResource>
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<mods:title id="425EE29362768BCB2A419727E16888E5">European Journal of Taxonomy</mods:title>
</mods:titleInfo>
<mods:part id="AAD890D8460D13127CFDA7201B79CF35">
<mods:date id="753CE01932DB141C2C75C6F83E8DD5A9">2021</mods:date>
<mods:detail id="42AF3741F7930A7BDF9AB0D61AD3AAA7" type="pubDate">
<mods:number id="9FA6E7D75D5E09B041476E8780C1C7FF">2021-09-28</mods:number>
</mods:detail>
<mods:detail id="11E500182A615BCDDF9DDEDC8695A63F" type="volume">
<mods:number id="8A9BC0111322CA53D047105DEFBF979D">773</mods:number>
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<mods:start id="4EB17DBD5B264CD88E8B809FEB431D7D">19</mods:start>
<mods:end id="C27BBEC9F75803889048CE154D262089">60</mods:end>
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<mods:url id="120B114DFA3DF70C26EF965062270142">http://dx.doi.org/10.5852/ejt.2021.773.1509</mods:url>
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<mods:classification id="89312C571398D1630A17642D7F1A61CD">journal article</mods:classification>
<mods:identifier id="ABB9433E5E94C923EF3EBDD910533D6F" type="CLB-Dataset">4131</mods:identifier>
<mods:identifier id="24B580399B1CEAE170C2E200F36A37C2" type="DOI">10.5852/ejt.2021.773.1509</mods:identifier>
<mods:identifier id="3C7C4E7948B5AF7CD2E16ED0B94858F2" type="GBIF-Dataset">30ee5d5d-c02a-4239-836e-f921911db9a2</mods:identifier>
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<mods:identifier id="B3534FAB954ED060BBD8C810E8C87E46" type="Zenodo-Dep">5536301</mods:identifier>
<mods:identifier id="5182069407D842672BA862CB28CA92C6" type="ZooBank">2A159A89-64D1-4352-8E0B-FA546983C99A</mods:identifier>
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<treatment id="E43BBB1DFFD4FFD826017EB47558F337" ID-DOI="http://doi.org/10.5281/zenodo.5536375" ID-GBIF-Taxon="188570454" ID-Zenodo-Dep="5536375" LSID="urn:lsid:zoobank.org:act:DAA613D6-5AB6-4F89-A0AD-0C718A60307E" httpUri="http://treatment.plazi.org/id/E43BBB1DFFD4FFD826017EB47558F337" lastPageId="35" lastPageNumber="54" pageId="32" pageNumber="51">
<subSubSection id="24885980FFD4FFDB26017EB4753DF7AA" box="[519,1068,1698,1725]" pageId="32" pageNumber="51" type="nomenclature">
<paragraph id="6C2D0A0BFFD4FFDB26017EB4753DF7AA" blockId="32.[356,1232,1698,1795]" box="[519,1068,1698,1725]" pageId="32" pageNumber="51">
<heading id="3765BD67FFD4FFDB26017EB4753DF7AA" box="[519,1068,1698,1725]" centered="true" fontSize="11" level="2" pageId="32" pageNumber="51" reason="2">
<taxonomicName id="AB927188FFD4FFDB26017EB4725CF7AA" ID-CoL="8QGMW" authority="Kantor &amp; Puillandre, 2021" authorityName="Kantor &amp; Puillandre" authorityYear="2021" box="[519,845,1698,1725]" class="Gastropoda" family="Cochlespiridae" genus="Sibogasyrinx" kingdom="Animalia" order="Neogastropoda" pageId="32" pageNumber="51" phylum="Mollusca" rank="species" species="pagodiformis" status="sp. nov.">
<emphasis id="5EE6D619FFD4FFDB26017EB4725CF7AA" bold="true" box="[519,845,1698,1725]" italics="true" pageId="32" pageNumber="51">Sibogasyrinx pagodiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45D56B62FFD4FFDB27527EB572BFF7AA" box="[852,942,1699,1725]" pageId="32" pageNumber="51" rank="species">sp. nov.</taxonomicNameLabel>
(= PSH 5)
</heading>
</paragraph>
</subSubSection>
<subSubSection id="24885980FFD4FFDB25627ED07266F614" pageId="32" pageNumber="51" type="description">
<paragraph id="6C2D0A0BFFD4FFDB25627ED075C1F7F6" blockId="32.[356,1232,1698,1795]" box="[356,1232,1734,1761]" pageId="32" pageNumber="51">
<uri id="18030609FFD4FFDB25627ED075C1F7F6" box="[356,1232,1734,1761]" pageId="32" pageNumber="51">
