403 lines
52 KiB
XML
403 lines
52 KiB
XML
<document id="6EACEAE3B53C40C2CF1A81CBB39AD7A1" ID-DOI="10.11646/zootaxa.3716.2.9" ID-GBIF-Dataset="cd3d13de-0da7-4dd6-acd0-38e59e99a9c7" ID-ISSN="1175-5326" ID-Zenodo-Dep="223737" ID-ZooBank="70D1FF8D-202C-4931-8537-AC2D5C266822" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460681376903" checkinUser="plazi" docAuthor="Ho, Hsuan-Ching" docDate="2013" docId="03C987BE0221FF9BFF3FFE458975FE64" docLanguage="en" docName="zt03716p300.pdf" docOrigin="Zootaxa 3716 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Malthopsis bradburyae Ho, 2013, n. sp." docType="treatment" docVersion="8" lastPageNumber="294" masterDocId="FFF0FFC60220FF9EFFA8FF888B25FFC8" masterDocTitle="Two new species of the batfish genus Malthopsis (Lophiiformes: Ogcocephalidae) from the Western Indian Ocean" masterLastPageNumber="300" masterPageNumber="289" pageNumber="290" updateTime="1698350518055" updateUser="plazi">
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<mods:titleInfo id="AD0CFB93FCD2B6822B679C0194D5CB00">
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<mods:title id="BF0F5A01B420A40751F3405331D3D471">Two new species of the batfish genus Malthopsis (Lophiiformes: Ogcocephalidae) from the Western Indian Ocean</mods:title>
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<mods:name id="F6495B879B8C6B7B50D1D50C344CE40A" type="personal">
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<mods:roleTerm id="6995005A4BA1319E097971BDE57060B0">Author</mods:roleTerm>
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<mods:namePart id="53FC6FEA3D090E5B6456FAB0501545C6">Ho, Hsuan-Ching</mods:namePart>
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<mods:typeOfResource id="530E843155D20226746CCEF85520D004">text</mods:typeOfResource>
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<mods:title id="B6633AD36973028243017ABE1036194D">Zootaxa</mods:title>
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<mods:part id="72F82B529619FE5D70C4C0B42D46C147">
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<mods:date id="ABE278659610E4BF565148EA1484031E">2013</mods:date>
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<mods:number id="9D645AFF722A335586E99B71780326AE">3716</mods:number>
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<mods:detail id="63FD0F824B0D2BC8011F4421C44E7216" type="issue">
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<mods:number id="46B6A814C94EF5E2E122CBD3E997CBB8">2</mods:number>
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<mods:classification id="B7C840EADCA61DB473BB6B370C3C0240">journal article</mods:classification>
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<mods:identifier id="A0977939E51E51D6FD1A51D72406EFAB" type="DOI">10.11646/zootaxa.3716.2.9</mods:identifier>
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<mods:identifier id="3FBAB03DFFCEDE25BF7775F5980E9FD2" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BA04FF9036EB70A69293B495EC09EDA5" type="Zenodo-Dep">223737</mods:identifier>
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<treatment id="03C987BE0221FF9BFF3FFE458975FE64" ID-DOI="http://doi.org/10.5281/zenodo.5696364" ID-GBIF-Taxon="119580066" ID-Zenodo-Dep="5696364" LSID="urn:lsid:plazi:treatment:03C987BE0221FF9BFF3FFE458975FE64" httpUri="http://treatment.plazi.org/id/03C987BE0221FF9BFF3FFE458975FE64" lastPageId="5" lastPageNumber="294" pageId="1" pageNumber="290">
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<subSubSection id="C37A65230221FF9FFF3FFE458AECFDE4" pageId="1" pageNumber="290" type="nomenclature">
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<paragraph id="8BDF36A80221FF9FFF3FFE458AD2FE2F" blockId="1.[151,661,461,557]" box="[151,503,461,487]" pageId="1" pageNumber="290">
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<heading id="D09781C40221FF9FFF3FFE458AD2FE2F" bold="true" box="[151,503,461,487]" fontSize="11" level="1" pageId="1" pageNumber="290" reason="1">
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<emphasis id="B914EABA0221FF9FFF3FFE458AD2FE2F" bold="true" box="[151,503,461,487]" pageId="1" pageNumber="290">
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<taxonomicName id="4C604D2B0221FF9FFF3FFE458A8BFE2F" ID-CoL="3XQSY" box="[151,430,461,487]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="species" species="bradburyae" status="sp. nov.">
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<emphasis id="B914EABA0221FF9FFF3FFE458A8BFE2F" bold="true" box="[151,430,461,487]" italics="true" pageId="1" pageNumber="290">Malthopsis bradburyae</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22757C10221FF9FFE1DFE458AD2FE2F" box="[437,503,461,487]" pageId="1" pageNumber="290" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDF36A80221FF9FFF3FFE788AECFDE4" blockId="1.[151,661,461,557]" pageId="1" pageNumber="290">
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Common name: Bradbury’s triangular batfish
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<figureCitation id="135B2A2D0221FF9FFF3FFD9C8BFBFDE4" box="[151,222,532,557]" captionStart="FIGURE 1" captionStartId="3.[151,250,1958,1980]" captionTargetBox="[352,1229,212,1915]" captionTargetId="figure@3.[338,1248,195,1934]" captionTargetPageId="3" captionText="FIGURE 1. Malthopsis bradburyae n. sp. A. Holotype, CAS 227226, 72.4 mm SL, dorsal view, preserved. B. holotype, ventral view, preserved. C. Paratype, CAS 39631, 1 of 7, 70.9 mm SL, dorsal view, preserved." httpUri="https://zenodo.org/record/223738/files/figure.png" pageId="1" pageNumber="290">Figs. 1</figureCitation>
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A–C, 2A–E;
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<tableCitation id="C6E203130221FF9FFEDEFD9C8AECFDE4" box="[374,457,532,557]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1401,238,1262]" captionTargetPageId="2" captionText="TABLE 1. Morphometric and meristic data of two new Malthopsis species described in present study. Pectoral-fin rays are counted on both sides." httpUri="http://table.plazi.org/id/DF1F66200222FF9CFF3FFF108A88FF1A" pageId="1" pageNumber="290" tableUuid="DF1F66200222FF9CFF3FFF108A88FF1A">Table 1</tableCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A65230221FF9FFF3FFDD48A25FD28" pageId="1" pageNumber="290" type="materials_examined">
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<paragraph id="8BDF36A80221FF9FFF3FFDD4880CFD50" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
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<emphasis id="B914EABA0221FF9FFF3FFDD48A2EFDBD" bold="true" box="[151,267,604,629]" pageId="1" pageNumber="290">
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<typeStatus id="54DB880A0221FF9FFF3FFDD48A23FDBD" box="[151,262,604,629]" pageId="1" pageNumber="290" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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CAS 227226 (
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<quantity id="4C989B4D0221FF9FFE19FDD48933FDBC" box="[433,534,604,628]" metricMagnitude="-2" metricUnit="m" metricValue="7.24" pageId="1" pageNumber="290" unit="mm" value="72.4">72.4 mm</quantity>
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SL), R/V
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<emphasis id="B914EABA0221FF9FFD22FDD4883CFDBC" box="[650,793,604,628]" italics="true" pageId="1" pageNumber="290">Anton Bruun</emphasis>
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, cruise 9, station 22,
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<geoCoordinate id="EE54506F0221FF9FFBAFFDD48F7DFDBD" box="[1031,1112,604,629]" direction="south" orientation="latitude" pageId="1" pageNumber="290" precision="925" value="-6.85">6°51’S</geoCoordinate>
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,
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<geoCoordinate id="EE54506F0221FF9FFBCBFDD48FE1FDBD" box="[1123,1220,604,629]" direction="east" orientation="longitude" pageId="1" pageNumber="290" precision="925" value="39.9">39°54’E</geoCoordinate>
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,
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<collectingCountry id="F37776380221FF9FFB66FDD48E13FDBD" box="[1230,1334,604,629]" name="Tanzania" pageId="1" pageNumber="290">Tanzania</collectingCountry>
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, Western Indian Ocean,
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<quantity id="4C989B4D0221FF9FFE93FD088AA1FD5F" box="[315,388,640,664]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="290" unit="m" value="100.0">100 m</quantity>
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, coll. A. H. Fehlmann,
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<date id="FFDE10680221FF9FFD22FD088800FD50" box="[650,805,639,664]" pageId="1" pageNumber="290" value="1964-11-19">19 Nov. 1964</date>
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.
