treatments-xml/data/4B/73/87/4B7387B5FFBCFFFB0CDDD204A9034EF4.xml

<document id="84D497987F27E030D02D65F265ADA70A" ID-CLB-Dataset="29778" ID-DOI="10.11646/zootaxa.4441.2.4" ID-GBIF-Dataset="8c18255f-0cb3-45f7-a763-5eba38606f83" ID-ISSN="1175-5326" ID-Zenodo-Dep="1301825" ID-ZooBank="293FEFDD-21D8-4C3A-A4B6-93179C2DE585" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530159416834" checkinUser="plazi" docAuthor="Ahmad, Wasim, Ahad, Sumaya, Islam, Md. Niraul &amp; Sturhan, Dieter" docDate="2018" docId="4B7387B5FFBCFFFB0CDDD204A9034EF4" docLanguage="en" docName="zootaxa.4441.2.4.pdf" docOrigin="Zootaxa 4441 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Doryllium enigmatum Ahmad &amp; Ahad &amp; Islam &amp; Sturhan 2018, sp. n." docType="treatment" docVersion="6" lastPageNumber="270" masterDocId="B74AFFCDFFBAFFF20C4AD75FAC45497F" masterDocTitle="Taxonomy of the genus Doryllium Cobb, 1920 (Nematoda: Dorylaimida) with description of two new and a known species" masterLastPageNumber="278" masterPageNumber="261" pageNumber="267" updateTime="1698551357680" updateUser="ExternalLinkService">
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<mods:title id="F7185CB1B125CA543852F77D27580E8C">Taxonomy of the genus Doryllium Cobb, 1920 (Nematoda: Dorylaimida) with description of two new and a known species</mods:title>
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<mods:namePart id="73963ABDEC40599CCBFFCCAC0B10719E">Ahad, Sumaya</mods:namePart>
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<mods:namePart id="446AE28B404528ECBA46ABB1B18CDBAF">Islam, Md. Niraul</mods:namePart>
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<subSubSection id="8BC06528FFBCFFF40CDDD204AD624CE8" pageId="6" pageNumber="267" type="nomenclature">
<paragraph id="C36536A3FFBCFFF40CDDD204ADA94C09" blockId="6.[151,492,1371,1431]" box="[151,492,1371,1398]" pageId="6" pageNumber="267">
<heading id="982D81CFFFBCFFF40CDDD204ADA94C09" bold="true" box="[151,492,1371,1398]" fontSize="11" level="1" pageId="6" pageNumber="267" reason="1">
<emphasis id="F1AEEAB1FFBCFFF40CDDD204ADA94C09" bold="true" box="[151,492,1371,1398]" pageId="6" pageNumber="267">
<taxonomicName id="04DA4D20FFBCFFF40CDDD204ADE64C0A" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[151,419,1371,1397]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="6" pageNumber="267" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFBCFFF40CDDD204ADE64C0A" bold="true" box="[151,419,1371,1397]" italics="true" pageId="6" pageNumber="267">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFBCFFF40DE0D203ADA94C09" box="[426,492,1372,1398]" pageId="6" pageNumber="267" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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<paragraph id="C36536A3FFBCFFF40CDDD221AD624CE8" blockId="6.[151,492,1371,1431]" box="[151,295,1406,1431]" pageId="6" pageNumber="267">
(
<figureCitation id="5BE12A26FFBCFFF40CD5D221ACA94CE8" box="[159,236,1406,1431]" captionStart="FIGURE 4" captionStartId="8.[151,250,1702,1724]" captionTargetBox="[252,1346,245,1561]" captionTargetId="figure@8.[235,1351,194,1680]" captionTargetPageId="8" captionText="FIGURE 4. Dorylllium enigmatum sp. n. A. Entire female; B. Anterior region; C. Anterior end showing amphid; D. Pharyngeal region; E. Pharyngeal bulb; F. Female genital system; G. Female posterior region." httpUri="https://zenodo.org/record/1301833/files/figure.png" pageId="6" pageNumber="267">Figs. 4</figureCitation>
&amp; 
<figureCitation id="5BE12A26FFBCFFF40D5AD221AD5B4CE8" box="[272,286,1406,1431]" captionStart="FIGURE 5" captionStartId="9.[151,250,1598,1620]" captionTargetBox="[381,1219,440,1577]" captionTargetId="figure@9.[357,1234,440,1605]" captionTargetPageId="9" captionText="FIGURE 5. Doryllium enigmatum sp. n. A. Anterior region; B. Anterior region showing basal flanges; C. Anterior end showing amphid; D. &amp; E. Pharyngeal bulb; F. Female genital system; G, H, I. Female posterior region. (Scale bar A–E, H, I= 10 µm; F, G = 20 µm)." httpUri="https://zenodo.org/record/1301835/files/figure.png" pageId="6" pageNumber="267">5</figureCitation>
)
</paragraph>
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<paragraph id="C36536A3FFBCFFF40CDDD299AFD34CA0" blockId="6.[151,1437,1478,2007]" box="[151,918,1478,1503]" pageId="6" pageNumber="267">
<emphasis id="F1AEEAB1FFBCFFF40CDDD299ADC54CA0" bold="true" box="[151,384,1478,1503]" pageId="6" pageNumber="267">Material examined.</emphasis>
Fourteen females in good state of preservation.
