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<document id="415D815EBE63E3CF50DC3704E96A23BD" ID-CLB-Dataset="26946" ID-DOI="10.11646/zootaxa.4590.4.1" ID-GBIF-Dataset="64c1a738-381b-4ba2-bc35-317d756e6a29" ID-ISSN="1175-5326" ID-Zenodo-Dep="2656164" ID-ZooBank="CD2A364C-6CF9-4FF5-AD28-7AF388868B15" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1556698917896" checkinUser="plazi" docAuthor="Mohrig, Werner, Kauschke, Ellen &amp; Broadley, Adam" docDate="2019" docId="03C987F6FFD1FFA2BD885841FD77729F" docLanguage="en" docName="zootaxa.4590.4.1.pdf" docOrigin="Zootaxa 4590 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Bradysia bishopi Steffan 1973" docType="treatment" docVersion="7" lastPageNumber="406" masterDocId="FFF0FF8EFFD4FFA7BD1F590EFFB97777" masterDocTitle="Revision of black fungus gnat species (Diptera, Sciaridae) described from the Hawaiian Islands by D. E. Hardy and W. A. Steffan, and a contribution to the knowledge of the sciarid fauna of the Galápagos Islands" masterLastPageNumber="439" masterPageNumber="401" pageNumber="406" updateTime="1698724290689" updateUser="ExternalLinkService">
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<mods:title id="F80C57BCAA3A01EA355349718F460E7E">Revision of black fungus gnat species (Diptera, Sciaridae) described from the Hawaiian Islands by D. E. Hardy and W. A. Steffan, and a contribution to the knowledge of the sciarid fauna of the Galápagos Islands</mods:title>
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<mods:namePart id="4863391557D3251B8275DDCED339805D">Kauschke, Ellen</mods:namePart>
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<subSubSection id="C37A656BFFD1FFA2BD885841FDB3761D" box="[151,522,335,362]" pageId="5" pageNumber="406" type="nomenclature">
<paragraph id="8BDF36E0FFD1FFA2BD885841FDB3761D" blockId="5.[151,522,335,395]" box="[151,522,335,362]" pageId="5" pageNumber="406">
<heading id="D097818CFFD1FFA2BD885841FDB3761D" bold="true" box="[151,522,335,362]" fontSize="11" level="1" pageId="5" pageNumber="406" reason="1">
<taxonomicName id="4C604D63FFD1FFA2BD885841FDB3761D" ID-CoL="N5HZ" authority="Steffan, 1973" authorityName="Steffan" authorityYear="1973" box="[151,522,335,362]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bishopi">
<emphasis id="B914EAF2FFD1FFA2BD885841FDB3761D" bold="true" box="[151,522,335,362]" pageId="5" pageNumber="406">
<emphasis id="B914EAF2FFD1FFA2BD885841FEDB761E" bold="true" box="[151,354,335,361]" italics="true" pageId="5" pageNumber="406">Bradysia bishopi</emphasis>
Steffan, 1973
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C37A656BFFD1FFA2BD88587CFD327533" pageId="5" pageNumber="406" type="description">
<paragraph id="8BDF36E0FFD1FFA2BD88587CFE9176FC" blockId="5.[151,522,335,395]" box="[151,296,370,395]" pageId="5" pageNumber="406">
(
<figureCitation id="135B2A65FFD1FFA2BD80587CFF5B76FC" box="[159,226,370,395]" captionStart="FIGURE 3" captionStartId="6.[151,250,1959,1981]" captionTargetBox="[366,1240,520,1882]" captionTargetId="figure@6.[340,1247,500,1937]" captionTargetPageId="6" captionText="FIGURE 3 AD. Bradysia bishopi Steffan, 1973. Holotype. A. Hypopygium; B. Flagellomeres 45; C. Tibial organ of fore tibia; D. Wing." httpUri="https://zenodo.org/record/2656170/files/figure.png" pageId="5" pageNumber="406">Fig. 3</figureCitation>
AD)
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BD8858BAFCE376BD" blockId="5.[151,1437,436,580]" box="[151,858,436,458]" pageId="5" pageNumber="406">
<taxonomicName id="4C604D63FFD1FFA2BD8858BAFE6876BD" authority="Steffan, 1973" authorityName="Steffan" authorityYear="1973" box="[151,465,436,458]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bishopi">
<emphasis id="B914EAF2FFD1FFA2BD8858BAFEFD76BD" box="[151,324,436,458]" italics="true" pageId="5" pageNumber="406">Bradysia bishopi</emphasis>
Steffan, 1973
</taxonomicName>
[Steffan (1973a): 353
<emphasis id="B914EAF2FFD1FFA2BFAF58BAFD0476BD" bold="true" box="[688,701,436,458]" pageId="5" pageNumber="406"></emphasis>
355, fig. 1 a
<emphasis id="B914EAF2FFD1FFA2BE2958BAFCFB76BD" bold="true" box="[822,834,436,458]" pageId="5" pageNumber="406"></emphasis>
f].