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</paragraph>
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<paragraph id="8BDF36A80221FF9FFF6FFD2C8A25FD28" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
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<emphasis id="B914EABA0221FF9FFF6FFD2C8A63FD75" bold="true" box="[199,326,676,701]" pageId="1" pageNumber="290">
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<typeStatus id="54DB880A0221FF9FFF6FFD2C8A64FD75" box="[199,321,676,701]" pageId="1" pageNumber="290" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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CAS 39631 (
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<specimenCount id="9D66FD210221FF9FFE40FD2C8952FD75" box="[488,631,676,701]" pageId="1" pageNumber="290" type="generic">7 specimens</specimenCount>
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,
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<quantity id="4C989B4D0221FF9FFD2EFD2C880EFD74" box="[646,811,676,701]" metricMagnitude="-2" metricUnit="m" metricValue="5.325" metricValueMax="7.09" metricValueMin="3.56" pageId="1" pageNumber="290" unit="mm" value="53.25" valueMax="70.9" valueMin="35.6">35.6–70.9 mm</quantity>
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SL) and NMMB-P17824 (1, 59.0), collected with the
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<typeStatus id="54DB880A0221FF9FFF3FFD4F8BD8FD28" box="[151,253,711,736]" pageId="1" pageNumber="290" type="holotype">holotype</typeStatus>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A65230221FF9FFF6FFD648AA8FC15" pageId="1" pageNumber="290" type="diagnosis">
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<paragraph id="8BDF36A80221FF9FFF6FFD648AA8FC15" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
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<emphasis id="B914EABA0221FF9FFF6FFD648A64FCCD" bold="true" box="[199,321,748,773]" pageId="1" pageNumber="290">Diagnosis.</emphasis>
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A species of
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<taxonomicName id="4C604D2B0221FF9FFE7EFD648976FCCD" box="[470,595,748,773]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="1" pageNumber="290" phylum="Chordata" rank="genus">
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<emphasis id="B914EABA0221FF9FFE7EFD648976FCCD" box="[470,595,748,773]" italics="true" pageId="1" pageNumber="290">Malthopsis</emphasis>
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</taxonomicName>
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distinguished by the following combination of characters: body covered by blunt, smooth bucklers and numerous prickles; rostrum moderately long (mean 7.7% SL) and stout, with a broad base, directed forward horizontally; three irregular rows of bucklers on top of head; ventral surface of disk densely covered by prickles and scattered small bucklers, with some slightly larger bucklers on breast; subopercular buckler blunt, without enlarged spines; orbit small (mean 12.4% SL); interorbital space broad (mean 8.5% SL), forming a deep groove; OD/RL = 1.3–2.2; OD/IO = 1.3–1.7; and anal fin reaches or nearly to the caudal-fin base when fully depressed.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C37A65230221FF9CFF6FFC6F88B6F87F" lastPageId="2" lastPageNumber="291" pageId="1" pageNumber="290" type="description">
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||
<paragraph id="8BDF36A80221FF9FFF6FFC6F88BBFBED" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
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<emphasis id="B914EABA0221FF9FFF6FFC6F8A7DFBC8" bold="true" box="[199,344,999,1024]" pageId="1" pageNumber="290">Description.</emphasis>
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Morphometric and meristic data are provided in
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<tableCitation id="C6E203130221FF9FFC0AFC6F88DFFBC8" box="[930,1018,999,1024]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1401,238,1262]" captionTargetPageId="2" captionText="TABLE 1. Morphometric and meristic data of two new Malthopsis species described in present study. Pectoral-fin rays are counted on both sides." httpUri="http://table.plazi.org/id/DF1F66200222FF9CFF3FFF108A88FF1A" pageId="1" pageNumber="290" tableUuid="DF1F66200222FF9CFF3FFF108A88FF1A">Table 1</tableCitation>
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||
. Values are given for the
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<typeStatus id="54DB880A0221FF9FFA9BFC6F8EBDFBC8" box="[1331,1432,999,1024]" pageId="1" pageNumber="290" type="holotype">holotype</typeStatus>
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, followed by the range and/or mean values of all
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<typeStatus id="54DB880A0221FF9FFD1DFB8589D5FBED" box="[693,752,1037,1061]" pageId="1" pageNumber="290">types</typeStatus>
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in parentheses.