</paragraph>
</subSubSection>
<subSubSection id="8BC06528FFBCFFF50C8DD2B5ADCE48F7" lastPageId="7" lastPageNumber="268" pageId="6" pageNumber="267" type="description">
<paragraph id="C36536A3FFBCFFF40C8DD2B5AECB4F7C" blockId="6.[151,1437,1478,2007]" box="[199,654,1514,1539]" pageId="6" pageNumber="267">
<emphasis id="F1AEEAB1FFBCFFF40C8DD2B5AD1C4F7C" bold="true" box="[199,345,1514,1539]" pageId="6" pageNumber="267">Description.</emphasis>
Measurements, see 
<tableCitation id="8E580318FFBCFFF40E71D2B5AECB4F7C" box="[571,654,1514,1539]" captionStart="TABLE 2" captionStartId="7.[151,239,728,751]" captionText="TABLE 2. Measurements of Doryllium enigmatum sp. n. (All measurements in µm)" pageId="6" pageNumber="267">Table 2</tableCitation>
</paragraph>
<paragraph id="C36536A3FFBCFFF50C8DD151ADA6481A" blockId="6.[151,1437,1478,2007]" lastBlockId="7.[151,1436,151,680]" lastPageId="7" lastPageNumber="268" pageId="6" pageNumber="267">
<emphasis id="F1AEEAB1FFBCFFF40C8DD151AD614F58" box="[199,292,1550,1575]" italics="true" pageId="6" pageNumber="267">Female:</emphasis>
Small sized nematodes, slightly curved upon fixation; body cylindrical tapering gradually towards both the extremities. Cuticle with two distinct layers, 3–4 µm thick at midbody and 4–5 µm on tail. Outer cuticle finely striated; inner layer smooth. Lateral chords occupying about one-fourth to one-third of the midbody diameter. Lateral, dorsal and ventral body pores indistinct. Lip region continuous, 1.6–2.0 times as wide as high or about one-third of the body diameter at neck base; lips rounded, amalgamated; papillae barely raised. Amphids cup-shaped, their aperture occupying about one-half of lip region diameter. Stoma a truncate cone. Odontostyle asymmetrical, 0.8–1.0 times the lip region diameter in length, with distinct lumen and aperture. Odontophore flanged, 1.8–2.1 times the odontostyle length. Guiding ring simple, refractive, at 0.6 times lip region diameter from anterior end. Pharynx consists of a slender and weakly muscular anterior part, expanding into a slightly constricted pyriform basal bulb, occupying about 16–18% of total neck length. Cardia rounded-conoid, about one-third of the corresponding body diameter long. Nerve ring located at 54–57% of neck length from anterior end.Genital system mono-opisthodelphic. Ovary reflexed, measuring 51–78 µm long, reaching the oviduct-uterus junction (surpassing the oviduct-uterus junction in one specimen); oocytes arranged in single row except near tip. Oviduct joining the ovary subterminally, measuring 65–76 µm, proximal and distal parts not differentiated. Oviduct-uterus junction marked by weak sphincter. Uterus short and tubular, measuring 15–20 µm long.Anterior genital branch small, 0.3– 0.5 times midbody diameter long.Vagina cylindrical; 
<emphasis id="F1AEEAB1FFBDFFF50F4FD7BEA85B4987" box="[773,1054,223,248]" italics="true" pageId="7" pageNumber="268">pars proximalis vaginae</emphasis>
5–6 µm long, wall encircled by muscles; 
<emphasis id="F1AEEAB1FFBDFFF50D4ED65AADB34863" box="[260,502,260,285]" italics="true" pageId="7" pageNumber="268">pars distalis vaginae</emphasis>
short, 3–4 µm long with slightly curved walls; 
<emphasis id="F1AEEAB1FFBDFFF50864D65AA8984863" box="[1070,1245,260,284]" italics="true" pageId="7" pageNumber="268">pars refringens</emphasis>
absent. Vulva a transverse slit.Prerectum 2.3–2.5 and rectum 0.9–1.2 anal body diameters long. Tail rounded-hemispheroid, 1.2– 1.5 anal body diameters long.
</paragraph>
<paragraph id="C36536A3FFBDFFF50C8DD630ADCE48F7" blockId="7.[151,1436,151,680]" box="[199,395,367,392]" pageId="7" pageNumber="268">
<emphasis id="F1AEEAB1FFBDFFF50C8DD630AD4F48F7" box="[199,266,367,392]" italics="true" pageId="7" pageNumber="268">Male:</emphasis>
Not found.
</paragraph>
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<subSubSection id="8BC06528FFBDFFF50C8DD6CBAD484B1F" pageId="7" pageNumber="268" type="materials_examined">
<paragraph id="C36536A3FFBDFFF50C8DD6CBAF6E48B0" blockId="7.[151,1436,151,680]" pageId="7" pageNumber="268">
<materialsCitation id="73B23CFEFFBDFFF50C8DD6CBAF6E48B0" ID-GBIF-Occurrence="1950907355" collectingDate="1985-05-29" collectorName="Dr. Dieter Sturhan" country="Germany" latitude="51.31278" location="Germany" longLatPrecision="17" longitude="9.481388" pageId="7" pageNumber="268" specimenCount="1" typeStatus="holotype">
<emphasis id="F1AEEAB1FFBDFFF50C8DD6CBADB148D2" bold="true" box="[199,500,404,429]" pageId="7" pageNumber="268">
<typeStatus id="1C618801FFBDFFF50C8DD6CBAD4748D2" box="[199,258,404,429]" pageId="7" pageNumber="268">Type</typeStatus>
habitat and locality.
</emphasis>
Meadow soil, Nienhagen near Kassel, 
<collectingCountry id="BBCD7633FFBDFFF50FE2D6CBA85148D3" box="[936,1044,404,428]" name="Germany" pageId="7" pageNumber="268">Germany</collectingCountry>
; collected by 
<collectorName id="6E2F5375FFBDFFF508E7D6CAA93C48D2" box="[1197,1401,404,429]" pageId="7" pageNumber="268">Dr. Dieter Sturhan</collectorName>
on 
<collectingDate id="A720E98BFFBDFFF50CDDD6E8AD5C48AF" box="[151,281,439,464]" pageId="7" pageNumber="268" value="1985-05-29">
<date id="B7641063FFBDFFF50CDDD6E8AD5C48AF" box="[151,281,439,464]" pageId="7" pageNumber="268" value="1985-05-29">29.05.1985</date>
</collectingDate>
. GPS Coordinates 
<geoCoordinate id="A6EE5064FFBDFFF50DA6D6E8AE3048B0" box="[492,629,439,464]" direction="north" orientation="latitude" pageId="7" pageNumber="268" precision="15" value="51.31278">51°18'46'' N</geoCoordinate>
and 
<geoCoordinate id="A6EE5064FFBDFFF50EE1D6E8AF6248B0" box="[683,807,439,464]" direction="east" orientation="longitude" pageId="7" pageNumber="268" precision="15" value="9.481388">9°28'53'' E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="C36536A3FFBDFFF50C8DD683AD484B1F" blockId="7.[151,1436,151,680]" pageId="7" pageNumber="268">
<emphasis id="F1AEEAB1FFBDFFF50C8DD683ADCA488A" bold="true" box="[199,399,476,501]" pageId="7" pageNumber="268">Type specimens.</emphasis>
Holotype female on slide 
<taxonomicName id="04DA4D20FFBDFFF50E98D683AF8A488B" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[722,975,476,501]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="268" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFBDFFF50E98D683AF8A488B" box="[722,975,476,501]" italics="true" pageId="7" pageNumber="268">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFBDFFF50F91D683A865488B" box="[987,1056,476,501]" pageId="7" pageNumber="268" rank="species">
<emphasis id="F1AEEAB1FFBDFFF50F91D683A85D488A" bold="true" box="[987,1048,476,501]" pageId="7" pageNumber="268">sp. n</emphasis>
.