urn:lsid:zoobank.org:act:
<uuid id="183430DEFFD4FFDB26847ED075C1F7F6" box="[642,1232,1734,1761]" pageId="32" pageNumber="51">DAA613D6-5AB6-4F89-A0AD-0C718A60307E</uuid>
</uri>
</paragraph>
<paragraph id="6C2D0A0BFFD4FFDB26BA7EFF7266F614" blockId="32.[356,1232,1698,1795]" box="[700,887,1769,1795]" pageId="32" pageNumber="51">
<figureCitation id="F4A9168EFFD4FFDB26BA7EFF7204F614" box="[700,789,1769,1795]" captionStart="Fig" captionStartId="23.[189,232,1750,1776]" captionTargetBox="[196,1389,265,1726]" captionTargetId="figure-126@23.[196,1389,265,1726]" captionTargetPageId="23" captionText="Fig. 8. Radulae of species of Sibogasyrinx Powell, 1969. A–B. Sibogasyrinx sangeri Kantor, Fedosov &amp; Puillandre, 2018, MNHN-IM-2009-16995 (shell seen on Fig. 7C). A. Part of radula with fully formed marginal teeth; white hollow arrow indicates overlapping edges of the tooth at its tip. B. Part of radula showing transition between unfolded (white arrow) and completely longitudinally folded (black arrow) marginal teeth (ae = anterior tooth edge; pe = posterior tooth edge). C. Sibogasyrinx filosa Ardovini, 2021, Solomon Islands, MNHN-IM-2009-16831 (shell seen Fig. 9E–F). D. Sibogasyrinx lolae sp. nov., MNHN-IM-2009-29311. E. Sibogasyrinx maximei sp. nov., holotype, MNHN-IM-2013-45883, anterior end of radula. F. Sibogasyrinx pagodiformis sp. nov., MNHN-IM-2009-11327." figureDoi="http://doi.org/10.5281/zenodo.5536321" httpUri="https://zenodo.org/record/5536321/files/figure.png" pageId="32" pageNumber="51">Figs 8F</figureCitation>
,
<figureCitation id="F4A9168EFFD4FFDB27277EFF7266F614" box="[801,887,1769,1795]" captionStart="Fig" captionStartId="34.[189,232,1892,1918]" captionTargetBox="[189,1399,396,1868]" captionTargetId="figure-117@34.[189,1399,396,1868]" captionTargetPageId="34" captionText="Fig. 12. A–D. Sibogasyrinx pagodiformis sp. nov. A–B. Solomon Islands, holotype, MNHN- IM-2009-16825, SL 28.8 mm. C–D. Vanuatu. MNHN-IM-2009-11327, SL 24.3 mm. E–H. Sibogasyrinx elbakyanae Kantor, Puillandre &amp; Bouchet sp. nov., holotype, Solomon Islands, MNHN-IM-2009-16834, SL 27.3 mm. H. Enlarged fragment of penultimate and antepenultimate whorls. A–G to same scale." figureDoi="http://doi.org/10.5281/zenodo.5536329" httpUri="https://zenodo.org/record/5536329/files/figure.png" pageId="32" pageNumber="51">12A–D</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="24885980FFD4FFDB24BB7F24750AF6F3" pageId="32" pageNumber="51" type="diagnosis">
<paragraph id="6C2D0A0BFFD4FFDB24BB7F247027F65B" blockId="32.[189,310,1842,1868]" box="[189,310,1842,1868]" pageId="32" pageNumber="51">
<heading id="3765BD67FFD4FFDB24BB7F247027F65B" bold="true" box="[189,310,1842,1868]" fontSize="11" level="3" pageId="32" pageNumber="51" reason="3">
<emphasis id="5EE6D619FFD4FFDB24BB7F247027F65B" bold="true" box="[189,310,1842,1868]" pageId="32" pageNumber="51">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="6C2D0A0BFFD4FFDB24BB7F77750AF6F3" blockId="32.[189,1399,1889,2020]" pageId="32" pageNumber="51">
Shell small, reaching
<quantity id="AB6AA7EEFFD4FFDB25BC7F77731DF66C" box="[442,524,1889,1915]" metricMagnitude="-2" metricUnit="m" metricValue="2.9" pageId="32" pageNumber="51" unit="mm" value="29.0">29 mm</quantity>
in length, pagodiform, with strongly concave subsutural ramp bearing two low spiral cords and distinct subsutural nodules. Axial sculpture of distinct opisthocline ribs extending to shell base, 10–11 per whorl. Spiral sculpture of closely spaced cords on and below shoulder. Radula lacking central tooth, marginal teeth trough-shaped with thickened edges.