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</paragraph>
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<paragraph id="8BDF36A80221FF9FFF6FFBA78800FAE8" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
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Body depressed, disk markedly triangular in dorsal view, head elevated above rest of disk; tail base broad; caudal peduncle thick and broad, semi-cylindrical, flattened ventrally and tapering posteriorly; rostrum thick and pointed, directed forward horizontally (
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<figureCitation id="135B2A2D0221FF9FFDF0FBFF8982FB58" box="[600,679,1143,1168]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Figs. 2</figureCitation>
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A–B), with a broad base, distinctly overhanging illicial cavity and mouth; rostrum moderately long, 6.2% SL (6.2–9.4% SL, mean 7.7% SL), shorter than orbital diameter; orbit small, 13.5% SL (11.1–13.5% SL, mean 12.4% SL), eyes directed dorsolaterally; no pupillary operculum; interorbital space wide 8.1% SL (7.6–9.8% SL, mean 8.5% SL), forming a deep groove (
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<figureCitation id="135B2A2D0221FF9FFB3AFB6C8FF2FB34" box="[1170,1239,1252,1277]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
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A); frontal ridges strongly convex, elevated above the rostral base (
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<figureCitation id="135B2A2D0221FF9FFD17FA8F8821FAE8" box="[703,772,1287,1312]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
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B).
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</paragraph>
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<paragraph id="8BDF36A80221FF9FFF6FFAA48F95FA45" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">Illicial cavity large, triangular, higher than wide; esca a single bulb, bearing 2 small cirri on dorsal margin; mouth small, terminal; small villiform teeth on jaws forming narrow bands, those on fifth ceratobranchial forming 2 large, elongated patches close together; teeth on vomer and palatines in quadrangular patch.</paragraph>
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<paragraph id="8BDF36A80221FF9FFF6FFA1F8967F93D" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
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Dorsal disk squamation well developed, consisting mainly of blunt, medium-sized, smoothly conical bucklers (
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<figureCitation id="135B2A2D0221FF9FFF37FA348BC0FA1C" box="[159,229,1468,1493]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
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C); interspaces between large bucklers covered by small bucklers and prickles; bucklers on frontal ridge small and blunt, 2 enlarged preorbital bucklers which overlap front border of orbit, upper buckler larger, fused to base of rostrum; usually 4 (4–5) smaller, subequal-sized blunt bucklers on supraorbit (
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<figureCitation id="135B2A2D0221FF9FFBC2F98C8F8AF9D4" box="[1130,1199,1540,1565]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
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B), not well-elevated from the margin; interorbital space densely covered by small bucklers; supraorbital membrane covered with several irregular rows of small bucklers on upper margin of orbit, naked elsewhere. Dorsal surface of head covered by about 3 irregular rows of 3 or 4 slightly larger bucklers on each row, with many smaller ones adjacent (
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<figureCitation id="135B2A2D0221FF9FFA9CF9E78E5DF940" box="[1332,1400,1647,1672]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
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A), followed by two irregular median rows of large bucklers predorsally, ending before dorsal-fin origin as a rectangular cluster of 5 large bucklers (
|
||
<figureCitation id="135B2A2D0221FF9FFDC1F93F8994F918" box="[617,689,1719,1744]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
|
||
C); shoulder with small scattered naked areas, extending to gill openings and base of pectoral elbows.
|
||
</paragraph>
|
||
<paragraph id="8BDF36A80221FF9FFF6FF9778FD4F839" blockId="1.[151,1436,604,2033]" pageId="1" pageNumber="290">
|
||
Disk margin with cluster of suborbital bucklers anteriorly, forming 3 well-defined rows posteriorly; uppermost row of bucklers elevated, broad based; middle and lower rows of bucklers depressed and associated with lateral line, all blunt; bucklers on middle row directed laterally, lower row directed ventrally and inferior; neuromasts well defined. Subopercular buckler enlarged, extended slightly beyond disk margin laterally; terminating on uppermost and middle rows of disk-margin bucklers; small spinelets (variable in size and coverage) at tip of subopercular buckler; two well-defined, blunt, post-subopercular bucklers (
|
||
<figureCitation id="135B2A2D0221FF9FFCF0F83C88B8F804" box="[856,925,1972,1997]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="1" pageNumber="290">Fig. 2</figureCitation>
|
||
D). Pectoral-fin base at rear of disk; covered dorsally with well-developed bucklers; 4 (3–5) fine bucklers over anterior fin ray, naked elsewhere.
|
||
</paragraph>
|
||
<caption id="DF1F66200222FF9CFF3FFF108A88FF1A" ID-Table-UUID="DF1F66200222FF9CFF3FFF108A88FF1A" httpUri="http://table.plazi.org/id/DF1F66200222FF9CFF3FFF108A88FF1A" pageId="2" pageNumber="291" targetBox="[159,1401,238,1262]" targetIsTable="true" targetPageId="2">
|
||
<paragraph id="8BDF36A80222FF9CFF3FFF108A88FF1A" blockId="2.[151,1435,151,210]" pageId="2" pageNumber="291">
|
||
<emphasis id="B914EABA0222FF9CFF3FFF108A2FFF67" bold="true" box="[151,266,152,175]" pageId="2" pageNumber="291">TABLE 1.</emphasis>