</taxonomicNameLabel>
/ 1; paratype females on slides 
<taxonomicName id="04DA4D20FFBDFFF50CDDD6A0ADCA4B67" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[151,399,511,536]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="268" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFBDFFF50CDDD6A0ADCA4B67" box="[151,399,511,536]" italics="true" pageId="7" pageNumber="268">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFBDFFF50DDDD6A0AD914B67" box="[407,468,511,536]" pageId="7" pageNumber="268" rank="species">
<emphasis id="F1AEEAB1FFBDFFF50DDDD6A0AD8A4B67" bold="true" box="[407,463,511,536]" pageId="7" pageNumber="268">sp. n</emphasis>
.
</taxonomicNameLabel>
/ 2–8; deposited with the nematode collection of the Department of Zoology, Aligarh Muslim University, India and on slide 
<taxonomicName id="04DA4D20FFBDFFF50E2ED57BAF254B43" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[612,864,548,573]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="268" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFBDFFF50E2ED57BAF254B43" box="[612,864,548,573]" italics="true" pageId="7" pageNumber="268">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFBDFFF50F26D57BAFE84B43" box="[876,941,548,573]" pageId="7" pageNumber="268" rank="species">
<emphasis id="F1AEEAB1FFBDFFF50F26D57BAFED4B42" bold="true" box="[876,936,548,573]" pageId="7" pageNumber="268">sp. n</emphasis>
.
</taxonomicNameLabel>
/ 9; deposited with the German nematode collection.
</paragraph>
</subSubSection>
<subSubSection id="8BC06528FFBDFFF50C8DD533AEA24BD8" pageId="7" pageNumber="268" type="etymology">
<paragraph id="C36536A3FFBDFFF50C8DD533AEA24BD8" blockId="7.[151,1436,151,680]" pageId="7" pageNumber="268">
<emphasis id="F1AEEAB1FFBDFFF50C8DD533AD0B4BFA" bold="true" box="[199,334,620,645]" pageId="7" pageNumber="268">Etymology.</emphasis>
The new species is named 
<taxonomicName id="04DA4D20FFBDFFF50EDBD532AF764BFB" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[657,819,621,644]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="268" phylum="Nematoda" rank="species" species="enigmatum">
<emphasis id="F1AEEAB1FFBDFFF50EDBD532AF764BFB" box="[657,819,621,644]" italics="true" pageId="7" pageNumber="268">D. enigmatum</emphasis>
</taxonomicName>
because of its unusual asymmetrical odontostyle and flanged odontophore, which is unique for 
<taxonomicName id="04DA4D20FFBDFFF50E27D5D0AEA54BD7" authorityName="Cobb" authorityYear="1920" box="[621,736,655,680]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="268" phylum="Nematoda" rank="genus">
<emphasis id="F1AEEAB1FFBDFFF50E27D5D0AEA54BD7" box="[621,736,655,680]" italics="true" pageId="7" pageNumber="268">Doryllium</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="97A5662BFFBDFFF50CDDD587A8584B90" box="[151,1053,727,751]" pageId="7" pageNumber="268">
<paragraph id="C36536A3FFBDFFF50CDDD587A8584B90" blockId="7.[151,1053,727,751]" box="[151,1053,727,751]" pageId="7" pageNumber="268">
<emphasis id="F1AEEAB1FFBDFFF50CDDD587AD4C4B90" bold="true" box="[151,265,728,751]" pageId="7" pageNumber="268">TABLE 2.</emphasis>
Measurements of 
<taxonomicName id="04DA4D20FFBDFFF50D85D588AEFF4B90" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[463,698,727,751]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="7" pageNumber="268" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFBDFFF50D85D588AEFF4B90" box="[463,698,727,751]" italics="true" pageId="7" pageNumber="268">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFBDFFF50E8BD587AEB84B90" box="[705,765,728,751]" pageId="7" pageNumber="268" rank="species">
<emphasis id="F1AEEAB1FFBDFFF50E8BD587AEB84B90" bold="true" box="[705,765,728,751]" pageId="7" pageNumber="268">sp. n.</emphasis>
</taxonomicNameLabel>
(All measurements in µm)
</paragraph>
</caption>
<paragraph id="C36536A3FFBDFFF50CD5D455A8D64EAB" pageId="7" pageNumber="268">
<table id="B1DAC403FFBD000D0CD5D455A8974EAB" box="[159,1234,778,2004]" gridcols="3" gridrows="30" pageId="7" pageNumber="268">
<tr id="7DEA34E1FFBD000D0CD5D455A8974A5F" box="[159,1234,778,800]" gridrow="0" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D455AD9D4A5F" box="[159,472,778,800]" gridcol="0" gridrow="0" pageId="7" pageNumber="268">Characters</th>
<th id="3E3B5D9DFFBD000D0EACD455AFCA4A5F" box="[742,911,778,800]" gridcol="1" gridrow="0" pageId="7" pageNumber="268">Holotype female</th>
<th id="3E3B5D9DFFBD000D0FA8D455A8974A5F" box="[994,1234,778,800]" gridcol="2" gridrow="0" pageId="7" pageNumber="268">Paratype females</th>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D46BA8974A35" box="[159,1234,820,842]" gridrow="1" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D46BAD9D4A35" box="[159,472,820,842]" gridcol="0" gridrow="1" pageId="7" pageNumber="268">n</th>
<td id="3E3B5D9DFFBD000D0EACD46BAFCA4A35" box="[742,911,820,842]" gridcol="1" gridrow="1" pageId="7" pageNumber="268">1</td>
<td id="3E3B5D9DFFBD000D0FA8D46BA8974A35" box="[994,1234,820,842]" gridcol="2" gridrow="1" pageId="7" pageNumber="268">13</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D402A8974A0C" box="[159,1234,861,883]" gridrow="2" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D402AD9D4A0C" box="[159,472,861,883]" gridcol="0" gridrow="2" pageId="7" pageNumber="268">L</th>