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BD8858DCFD327533" blockId="5.[151,1437,436,580]" pageId="5" pageNumber="406">
Selected literature:
<bibRefCitation id="EFF14B11FFD1FFA2BC7E58DCFE49769F" author="Hardy, D. E." box="[353,496,466,488]" pageId="5" pageNumber="406" pagination="208 - 235" refId="ref15113" refString="Hardy, D. E. (1960) Family Sciaridae. In: Zimmerman, E. C. (Ed.), Insects of Hawaii. A Manual of the Insects of the Hawaiian Islands, including on Enumeration of the Species and Notes on their Origin, Distribution, Hosts, Parasites etc. 10. University Hawaii Press, Honolulu, pp. 208 - 235." type="book chapter" year="1960">Hardy (1960)</bibRefCitation>
: 227
<emphasis id="B914EAF2FFD1FFA2BF3D58DCFD97769F" bold="true" box="[546,558,466,488]" pageId="5" pageNumber="406"></emphasis>
228, fig. 75 c
<emphasis id="B914EAF2FFD1FFA2BFA358DCFD70769F" bold="true" box="[700,713,466,488]" pageId="5" pageNumber="406"></emphasis>
e [as
<taxonomicName id="4C604D63FFD1FFA2BE1C58DDFB3B769F" authority="Brunetti" authorityName="Brunetti. Hardy" authorityYear="1960" box="[771,1154,466,488]" class="Insecta" family="Sciaridae" genus="Sciara" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="radicum" subGenus="Lycoriella">
<emphasis id="B914EAF2FFD1FFA2BE1C58DDFCFC769F" box="[771,837,467,488]" italics="true" pageId="5" pageNumber="406">Sciara</emphasis>
(
<emphasis id="B914EAF2FFD1FFA2BE4858DCFC07769F" box="[855,958,466,488]" italics="true" pageId="5" pageNumber="406">Lycoriella</emphasis>
)
<emphasis id="B914EAF2FFD1FFA2BED058DCFB9A769F" box="[975,1059,466,488]" italics="true" pageId="5" pageNumber="406">radicum</emphasis>
Brunetti
</taxonomicName>
, misinterpretation];
<bibRefCitation id="EFF14B11FFD1FFA2B84B58DCFEB77570" author="Steffan, W. A." pageId="5" pageNumber="406" pagination="669 - 732" refId="ref17595" refString="Steffan, W. A. (1969) Insects of Micronesia. Diptera: Sciaridae. Insects of Micronesia, 12 (7), 669 - 732." type="journal article" year="1969">Steffan (1969)</bibRefCitation>
: 719
<emphasis id="B914EAF2FFD1FFA2BC5F58FFFEF47570" bold="true" box="[320,333,497,519]" pageId="5" pageNumber="406"></emphasis>
722, fig. 20 a
<emphasis id="B914EAF2FFD1FFA2BCC458FFFE5E7570" bold="true" box="[475,487,497,519]" pageId="5" pageNumber="406"></emphasis>
g [as
<taxonomicName id="4C604D63FFD1FFA2BF3C58FFFCF27570" authority="(Brunetti)" authorityName="Brunetti" baseAuthorityName="Brunetti" baseAuthorityYear="1912" box="[547,843,497,519]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="radicum">
<emphasis id="B914EAF2FFD1FFA2BF3C58FFFD627570" box="[547,731,497,519]" italics="true" pageId="5" pageNumber="406">Bradysia radicum</emphasis>
(Brunetti)
</taxonomicName>
, misinterpretation];
<bibRefCitation id="EFF14B11FFD1FFA2B90258FFFAA37570" author="Menzel, F. &amp; Smith, J. E." box="[1053,1306,497,519]" pageId="5" pageNumber="406" pagination="19 - 45" refId="ref15871" refString="Menzel, F. &amp; Smith, J. E. (2009) Family Sciaridae. In: Gerlach, J. (Ed.), The Diptera of the Seychelles islands. Pensoft Series Faunistica 85. Pensoft Publishers, Sofia &amp; Moscow, pp. 19 - 45." type="book chapter" year="2009">Menzel &amp; Smith (2009)</bibRefCitation>
: 24
<emphasis id="B914EAF2FFD1FFA2B85E58FFFAF77570" bold="true" box="[1345,1358,497,519]" pageId="5" pageNumber="406"></emphasis>
26, figs 4
<emphasis id="B914EAF2FFD1FFA2BDCB5B1EFF597551" bold="true" box="[212,224,528,550]" pageId="5" pageNumber="406"></emphasis>
6;