|
||
Morphometric and meristic data of two new
|
||
<taxonomicName id="4C604D2B0222FF9CFD5CFF1F884FFF67" box="[756,874,151,175]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="genus">
|
||
<emphasis id="B914EABA0222FF9CFD5CFF1F884FFF67" box="[756,874,151,175]" italics="true" pageId="2" pageNumber="291">Malthopsis</emphasis>
|
||
</taxonomicName>
|
||
species described in present study. Pectoral-fin rays are counted on both sides.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDF36A80222FF9CFD83FF678E07FB26" pageId="2" pageNumber="291">
|
||
<table id="F960C40802220061FF37FF668E5CFB26" box="[159,1401,238,1262]" gridcols="3" gridrows="23" pageId="2" pageNumber="291">
|
||
<tr id="355034EA02220061FF37FF668E5CFECC" box="[159,1401,238,260]" gridrow="0" pageId="2" pageNumber="291" rowspan-0="1">
|
||
<th id="76815D9602220061FE1DFF668846FECC" box="[437,867,238,260]" gridcol="1" gridrow="0" pageId="2" pageNumber="291">
|
||
<taxonomicName id="4C604D2B0222FF9CFD83FF6789E7FECC" box="[555,706,238,260]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="species" species="bradburyae" status="sp. nov.">
|
||
<emphasis id="B914EABA0222FF9CFD83FF6789E7FECC" box="[555,706,238,260]" italics="true" pageId="2" pageNumber="291">M. bradburyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22757C10222FF9CFD61FF678825FECC" box="[713,768,239,260]" pageId="2" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
|
||
</th>
|
||
<th id="76815D9602220061FC6CFF668E5CFECC" box="[964,1401,238,260]" gridcol="2" gridrow="0" pageId="2" pageNumber="291">
|
||
<taxonomicName id="4C604D2B0222FF9CFB9CFF678FFAFECC" box="[1076,1247,238,260]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="2" pageNumber="291" phylum="Chordata" rank="species" species="austrafricana" status="sp. nov.">
|
||
<emphasis id="B914EABA0222FF9CFB9CFF678FFAFECC" box="[1076,1247,238,260]" italics="true" pageId="2" pageNumber="291">M. austrafricana</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22757C10222FF9CFB4DFF678E39FECC" box="[1253,1308,239,260]" pageId="2" pageNumber="291" rank="species">n. sp.</taxonomicNameLabel>
|
||
</th>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FE978E5CFEFD" box="[159,1401,287,309]" gridrow="1" pageId="2" pageNumber="291" rowspan-0="1">
|
||
<td id="76815D9602220061FE1DFE978846FEFD" box="[437,867,287,309]" gridcol="1" gridrow="1" pageId="2" pageNumber="291">Holotype All types</td>
|
||
<td id="76815D9602220061FC6CFE978E5CFEFD" box="[964,1401,287,309]" gridcol="2" gridrow="1" pageId="2" pageNumber="291">Holotype All types</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FED98E5CFEAF" box="[159,1401,337,359]" gridrow="2" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FED98A47FEAF" box="[159,354,337,359]" gridcol="0" gridrow="2" pageId="2" pageNumber="291">SL</th>
|
||
<td id="76815D9602220061FE1DFED98846FEAF" box="[437,867,337,359]" gridcol="1" gridrow="2" pageId="2" pageNumber="291">72.4 35.6–72.4 (n = 9)</td>
|
||
<td id="76815D9602220061FC6CFED98E5CFEAF" box="[964,1401,337,359]" gridcol="2" gridrow="2" pageId="2" pageNumber="291">70.0 45.2–84.0 (n = 17)</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FE0B8E5CFE51" box="[159,1401,387,409]" gridrow="3" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FE0B8A47FE51" box="[159,354,387,409]" gridcol="0" gridrow="3" pageId="2" pageNumber="291">% SL</th>
|
||
<td id="76815D9602220061FE1DFE0B8846FE51" box="[437,867,387,409]" gridcol="1" gridrow="3" pageId="2" pageNumber="291">Mean (Range) SD</td>
|
||
<td id="76815D9602220061FC6CFE0B8E5CFE51" box="[964,1401,387,409]" gridcol="2" gridrow="3" pageId="2" pageNumber="291">Mean (Range) SD</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FE278E5CFE0D" box="[159,1401,431,453]" gridrow="4" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FE278A47FE0D" box="[159,354,431,453]" gridcol="0" gridrow="4" pageId="2" pageNumber="291">Head length</th>
|
||
<td id="76815D9602220061FE1DFE278846FE0D" box="[437,867,431,453]" gridcol="1" gridrow="4" pageId="2" pageNumber="291">27.2 27.8 (26.0–29.5) 1.1</td>
|
||
<td id="76815D9602220061FC6CFE278E5CFE0D" box="[964,1401,431,453]" gridcol="2" gridrow="4" pageId="2" pageNumber="291">27.4 27.1 (24.8–30.3) 1.3</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FE538E5CFE39" box="[159,1401,475,497]" gridrow="5" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FE538A47FE39" box="[159,354,475,497]" gridcol="0" gridrow="5" pageId="2" pageNumber="291">Head width</th>
|
||
<td id="76815D9602220061FE1DFE538846FE39" box="[437,867,475,497]" gridcol="1" gridrow="5" pageId="2" pageNumber="291">21.5 22.5 (20.8–25.0) 1.2</td>
|
||
<td id="76815D9602220061FC6CFE538E5CFE39" box="[964,1401,475,497]" gridcol="2" gridrow="5" pageId="2" pageNumber="291">22.9 23.0 (20.6–25.8) 1.4</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FD8F8E5CFDD5" box="[159,1401,519,541]" gridrow="6" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FD8F8A47FDD5" box="[159,354,519,541]" gridcol="0" gridrow="6" pageId="2" pageNumber="291">Head depth</th>
|
||
<td id="76815D9602220061FE1DFD8F8846FDD5" box="[437,867,519,541]" gridcol="1" gridrow="6" pageId="2" pageNumber="291">22.8 23.1 (21.2–25.9) 1.5</td>
|
||
<td id="76815D9602220061FC6CFD8F8E5CFDD5" box="[964,1401,519,541]" gridcol="2" gridrow="6" pageId="2" pageNumber="291">21.4 22.2 (19.8–24.0) 1.4</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FDBB8E5CFD81" box="[159,1401,563,585]" gridrow="7" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FDBB8A47FD81" box="[159,354,563,585]" gridcol="0" gridrow="7" pageId="2" pageNumber="291">Orbital width</th>
|
||
<td id="76815D9602220061FE1DFDBB8846FD81" box="[437,867,563,585]" gridcol="1" gridrow="7" pageId="2" pageNumber="291">13.5 12.4 (11.2–13.5) 0.8</td>
|
||