<td id="3E3B5D9DFFBD000D0EACD402AFCA4A0C" box="[742,911,861,883]" gridcol="1" gridrow="2" pageId="7" pageNumber="268">480</td>
<td id="3E3B5D9DFFBD000D0FA8D402A8974A0C" box="[994,1234,861,883]" gridcol="2" gridrow="2" pageId="7" pageNumber="268">495.6 ± 42.2 (409–566)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D4D9A8974AE3" box="[159,1234,902,924]" gridrow="3" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D4D9AD9D4AE3" box="[159,472,902,924]" gridcol="0" gridrow="3" pageId="7" pageNumber="268">Body diameter at neck base</th>
<td id="3E3B5D9DFFBD000D0EACD4D9AFCA4AE3" box="[742,911,902,924]" gridcol="1" gridrow="3" pageId="7" pageNumber="268">17</td>
<td id="3E3B5D9DFFBD000D0FA8D4D9A8974AE3" box="[994,1234,902,924]" gridcol="2" gridrow="3" pageId="7" pageNumber="268">18.3±1.13 (16–20)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D4EFA8974AB9" box="[159,1234,944,966]" gridrow="4" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D4EFAD9D4AB9" box="[159,472,944,966]" gridcol="0" gridrow="4" pageId="7" pageNumber="268">Body diameter at mid body</th>
<td id="3E3B5D9DFFBD000D0EACD4EFAFCA4AB9" box="[742,911,944,966]" gridcol="1" gridrow="4" pageId="7" pageNumber="268">19</td>
<td id="3E3B5D9DFFBD000D0FA8D4EFA8974AB9" box="[994,1234,944,966]" gridcol="2" gridrow="4" pageId="7" pageNumber="268">20.3±1.4 (17–22)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D486A8974A90" box="[159,1234,985,1007]" gridrow="5" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D486AD9D4A90" box="[159,472,985,1007]" gridcol="0" gridrow="5" pageId="7" pageNumber="268">Body diameter at anus</th>
<td id="3E3B5D9DFFBD000D0EACD486AFCA4A90" box="[742,911,985,1007]" gridcol="1" gridrow="5" pageId="7" pageNumber="268">12</td>
<td id="3E3B5D9DFFBD000D0FA8D486A8974A90" box="[994,1234,985,1007]" gridcol="2" gridrow="5" pageId="7" pageNumber="268">13±0.9 (11–14)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D35DA8974D67" box="[159,1234,1026,1048]" gridrow="6" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D35DAD9D4D67" box="[159,472,1026,1048]" gridcol="0" gridrow="6" pageId="7" pageNumber="268">a</th>
<td id="3E3B5D9DFFBD000D0EACD35DAFCA4D67" box="[742,911,1026,1048]" gridcol="1" gridrow="6" pageId="7" pageNumber="268">25.7</td>
<td id="3E3B5D9DFFBD000D0FA8D35DA8974D67" box="[994,1234,1026,1048]" gridcol="2" gridrow="6" pageId="7" pageNumber="268">24.3±1.4 (20.4–26.9)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D373A8974D3D" box="[159,1234,1068,1090]" gridrow="7" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D373AD9D4D3D" box="[159,472,1068,1090]" gridcol="0" gridrow="7" pageId="7" pageNumber="268">b</th>
<td id="3E3B5D9DFFBD000D0EACD373AFCA4D3D" box="[742,911,1068,1090]" gridcol="1" gridrow="7" pageId="7" pageNumber="268">4.1</td>
<td id="3E3B5D9DFFBD000D0FA8D373A8974D3D" box="[994,1234,1068,1090]" gridcol="2" gridrow="7" pageId="7" pageNumber="268">4.1±0.32 (3.3–4.7)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D30AA8974D14" box="[159,1234,1109,1131]" gridrow="8" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D30AAD9D4D14" box="[159,472,1109,1131]" gridcol="0" gridrow="8" pageId="7" pageNumber="268">c</th>
<td id="3E3B5D9DFFBD000D0EACD30AAFCA4D14" box="[742,911,1109,1131]" gridcol="1" gridrow="8" pageId="7" pageNumber="268">32</td>
<td id="3E3B5D9DFFBD000D0FA8D30AA8974D14" box="[994,1234,1109,1131]" gridcol="2" gridrow="8" pageId="7" pageNumber="268">27.8±2.5 (24.0–31.2)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D321A8974DEB" box="[159,1234,1150,1172]" gridrow="9" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D321AD9D4DEB" box="[159,472,1150,1172]" gridcol="0" gridrow="9" pageId="7" pageNumber="268">c`</th>
<td id="3E3B5D9DFFBD000D0EACD321AFCA4DEB" box="[742,911,1150,1172]" gridcol="1" gridrow="9" pageId="7" pageNumber="268">1.2</td>
<td id="3E3B5D9DFFBD000D0FA8D321A8974DEB" box="[994,1234,1150,1172]" gridcol="2" gridrow="9" pageId="7" pageNumber="268">1.3±0.09 (1.2–1.5)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D3F7A8974DC1" box="[159,1234,1192,1214]" gridrow="10" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D3F7AD9D4DC1" box="[159,472,1192,1214]" gridcol="0" gridrow="10" pageId="7" pageNumber="268">V</th>
<td id="3E3B5D9DFFBD000D0EACD3F7AFCA4DC1" box="[742,911,1192,1214]" gridcol="1" gridrow="10" pageId="7" pageNumber="268">43.1</td>
<td id="3E3B5D9DFFBD000D0FA8D3F7A8974DC1" box="[994,1234,1192,1214]" gridcol="2" gridrow="10" pageId="7" pageNumber="268">45.1±3.4(40.2–54.0)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D38EA8974D94" box="[159,1234,1233,1259]" gridrow="11" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D38EAD9D4D94" box="[159,472,1233,1259]" gridcol="0" gridrow="11" pageId="7" pageNumber="268">G1</th>
<td id="3E3B5D9DFFBD000D0EACD38EAFCA4D94" box="[742,911,1233,1259]" gridcol="1" gridrow="11" pageId="7" pageNumber="268">1.6</td>
<td id="3E3B5D9DFFBD000D0FA8D38EA8974D94" box="[994,1234,1233,1259]" gridcol="2" gridrow="11" pageId="7" pageNumber="268">1.6±0.3(0.9–2.2)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D3A2A8974C69" box="[159,1234,1277,1302]" gridrow="12" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D3A2AD9D4C69" box="[159,472,1277,1302]" gridcol="0" gridrow="12" pageId="7" pageNumber="268">G2</th>