</paragraph>
</subSubSection>
<subSubSection id="24885980FFD5FFDA24BB791F72FEF046" box="[189,1007,265,338]" pageId="33" pageNumber="52" type="etymology">
<paragraph id="6C2D0A0BFFD5FFDA24BB791F7055F034" blockId="33.[189,324,265,291]" box="[189,324,265,291]" pageId="33" pageNumber="52">
<heading id="3765BD67FFD5FFDA24BB791F7055F034" bold="true" box="[189,324,265,291]" fontSize="11" level="3" pageId="33" pageNumber="52" reason="3">
<emphasis id="5EE6D619FFD5FFDA24BB791F7055F034" bold="true" box="[189,324,265,291]" pageId="33" pageNumber="52">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB792172FEF046" blockId="33.[189,1007,311,338]" box="[189,1007,311,338]" pageId="33" pageNumber="52">The name reflects the characteristic shell shape, resembling a pagoda.</paragraph>
</subSubSection>
<subSubSection id="24885980FFD5FFDA24BB797870F2F36D" pageId="33" pageNumber="52" type="materials_examined">
<paragraph id="6C2D0A0BFFD5FFDA24BB797870BBF09F" blockId="33.[189,426,366,392]" box="[189,426,366,392]" pageId="33" pageNumber="52">
<heading id="3765BD67FFD5FFDA24BB797870BBF09F" bold="true" box="[189,426,366,392]" fontSize="11" level="3" pageId="33" pageNumber="52" reason="3">
<emphasis id="5EE6D619FFD5FFDA24BB797870BBF09F" bold="true" box="[189,426,366,392]" pageId="33" pageNumber="52">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24EC798B7388F0E9" blockId="33.[189,1398,413,510]" pageId="33" pageNumber="52">
<materialsCitation id="DCFA0056FFD5FFDA24EC798B7388F0E9" ID-GBIF-Occurrence="3381483305" country="Solomon Islands" elevation="1326" latitude="-8.466666" location="SOLOMON ISLANDS" longLatPrecision="1301" longitude="160.53334" pageId="33" pageNumber="52" specimenCode="MNHN-IM-2009-16825" specimenCount="1" stateProvince="Malaita" typeStatus="holotype">
<emphasis id="5EE6D619FFD5FFDA24EC798B704DF0A0" bold="true" box="[234,348,413,439]" pageId="33" pageNumber="52">
<typeStatus id="B329B4A9FFD5FFDA24EC798B704DF0A0" box="[234,348,413,439]" pageId="33" pageNumber="52" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="14854A9BFFD5FFDA24BB79D670C0F0CD" box="[189,465,448,474]" name="Solomon Islands" pageId="33" pageNumber="52">SOLOMON ISLANDS</collectingCountry>
•
<locationDeviation id="7D9F1329FFD5FFDA25EE79D673A1F0CD" box="[488,688,448,474]" pageId="33" pageNumber="52">NW of Malaita I.</locationDeviation>
;
<geoCoordinate id="09A66CCCFFD5FFDA26BB79A87232F0CD" box="[701,803,446,474]" degrees="08" direction="south" minutes="28" orientation="latitude" pageId="33" pageNumber="52" precision="925" value="-8.466666">08°28′ S</geoCoordinate>
,
<geoCoordinate id="09A66CCCFFD5FFDA272979A872B7F0CD" box="[815,934,446,474]" degrees="160" direction="east" minutes="32" orientation="longitude" pageId="33" pageNumber="52" precision="925" value="160.53334">160°32′ E</geoCoordinate>
; depth
<quantity id="AB6AA7EEFFD5FFDA27FF79A9758FF0CD" box="[1017,1182,447,474]" metricMagnitude="3" metricUnit="m" metricValue="1.326" metricValueMax="1.402" metricValueMin="1.25" pageId="33" pageNumber="52" unit="m" value="1326.0" valueMax="1402.0" valueMin="1250.0">
<elevation id="E7BFED38FFD5FFDA27FF79A9758FF0CD" box="[1017,1182,447,474]" metricMagnitude="3" metricUnit="m" metricValue="1.326" metricValueMax="1.402" metricValueMin="1.25" pageId="33" pageNumber="52" unit="m" value="1326.0" valueMax="1402.0" valueMin="1250.0">1250–1402 m</elevation>
</quantity>
; SALOMONBOA 3, stn CP2789;
<specimenCode id="3C34A270FFD5FFDA257479F57385F0E9" box="[370,660,483,510]" pageId="33" pageNumber="52">MNHN-IM-2009-16825</specimenCode>
.