<td id="76815D9602220061FC6CFDBB8E5CFD81" box="[964,1401,563,585]" gridcol="2" gridrow="7" pageId="2" pageNumber="291">12.1 12.5 (10.7–14.5) 0.9</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FDD78E5CFDBD" box="[159,1401,607,629]" gridrow="8" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FDD78A47FDBD" box="[159,354,607,629]" gridcol="0" gridrow="8" pageId="2" pageNumber="291">Interorbiral width</th>
|
||
<td id="76815D9602220061FE1DFDD78846FDBD" box="[437,867,607,629]" gridcol="1" gridrow="8" pageId="2" pageNumber="291">8.1 8.5 (7.6–9.8) 0.7</td>
|
||
<td id="76815D9602220061FC6CFDD78E5CFDBD" box="[964,1401,607,629]" gridcol="2" gridrow="8" pageId="2" pageNumber="291">9.4 8.4 (7.4–9.7) 0.7</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FD038E5CFD69" box="[159,1401,651,673]" gridrow="9" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FD038A47FD69" box="[159,354,651,673]" gridcol="0" gridrow="9" pageId="2" pageNumber="291">Rostral length</th>
|
||
<td id="76815D9602220061FE1DFD038846FD69" box="[437,867,651,673]" gridcol="1" gridrow="9" pageId="2" pageNumber="291">6.2 7.7 (6.2–9.4) 1.1</td>
|
||
<td id="76815D9602220061FC6CFD038E5CFD69" box="[964,1401,651,673]" gridcol="2" gridrow="9" pageId="2" pageNumber="291">9.4 8.3 (6.0–10.7) 1.7</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FD3F8E5CFD05" box="[159,1401,695,717]" gridrow="10" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FD3F8A47FD05" box="[159,354,695,717]" gridcol="0" gridrow="10" pageId="2" pageNumber="291">Predorsal length</th>
|
||
<td id="76815D9602220061FE1DFD3F8846FD05" box="[437,867,695,717]" gridcol="1" gridrow="10" pageId="2" pageNumber="291">66.2 67.5 (65.3–69.8) 1.8</td>
|
||
<td id="76815D9602220061FC6CFD3F8E5CFD05" box="[964,1401,695,717]" gridcol="2" gridrow="10" pageId="2" pageNumber="291">64.3 12.9 (10.7–14.5) 1.0</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FD6B8E5CFD31" box="[159,1401,739,761]" gridrow="11" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FD6B8A47FD31" box="[159,354,739,761]" gridcol="0" gridrow="11" pageId="2" pageNumber="291">Preanus length</th>
|
||
<td id="76815D9602220061FE1DFD6B8846FD31" box="[437,867,739,761]" gridcol="1" gridrow="11" pageId="2" pageNumber="291">53.0 55.3 (53.0–58.2) 1.9</td>
|
||
<td id="76815D9602220061FC6CFD6B8E5CFD31" box="[964,1401,739,761]" gridcol="2" gridrow="11" pageId="2" pageNumber="291">54.3 66.5 (63.8–71.4) 2.0</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FC878E5CFCED" box="[159,1401,783,805]" gridrow="12" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FC878A47FCED" box="[159,354,783,805]" gridcol="0" gridrow="12" pageId="2" pageNumber="291">Preanal length</th>
|
||
<td id="76815D9602220061FE1DFC878846FCED" box="[437,867,783,805]" gridcol="1" gridrow="12" pageId="2" pageNumber="291">79.8 81.0 (77.5–83.0) 1.8</td>
|
||
<td id="76815D9602220061FC6CFC878E5CFCED" box="[964,1401,783,805]" gridcol="2" gridrow="12" pageId="2" pageNumber="291">82.4 54.8 (52.6–57.5) 1.5</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FCB38E5CFC99" box="[159,1401,827,849]" gridrow="13" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FCB38A47FC99" box="[159,354,827,849]" gridcol="0" gridrow="13" pageId="2" pageNumber="291">Disk margin length</th>
|
||
<td id="76815D9602220061FE1DFCB38846FC99" box="[437,867,827,849]" gridcol="1" gridrow="13" pageId="2" pageNumber="291">43.0 44.1 (40.1–47.2) 2.7</td>
|
||
<td id="76815D9602220061FC6CFCB38E5CFC99" box="[964,1401,827,849]" gridcol="2" gridrow="13" pageId="2" pageNumber="291">44.9 80.5 (78.1–82.9) 1.3</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FCEF8E5CFCB5" box="[159,1401,871,893]" gridrow="14" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FCEF8A47FCB5" box="[159,354,871,893]" gridcol="0" gridrow="14" pageId="2" pageNumber="291">Mouth width</th>
|
||
<td id="76815D9602220061FE1DFCEF8846FCB5" box="[437,867,871,893]" gridcol="1" gridrow="14" pageId="2" pageNumber="291">14.4 14.4 (13.1–15.9) 0.9</td>
|
||
<td id="76815D9602220061FC6CFCEF8E5CFCB5" box="[964,1401,871,893]" gridcol="2" gridrow="14" pageId="2" pageNumber="291">12.9 44.5 (39.6–48.1) 2.7</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FC1B8E5CFC61" box="[159,1401,915,937]" gridrow="15" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FC1B8A47FC61" box="[159,354,915,937]" gridcol="0" gridrow="15" pageId="2" pageNumber="291">Dorsal-fin length</th>
|
||
<td id="76815D9602220061FE1DFC1B8846FC61" box="[437,867,915,937]" gridcol="1" gridrow="15" pageId="2" pageNumber="291">18.9 18.6 (16.8–20.2) 1.2</td>
|
||
<td id="76815D9602220061FC6CFC1B8E5CFC61" box="[964,1401,915,937]" gridcol="2" gridrow="15" pageId="2" pageNumber="291">17.4 22.3 (19.6–24.8) 1.4</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FC378E5CFC1D" box="[159,1401,959,981]" gridrow="16" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FC378A47FC1D" box="[159,354,959,981]" gridcol="0" gridrow="16" pageId="2" pageNumber="291">Anal-fin length</th>
|
||
<td id="76815D9602220061FE1DFC378846FC1D" box="[437,867,959,981]" gridcol="1" gridrow="16" pageId="2" pageNumber="291">15.1 17.7 (15.1–20.5) 1.9</td>
|
||
<td id="76815D9602220061FC6CFC378E5CFC1D" box="[964,1401,959,981]" gridcol="2" gridrow="16" pageId="2" pageNumber="291">14.3 17.4 (14.3–19.3) 1.4</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FC638E5CFBC9" box="[159,1401,1003,1025]" gridrow="17" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FC638A47FBC9" box="[159,354,1003,1025]" gridcol="0" gridrow="17" pageId="2" pageNumber="291">Pectoral-fin length</th>
|
||
<td id="76815D9602220061FE1DFC638846FBC9" box="[437,867,1003,1025]" gridcol="1" gridrow="17" pageId="2" pageNumber="291">22.0 22.5 (20.9–24.2) 1.1</td>
|
||
<td id="76815D9602220061FC6CFC638E5CFBC9" box="[964,1401,1003,1025]" gridcol="2" gridrow="17" pageId="2" pageNumber="291">22.6 17.7 (14.3–20.3) 1.7</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FB9F8E5CFBE5" box="[159,1401,1047,1069]" gridrow="18" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FB9F8A47FBE5" box="[159,354,1047,1069]" gridcol="0" gridrow="18" pageId="2" pageNumber="291">Caudal-fin length</th>