<td id="3E3B5D9DFFBD000D0EACD3A2AFCA4C69" box="[742,911,1277,1302]" gridcol="1" gridrow="12" pageId="7" pageNumber="268">18.3</td>
<td id="3E3B5D9DFFBD000D0FA8D3A2A8974C69" box="[994,1234,1277,1302]" gridcol="2" gridrow="12" pageId="7" pageNumber="268">18.6±3.7 (12.8–28.0)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D276A8974C40" box="[159,1234,1321,1343]" gridrow="13" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D276AD9D4C40" box="[159,472,1321,1343]" gridcol="0" gridrow="13" pageId="7" pageNumber="268">Lip region diameter</th>
<td id="3E3B5D9DFFBD000D0EACD276AFCA4C40" box="[742,911,1321,1343]" gridcol="1" gridrow="13" pageId="7" pageNumber="268">5</td>
<td id="3E3B5D9DFFBD000D0FA8D276A8974C40" box="[994,1234,1321,1343]" gridcol="2" gridrow="13" pageId="7" pageNumber="268">6±0 (6–6)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D20DA8974C17" box="[159,1234,1362,1384]" gridrow="14" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D20DAD9D4C17" box="[159,472,1362,1384]" gridcol="0" gridrow="14" pageId="7" pageNumber="268">Lip region height</th>
<td id="3E3B5D9DFFBD000D0EACD20DAFCA4C17" box="[742,911,1362,1384]" gridcol="1" gridrow="14" pageId="7" pageNumber="268">3</td>
<td id="3E3B5D9DFFBD000D0FA8D20DA8974C17" box="[994,1234,1362,1384]" gridcol="2" gridrow="14" pageId="7" pageNumber="268">2.9±0.1 (2.5–3)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D223A8974CED" box="[159,1234,1404,1426]" gridrow="15" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D223AD9D4CED" box="[159,472,1404,1426]" gridcol="0" gridrow="15" pageId="7" pageNumber="268">Amphid aperture</th>
<td id="3E3B5D9DFFBD000D0EACD223AFCA4CED" box="[742,911,1404,1426]" gridcol="1" gridrow="15" pageId="7" pageNumber="268">4</td>
<td id="3E3B5D9DFFBD000D0FA8D223A8974CED" box="[994,1234,1404,1426]" gridcol="2" gridrow="15" pageId="7" pageNumber="268">4.4±0.5 (3.5–5)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D2FAA8974CC4" box="[159,1234,1445,1467]" gridrow="16" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D2FAAD9D4CC4" box="[159,472,1445,1467]" gridcol="0" gridrow="16" pageId="7" pageNumber="268">Odontostyle length</th>
<td id="3E3B5D9DFFBD000D0EACD2FAAFCA4CC4" box="[742,911,1445,1467]" gridcol="1" gridrow="16" pageId="7" pageNumber="268">5</td>
<td id="3E3B5D9DFFBD000D0FA8D2FAA8974CC4" box="[994,1234,1445,1467]" gridcol="2" gridrow="16" pageId="7" pageNumber="268">5.2±0.3 (5–6)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D291A8974C9B" box="[159,1234,1486,1508]" gridrow="17" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D291AD9D4C9B" box="[159,472,1486,1508]" gridcol="0" gridrow="17" pageId="7" pageNumber="268">Odontophore length</th>
<td id="3E3B5D9DFFBD000D0EACD291AFCA4C9B" box="[742,911,1486,1508]" gridcol="1" gridrow="17" pageId="7" pageNumber="268">9</td>
<td id="3E3B5D9DFFBD000D0FA8D291A8974C9B" box="[994,1234,1486,1508]" gridcol="2" gridrow="17" pageId="7" pageNumber="268">9.8±0.6 (9–11)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D2A8A8974F72" box="[159,1234,1527,1549]" gridrow="18" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D2A8AD9D4F72" box="[159,472,1527,1549]" gridcol="0" gridrow="18" pageId="7" pageNumber="268">Guiding ring from anterior end</th>
<td id="3E3B5D9DFFBD000D0EACD2A8AFCA4F72" box="[742,911,1527,1549]" gridcol="1" gridrow="18" pageId="7" pageNumber="268">3</td>
<td id="3E3B5D9DFFBD000D0FA8D2A8A8974F72" box="[994,1234,1527,1549]" gridcol="2" gridrow="18" pageId="7" pageNumber="268">3.6±0.4 (3–4)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D17EA8974F48" box="[159,1234,1569,1591]" gridrow="19" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D17EAD9D4F48" box="[159,472,1569,1591]" gridcol="0" gridrow="19" pageId="7" pageNumber="268">Nerve ring from anterior end</th>
<td id="3E3B5D9DFFBD000D0EACD17EAFCA4F48" box="[742,911,1569,1591]" gridcol="1" gridrow="19" pageId="7" pageNumber="268">64</td>
<td id="3E3B5D9DFFBD000D0FA8D17EA8974F48" box="[994,1234,1569,1591]" gridcol="2" gridrow="19" pageId="7" pageNumber="268">64.2±2.7 (59–70)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D115A8974F1F" box="[159,1234,1610,1632]" gridrow="20" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D115AD9D4F1F" box="[159,472,1610,1632]" gridcol="0" gridrow="20" pageId="7" pageNumber="268">Neck length</th>
<td id="3E3B5D9DFFBD000D0EACD115AFCA4F1F" box="[742,911,1610,1632]" gridcol="1" gridrow="20" pageId="7" pageNumber="268">119</td>
<td id="3E3B5D9DFFBD000D0FA8D115A8974F1F" box="[994,1234,1610,1632]" gridcol="2" gridrow="20" pageId="7" pageNumber="268">118.6±4.1 (110–124)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D12BA8974FF5" box="[159,1234,1652,1674]" gridrow="21" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D12BAD9D4FF5" box="[159,472,1652,1674]" gridcol="0" gridrow="21" pageId="7" pageNumber="268">Expanded part of pharynx</th>
<td id="3E3B5D9DFFBD000D0EACD12BAFCA4FF5" box="[742,911,1652,1674]" gridcol="1" gridrow="21" pageId="7" pageNumber="268">21</td>
<td id="3E3B5D9DFFBD000D0FA8D12BA8974FF5" box="[994,1234,1652,1674]" gridcol="2" gridrow="21" pageId="7" pageNumber="268">19.5±1.08 (18–21)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D1C2A8974FCC" box="[159,1234,1693,1715]" gridrow="22" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D1C2AD9D4FCC" box="[159,472,1693,1715]" gridcol="0" gridrow="22" pageId="7" pageNumber="268">Cardia length</th>