<bibRefCitation id="EFF14B11FFD1FFA2BDE05B1EFDB47551" author="Kohler, A. &amp; Menzel, F." box="[255,525,528,550]" pageId="5" pageNumber="406" pagination="63 - 72" refId="ref15455" refString="Kohler, A. &amp; Menzel, F. (2013) New records of Black Fungus Gnats (Diptera: Sciaridae) from New Caledonia, with the description of two new Bradysia species and an updated checklist. Zootaxa 3718 (1), 63 - 72. http: // dx. doi. org / 10.11646 / zootaxa. 3718.1.5" type="journal article" year="2013">Köhler &amp; Menzel (2013)</bibRefCitation>
: 68; Mohrig (2016): 12
<emphasis id="B914EAF2FFD1FFA2BE135B1EFCA07551" bold="true" box="[780,793,528,550]" pageId="5" pageNumber="406"></emphasis>
13, fig 12 a
<emphasis id="B914EAF2FFD1FFA2BE865B1EFC1F7551" bold="true" box="[921,934,528,550]" pageId="5" pageNumber="406"></emphasis>
c (as
<taxonomicName id="4C604D63FFD1FFA2BEFB5B1EFB277551" authorityName="Mohrig" authorityYear="2016" box="[996,1182,528,550]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="mutuata">
<emphasis id="B914EAF2FFD1FFA2BEFB5B1EFB277551" box="[996,1182,528,550]" italics="true" pageId="5" pageNumber="406">Bradysia mutuata</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFF14B11FFD1FFA2B9A75B1EFA207551" author="Mohrig, W. &amp; Kauschke, E. &amp; Broadley, A." box="[1208,1433,528,550]" pageId="5" pageNumber="406" pagination="203 - 241" refId="ref16648" refString="Mohrig, W., Kauschke, E. &amp; Broadley, A. (2018) New black fungus gnats (Diptera: Sciaridae) from Eastern Australia. Zootaxa, 4450 (2), 203 - 241. https: // doi. org / 10.11646 / zootaxa. 4450.2.3" type="journal article" year="2018">
Mohrig
<emphasis id="B914EAF2FFD1FFA2B8105B1FFAFE7551" box="[1295,1351,528,550]" italics="true" pageId="5" pageNumber="406">et al.</emphasis>
(2018)
</bibRefCitation>
: 212
<emphasis id="B914EAF2FFD1FFA2BDF25B20FF407533" bold="true" box="[237,249,558,580]" pageId="5" pageNumber="406"></emphasis>
213, fig 7 A
<emphasis id="B914EAF2FFD1FFA2BC6B5B20FE397533" bold="true" box="[372,384,558,580]" pageId="5" pageNumber="406"></emphasis>
C (as
<taxonomicName id="4C604D63FFD1FFA2BCA35B20FDC47533" authorityName="Vilkamaa, Hippa &amp; Mohrig" authorityYear="2012" box="[444,637,558,580]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="centidens">
<emphasis id="B914EAF2FFD1FFA2BCA35B20FDC47533" box="[444,637,558,580]" italics="true" pageId="5" pageNumber="406">Bradysia centidens</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C37A656BFFD1FFA2BD885B64FC96747B" pageId="5" pageNumber="406" type="materials_examined">
<paragraph id="8BDF36E0FFD1FFA2BD885B64FA9575B9" blockId="5.[151,1436,618,718]" pageId="5" pageNumber="406">
<emphasis id="B914EAF2FFD1FFA2BD885B64FE5975F4" bold="true" box="[151,480,618,643]" pageId="5" pageNumber="406">
Material studied:
<materialsCitation id="3B083CBDFFD1FFA2BC6C5B64FE5975F4" ID-GBIF-Occurrence="2237390858" box="[371,480,618,643]" collectingDate="1968-11-11" collectionCode="BPBM, PWMP" collectorName="W. A. Steffan &amp; Conservation" county="Male" location="Embedded in Euparal. Body" municipality="Kailua" pageId="5" pageNumber="406" specimenCount="3" specimenCount-male="3" stateProvince="Hawaii" typeStatus="holotype">
<typeStatus id="54DB8842FFD1FFA2BC6C5B64FE5975F4" box="[371,480,618,643]" pageId="5" pageNumber="406" type="holotype">Holotype</typeStatus>
</materialsCitation>
</emphasis>
:
<collectingCounty id="62BE4E6CFFD1FFA2BCEE5B64FD9475F4" box="[497,557,618,643]" pageId="5" pageNumber="406">Male</collectingCounty>
,
<collectingRegion id="49A4F802FFD1FFA2BF245B64FD2175F4" box="[571,664,618,643]" country="United States of America" name="Hawaii" pageId="5" pageNumber="406">Hawaii</collectingRegion>
, Oahu I., HW 162, F
<subScript id="17E434A5FFD1FFA2BE865B77FC1875F0" attach="left" box="[921,929,633,647]" fontSize="6" pageId="5" pageNumber="406">9</subScript>
, leg. unknown (BPBM No. 9943).