</materialsCitation>
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24EC7A0F732AF322" blockId="33.[189,1398,537,634]" box="[234,571,537,565]" pageId="33" pageNumber="52">
<emphasis id="5EE6D619FFD5FFDA24EC7A0F70B7F324" bold="true" box="[234,422,537,563]" pageId="33" pageNumber="52">Other material</emphasis>
(sequenced)
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB7A2B70F2F36D" blockId="33.[189,1398,537,634]" pageId="33" pageNumber="52">
<materialsCitation id="DCFA0056FFD5FFDA24BB7A2B70CFF36D" ID-GBIF-Occurrence="3381483345" country="Vanuatu" elevation="1274" latitude="-16.0" location="VANUATU" longLatPrecision="1283" longitude="167.63333" pageId="33" pageNumber="52" specimenCode="MNHN-IM-2009-11327" specimenCount="1" specimenCount-larva="1">
<collectingCountry id="14854A9BFFD5FFDA24BB7A2B7056F340" box="[189,327,573,599]" name="Vanuatu" pageId="33" pageNumber="52">VANUATU</collectingCountry>
•
<specimenCount id="7A94C182FFD5FFDA255B7A2B709DF340" box="[349,396,573,599]" pageId="33" pageNumber="52" type="larva">1 lv</specimenCount>
;
<locationDeviation id="7D9F1329FFD5FFDA25917A2B7345F340" box="[407,596,573,599]" pageId="33" pageNumber="52">E of Malekula I.</locationDeviation>
;
<geoCoordinate id="09A66CCCFFD5FFDA26677A2D73D7F340" box="[609,710,571,599]" degrees="16" direction="south" minutes="00" orientation="latitude" pageId="33" pageNumber="52" precision="925" value="-16.0">16°00′ S</geoCoordinate>
,
<geoCoordinate id="09A66CCCFFD5FFDA26D77A2D7259F340" box="[721,840,571,599]" degrees="167" direction="east" minutes="38" orientation="longitude" pageId="33" pageNumber="52" precision="925" value="167.63333">167°38′ E</geoCoordinate>
; depth
<quantity id="AB6AA7EEFFD5FFDA279C7A2B752FF340" box="[922,1086,573,600]" metricMagnitude="3" metricUnit="m" metricValue="1.2734999999999999" metricValueMax="1.285" metricValueMin="1.262" pageId="33" pageNumber="52" unit="m" value="1273.5" valueMax="1285.0" valueMin="1262.0">
<elevation id="E7BFED38FFD5FFDA279C7A2B752FF340" box="[922,1086,573,600]" metricMagnitude="3" metricUnit="m" metricValue="1.2734999999999999" metricValueMax="1.285" metricValueMin="1.262" pageId="33" pageNumber="52" unit="m" value="1273.5" valueMax="1285.0" valueMin="1262.0">1262–1285 m</elevation>
</quantity>
; SANTO 2006, stn AT139;
<specimenCode id="3C34A270FFD5FFDA24BB7A7670CFF36D" box="[189,478,608,634]" pageId="33" pageNumber="52">MNHN-IM-2009-11327</specimenCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="24885980FFD5FFDA24BB7A817249F649" pageId="33" pageNumber="52" type="description">
<paragraph id="6C2D0A0BFFD5FFDA24BB7A81705FF3A6" blockId="33.[189,334,663,689]" box="[189,334,663,689]" pageId="33" pageNumber="52">
<heading id="3765BD67FFD5FFDA24BB7A81705FF3A6" bold="true" box="[189,334,663,689]" fontSize="11" level="3" pageId="33" pageNumber="52" reason="3">
<emphasis id="5EE6D619FFD5FFDA24BB7A81705FF3A6" bold="true" box="[189,334,663,689]" pageId="33" pageNumber="52">Description</emphasis>
</heading>
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB7AD0703FF214" blockId="33.[189,1398,709,771]" pageId="33" pageNumber="52">
MEASUREMENTS (
<typeStatus id="B329B4A9FFD5FFDA25807AD370E1F3F7" box="[390,496,709,736]" pageId="33" pageNumber="52" type="holotype">holotype</typeStatus>
). SL
<quantity id="AB6AA7EEFFD5FFDA26327AD073B3F3F7" box="[564,674,710,736]" metricMagnitude="-2" metricUnit="m" metricValue="2.88" pageId="33" pageNumber="52" unit="mm" value="28.8">28.8 mm</quantity>
, AL (with canal)
<quantity id="AB6AA7EEFFD5FFDA27707AD372F5F3F7" box="[886,996,709,736]" metricMagnitude="-2" metricUnit="m" metricValue="1.52" pageId="33" pageNumber="52" unit="mm" value="15.2">15.2 mm</quantity>
, AL (without canal)
<quantity id="AB6AA7EEFFD5FFDA20D87AD0742CF3F7" box="[1246,1341,710,736]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="33" pageNumber="52" unit="mm" value="9.9">9.9 mm</quantity>
, SW
<quantity id="AB6AA7EEFFD5FFDA24B97AFF703BF214" box="[191,298,745,771]" metricMagnitude="-2" metricUnit="m" metricValue="1.19" pageId="33" pageNumber="52" unit="mm" value="11.9">11.9 mm</quantity>
.