<td id="3E3B5D9DFFBD000D0EACD1C2AFCA4FCC" box="[742,911,1693,1715]" gridcol="1" gridrow="22" pageId="7" pageNumber="268">6</td>
<td id="3E3B5D9DFFBD000D0FA8D1C2A8974FCC" box="[994,1234,1693,1715]" gridcol="2" gridrow="22" pageId="7" pageNumber="268">5.8±0.7 (5–7)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D199A8974FA3" box="[159,1234,1734,1756]" gridrow="23" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D199AD9D4FA3" box="[159,472,1734,1756]" gridcol="0" gridrow="23" pageId="7" pageNumber="268">Anterior genital branch</th>
<td id="3E3B5D9DFFBD000D0EACD199AFCA4FA3" box="[742,911,1734,1756]" gridcol="1" gridrow="23" pageId="7" pageNumber="268">8</td>
<td id="3E3B5D9DFFBD000D0FA8D199A8974FA3" box="[994,1234,1734,1756]" gridcol="2" gridrow="23" pageId="7" pageNumber="268">8±1.4 (5–10)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D1B0A8974E7A" box="[159,1234,1775,1797]" gridrow="24" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D1B0AD9D4E7A" box="[159,472,1775,1797]" gridcol="0" gridrow="24" pageId="7" pageNumber="268">Posterior genital branch</th>
<td id="3E3B5D9DFFBD000D0EACD1B0AFCA4E7A" box="[742,911,1775,1797]" gridcol="1" gridrow="24" pageId="7" pageNumber="268">90</td>
<td id="3E3B5D9DFFBD000D0FA8D1B0A8974E7A" box="[994,1234,1775,1797]" gridcol="2" gridrow="24" pageId="7" pageNumber="268">92.6±21.7 (64–147)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D046A8974E50" box="[159,1234,1817,1839]" gridrow="25" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D046AD9D4E50" box="[159,472,1817,1839]" gridcol="0" gridrow="25" pageId="7" pageNumber="268">Vaginal length</th>
<td id="3E3B5D9DFFBD000D0EACD046AFCA4E50" box="[742,911,1817,1839]" gridcol="1" gridrow="25" pageId="7" pageNumber="268">10</td>
<td id="3E3B5D9DFFBD000D0FA8D046A8974E50" box="[994,1234,1817,1839]" gridcol="2" gridrow="25" pageId="7" pageNumber="268">9.1±0.5 (8–10)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D01DA8974E27" box="[159,1234,1858,1880]" gridrow="26" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D01DAD9D4E27" box="[159,472,1858,1880]" gridcol="0" gridrow="26" pageId="7" pageNumber="268">Vulva from anterior end</th>
<td id="3E3B5D9DFFBD000D0EACD01DAFCA4E27" box="[742,911,1858,1880]" gridcol="1" gridrow="26" pageId="7" pageNumber="268">207</td>
<td id="3E3B5D9DFFBD000D0FA8D01DA8974E27" box="[994,1234,1858,1880]" gridcol="2" gridrow="26" pageId="7" pageNumber="268">222.6±119 (197–239)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D033A8974EFD" box="[159,1234,1900,1922]" gridrow="27" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D033AD9D4EFD" box="[159,472,1900,1922]" gridcol="0" gridrow="27" pageId="7" pageNumber="268">Prerectum length</th>
<td id="3E3B5D9DFFBD000D0EACD033AFCA4EFD" box="[742,911,1900,1922]" gridcol="1" gridrow="27" pageId="7" pageNumber="268">30</td>
<td id="3E3B5D9DFFBD000D0FA8D033A8974EFD" box="[994,1234,1900,1922]" gridcol="2" gridrow="27" pageId="7" pageNumber="268">24±3.7 (20–31)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D0CAA8974ED4" box="[159,1234,1941,1963]" gridrow="28" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D0CAAD9D4ED4" box="[159,472,1941,1963]" gridcol="0" gridrow="28" pageId="7" pageNumber="268">Rectum length</th>
<td id="3E3B5D9DFFBD000D0EACD0CAAFCA4ED4" box="[742,911,1941,1963]" gridcol="1" gridrow="28" pageId="7" pageNumber="268">12</td>
<td id="3E3B5D9DFFBD000D0FA8D0CAA8974ED4" box="[994,1234,1941,1963]" gridcol="2" gridrow="28" pageId="7" pageNumber="268">11.4±1.7 (9–15)</td>
</tr>
<tr id="7DEA34E1FFBD000D0CD5D0E1A8974EAB" box="[159,1234,1982,2004]" gridrow="29" pageId="7" pageNumber="268">
<th id="3E3B5D9DFFBD000D0CD5D0E1AD9D4EAB" box="[159,472,1982,2004]" gridcol="0" gridrow="29" pageId="7" pageNumber="268">Tail length</th>
<td id="3E3B5D9DFFBD000D0EACD0E1AFCA4EAB" box="[742,911,1982,2004]" gridcol="1" gridrow="29" pageId="7" pageNumber="268">15</td>
<td id="3E3B5D9DFFBD000D0FA8D0E1A8974EAB" box="[994,1234,1982,2004]" gridcol="2" gridrow="29" pageId="7" pageNumber="268">18.0±1.8 (15–21)</td>
</tr>
</table>
</paragraph>
<caption id="97A5662BFFB2FFFA0CDDD1F9A80C4FA5" httpUri="https://zenodo.org/record/1301833/files/figure.png" pageId="8" pageNumber="269" startId="8.[151,250,1702,1724]" targetBox="[252,1346,245,1561]" targetPageId="8">
<paragraph id="C36536A3FFB2FFFA0CDDD1F9A80C4FA5" blockId="8.[151,1436,1701,1754]" pageId="8" pageNumber="269">
<emphasis id="F1AEEAB1FFB2FFFA0CDDD1F9AE5C4FC4" bold="true" box="[151,537,1701,1724]" pageId="8" pageNumber="269">
FIGURE 4. 
<taxonomicName id="04DA4D20FFB2FFFA0D6ED1FAAE5C4FC4" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[292,537,1701,1724]" genus="Dorylllium" pageId="8" pageNumber="269" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFB2FFFA0D6ED1FAAE5C4FC4" bold="true" box="[292,537,1701,1724]" italics="true" pageId="8" pageNumber="269">Dorylllium enigmatum</emphasis>
</taxonomicName>
</emphasis>
<taxonomicNameLabel id="EA9D57CAFFB2FFFA0E6ED1F9AE274FC3" box="[548,610,1702,1724]" pageId="8" pageNumber="269" rank="species">
<emphasis id="F1AEEAB1FFB2FFFA0E6ED1F9AE194FC4" bold="true" box="[548,604,1702,1723]" pageId="8" pageNumber="269">sp. n</emphasis>
.