|
||
<td id="76815D9602220061FE1DFB9F8846FBE5" box="[437,867,1047,1069]" gridcol="1" gridrow="18" pageId="2" pageNumber="291">25.8 26.1 (24.0–29.2) 1.9</td>
|
||
<td id="76815D9602220061FC6CFB9F8E5CFBE5" box="[964,1401,1047,1069]" gridcol="2" gridrow="18" pageId="2" pageNumber="291">25.7 25.2 (23.1–27.3) 1.3</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FBCB8E5CFB91" box="[159,1401,1091,1113]" gridrow="19" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FBCB8A47FB91" box="[159,354,1091,1113]" gridcol="0" gridrow="19" pageId="2" pageNumber="291">Meristics</th>
|
||
<td id="76815D9602220061FE1DFBCB8846FB91" box="[437,867,1091,1113]" gridcol="1" gridrow="19" pageId="2" pageNumber="291">Value (Frequency)</td>
|
||
<td id="76815D9602220061FC6CFBCB8E5CFB91" box="[964,1401,1091,1113]" gridcol="2" gridrow="19" pageId="2" pageNumber="291">Value (Frequency)</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FBFC8E5CFB42" box="[159,1401,1140,1162]" gridrow="20" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FBFC8A47FB42" box="[159,354,1140,1162]" gridcol="0" gridrow="20" pageId="2" pageNumber="291">Dorsal-fin rays</th>
|
||
<td id="76815D9602220061FE1DFBFC8846FB42" box="[437,867,1140,1162]" gridcol="1" gridrow="20" pageId="2" pageNumber="291">5 5 (7), 6 (2)</td>
|
||
<td id="76815D9602220061FC6CFBFC8E5CFB42" box="[964,1401,1140,1162]" gridcol="2" gridrow="20" pageId="2" pageNumber="291">5 5 (12), 6 (5)</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FB2E8E5CFB74" box="[159,1401,1190,1212]" gridrow="21" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FB2E8A47FB74" box="[159,354,1190,1212]" gridcol="0" gridrow="21" pageId="2" pageNumber="291">Pectoral-fin rays</th>
|
||
<td id="76815D9602220061FE1DFB2E8846FB74" box="[437,867,1190,1212]" gridcol="1" gridrow="21" pageId="2" pageNumber="291">12/13 11 (2), 12 (15), 13 (1)</td>
|
||
<td id="76815D9602220061FC6CFB2E8E5CFB74" box="[964,1401,1190,1212]" gridcol="2" gridrow="21" pageId="2" pageNumber="291">13/13 12 (30), 13 (4)</td>
|
||
</tr>
|
||
<tr id="355034EA02220061FF37FB508E5CFB26" box="[159,1401,1240,1262]" gridrow="22" pageId="2" pageNumber="291">
|
||
<th id="76815D9602220061FF37FB508A47FB26" box="[159,354,1240,1262]" gridcol="0" gridrow="22" pageId="2" pageNumber="291">Anal-fin rays</th>
|
||
<td id="76815D9602220061FE1DFB508846FB26" box="[437,867,1240,1262]" gridcol="1" gridrow="22" pageId="2" pageNumber="291">3 3 (3), 4 (5)</td>
|
||
<td id="76815D9602220061FC6CFB508E5CFB26" box="[964,1401,1240,1262]" gridcol="2" gridrow="22" pageId="2" pageNumber="291">4 3(2), 4 (15)</td>
|
||
</tr>
|
||
</table>
|
||
</paragraph>
|
||
<paragraph id="8BDF36A80222FF9CFF6FFAB288D7F9CF" blockId="2.[151,1436,1338,2011]" pageId="2" pageNumber="291">Dorsal surface of tail strongly armoured, entirely covered with large blunt bucklers; a row of 4–5 large dorsolateral bucklers, extending from last pair of predorsal bucklers below dorsal fin; a longer, highly irregular, semi-oblique row of slightly smaller bucklers extending along lateral margin of peduncle to caudal-fin base; a single irregular row of flattened bucklers on dorsal midline before caudal fin; dorsal tail rows uniting to form a single, slightly elevated buckler at caudal-fin base. Lateral margin of tail with two rows of small, low bucklers associated with lateral line, similar in appearance to bucklers of disk margin.</paragraph>
|
||
<paragraph id="8BDF36A80222FF9CFF6FF99A8BF5F973" blockId="2.[151,1436,1338,2011]" pageId="2" pageNumber="291">
|
||
Ventral surface of disk predominantly covered with small, flat bucklers; no apical spines on these bucklers; belly densely covered with prickles and scattered small bucklers, some slightly larger on breast than posteiorly (
|
||
<figureCitation id="135B2A2D0222FF9CFF37F9D28BC6F9BB" box="[159,227,1626,1651]" captionStart="FIGURE 2" captionStartId="4.[151,250,1627,1649]" captionTargetBox="[154,1436,199,1599]" captionTargetId="figure@4.[154,1439,195,1608]" captionTargetPageId="4" captionText="FIGURE 2. Close-up view of body portions of Malthopsis bradburyae n. sp., from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale." httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="2" pageNumber="291">Fig. 2</figureCitation>
|
||
E); rear margin of anus surrounded by 5 (4–5) blunt bucklers; large naked area outside gill chamber; ventral surface of tail with a single regular row of blunt bucklers on each side, coalescing to bulbous buckler at caudal-fin base.
|
||
</paragraph>
|
||
<paragraph id="8BDF36A80222FF9CFF6FF94E8A65F883" blockId="2.[151,1436,1338,2011]" pageId="2" pageNumber="291">Fins generally naked, without bucklers, sometimes with small bucklers on rays near caudal-fin base; interradials of pectoral fins thin, transparent; dermal cirri short, thin, flap-like, present on disk margin and lateral sides of tail and associated with lateral-line neuromasts. Anal fin reaches to or slightly short of caudal-fin base when fully laid back.</paragraph>
|
||
<paragraph id="8BDF36A80222FF9CFF6FF8DE88E0F85B" blockId="2.[151,1436,1338,2011]" pageId="2" pageNumber="291">
|
||
<emphasis id="B914EABA0222FF9CFF6FF8DE8A6AF8A7" bold="true" box="[199,335,1878,1903]" pageId="2" pageNumber="291">Coloration.</emphasis>
|
||
Fresh color unknown. Preserved specimens (
|
||
<figureCitation id="135B2A2D0222FF9CFCF9F8DE88BEF8A7" box="[849,923,1878,1903]" captionStart="FIGURE 1" captionStartId="3.[151,250,1958,1980]" captionTargetBox="[352,1229,212,1915]" captionTargetId="figure@3.[338,1248,195,1934]" captionTargetPageId="3" captionText="FIGURE 1. Malthopsis bradburyae n. sp. A. Holotype, CAS 227226, 72.4 mm SL, dorsal view, preserved. B. holotype, ventral view, preserved. C. Paratype, CAS 39631, 1 of 7, 70.9 mm SL, dorsal view, preserved." httpUri="https://zenodo.org/record/223738/files/figure.png" pageId="2" pageNumber="291">Figs. 1</figureCitation>
|
||
A, C) uniformly creamy yellowish.