<specimenCount id="9D66FD69FFD1FFA2B8235B65FA2175F4" box="[1340,1432,618,643]" pageId="5" pageNumber="406" type="male">3 males</specimenCount>
,
<collectingRegion id="49A4F802FFD1FFA2BD885B9FFF4D75DD" box="[151,244,657,682]" country="United States of America" name="Hawaii" pageId="5" pageNumber="406">Hawaii</collectingRegion>
, Oahu,
<collectingMunicipality id="6BBBAC9AFFD1FFA2BC4C5B9FFE1975DD" box="[339,416,657,682]" pageId="5" pageNumber="406">Kailua</collectingMunicipality>
,
<date id="FFDE1020FFD1FFA2BCB35B9FFD9D75DD" box="[428,548,657,682]" pageId="5" pageNumber="406" value="1968-11-11">
<collectingDate id="EF9AE9C8FFD1FFA2BCB35B9FFD9D75DD" box="[428,548,657,682]" pageId="5" pageNumber="406" value="1968-11-11">11.xi.1968</collectingDate>
</date>
, blacklight, leg.
<collectorName id="26955336FFD1FFA2BFC05B9CFCC975DD" box="[735,880,657,682]" pageId="5" pageNumber="406">W.A. Steffan</collectorName>
(
<quantity id="4C989B05FFD1FFA2BE9D5B9CFC1275DD" box="[898,939,657,682]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="5" pageNumber="406" unit="in" value="1.0">1 in</quantity>
PWMP).
<emphasis id="B914EAF2FFD1FFA2BDD85BBBFE0475B9" bold="true" box="[199,445,693,718]" pageId="5" pageNumber="406">
<collectorName id="26955336FFD1FFA2BDD85BBBFEDE75B9" box="[199,359,693,718]" pageId="5" pageNumber="406">Conservation</collectorName>
status:
</emphasis>
<location id="8EBF603BFFD1FFA2BCDB5BBBFCBB75B9" LSID="urn:lsid:plazi:treatment:03C987F6FFD1FFA2BD885841FD77729F:8EBF603BFFD1FFA2BCDB5BBBFCBB75B9" box="[452,770,693,718]" county="Male" municipality="Kailua" name="Embedded in Euparal. Body" pageId="5" pageNumber="406" stateProvince="Hawaii">Embedded in Euparal. Body</location>
and head separated, all details in good condition.
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BD885BF8FC96747B" blockId="5.[151,1123,758,811]" box="[151,815,758,780]" pageId="5" pageNumber="406">
<emphasis id="B914EAF2FFD1FFA2BD885BF8FC96747B" bold="true" box="[151,815,758,780]" pageId="5" pageNumber="406">
=
<taxonomicName id="4C604D63FFD1FFA2BDBA5BF8FD67747B" authority="Vilkamaa, Hippa &amp; Mohrig, 2012" authorityName="Vilkamaa, Hippa &amp; Mohrig" authorityYear="2012" box="[165,734,758,780]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="centidens">
<emphasis id="B914EAF2FFD1FFA2BDBA5BF8FED2747B" bold="true" box="[165,363,758,780]" italics="true" pageId="5" pageNumber="406">Bradysia centidens</emphasis>
Vilkamaa, Hippa &amp; Mohrig, 2012
</taxonomicName>
,
<taxonomicNameLabel id="A2275789FFD1FFA2BFF65BF8FC96747B" box="[745,815,758,780]" pageId="5" pageNumber="406">syn. n.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C37A656BFFD1FFA2BD885A1BFD0B74F0" pageId="5" pageNumber="406" type="description">
<paragraph id="8BDF36E0FFD1FFA2BD885A1BFBDA745C" blockId="5.[151,1123,758,811]" box="[151,1123,789,811]" pageId="5" pageNumber="406">
<taxonomicName id="4C604D63FFD1FFA2BD885A1BFD0F745C" authority="Vilkamaa, Hippa &amp; Mohrig, 2012" authorityName="Vilkamaa, Hippa &amp; Mohrig" authorityYear="2012" box="[151,694,789,811]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="centidens">
<emphasis id="B914EAF2FFD1FFA2BD885A1BFEE0745C" box="[151,345,789,811]" italics="true" pageId="5" pageNumber="406">Bradysia centidens</emphasis>
Vilkamaa, Hippa &amp; Mohrig, 2012
</taxonomicName>
[Vilkamaa
<emphasis id="B914EAF2FFD1FFA2BE345A18FCE1745C" box="[811,856,789,811]" italics="true" pageId="5" pageNumber="406">et al</emphasis>
. (2012c): 28, fig. 2 A
<emphasis id="B914EAF2FFD1FFA2B92B5A1BFBF9745C" bold="true" box="[1076,1088,789,811]" pageId="5" pageNumber="406"></emphasis>
D].
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BD885A5AFB87741E" blockId="5.[151,1086,850,903]" box="[151,1086,850,873]" pageId="5" pageNumber="406">
=
<taxonomicName id="4C604D63FFD1FFA2BDB45A5CFE44741E" authority="Mohrig, 2016" authorityName="Mohrig" authorityYear="2016" box="[171,509,850,873]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="mutuata">
<emphasis id="B914EAF2FFD1FFA2BDB45A5CFE44741E" bold="true" box="[171,509,850,873]" pageId="5" pageNumber="406">
<emphasis id="B914EAF2FFD1FFA2BDB45A5CFEDC741F" bold="true" box="[171,357,850,872]" italics="true" pageId="5" pageNumber="406">Bradysia mutuata</emphasis>
Mohrig, 2016
</emphasis>
</taxonomicName>
(syn. to
<taxonomicName id="4C604D63FFD1FFA2BF4A5A5AFB8B741E" authority="Vilkamaa, Hippa &amp; Mohrig, 2012" authorityName="Vilkamaa, Hippa &amp; Mohrig" authorityYear="2012" box="[597,1074,851,873]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="centidens">
<emphasis id="B914EAF2FFD1FFA2BF4A5A5AFD68741E" box="[597,721,851,873]" italics="true" pageId="5" pageNumber="406">B. centidens</emphasis>
Vilkamaa, Hippa &amp; Mohrig, 2012
</taxonomicName>
).
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BD885A7FFD0B74F0" blockId="5.[151,1086,850,903]" box="[151,690,881,903]" pageId="5" pageNumber="406">
Literature:
<bibRefCitation id="EFF14B11FFD1FFA2BC175A7FFE6274F0" author="Mohrig, W. &amp; Kauschke, E. &amp; Broadley, A." box="[264,475,881,903]" pageId="5" pageNumber="406" pagination="203 - 241" refId="ref16648" refString="Mohrig, W., Kauschke, E. &amp; Broadley, A. (2018) New black fungus gnats (Diptera: Sciaridae) from Eastern Australia. Zootaxa, 4450 (2), 203 - 241. https: // doi. org / 10.11646 / zootaxa. 4450.2.3" type="journal article" year="2018">
Mohrig
<emphasis id="B914EAF2FFD1FFA2BC455A7CFE3E74F0" box="[346,391,881,903]" italics="true" pageId="5" pageNumber="406">et al</emphasis>
. (2018)
</bibRefCitation>
: 212-213, fig.7 AC.
</paragraph>
</subSubSection>
<subSubSection id="C37A656BFFD1FFA2BDD85AA3FD5972D7" pageId="5" pageNumber="406" type="discussion">
<paragraph id="8BDF36E0FFD1FFA2BDD85AA3FAE17342" blockId="5.[151,1437,941,1512]" pageId="5" pageNumber="406">
<emphasis id="B914EAF2FFD1FFA2BDD85AA3FEFE74B1" bold="true" box="[199,327,941,966]" pageId="5" pageNumber="406">Comments</emphasis>
. Two specimens from the Hawaiian Islands, Oahu, Kailua,
<date id="FFDE1020FFD1FFA2BEE15AA3FBD074B1" box="[1022,1129,941,966]" pageId="5" pageNumber="406" value="1968-11-06">6.xi.1968</date>
, 157, blacklight, leg. W.A. Steffan and one specimen from Oahu, Kailua,
<date id="FFDE1020FFD1FFA2BFAC5ADFFCA7749D" box="[691,798,977,1002]" pageId="5" pageNumber="406" value="1968-11-02">2.xi.1968</date>
, HW 162, F
<subScript id="17E434A5FFD1FFA2BEB35AEEFC0A7499" attach="left" box="[940,947,992,1006]" fontSize="6" pageId="5" pageNumber="406">9</subScript>
, leg. W.A. Steffan, were misinterpreted by Steffan as
<taxonomicName id="4C604D63FFD1FFA2BC155AF7FE3C7367" baseAuthorityName="Brunetti" baseAuthorityYear="1912" box="[266,389,1016,1040]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="radicum">
<emphasis id="B914EAF2FFD1FFA2BC155AF7FE3C7367" box="[266,389,1016,1040]" italics="true" pageId="5" pageNumber="406">B. radicum</emphasis>
</taxonomicName>
. The illustration of the hypopygium of
<taxonomicName id="4C604D63FFD1FFA2BE275AF7FC0A7367" baseAuthorityName="Brunetti" baseAuthorityYear="1912" box="[824,947,1016,1040]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="radicum">
<emphasis id="B914EAF2FFD1FFA2BE275AF7FC0A7367" box="[824,947,1016,1040]" italics="true" pageId="5" pageNumber="406">B. radicum</emphasis>
</taxonomicName>
(
<figureCitation id="135B2A65FFD1FFA2BEDE5AF6FB977367" box="[961,1070,1016,1041]" captionStart="FIGURE 20" captionStartId="30.[151,250,1572,1594]" captionTargetBox="[168,1397,324,1499]" captionTargetId="figure@30.[151,1435,193,1551]" captionTargetPageId="30" captionText="FIGURE 20 AD. Scatopsciara hoyti (Hardy, 1956). Holotype. A. Wing; B. Basal segments of antenna; C. Palpus; D. Apex of fore tibia." httpUri="https://zenodo.org/record/2656202/files/figure.png" pageId="5" pageNumber="406">Figure 20</figureCitation>
) in
<bibRefCitation id="EFF14B11FFD1FFA2B9465AF6FB417366" author="Steffan, W. A." box="[1113,1272,1016,1041]" pageId="5" pageNumber="406" pagination="669 - 732" refId="ref17595" refString="Steffan, W. A. (1969) Insects of Micronesia. Diptera: Sciaridae. Insects of Micronesia, 12 (7), 669 - 732." type="journal article" year="1969">Steffan (1969)</bibRefCitation>
is very similar to the illustration of
<taxonomicName id="4C604D63FFD1FFA2BC675D13FD877342" authority="Steffan" authorityName="Steffan" authorityYear="1973" box="[376,574,1052,1077]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bishopi">
<emphasis id="B914EAF2FFD1FFA2BC675D13FE517343" box="[376,488,1052,1076]" italics="true" pageId="5" pageNumber="406">B. bishopi</emphasis>
Steffan
</taxonomicName>
(
<figureCitation id="135B2A65FFD1FFA2BF535D12FD007342" box="[588,697,1052,1077]" captionStart="FIGURE 1" captionStartId="2.[151,250,1992,2014]" captionTargetBox="[226,1359,193,1979]" captionTargetId="figure@2.[226,1359,193,1979]" captionTargetPageId="2" captionText="FIGURE 1. Elmo Hardy in 1978. Photo courtesy of Neal Evenhuis." httpUri="https://zenodo.org/record/2656166/files/figure.png" pageId="5" pageNumber="406">Figure 1d</figureCitation>
) in Steffan (1973a) and
<figureCitation id="135B2A65FFD1FFA2BED85D12FB9D7342" box="[967,1060,1052,1077]" captionStart="FIGURE 4" captionStartId="7.[151,250,1562,1584]" captionTargetBox="[362,1247,193,1504]" captionTargetId="figure@7.[340,1247,193,1541]" captionTargetPageId="7" captionText="FIGURE 4 AC. Bradysia crassicornis (Skuse, 1890). Specimen from Barbers Point, Oahu. A. Hypopygium; B. Head with basal segments of flagellomeres; C. Tibial organ of fore tibia." httpUri="https://zenodo.org/record/2656174/files/figure.png" pageId="5" pageNumber="406">
Figure
<quantity id="4C989B05FFD1FFA2B9095D13FB9D7342" box="[1046,1060,1053,1077]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="5" pageNumber="406" unit="in" value="4.0">4</quantity>
</figureCitation>
in
<bibRefCitation id="EFF14B11FFD1FFA2B9585D12FAED7342" author="Menzel, F. &amp; Smith, J. E." box="[1095,1364,1052,1077]" pageId="5" pageNumber="406" pagination="19 - 45" refId="ref15871" refString="Menzel, F. &amp; Smith, J. E. (2009) Family Sciaridae. In: Gerlach, J. (Ed.), The Diptera of the Seychelles islands. Pensoft Series Faunistica 85. Pensoft Publishers, Sofia &amp; Moscow, pp. 19 - 45." type="book chapter" year="2009">Menzel &amp; Smith (2009)</bibRefCitation>
.
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BDD85D4EFBB773D6" blockId="5.[151,1437,941,1512]" pageId="5" pageNumber="406">
The species is characterized by 2 dorsally inserted spines distinctly separated from the 45 apical spines of the gonostylus. A comparison with the
<typeStatus id="54DB8842FFD1FFA2BF2F5D6BFD29730A" box="[560,656,1125,1149]" pageId="5" pageNumber="406" type="paratype">paratype</typeStatus>
of
<taxonomicName id="4C604D63FFD1FFA2BFA65D6AFAA6730A" authority="Vilkamaa, Hippa &amp; Mohrig, 2012" authorityName="Vilkamaa, Hippa &amp; Mohrig" authorityYear="2012" box="[697,1311,1124,1149]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="centidens">
<emphasis id="B914EAF2FFD1FFA2BFA65D6AFC28730B" box="[697,913,1124,1148]" italics="true" pageId="5" pageNumber="406">Bradysia centidens</emphasis>
Vilkamaa, Hippa &amp; Mohrig, 2012
</taxonomicName>
from
<collectingCountry id="F3777670FFD1FFA2B8785D6BFEB073D6" name="New Caledonia" pageId="5" pageNumber="406">New Caledonia</collectingCountry>
showed that
<emphasis id="B914EAF2FFD1FFA2BC825D87FD1F73D7" box="[413,678,1160,1184]" italics="true" pageId="5" pageNumber="406">
<taxonomicName id="4C604D63FFD1FFA2BC825D87FDA873D7" authorityName="Steffan" authorityYear="1973" box="[413,529,1160,1184]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bishopi">B. bishopi</taxonomicName>
,
<taxonomicName id="4C604D63FFD1FFA2BF035D87FD1F73D7" authorityName="Vilkamaa, Hippa &amp; Mohrig" authorityYear="2012" box="[540,678,1160,1184]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="centidens">B. centidens</taxonomicName>
</emphasis>
and
<taxonomicName id="4C604D63FFD1FFA2BFC35D87FCEE73D7" authorityName="Mohrig" authorityYear="2016" box="[732,855,1161,1184]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="mutuata">
<emphasis id="B914EAF2FFD1FFA2BFC35D87FCEE73D7" box="[732,855,1161,1184]" italics="true" pageId="5" pageNumber="406">B. mutuata</emphasis>
</taxonomicName>
are conspecific.
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BDD85DA2FBC6727A" blockId="5.[151,1437,941,1512]" pageId="5" pageNumber="406">
The slide labelled
<typeStatus id="54DB8842FFD1FFA2BC845DA2FE4473B2" box="[411,509,1196,1221]" pageId="5" pageNumber="406" type="holotype">holotype</typeStatus>
bears the number BPBM 9943 which differs from the number listed in Steffans original description and in his types catalog (BPBM 9942) (Steffan 1973
<bibRefCitation id="EFF14B11FFD1FFA2BED55DDEFBCF739E" author="Steffan, W. A." box="[970,1142,1232,1257]" pageId="5" pageNumber="406" pagination="47 - 50" refId="ref17917" refString="Steffan, W. A. (1976) Catalog of entomological types in the Bernice P. Bishop Museum. Diptera: Sciaridae. Pacific Insects, 17 (1), 47 - 50." type="journal article" year="1976">a, Steffan 1976</bibRefCitation>
). Steffan (1973a) reported that the types were generation F
<subScript id="17E434A5FFD1FFA2BCE25C0DFDBC7266" attach="left" box="[509,517,1283,1297]" fontSize="6" pageId="5" pageNumber="406">9</subScript>
, reared from a single female captured on the
<date id="FFDE1020FFD1FFA2B91E5DFAFBC5727A" box="[1025,1148,1268,1293]" pageId="5" pageNumber="406" value="1968-11-11">11.xi.1968</date>
.
</paragraph>
<paragraph id="8BDF36E0FFD1FFA2BDD85C15FD5972D7" blockId="5.[151,1437,941,1512]" pageId="5" pageNumber="406">
The distribution of
<taxonomicName id="4C604D63FFD1FFA2BC805C12FDB67244" authorityName="Steffan" authorityYear="1973" box="[415,527,1307,1331]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bishopi">
<emphasis id="B914EAF2FFD1FFA2BC805C12FDB67244" box="[415,527,1307,1331]" italics="true" pageId="5" pageNumber="406">B. bishopi</emphasis>
</taxonomicName>
was reported by
<bibRefCitation id="EFF14B11FFD1FFA2BFD05C15FC627244" author="Menzel, F. &amp; Smith, J. E." box="[719,987,1307,1332]" pageId="5" pageNumber="406" pagination="19 - 45" refId="ref15871" refString="Menzel, F. &amp; Smith, J. E. (2009) Family Sciaridae. In: Gerlach, J. (Ed.), The Diptera of the Seychelles islands. Pensoft Series Faunistica 85. Pensoft Publishers, Sofia &amp; Moscow, pp. 19 - 45." type="book chapter" year="2009">Menzel &amp; Smith (2009)</bibRefCitation>
and
<bibRefCitation id="EFF14B11FFD1FFA2B90F5C15FA9E7244" author="Kohler, A. &amp; Menzel, F." box="[1040,1319,1307,1332]" pageId="5" pageNumber="406" pagination="63 - 72" refId="ref15455" refString="Kohler, A. &amp; Menzel, F. (2013) New records of Black Fungus Gnats (Diptera: Sciaridae) from New Caledonia, with the description of two new Bradysia species and an updated checklist. Zootaxa 3718 (1), 63 - 72. http: // dx. doi. org / 10.11646 / zootaxa. 3718.1.5" type="journal article" year="2013">Köhler &amp; Menzel (2013)</bibRefCitation>
to include
<collectingCountry id="F3777670FFD1FFA2BD885C31FEE0722F" box="[151,345,1343,1368]" name="French Polynesia" pageId="5" pageNumber="406">French Polynesia</collectingCountry>
(Tahiti,
<collectingRegion id="49A4F802FFD1FFA2BCA55C4EFD37722F" box="[442,654,1343,1368]" country="French Polynesia" name="Iles Marquises" pageId="5" pageNumber="406">Marquesas Islands</collectingRegion>
),
<collectingCountry id="F3777670FFD1FFA2BFBD5C31FCDC7220" box="[674,869,1343,1368]" name="American Samoa" pageId="5" pageNumber="406">American Samoa</collectingCountry>
(Tutuila),
<collectingCountry id="F3777670FFD1FFA2BEC15C31FB937220" box="[990,1066,1343,1367]" name="Samoa" pageId="5" pageNumber="406">Samoa</collectingCountry>
(Upolu, Savaii), and
<collectingCountry id="F3777670FFD1FFA2B8015C31FAFC722F" box="[1310,1349,1343,1368]" name="Fiji" pageId="5" pageNumber="406">Fiji</collectingCountry>
(Vanua Levu); however these were based on erroneous records by Edwards (see comments for
<taxonomicName id="4C604D63FFD1FFA2B96E5C6AFB57720C" baseAuthorityName="Brunetti" baseAuthorityYear="1912" box="[1137,1262,1379,1403]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="radicum">
<emphasis id="B914EAF2FFD1FFA2B96E5C6AFB57720C" box="[1137,1262,1379,1403]" italics="true" pageId="5" pageNumber="406">B. radicum</emphasis>
</taxonomicName>
). The presence of
<taxonomicName id="4C604D63FFD1FFA2BDAA5C86FE9F72E8" authorityName="Steffan" authorityYear="1973" box="[181,294,1415,1439]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="5" pageNumber="406" phylum="Arthropoda" rank="species" species="bishopi">
<emphasis id="B914EAF2FFD1FFA2BDAA5C86FE9F72E8" box="[181,294,1415,1439]" italics="true" pageId="5" pageNumber="406">B. bishopi</emphasis>
</taxonomicName>
on these islands requires confirmation.
</paragraph>
</subSubSection>
<subSubSection id="C37A656BFFD1FFA2BDD85CA5FD77729F" pageId="5" pageNumber="406" type="distribution">
<paragraph id="8BDF36E0FFD1FFA2BDD85CA5FD77729F" blockId="5.[151,1437,941,1512]" pageId="5" pageNumber="406">
<emphasis id="B914EAF2FFD1FFA2BDD85CA5FEE072B3" bold="true" box="[199,345,1451,1476]" pageId="5" pageNumber="406">Distribution</emphasis>
. Widespread in the Southern Hemisphere. Hawaii;
<collectingCountry id="F3777670FFD1FFA2BEF65CA5FBEA72B3" box="[1001,1107,1451,1476]" name="Australia" pageId="5" pageNumber="406">Australia</collectingCountry>
;
<collectingCountry id="F3777670FFD1FFA2B9775CA2FA9A72B3" box="[1128,1315,1451,1476]" name="New Caledonia" pageId="5" pageNumber="406">New Caledonia</collectingCountry>
;
<collectingCountry id="F3777670FFD1FFA2B8265CA5FA2472B3" box="[1337,1437,1451,1476]" name="Thailand" pageId="5" pageNumber="406">Thailand</collectingCountry>
(
<collectingRegion id="49A4F802FFD1FFA2BD805CC1FEB2729F" box="[159,267,1487,1512]" country="Thailand" name="Krung Thep Mahanakhon" pageId="5" pageNumber="406">Bangkok</collectingRegion>
);
<collectingCountry id="F3777670FFD1FFA2BC025CC1FE2C729F" box="[285,405,1487,1512]" name="Seychelles" pageId="5" pageNumber="406">Seychelles</collectingCountry>
Islands; Galápagos Islands.
</paragraph>
</subSubSection>
</treatment>
</document>