</taxonomicNameLabel>
A. Entire female; B. Anterior region; C. Anterior end showing amphid; D. Pharyngeal region; E. Pharyngeal bulb; F. Female genital system; G. Female posterior region.
</paragraph>
</caption>
<subSubSection id="8BC06528FFB2FFFB0C8DD05DAECD4863" lastPageId="9" lastPageNumber="270" pageId="8" pageNumber="269" type="diagnosis">
<paragraph id="C36536A3FFB2FFFA0C8DD05DAFAA4ED4" blockId="8.[151,1437,1794,2035]" pageId="8" pageNumber="269">
<emphasis id="F1AEEAB1FFB2FFFA0C8DD05DAE504E64" bold="true" box="[199,533,1794,1819]" pageId="8" pageNumber="269">Diagnosis and relationships.</emphasis>
<taxonomicName id="04DA4D20FFB2FFFA0E56D05DAF574E65" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[540,786,1794,1819]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="8" pageNumber="269" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFB2FFFA0E56D05DAF574E65" box="[540,786,1794,1819]" italics="true" pageId="8" pageNumber="269">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFB2FFFA0F52D05DAF124E65" box="[792,855,1794,1819]" pageId="8" pageNumber="269" rank="species">
<emphasis id="F1AEEAB1FFB2FFFA0F52D05DAF0A4E64" bold="true" box="[792,847,1794,1819]" pageId="8" pageNumber="269">sp. n</emphasis>
.
</taxonomicNameLabel>
is characterized by a body length of 
<quantity id="04229B46FFB2FFFA08B8D05DA9DD4E65" box="[1266,1432,1794,1819]" metricMagnitude="-4" metricUnit="m" metricValue="4.6" metricValueMax="5.2" metricValueMin="4.0" pageId="8" pageNumber="269" unit="mm" value="0.46" valueMax="0.52" valueMin="0.4">0.40–0.52 mm</quantity>
; continuous lip region; rounded lips; asymmetrical odontostyle, 5–6 µm long, flanged odontophore 9–11 µm long (total spear length: 14–17 µm); pharynx with slender anterior part, which expands into a slightly constricted pyriform basal bulb occupying about 16–18% of total neck length; mono-opisthodelphic female genital system, anterior uterine sac 7–9 µm; transverse vulva and rounded-hemispheroidtail.
</paragraph>
<paragraph id="C36536A3FFB2FFFA0C8DD0EAACAA4E8C" blockId="8.[151,1437,1794,2035]" pageId="8" pageNumber="269">
The new species is distinctive because of its asymmetrical odontostyle which is unusual for a 
<emphasis id="F1AEEAB1FFB2FFFA097ED0EAA9D94EB1" box="[1332,1436,1973,1998]" italics="true" pageId="8" pageNumber="269">Doyllium</emphasis>
species.
</paragraph>
<paragraph id="C36536A3FFB3FFFB0C8DD7C7AECD4863" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="270">
In the shape of its lip region and nature of pharyngeal expansion, the new species comes close to 
<taxonomicName id="04DA4D20FFB3FFFB095ED7C6AD5E49AB" authority="Cobb, 1920" authorityName="Cobb" authorityYear="1920" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="270" phylum="Nematoda" rank="species" species="uniforme">
<emphasis id="F1AEEAB1FFB3FFFB095ED7C6A9D949CF" box="[1300,1436,152,176]" italics="true" pageId="9" pageNumber="270">D. uniforme</emphasis>
<bibRefCitation id="A74B4B52FFB3FFFB0CDDD7E3AD5E49AB" box="[151,283,188,213]" pageId="9" pageNumber="270" refId="ref4627" refString="Cobb, N. A. (1920) One hundred new nemas (Type species of 100 new genera). Contribution to Science of Nematology, 9, 217 - 343." type="journal article">Cobb, 1920</bibRefCitation>
</taxonomicName>
but differs in its smaller body size (
<emphasis id="F1AEEAB1FFB3FFFB0EF2D7E2AE8A49AB" box="[696,719,189,212]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
L = 
<quantity id="04229B46FFB3FFFB0F43D7E3AFF749AB" box="[777,946,188,213]" metricMagnitude="-4" metricUnit="m" metricValue="8.149999999999999" metricValueMax="8.5" metricValueMin="7.8" pageId="9" pageNumber="270" unit="mm" value="0.815" valueMax="0.85" valueMin="0.78">0.78–0.85 mm</quantity>
); more posterior vulva position (
<emphasis id="F1AEEAB1FFB3FFFB096CD7E2A97849AB" box="[1318,1341,189,212]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
V= 33– 35); very short anterior uterine sac (
<emphasis id="F1AEEAB1FFB3FFFB0E61D7BEAE074987" box="[555,578,225,248]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
anterior uterine branch 32 µm); comparatively shorter tail (c= 24–32 vs 38– 41) and in the absence of males (
<emphasis id="F1AEEAB1FFB3FFFB0E42D65AAE5A4863" box="[520,543,261,284]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
present).
</paragraph>
</subSubSection>
<subSubSection id="8BC06528FFB3FFFB0C8DD676A9034EF4" pageId="9" pageNumber="270" type="reference_group">
<paragraph id="C36536A3FFB3FFFB0C8DD676AF4348F7" blockId="9.[151,1436,151,392]" pageId="9" pageNumber="270">
<taxonomicName id="04DA4D20FFB3FFFB0C8DD676AD9B483F" authority="Timm, 1967" authorityName="Timm" authorityYear="1967" box="[199,478,295,320]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="270" phylum="Nematoda" rank="species" species="aestuarii">
<emphasis id="F1AEEAB1FFB3FFFB0C8DD676AD08483F" box="[199,333,297,320]" italics="true" pageId="9" pageNumber="270">D. aestuarii</emphasis>
<bibRefCitation id="A74B4B52FFB3FFFB0D1ED678AD9B483F" box="[340,478,295,320]" pageId="9" pageNumber="270" refId="ref5474" refString="Timm, R. W. (1967) Some estuarine nematodes from the Sunderbans. Proceedings of the Pakistan Academy of Science s, 4, 1 - 13." type="journal article">Timm, 1967</bibRefCitation>
</taxonomicName>
also has a continuous lip region and constricted pharyngeal bulb but the new species distinctly differs from it in being smaller in size (
<emphasis id="F1AEEAB1FFB3FFFB0E8AD612AE92481B" box="[704,727,333,356]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
L= 
<quantity id="04229B46FFB3FFFB0F4CD613AFEB481B" box="[774,942,332,357]" metricMagnitude="-3" metricUnit="m" metricValue="1.045" metricValueMax="1.13" metricValueMin="0.96" pageId="9" pageNumber="270" unit="mm" value="1.045" valueMax="1.13" valueMin="0.96">0.96–1.13 mm</quantity>
); having a more posterior vulva position (
<emphasis id="F1AEEAB1FFB3FFFB09CED612A9DE481B" box="[1412,1435,333,356]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
V= 23.7–24.7) and in tail shape (
<emphasis id="F1AEEAB1FFB3FFFB0E43D62EAE6548F7" box="[521,544,369,392]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
tail bluntly conoid).