|
||
<typeStatus id="54DB880A0222FF9CFA9CF8DE8EB9F8A7" box="[1332,1436,1878,1903]" pageId="2" pageNumber="291" type="holotype">Holotype</typeStatus>
|
||
and some
|
||
<typeStatus id="54DB880A0222FF9CFEA0F8F38A56F85B" box="[264,371,1915,1939]" pageId="2" pageNumber="291" type="paratype">paratypes</typeStatus>
|
||
with very fine brown reticulations on dorsal surface.
|
||
</paragraph>
|
||
<paragraph id="8BDF36A80222FF9CFF6FF81688B6F87F" blockId="2.[151,1436,1338,2011]" box="[199,915,1950,1975]" pageId="2" pageNumber="291">
|
||
<emphasis id="B914EABA0222FF9CFF6FF8168BDBF87F" bold="true" box="[199,254,1950,1975]" pageId="2" pageNumber="291">Size.</emphasis>
|
||
A medium-sized species. The largest adult is
|
||
<quantity id="4C989B4D0222FF9CFCA9F8178840F87F" box="[769,869,1951,1975]" metricMagnitude="-2" metricUnit="m" metricValue="7.24" pageId="2" pageNumber="291" unit="mm" value="72.4">72.4 mm</quantity>
|
||
SL.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A65230222FF9CFF6FF84A8F53F812" box="[199,1142,1986,2011]" pageId="2" pageNumber="291" type="distribution">
|
||
<paragraph id="8BDF36A80222FF9CFF6FF84A8F53F812" blockId="2.[151,1436,1338,2011]" box="[199,1142,1986,2011]" pageId="2" pageNumber="291">
|
||
<emphasis id="B914EABA0222FF9CFF6FF84A8A45F813" bold="true" box="[199,352,1986,2011]" pageId="2" pageNumber="291">Distribution.</emphasis>
|
||
Known only from the
|
||
<typeStatus id="54DB880A0222FF9CFDC8F84B89B5F813" box="[608,656,1987,2011]" pageId="2" pageNumber="291">type</typeStatus>
|
||
series taken off
|
||
<collectingCountry id="F37776380222FF9CFCE1F84A8888F813" box="[841,941,1986,2011]" name="Tanzania" pageId="2" pageNumber="291">Tanzania</collectingCountry>
|
||
at a depth
|
||
<specimenCount id="9D66FD210222FF9CFB80F84B8F53F812" box="[1064,1142,1987,2011]" pageId="2" pageNumber="291" type="generic">
|
||
<quantity id="4C989B4D0222FF9CFB80F84B8F57F812" box="[1064,1138,1987,2011]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="2" pageNumber="291" unit="m" value="100.0">100 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF1F66200223FF9DFF3FF82E8F21F813" httpUri="https://zenodo.org/record/223738/files/figure.png" pageId="3" pageNumber="292" targetBox="[352,1229,212,1915]" targetPageId="3">
|
||
<paragraph id="8BDF36A80223FF9DFF3FF82E8F21F813" blockId="3.[151,1435,1958,2011]" pageId="3" pageNumber="292">
|
||
<emphasis id="B914EABA0223FF9DFF3FF82E8A37F874" bold="true" box="[151,274,1958,1980]" pageId="3" pageNumber="292">FIGURE 1.</emphasis>
|
||
<taxonomicName id="4C604D2B0223FF9DFEB0F82E8921F874" box="[280,516,1958,1980]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="3" pageNumber="292" phylum="Chordata" rank="species" species="bradburyae" status="sp. nov.">
|
||
<emphasis id="B914EABA0223FF9DFEB0F82E8921F874" box="[280,516,1958,1980]" italics="true" pageId="3" pageNumber="292">Malthopsis bradburyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22757C10223FF9DFDA2F82F8965F874" box="[522,576,1959,1980]" pageId="3" pageNumber="292" rank="species">n. sp.</taxonomicNameLabel>
|
||
A. Holotype, CAS 227226, 72.4 mm SL, dorsal view, preserved. B. holotype, ventral view, preserved. C. Paratype, CAS 39631, 1 of 7, 70.9 mm SL, dorsal view, preserved.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1F66200224FF9AFF3FF9D38E63F966" httpUri="https://zenodo.org/record/223739/files/figure.png" pageId="4" pageNumber="293" targetBox="[154,1436,199,1599]" targetPageId="4">
|
||
<paragraph id="8BDF36A80224FF9AFF3FF9D38E63F966" blockId="4.[151,1436,1627,1710]" pageId="4" pageNumber="293">
|
||
<emphasis id="B914EABA0224FF9AFF3FF9D38A31F9B9" bold="true" box="[151,276,1627,1649]" pageId="4" pageNumber="293">FIGURE 2.</emphasis>
|
||
Close-up view of body portions of
|
||
<taxonomicName id="4C604D2B0224FF9AFD2DF9D38856F9B9" box="[645,883,1627,1649]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="bradburyae" status="sp. nov.">
|
||
<emphasis id="B914EABA0224FF9AFD2DF9D38856F9B9" box="[645,883,1627,1649]" italics="true" pageId="4" pageNumber="293">Malthopsis bradburyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22757C10224FF9AFCD3F9D48891F9B9" box="[891,948,1628,1649]" pageId="4" pageNumber="293" rank="species">n. sp.</taxonomicNameLabel>
|
||
, from the holotype. A. Dorsal view of head. B. Lateral view of head. C. Dorsal view of dorsal surface between dorsal fin and cranium. D. Dorsal view of right portion of disk. E. Ventral view of surface of central disk. Anterior to left in Figs. A, D, E. Anterior to right in Figs. B, C. Not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C37A65230224FF9AFF6FF9538870F8D0" pageId="4" pageNumber="293" type="etymology">
|
||
<paragraph id="8BDF36A80224FF9AFF6FF9538870F8D0" blockId="4.[151,1436,1755,2032]" pageId="4" pageNumber="293">
|
||
<emphasis id="B914EABA0224FF9AFF6FF9538A6BF93C" bold="true" box="[199,334,1755,1780]" pageId="4" pageNumber="293">Etymology.</emphasis>
|
||
The species is named after the late Dr. Margaret G. Bradbury, in recognition of her great contribution to our knowledge of batfishes, and her friendship.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A65230224FF9BFF6FF8AB8975FE64" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293" type="discussion">
|
||
<paragraph id="8BDF36A80224FF9BFF6FF8AB8EAEFF30" blockId="4.[151,1436,1755,2032]" lastBlockId="5.[151,1436,151,429]" lastPageId="5" lastPageNumber="294" pageId="4" pageNumber="293">
|
||
<emphasis id="B914EABA0224FF9AFF6FF8AB8A1EF8F4" bold="true" box="[199,315,1827,1852]" pageId="4" pageNumber="293">Remarks.</emphasis>
|
||
<taxonomicName id="4C604D2B0224FF9AFEEBF8AB896EF8F4" box="[323,587,1827,1852]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="bradburyae" status="sp. nov.">
|
||