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB7B36751CF4A4" blockId="33.[186,1401,799,1460]" pageId="33" pageNumber="52">
SHELL (
<typeStatus id="B329B4A9FFD5FFDA25107B097091F22D" box="[278,384,799,826]" pageId="33" pageNumber="52" type="holotype">holotype</typeStatus>
). Moderately thick, except for fragile, partially chipped outer aperture lip; pagodiform, with moderately high spire and moderately long, narrow siphonal canal inclined to abaxially left. Protoconch missing (rendering exact an whorl count and protoconch measurements impossible), upper teleoconch whorls eroded. Teleoconch whorls rapidly increasing and strongly angled at shoulder, more than 8 remaining. Suture shallowly impressed, subsutural ramp broad, concave. A row of distinct, sharp, subsutural nodules on upper teleoconch whorls. Nodules less distinct on last three whorls and situated on short, slightly raised, prosocline axial ribs. Number of ribs on uppermost whorls indeterminate due to shell surface erosion, number of nodules per whorl increasing from 16–17 on first intact whorl to 22 on penultimate and 26 on last whorl. Subsutural ramp smooth except for nodules and two low, indistinct subsutural cords. Shoulder with pronounced, thickened, opisthocline axial ribs on all teleoconch whorls. Axial ribs on upper teleoconch whorls situated just above suture, extending to shell base on last whorl, number of ribs constant, 11–12 per whorl. Ribs intersected by closely spaced, rounded spiral cords, number increasing from 5 on uppermost remaining whorl to 10 on penultimate whorl. Last whorl with total of 38 cords, including 18 on canal. Cords narrow, their intervals 0.5–1.5 times cord width. Shell base rapidly narrowing towards narrow, moderately long siphonal canal. Aperture narrow, constricted posteriorly, with thin, moderately broad parietal callus, outer lip partially broken, convex on subsutural ramp, with rounded angle at shoulder, convex below shoulder and weakly concave at transition to canal. Anal sinus judging from growth lines shallow, subsutural, broadly arcuate. Shell flesh-coloured, with darker base, glossy. Periostracum thin, persisting in some cord intervals.
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB7DC6703BF760" blockId="33.[189,1400,1488,1655]" pageId="33" pageNumber="52">ANATOMY (n = 1; MNHN-IM-2009-11327). Female. Proboscis relatively short, straight, cylindrical. Proboscis retractors large, in two bundles attached at proboscis base. Buccal mass within proboscis, rather long, about half length of proboscis, radular sac opening on right. Venom gland very thick, not long, opening with very short constriction just posterior to nerve ring. Venom bulb moderately large, elongate.</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB7E827249F649" blockId="33.[188,1399,1683,1886]" pageId="33" pageNumber="52">
RADULA (n = 2; MNHN-IM-2009-16825, AL
<quantity id="AB6AA7EEFFD5FFDA26D17E827220F7B9" box="[727,817,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="9.9" pageId="33" pageNumber="52" unit="mm" value="9.9">9.9 mm</quantity>
, IM-2009-11327, AL ca
<quantity id="AB6AA7EEFFD5FFDA20507E82758BF7B9" box="[1110,1178,1684,1710]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="33" pageNumber="52" unit="mm" value="9.0">9 mm</quantity>
). Relatively short, comprising 20–24 rows of teeth, 8–9 nascent. Radula length
<quantity id="AB6AA7EEFFD5FFDA277D7EA17519F7C6" box="[891,1032,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.35" metricValueMax="1.6" metricValueMin="1.1" pageId="33" pageNumber="52" unit="mm" value="1.35" valueMax="1.6" valueMin="1.1">1.1–1.6 mm</quantity>
(12–16% of AL without canal), tooth length 150 μm (1.7% of AL without canal). Central tooth absent. Marginal teeth trough-shaped when mature (
<figureCitation id="F4A9168EFFD5FFDA25627EEB70ADF600" box="[356,444,1789,1815]" captionStart="Fig" captionStartId="23.[189,232,1750,1776]" captionTargetBox="[196,1389,265,1726]" captionTargetId="figure-126@23.[196,1389,265,1726]" captionTargetPageId="23" captionText="Fig. 8. Radulae of species of Sibogasyrinx Powell, 1969. A–B. Sibogasyrinx sangeri Kantor, Fedosov &amp; Puillandre, 2018, MNHN-IM-2009-16995 (shell seen on Fig. 7C). A. Part of radula with fully formed marginal teeth; white hollow arrow indicates overlapping edges of the tooth at its tip. B. Part of radula showing transition between unfolded (white arrow) and completely longitudinally folded (black arrow) marginal teeth (ae = anterior tooth edge; pe = posterior tooth edge). C. Sibogasyrinx filosa Ardovini, 2021, Solomon Islands, MNHN-IM-2009-16831 (shell seen Fig. 9E–F). D. Sibogasyrinx lolae sp. nov., MNHN-IM-2009-29311. E. Sibogasyrinx maximei sp. nov., holotype, MNHN-IM-2013-45883, anterior end of radula. F. Sibogasyrinx pagodiformis sp. nov., MNHN-IM-2009-11327." figureDoi="http://doi.org/10.5281/zenodo.5536321" httpUri="https://zenodo.org/record/5536321/files/figure.png" pageId="33" pageNumber="52">Fig. 8F</figureCitation>
), with slightly thickened edges and sharp pointed tips. Teeth initially forming as a flat, elongate-oval plate, its edges gradually thickening along radular membrane and then by longitudinal folding of each tooth on 10–11
<superScript id="9BE7A743FFD5FFDA26207F557322F646" attach="left" box="[550,563,1859,1873]" fontSize="6" pageId="33" pageNumber="52">th</superScript>
row, counting from rear.
</paragraph>
</subSubSection>
<subSubSection id="24885980FFD5FFD824BB7F6D72DFF0A7" lastPageId="35" lastPageNumber="54" pageId="33" pageNumber="52" type="discussion">
<paragraph id="6C2D0A0BFFD5FFDA24BB7F6D703EF682" blockId="33.[189,303,1915,1941]" box="[189,303,1915,1941]" pageId="33" pageNumber="52">
<heading id="3765BD67FFD5FFDA24BB7F6D703EF682" bold="true" box="[189,303,1915,1941]" fontSize="11" level="3" pageId="33" pageNumber="52" reason="3">
<emphasis id="5EE6D619FFD5FFDA24BB7F6D703EF682" bold="true" box="[189,303,1915,1941]" pageId="33" pageNumber="52">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="6C2D0A0BFFD5FFDA24BB7FBF703DF6F1" blockId="33.[189,1399,1961,2022]" pageId="33" pageNumber="52">
The second live collected specimen has a broken shell base. In sculpture pattern it is very similar to the
<typeStatus id="B329B4A9FFD5FFDA24BB7FDA7039F6F1" box="[189,296,1996,2022]" pageId="33" pageNumber="52" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="6C2D0A0BFFD6FFD824BB791F72DFF0A7" blockId="34.[188,1398,265,362]" lastBlockId="35.[189,1400,265,432]" lastPageId="35" lastPageNumber="54" pageId="34" pageNumber="53">
The species is distinct from all congeners in having a relatively broad, pagodiform shell with a shorter canal. Conchologically the species is quite similar to
<taxonomicName id="AB927188FFD6FFD92725793B75B9F051" authority="(Watson, 1881)" baseAuthorityName="Watson" baseAuthorityYear="1881" box="[803,1192,300,327]" class="Gastropoda" family="Pseudomelatomidae" genus="Comitas" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="53" phylum="Mollusca" rank="species" species="paupera">
<emphasis id="5EE6D619FFD6FFD92725793B72FCF051" box="[803,1005,301,327]" italics="true" pageId="34" pageNumber="53">Comitas paupera</emphasis>
(Watson, 1881)
</taxonomicName>
, a highly variable species of uncertain generic affinity, but differing in lacking distinct spiral cords on the subsutural ramp. The
<typeStatus id="B329B4A9FFD7FFD824F6791F704DF034" box="[240,348,265,291]" pageId="35" pageNumber="54" type="lectotype">lectotype</typeStatus>
and
<typeStatus id="B329B4A9FFD7FFD82596791F7328F034" box="[400,569,265,291]" pageId="35" pageNumber="54" type="paralectotype">paralectotypes</typeStatus>
of the latter were illustrated by
<bibRefCitation id="080377FAFFD7FFD827A7791F7552F034" author="Sysoev A. V." box="[929,1091,265,291]" pageId="35" pageNumber="54" pagination="1 - 30" refId="ref22508" refString="Sysoev A. V. 1996. Deep-sea conoidean gastropods collected by the John Murray Expedition, 1933 - 34. Bulletin of the Natural History Museum of London, Zoology 62 (1): 1 - 30. Available from https: // www. biodiversitylibrary. org / page / 41008506 [accessed 7 Sep. 2021]." type="journal article" year="1996">Sysoev (1996</bibRefCitation>
: figs 42–48). Superficially, the new species also strongly resembles a miniature copy of
<taxonomicName id="AB927188FFD7FFD8278B793B7451F050" authority="(Powell, 1969)" baseAuthorityName="Powell" baseAuthorityYear="1969" box="[909,1344,300,327]" class="Gastropoda" family="Pseudomelatomidae" genus="Leucosyrinx" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="54" phylum="Mollusca" rank="species" species="luzonica">
<emphasis id="5EE6D619FFD7FFD8278B793B7599F051" box="[909,1160,300,326]" italics="true" pageId="35" pageNumber="54">Leucosyrinx luzonica</emphasis>
(
<bibRefCitation id="080377FAFFD7FFD8209F793A7426F050" author="Powell A. W. B." box="[1177,1335,300,327]" pageId="35" pageNumber="54" pagination="215 - 416" refId="ref21901" refString="Powell A. W. B. 1969. The family Turridae in the Indo-Pacific. Part 2. The subfamily Turriculinae. Indo- Pacific Mollusca 2 (10): 215 - 416. Available from https: // www. biodiversitylibrary. org / page / 49824050 [accessed 24 Aug. 2021]." type="journal article" year="1969">Powell, 1969</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="080377FAFFD7FFD824B87946708AF07E" author="Kantor Y. I. &amp; Fedosov A. E. &amp; Puillandre N." box="[190,411,335,362]" pageId="35" pageNumber="54" pagination="47 - 82" refId="ref21780" refString="Kantor Y. I., Fedosov A. E. &amp; Puillandre N. 2018. New and unusual deep-water Conoidea revised with shell, radula and DNA characters. Ruthenica, Russian Malacological Journal 28: 47 - 82." type="journal article" year="2018">
Kantor
<emphasis id="5EE6D619FFD7FFD8251179467042F07E" box="[279,339,335,361]" italics="true" pageId="35" pageNumber="54">et al.</emphasis>
2018
</bibRefCitation>
: fig. 21), but differs in the shape of the marginal radular teeth and foregut anatomy. Another species of
<taxonomicName id="AB927188FFD7FFD825AC79657328F09B" authorityName="Dall" authorityYear="1889" box="[426,569,371,396]" class="Gastropoda" family="Pseudomelatomidae" genus="Leucosyrinx" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="54" phylum="Mollusca" rank="genus">
<emphasis id="5EE6D619FFD7FFD825AC79657328F09B" box="[426,569,371,396]" italics="true" pageId="35" pageNumber="54">Leucosyrinx</emphasis>
</taxonomicName>
from the
<collectingCountry id="14854A9BFFD7FFD826B079647291F09B" box="[694,896,370,396]" name="Solomon Islands" pageId="35" pageNumber="54">Solomon Islands</collectingCountry>
,
<taxonomicName id="AB927188FFD7FFD827897965757CF09A" box="[911,1133,371,397]" class="Gastropoda" family="Pseudomelatomidae" genus="Leucosyrinx" kingdom="Animalia" order="Neogastropoda" pageId="35" pageNumber="54" phylum="Mollusca" rank="species" species="undefined-d">
<emphasis id="5EE6D619FFD7FFD827897965750FF09B" box="[911,1054,371,396]" italics="true" pageId="35" pageNumber="54">Leucosyrinx</emphasis>
sp. D
</taxonomicName>
, is similar to the new species in shell outline (
<figureCitation id="F4A9168EFFD7FFD825D17980732EF0A7" box="[471,575,406,432]" captionStart="Fig" captionStartId="37.[189,232,1680,1706]" captionTargetBox="[244,1343,265,1653]" captionTargetId="figure-159@37.[244,1343,265,1653]" captionTargetPageId="37" captionText="Fig. 13. Shells of species of Leucosyrinx Dall, 1889. A–B. Leucosyrinx verrillii (Dall, 1881), French Guiana, MNHN-IM-2013-56840, SL 25.1 mm. C. L. verrillii, French Guiana, MNHN-IM-2013-56288, SL 31.9 mm (radula seen on Fig. 11C). D–E. Leucosyrinx sp. G. D. Papua New Guinea, MNHN- IM-2009-17089, SL 32.2 mm (radula seen on Fig. 11E). E. Bismarck Sea, MNHN-IM-2013-19689, SL 47.3 mm.F. Leucosyrinx sp. D, Solomon Islands, MNHN-IM-2009-16769, SL 25 mm.G. Leucosyrinx sp. E, Madagascar, MNHN-IM-2009-16897, SL 46.1 mm. H. Leucosyrinx sp. A, Philippines, MNHN- IM-2007-42445, SL 34.1 mm. I. Leucosyrinx sp. B, Solomon Islands, MNHN IM-2009-16764, SL 29.9 mm. J. Leucosyrinx sp. C, Solomon Islands, MNHN-IM-2007-42503, SL 41.2 mm (radula seen on Fig. 11F). K. Leucosyrinx sp. F, Coral Sea, Chesterfield Plateau, MNHN-IM-2007-17846, SL 27.3 mm (radula seen on Fig. 11D). All shells to same scale." figureDoi="http://doi.org/10.5281/zenodo.5536331" httpUri="https://zenodo.org/record/5536331/files/figure.png" pageId="35" pageNumber="54">Fig. 13F</figureCitation>
), differing in radular morphology.
</paragraph>
</subSubSection>
<caption id="38ED5A83FFD6FFD924BB7F72745AF6FF" ID-DOI="http://doi.org/10.5281/zenodo.5536329" ID-Zenodo-Dep="5536329" httpUri="https://zenodo.org/record/5536329/files/figure.png" pageId="34" pageNumber="53" startId="34.[189,232,1892,1918]" targetBox="[189,1399,396,1868]" targetPageId="34">
<paragraph id="6C2D0A0BFFD6FFD924BB7F72745AF6FF" blockId="34.[187,1398,1892,2024]" pageId="34" pageNumber="53">
<emphasis id="5EE6D619FFD6FFD924BB7F72707FF669" bold="true" box="[189,366,1892,1918]" pageId="34" pageNumber="53">Fig. 12. A–D</emphasis>
.
<taxonomicName id="AB927188FFD6FFD925817F7273DEF669" authority="Kantor &amp; Puillandre, 2021" authorityName="Kantor &amp; Puillandre" authorityYear="2021" box="[391,719,1892,1918]" class="Gastropoda" family="Cochlespiridae" genus="Sibogasyrinx" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="53" phylum="Mollusca" rank="species" species="pagodiformis" status="sp. nov.">
<emphasis id="5EE6D619FFD6FFD925817F7273DEF669" box="[391,719,1892,1918]" italics="true" pageId="34" pageNumber="53">Sibogasyrinx pagodiformis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="45D56B62FFD6FFD926E77F737256F668" box="[737,839,1893,1919]" pageId="34" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="5EE6D619FFD6FFD9275F7F727281F669" bold="true" box="[857,912,1892,1918]" pageId="34" pageNumber="53">A–B</emphasis>
. Solomon Islands, holotype, MNHN- IM-2009-16825, SL 28.8 mm.
<emphasis id="5EE6D619FFD6FFD926187F917346F6B6" bold="true" box="[542,599,1927,1953]" pageId="34" pageNumber="53">C–D</emphasis>
. Vanuatu. MNHN-IM-2009-11327, SL 24.3 mm.
<emphasis id="5EE6D619FFD6FFD920907F9175DEF6B6" bold="true" box="[1174,1231,1927,1953]" pageId="34" pageNumber="53">E–H</emphasis>
.
<taxonomicName id="AB927188FFD6FFD920DB7F9173B3F6D3" authority="Kantor, Puillandre &amp; Bouchet" authorityName="Kantor, Puillandre &amp; Bouchet" authorityYear="2021" class="Gastropoda" family="Cochlespiridae" genus="Sibogasyrinx" kingdom="Animalia" order="Neogastropoda" pageId="34" pageNumber="53" phylum="Mollusca" rank="species" species="elbakyanae" status="sp. nov.">
<emphasis id="5EE6D619FFD6FFD920DB7F917052F6D3" italics="true" pageId="34" pageNumber="53">Sibogasyrinx elbakyanae</emphasis>
Kantor, Puillandre &amp; Bouchet
</taxonomicName>
<taxonomicNameLabel id="45D56B62FFD6FFD926AE7FBD7212F6D2" box="[680,771,1963,1989]" pageId="34" pageNumber="53" rank="species">sp. nov.</taxonomicNameLabel>
, holotype, Solomon Islands, MNHN-IM-2009-16834, SL 27.3 mm.
<emphasis id="5EE6D619FFD6FFD9255A7FD87062F6FF" bold="true" box="[348,371,1998,2024]" pageId="34" pageNumber="53">H</emphasis>
. Enlarged fragment of penultimate and antepenultimate whorls. A–G to same scale.
</paragraph>
</caption>
<subSubSection id="24885980FFD7FFD824BB79C17558F337" box="[189,1097,471,545]" pageId="35" pageNumber="54" type="distribution">
<paragraph id="6C2D0A0BFFD7FFD824BB79C17047F0E6" blockId="35.[189,342,471,497]" box="[189,342,471,497]" pageId="35" pageNumber="54">
<heading id="3765BD67FFD7FFD824BB79C17047F0E6" bold="true" box="[189,342,471,497]" fontSize="11" level="3" pageId="35" pageNumber="54" reason="3">
<emphasis id="5EE6D619FFD7FFD824BB79C17047F0E6" bold="true" box="[189,342,471,497]" pageId="35" pageNumber="54">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="6C2D0A0BFFD7FFD824BB7A107558F337" blockId="35.[189,1097,518,545]" box="[189,1097,518,545]" pageId="35" pageNumber="54">
This species was found in the
<collectingCountry id="14854A9BFFD7FFD826197A1073F2F337" box="[543,739,518,544]" name="Solomon Islands" pageId="35" pageNumber="54">Solomon Islands</collectingCountry>
and
<collectingCountry id="14854A9BFFD7FFD8271D7A10726AF337" box="[795,891,518,544]" name="Vanuatu" pageId="35" pageNumber="54">Vanuatu</collectingCountry>
at
<quantity id="AB6AA7EEFFD7FFD827997A107554F337" box="[927,1093,518,545]" metricMagnitude="3" metricUnit="m" metricValue="1.2559999999999998" metricValueMax="1.262" metricValueMin="1.25" pageId="35" pageNumber="54" unit="m" value="1256.0" valueMax="1262.0" valueMin="1250.0">1250–1262 m</quantity>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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