</paragraph>
<caption id="97A5662BFFB3FFFB0CDDD161AD3E4FEE" httpUri="https://zenodo.org/record/1301835/files/figure.png" pageId="9" pageNumber="270" startId="9.[151,250,1598,1620]" targetBox="[381,1219,440,1577]" targetPageId="9">
<paragraph id="C36536A3FFB3FFFB0CDDD161AD3E4FEE" blockId="9.[151,1436,1597,1681]" pageId="9" pageNumber="270">
<emphasis id="F1AEEAB1FFB3FFFB0CDDD161AE164F2C" bold="true" box="[151,595,1597,1620]" pageId="9" pageNumber="270">
FIGURE 5. 
<taxonomicName id="04DA4D20FFB3FFFB0D6BD162AE494F2C" authority="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan, 2018" authorityName="Ahmad &amp; Ahad &amp; Islam &amp; Sturhan" authorityYear="2018" box="[289,524,1597,1620]" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="270" phylum="Nematoda" rank="species" species="enigmatum" status="sp. nov.">
<emphasis id="F1AEEAB1FFB3FFFB0D6BD162AE494F2C" bold="true" box="[289,524,1597,1620]" italics="true" pageId="9" pageNumber="270">Doryllium enigmatum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EA9D57CAFFB3FFFB0E5CD161AE164F2C" box="[534,595,1598,1619]" pageId="9" pageNumber="270" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
A. Anterior region; B. Anterior region showing basal flanges; C. Anterior end showing amphid; D. &amp; E. Pharyngeal bulb; F. Female genital system; G, H, I. Female posterior region. (Scale bar A–E, H, I= 10 µm; F, G = 20 µm).
</paragraph>
</caption>
<paragraph id="C36536A3FFB3FFFB0C8DD1E0A9034EF4" blockId="9.[151,1436,1725,1931]" pageId="9" pageNumber="270">
In its body size, lip region shape and nature of the anterior uterine sac, the new species is very close to 
<taxonomicName id="04DA4D20FFB3FFFB09CBD1E0AE7F4F84" authority="Goseco, Ferris &amp; Ferris, 1975" authorityName="Goseco, Ferris &amp; Ferris" authorityYear="1975" class="Adenophorea" family="Leptonchidae" genus="Gerthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="270" phylum="Nematoda" rank="species" species="jamesi">
<emphasis id="F1AEEAB1FFB3FFFB09CBD1E0ACA54F85" italics="true" pageId="9" pageNumber="270">D. jamesi</emphasis>
<bibRefCitation id="A74B4B52FFB3FFFB0CADD1BDAE7F4F84" box="[231,570,1762,1787]" pageId="9" pageNumber="270" refId="ref4661" refString="Goseco, C. G., Ferris, V. R &amp; Ferris, J. M. (1975) Revision in Leptonchoidea (Nematoda: Dorylaimida). Tylencholaimellus, Doryllium, Gerthus n. gen. and Agmodorus in Tylencholaimellidae; and Aulolaimoies and Adenolaimus in Aulolaimoididae. Research Bulletin, Agricultural Experiment Station, Purdue University, West Lafayette, 928, 1 - 40." type="journal article">Goseco, Ferris &amp; Ferris, 1975</bibRefCitation>
</taxonomicName>
but differs in having an asymmetrical odontostyle (
<emphasis id="F1AEEAB1FFB3FFFB08CFD1BCA8D94F85" box="[1157,1180,1763,1786]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
symmetrical); slightly constricted pharyngeal bulb (
<emphasis id="F1AEEAB1FFB3FFFB0D97D058ADB14E61" box="[477,500,1799,1822]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
not constricted) and longer tail (
<emphasis id="F1AEEAB1FFB3FFFB0F28D058AF3C4E61" box="[866,889,1799,1822]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
tail 7–9 µm). The new species is also close to 
<taxonomicName id="04DA4D20FFB3FFFB09CBD058ADDA4E3C" authority="Thorne, 1964" authorityName="Thorne" authorityYear="1964" class="Adenophorea" family="Leptonchidae" genus="Doryllium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Dorylaimida" pageId="9" pageNumber="270" phylum="Nematoda" rank="species" species="flangum">
<emphasis id="F1AEEAB1FFB3FFFB09CBD058ACB64E3C" italics="true" pageId="9" pageNumber="270">D. flangum</emphasis>
<bibRefCitation id="A74B4B52FFB3FFFB0D4BD075ADDA4E3C" box="[257,415,1834,1859]" pageId="9" pageNumber="270" refId="ref5424" refString="Thorne, G. (1964) Nematodes of Puerto Rico: Belondiroidea, new superfamily, Leptonchidae Thorne, 1935, and Belonenchidae new family (Nematoda, Adenophorea, Dorylaimida). University Puerto Rico Agriculture Experimental Station Technical Paper, No 39, 1 - 51." type="journal article">Thorne, 1964</bibRefCitation>
</taxonomicName>
but differs in its asymmetrical odontostyle and shorter odontophore (
<emphasis id="F1AEEAB1FFB3FFFB08B9D074A94F4E3D" box="[1267,1290,1835,1858]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
odontostyle symmetrical and odontophore three times odontostyle length); slightly constricted pharyngeal bulb (
<emphasis id="F1AEEAB1FFB3FFFB0918D010A92C4E19" box="[1362,1385,1871,1894]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
not constricted) and in the presence of the short anterior uterine sac (
<emphasis id="F1AEEAB1FFB3FFFB0F24D02CAFC04EF5" box="[878,901,1907,1930]" italics="true" pageId="9" pageNumber="270">vs</emphasis>
anterior uterine sac completely absent).
</paragraph>
</subSubSection>
</treatment>
</document>