<emphasis id="B914EABA0224FF9AFEEBF8AB896EF8F4" box="[323,587,1827,1852]" italics="true" pageId="4" pageNumber="293">Malthopsis bradburyae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22757C10224FF9AFDFBF8AB89B6F8F4" box="[595,659,1827,1852]" pageId="4" pageNumber="293" rank="species">
|
||
<emphasis id="B914EABA0224FF9AFDFBF8AB89B6F8F4" bold="true" box="[595,659,1827,1852]" pageId="4" pageNumber="293">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
belongs to a species group with the ventral surface densely covered by minute prickles (e.g.,
|
||
<taxonomicName id="4C604D2B0224FF9AFE12F8C08979F8A8" box="[442,604,1863,1888]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="kobayashii">
|
||
<emphasis id="B914EABA0224FF9AFE12F8C08979F8A8" box="[442,604,1863,1888]" italics="true" pageId="4" pageNumber="293">M. kobayashii</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C604D2B0224FF9AFDC5F8C089C3F8A8" box="[621,742,1863,1888]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="tiarella">
|
||
<emphasis id="B914EABA0224FF9AFDC5F8C089C3F8A8" box="[621,742,1863,1888]" italics="true" pageId="4" pageNumber="293">M. tiarella</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C604D2B0224FF9AFD50F8C088A7F897" box="[760,898,1864,1887]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="asperata">
|
||
<emphasis id="B914EABA0224FF9AFD50F8C088A7F897" box="[760,898,1864,1887]" italics="true" pageId="4" pageNumber="293">M. asperata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C604D2B0224FF9AFC17F8C08F16F897" box="[959,1075,1864,1887]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="gnoma">
|
||
<emphasis id="B914EABA0224FF9AFC17F8C08F16F897" box="[959,1075,1864,1887]" italics="true" pageId="4" pageNumber="293">M. gnoma</emphasis>
|
||
</taxonomicName>
|
||
). It differs from
|
||
<taxonomicName id="4C604D2B0224FF9AFB54F8C08E50F8A8" box="[1276,1397,1863,1888]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="tiarella">
|
||
<emphasis id="B914EABA0224FF9AFB54F8C08E50F8A8" box="[1276,1397,1863,1888]" italics="true" pageId="4" pageNumber="293">M. tiarella</emphasis>
|
||
</taxonomicName>
|
||
by having 5–6 dorsal-fin rays (modally 5, vs.
|
||
<quantity id="4C989B4D0224FF9AFD22F8E389F1F84C" box="[650,724,1899,1924]" metricMagnitude="-1" metricUnit="m" metricValue="1.651" metricValueMax="1.778" metricValueMin="1.524" pageId="4" pageNumber="293" unit="in" value="6.5" valueMax="7.0" valueMin="6.0">6–7 in</quantity>
|
||
<taxonomicName id="4C604D2B0224FF9AFD76F8E4887DF84C" box="[734,856,1899,1924]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="tiarella">
|
||
<emphasis id="B914EABA0224FF9AFD76F8E4887DF84C" box="[734,856,1899,1924]" italics="true" pageId="4" pageNumber="293">M. tiarella</emphasis>
|
||
</taxonomicName>
|
||
), bucklers on body surface blunt and smooth (vs. pointed and rough), and lacking brown bands on the caudal peduncle (vs. bands present on peduncle); from
|
||
<taxonomicName id="4C604D2B0224FF9AFAD5F8188A35F804" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="4" pageNumber="293" phylum="Chordata" rank="species" species="kobayashii">
|
||
<emphasis id="B914EABA0224FF9AFAD5F8188A35F804" italics="true" pageId="4" pageNumber="293">M. kobayashii</emphasis>
|
||
</taxonomicName>
|
||
in its rostral spine directed forward horizontally (vs. directed forward and upward), in having a relatively wide interorbital space (7.6–9.8% SL vs. 6.0–7.5% SL), and bucklers on body surface blunt and smooth (vs. relatively pointed and rough); from
|
||
<taxonomicName id="4C604D2B0225FF9BFDCBFF1189C9FF78" box="[611,748,153,176]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="asperata">
|
||
<emphasis id="B914EABA0225FF9BFDCBFF1189C9FF78" box="[611,748,153,176]" italics="true" pageId="5" pageNumber="294">M. asperata</emphasis>
|
||
</taxonomicName>
|
||
in having all bucklers blunt (vs. pyramid-like bucklers with spinelets); and from the western Atlantic species,
|
||
<taxonomicName id="4C604D2B0225FF9BFD7AFF358860FF1C" box="[722,837,189,212]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="gnoma">
|
||
<emphasis id="B914EABA0225FF9BFD7AFF358860FF1C" box="[722,837,189,212]" italics="true" pageId="5" pageNumber="294">M. gnoma</emphasis>
|
||
</taxonomicName>
|
||
, in lacking gill filaments on the fourth gill arch (vs. filaments on the fourth gill arch); and in the rostral spine directed forward horizontally (vs. upward and forward).
|
||
</paragraph>
|
||
<paragraph id="8BDF36A80225FF9BFF6FFE8D8975FE64" blockId="5.[151,1436,151,429]" pageId="5" pageNumber="294">
|
||
It is also similar to
|
||
<taxonomicName id="4C604D2B0225FF9BFE12FE8D897FFED5" box="[442,602,260,285]" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="annulifera">
|
||
<emphasis id="B914EABA0225FF9BFE12FE8D897FFED5" box="[442,602,260,285]" italics="true" pageId="5" pageNumber="294">M. annulifera</emphasis>
|
||
</taxonomicName>
|
||
from the western Pacific in having the rostral spine directed forward horizontally and principal bucklers on the body are relatively flat and smooth. It can be distinguished from
|
||
<taxonomicName id="4C604D2B0225FF9BFAD5FEA18A29FEAD" class="Actinopterygii" family="Ogcocephalidae" genus="Malthopsis" kingdom="Animalia" order="Lophiiformes" pageId="5" pageNumber="294" phylum="Chordata" rank="species" species="annulifera">
|
||
<emphasis id="B914EABA0225FF9BFAD5FEA18A29FEAD" italics="true" pageId="5" pageNumber="294">M. annulifera</emphasis>
|
||
</taxonomicName>
|
||
in having ventral surface covered by prickles (vs. no prickles); blunt subopercular bucklers without well-developed spines (vs. 1 well-developed spine directed forward), and lacking ring marks on the dorsal surface (vs. few to more than 20 rings present).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |