treatments-xml/data/D6/28/30/D628304F9621FFB8FF2C947BFDA9FD4F.xml
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<document id="C4119F1772019F59BB0AC1CF43824DEC" ID-CLB-Dataset="199057" ID-DOI="10.11646/zootaxa.5213.5.7" ID-GBIF-Dataset="e6d1800f-edfe-4599-9ba9-610594170389" ID-ISSN="1175-5326" ID-Zenodo-Dep="7381759" ID-ZooBank="918E8E7E-A6EC-4935-8693-A80052FF9199" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669811020408" checkinUser="plazi" docAuthor="Brown, John W." docDate="2022" docId="D628304F9621FFB8FF2C947BFDA9FD4F" docLanguage="en" docName="zootaxa.5213.5.7.pdf" docOrigin="Zootaxa 5213 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thalleulia bisexta Brown 2022, new species" docType="treatment" docUuid="0ACDDA61-B36C-4602-9D73-A7E79FC23663" docUuidSource="ZooBank" docVersion="7" lastPageNumber="590" masterDocId="2A1148379620FFBBFFBB9170FFFFFFCA" masterDocTitle="A new species of Thalleulia Razowski, 2004 from Costa Rica with the first reported female of the genus (Lepidoptera: Tortricidae: Euliina)" masterLastPageNumber="590" masterPageNumber="587" pageNumber="588" updateTime="1699412608208" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="86241D3D1146A7E6358FB616E9858FDC">A new species of Thalleulia Razowski, 2004 from Costa Rica with the first reported female of the genus (Lepidoptera: Tortricidae: Euliina)</mods:title>
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<mods:date id="69C91B1A645C95E0231F13EE039DFC34">2022</mods:date>
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<taxonomicName id="9981FADA9621FFBAFF2C947BFE31FAEC" authority="Brown" authorityName="Brown" authorityYear="2022" box="[151,462,1291,1318]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="bisexta" status="sp. nov.">
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Brown
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,
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,
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<elevation id="D5AC666A9621FFBAFF2C973BFF12F9AF" box="[151,237,1611,1637]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="1" pageNumber="588" unit="m" value="2600.0">2600 m</elevation>
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,
<date id="2A3FA7999621FFBAFF43973BFE95F9AF" box="[248,362,1611,1637]" pageId="1" pageNumber="588" value="2001-05">
<collectingDate id="3A7B5E719621FFBAFF43973BFE95F9AF" box="[248,362,1611,1637]" pageId="1" pageNumber="588" value="2001-05">May 2001</collectingDate>
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.
<collectorName id="F374E48F9621FFBAFE69973BFDCEF9AF" box="[466,561,1611,1637]" pageId="1" pageNumber="588">Delgado</collectorName>
(
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)
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<collectorName id="F374E48F9621FFBAFD7C973BFCC4F9AF" box="[711,827,1611,1637]" pageId="1" pageNumber="588">El Guarco</collectorName>
,
<location id="5B5ED7829621FFBAFCFE973BFC40F9AF" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFCFE973BFC40F9AF" box="[837,959,1611,1637]" country="Costa Rica" name="Villa Mills" pageId="1" pageNumber="588" stateProvince="Cartago">Villa Mills</location>
, CATIE,
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<collectingDate id="3A7B5E719621FFBAFB3C973BFAC5F9AF" box="[1159,1338,1611,1637]" pageId="1" pageNumber="588" value="2000-10-26" valueMax="2000-10-28" valueMin="2000-10-26">2628 Oct 2000</collectingDate>
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(
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),
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.
<collectorName id="F374E48F9621FFBAFF2C971FFF09F943" box="[151,246,1647,1673]" pageId="1" pageNumber="588">Delgado</collectorName>
(
<collectionCode id="3890199C9621FFBAFEBE971FFE83F943" box="[261,380,1647,1673]" pageId="1" pageNumber="588">MNCR-A</collectionCode>
)
</materialsCitation>
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<materialsCitation id="EEE98B049621FFBAFE34971FFCF2F967" ID-GBIF-Occurrence="3979530302" collectingDate="2005-04-05" collectingDateMax="2005-04-13" collectingDateMin="2005-04-05" collectionCode="MNCR-A" collectorName="K. Nishida" country="Costa Rica" elevation="1500" location="Finca Murrillo" pageId="1" pageNumber="588" specimenCount="2" specimenCount-male="2" stateProvince="Heredia" typeStatus="paratype">
<collectingRegion id="9C454FBB9621FFBAFE34971FFE13F943" box="[399,492,1647,1673]" country="Costa Rica" name="Heredia" pageId="1" pageNumber="588">Heredia</collectingRegion>
:
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NE Vara Blanca
</locationDeviation>
,
<location id="5B5ED7829621FFBAFD40971FFC5DF943" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFD40971FFC5DF943" box="[763,930,1647,1673]" country="Costa Rica" name="Finca Murrillo" pageId="1" pageNumber="588" stateProvince="Heredia">Finca Murrillo</location>
,
<quantity id="99792CBC9621FFBAFC15971EFBFAF943" box="[942,1029,1646,1673]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="m" value="1500.0">
<elevation id="D5AC666A9621FFBAFC15971EFBFAF943" box="[942,1029,1646,1673]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="2A3FA7999621FFBAFBAA971FFB6FF943" box="[1041,1168,1646,1673]" pageId="1" pageNumber="588" value="2005-04-08">
<collectingDate id="3A7B5E719621FFBAFBAA971FFB6FF943" box="[1041,1168,1646,1673]" pageId="1" pageNumber="588" value="2005-04-08">8 Apr 2005</collectingDate>
</date>
(
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),
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,
<date id="2A3FA7999621FFBAFADF971EFEFDF967" pageId="1" pageNumber="588" value="2005-04-05" valueMax="2005-04-13" valueMin="2005-04-05">
<collectingDate id="3A7B5E719621FFBAFADF971EFEFDF967" pageId="1" pageNumber="588" value="2005-04-05" valueMax="2005-04-13" valueMin="2005-04-05">513 Apr 2005</collectingDate>
</date>
(
<specimenCount id="48874AD09621FFBAFEA997E3FEC9F967" box="[274,310,1683,1709]" pageId="1" pageNumber="588" type="male">1♂</specimenCount>
), INBio-OET-ALAS transect (
<collectionCode id="3890199C9621FFBAFD3597E3FCFAF967" box="[654,773,1683,1709]" pageId="1" pageNumber="588">MNCR-A</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EEE98B049621FFBAFCAC97E3FCDDF91B" ID-GBIF-Occurrence="3979530306" collectingDate="2006-07-31" collectionCode="USNM" collectorName="J. B. Sullivan" country="Costa Rica" county="Calle Ziriqui" elevation="1500" location="St. Elena" municipality="San Ysidro" pageId="1" pageNumber="588" specimenCount="1" specimenCount-male="1" stateProvince="Heredia" typeStatus="paratype">
<collectingCounty id="B75FF9D59621FFBAFCAC97E3FC55F967" box="[791,938,1683,1709]" pageId="1" pageNumber="588">Calle Ziriqui</collectingCounty>
,
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<quantity id="99792CBC9621FFBAFC0E97E2FBFCF967" box="[949,1027,1682,1709]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="km" value="1.5">1.5 km</quantity>
E Porton Municipal
</locationDeviation>
,
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<location id="5B5ED7829621FFBAFAC597E3FF25F91B" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFAC597E3FF25F91B" country="Costa Rica" county="Calle Ziriqui" municipality="San Ysidro" name="St. Elena" pageId="1" pageNumber="588" stateProvince="Heredia">St. Elena</location>
,
<quantity id="99792CBC9621FFBAFF5C97C6FEC0F91B" box="[231,319,1718,1745]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="m" value="1500.0">
<elevation id="D5AC666A9621FFBAFF5C97C6FEC0F91B" box="[231,319,1718,1745]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="2A3FA7999621FFBAFEF697C7FE2DF91B" box="[333,466,1719,1745]" pageId="1" pageNumber="588" value="2006-07-31">
<collectingDate id="3A7B5E719621FFBAFEF697C7FE2DF91B" box="[333,466,1719,1745]" pageId="1" pageNumber="588" value="2006-07-31">31 Jul 2006</collectingDate>
</date>
(
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),
<collectorName id="F374E48F9621FFBAFD9B97C7FD4EF91B" box="[544,689,1719,1745]" pageId="1" pageNumber="588">J.B. Sullivan</collectorName>
(
<collectionCode id="3890199C9621FFBAFD7997C7FCE6F91B" box="[706,793,1719,1745]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="588" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EEE98B049621FFBAFC9697C7FD9EF93F" ID-GBIF-Occurrence="3979530304" collectingDate="1965-03-19" collectingDateMax="1965-03-20" collectingDateMin="1965-03-19" collectionCode="USNM" collectorName="S. S. &amp; W. D. Duckworth" country="Costa Rica" location="San Vito" municipality="Las Cruces" pageId="1" pageNumber="588" specimenCount="2" specimenCount-female="2" stateProvince="Puntarenas" typeStatus="paratype">
<collectingRegion id="9C454FBB9621FFBAFC9697C7FC53F91B" box="[813,940,1719,1745]" country="Costa Rica" name="Puntarenas" pageId="1" pageNumber="588">Puntarenas</collectingRegion>
:
<collectingMunicipality id="BE5A1B239621FFBAFC0297C7FBC4F91B" box="[953,1083,1719,1745]" pageId="1" pageNumber="588">Las Cruces</collectingMunicipality>
, nr.
<location id="5B5ED7829621FFBAFBD797C7FB2DF91B" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFBD797C7FB2DF91B" box="[1132,1234,1719,1745]" country="Costa Rica" municipality="Las Cruces" name="San Vito" pageId="1" pageNumber="588" stateProvince="Puntarenas">San Vito</location>
,
<date id="2A3FA7999621FFBAFB6597C7FA63F91B" box="[1246,1436,1718,1745]" pageId="1" pageNumber="588" value="1965-03-19" valueMax="1965-03-20" valueMin="1965-03-19">
<collectingDate id="3A7B5E719621FFBAFB6597C7FA63F91B" box="[1246,1436,1718,1745]" pageId="1" pageNumber="588" value="1965-03-19" valueMax="1965-03-20" valueMin="1965-03-19">1920 Mar 1965</collectingDate>
</date>
(
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),
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(
<collectionCode id="3890199C9621FFBAFDB997ABFDA6F93F" box="[514,601,1755,1781]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="588" type="Museum">USNM</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EEE98B049621FFBAFDD797ABFBF6F8D3" ID-GBIF-Occurrence="3979530303" collectingDate="1999-03-15" collectingDateMax="1999-03-24" collectingDateMin="1999-03-15" collectionCode="R, MNCR-A" collectorName="Barton &amp; Rodriguez &amp; Delgado" country="Costa Rica" elevation="1500" location="Est. Biol. Las Alturas" pageId="1" pageNumber="588" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Puntarenas" typeStatus="paratype">
A.C.L.
<collectorName id="F374E48F9621FFBAFD0397ABFD15F93F" box="[696,746,1755,1781]" name="J.A. Powell" pageId="1" pageNumber="588">A.P.</collectorName>
,
<collectorName id="F374E48F9621FFBAFD4397ABFBD2F93F" box="[760,1069,1755,1781]" pageId="1" pageNumber="588">Coto Brus, Z.P. Las Tablas</collectorName>
,
<location id="5B5ED7829621FFBAFB8297ABFACEF93F" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFB8297ABFACEF93F" box="[1081,1329,1755,1781]" country="Costa Rica" name="Est. Biol. Las Alturas" pageId="1" pageNumber="588" stateProvince="Puntarenas">Est. Biol. Las Alturas</location>
,
<quantity id="99792CBC9621FFBAFA8497AAFA68F93F" box="[1343,1431,1754,1781]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="m" value="1500.0">
<elevation id="D5AC666A9621FFBAFA8497AAFA68F93F" box="[1343,1431,1754,1781]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="1" pageNumber="588" unit="m" value="1500.0">1500 m</elevation>
</quantity>
,
<date id="2A3FA7999621FFBAFF2C978EFEAEF8D3" box="[151,337,1790,1817]" pageId="1" pageNumber="588" value="1999-03-15" valueMax="1999-03-24" valueMin="1999-03-15">
<collectingDate id="3A7B5E719621FFBAFF2C978EFEAEF8D3" box="[151,337,1790,1817]" pageId="1" pageNumber="588" value="1999-03-15" valueMax="1999-03-24" valueMin="1999-03-15">1524 Mar 1999</collectingDate>
</date>
(
<specimenCount id="48874AD09621FFBAFEDB978FFE79F8D3" box="[352,390,1791,1817]" pageId="1" pageNumber="588" type="male">2♂</specimenCount>
,
<specimenCount id="48874AD09621FFBAFE2A978FFE4DF8D3" box="[401,434,1791,1817]" pageId="1" pageNumber="588" type="female">1♀</specimenCount>
),
<collectionCode id="3890199C9621FFBAFE70978FFE1EF8D3" box="[459,481,1791,1817]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="588">R</collectionCode>
.
<collectorName id="F374E48F9621FFBAFE50978FFDC5F8D3" box="[491,570,1791,1817]" pageId="1" pageNumber="588">Barton</collectorName>
, G.
<collectorName id="F374E48F9621FFBAFDDD978FFD24F8D3" box="[614,731,1791,1817]" pageId="1" pageNumber="588">Rodriguez</collectorName>
&amp;
<collectionCode id="3890199C9621FFBAFD46978FFCECF8D3" box="[765,787,1791,1817]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="1" pageNumber="588">R</collectionCode>
.
<collectorName id="F374E48F9621FFBAFCA6978FFC83F8D3" box="[797,892,1791,1817]" pageId="1" pageNumber="588">Delgado</collectorName>
(
<collectionCode id="3890199C9621FFBAFC31978FFBFEF8D3" box="[906,1025,1791,1817]" pageId="1" pageNumber="588">MNCR-A</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EEE98B049621FFBAFBA8978FFE4CF8F7" ID-GBIF-Occurrence="3979530301" collectingDate="1979-06-25" collectingDateMax="1979-06-28" collectingDateMin="1979-06-25" collectionCode="MNCR-A" collectorName="D. H. Janzen" country="Costa Rica" location="Monteverde" pageId="1" pageNumber="588" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas" typeStatus="paratype">
<location id="5B5ED7829621FFBAFBA8978FFB62F8D3" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFBA8978FFB62F8D3" box="[1043,1181,1791,1817]" country="Costa Rica" name="Monteverde" pageId="1" pageNumber="588" stateProvince="Puntarenas">Monteverde</location>
,
<date id="2A3FA7999621FFBAFB1C978EFAA6F8D3" box="[1191,1369,1790,1817]" pageId="1" pageNumber="588" value="1979-06-25" valueMax="1979-06-28" valueMin="1979-06-25">
<collectingDate id="3A7B5E719621FFBAFB1C978EFAA6F8D3" box="[1191,1369,1790,1817]" pageId="1" pageNumber="588" value="1979-06-25" valueMax="1979-06-28" valueMin="1979-06-25">2528 Jun 1979</collectingDate>
</date>
(
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),
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(
<collectionCode id="3890199C9621FFBAFE8F9653FE54F8F7" box="[308,427,1827,1853]" pageId="1" pageNumber="588">MNCR-A</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="EEE98B049621FFBAFE079653FD86F8AB" ID-GBIF-Occurrence="3979530311" collectingDate="1996-04-19" collectingDateMax="1996-04-20" collectingDateMin="1996-04-19" collectionCode="NMCR-A" collectorName="A. Picado &amp; B. Gamboa" country="Costa Rica" county="Est. Cuerci" elevation="2640" location="Mirador" municipality="Sendero" pageId="1" pageNumber="588" specimenCount="2" specimenCount-male="2" stateProvince="San Jose" typeStatus="paratype">
<collectingRegion id="9C454FBB9621FFBAFE079653FDDEF8F7" box="[444,545,1827,1853]" country="Costa Rica" name="San Jose" pageId="1" pageNumber="588">San José</collectingRegion>
:
<collectingCounty id="B75FF9D59621FFBAFD909653FD55F8F7" box="[555,682,1827,1853]" pageId="1" pageNumber="588">Est. Cuerci</collectingCounty>
,
<collectingMunicipality id="BE5A1B239621FFBAFD0F9653FCEFF8F7" box="[692,784,1827,1853]" pageId="1" pageNumber="588">Sendero</collectingMunicipality>
al
<location id="5B5ED7829621FFBAFC8B9653FC71F8F7" LSID="urn:lsid:plazi:treatment:D628304F9621FFB8FF2C947BFDA9FD4F:5B5ED7829621FFBAFC8B9653FC71F8F7" box="[816,910,1827,1853]" country="Costa Rica" county="Est. Cuerci" municipality="Sendero" name="Mirador" pageId="1" pageNumber="588" stateProvince="San Jose">Mirador</location>
,
<locationDeviation id="4F8C987B9621FFBAFC239653FB83F8F7" box="[920,1148,1827,1853]" location="Villa Mills" pageId="1" pageNumber="588">
<quantity id="99792CBC9621FFBAFC239653FC1AF8F7" box="[920,997,1827,1853]" metricMagnitude="3" metricUnit="m" metricValue="4.6" pageId="1" pageNumber="588" unit="km" value="4.6">4.6 km</quantity>
E Villa Mills
</locationDeviation>
,
<quantity id="99792CBC9621FFBAFB3C9653FB22F8F7" box="[1159,1245,1827,1853]" metricMagnitude="3" metricUnit="m" metricValue="2.6399999999999997" pageId="1" pageNumber="588" unit="m" value="2640.0">
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</quantity>
,
<date id="2A3FA7999621FFBAFB539653FA63F8F7" box="[1256,1436,1827,1853]" pageId="1" pageNumber="588" value="1996-04-19" valueMax="1996-04-20" valueMin="1996-04-19">
<collectingDate id="3A7B5E719621FFBAFB539653FA63F8F7" box="[1256,1436,1827,1853]" pageId="1" pageNumber="588" value="1996-04-19" valueMax="1996-04-20" valueMin="1996-04-19">1920 Apr 1996</collectingDate>
</date>
(
<specimenCount id="48874AD09621FFBAFF1B9637FF39F8AB" box="[160,198,1863,1889]" pageId="1" pageNumber="588" type="male">2♂</specimenCount>
),
<collectorName id="F374E48F9621FFBAFF629637FEB7F8AB" box="[217,328,1863,1889]" pageId="1" pageNumber="588">A. Picado</collectorName>
&amp;
<collectorName id="F374E48F9621FFBAFED79637FE14F8AB" box="[364,491,1863,1889]" pageId="1" pageNumber="588">B. Gamboa</collectorName>
(
<collectionCode id="3890199C9621FFBAFE419637FD8EF8AB" box="[506,625,1863,1889]" pageId="1" pageNumber="588">NMCR-A</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="169BD2D29621FFB9FF7C961AFF1AFF33" lastPageId="2" lastPageNumber="589" pageId="1" pageNumber="588" type="diagnosis">
<paragraph id="5E3E81599621FFB9FF7C961AFF1AFF33" blockId="1.[151,1437,1430,2033]" lastBlockId="2.[151,1436,151,249]" lastPageId="2" lastPageNumber="589" pageId="1" pageNumber="588">
<emphasis id="6CF55D4B9621FFBAFF7C961AFEC5F84E" bold="true" box="[199,314,1898,1924]" pageId="1" pageNumber="588">Diagnosis</emphasis>
.
<taxonomicName id="9981FADA9621FFBAFEF7961BFDE7F84F" authorityName="Brown" authorityYear="2022" box="[332,536,1899,1925]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="bisexta">
<emphasis id="6CF55D4B9621FFBAFEF7961BFDE7F84F" box="[332,536,1899,1925]" italics="true" pageId="1" pageNumber="588">Thalleulia bisexta</emphasis>
</taxonomicName>
is superficially most similar to
<taxonomicName id="9981FADA9621FFBAFC2B961BFBE0F84F" authorityName="Razowski and Wojtusiak" authorityYear="2009" box="[912,1055,1899,1925]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pondoana">
<emphasis id="6CF55D4B9621FFBAFC2B961BFBE0F84F" box="[912,1055,1899,1925]" italics="true" pageId="1" pageNumber="588">T. pondoana</emphasis>
</taxonomicName>
from
<collectingCountry id="2696C1C99621FFBAFBD1961BFB36F84F" box="[1130,1225,1899,1925]" name="Ecuador" pageId="1" pageNumber="588">Ecuador</collectingCountry>
, with a somewhat uniform brown forewing lacking distinct pattern elements.
<taxonomicName id="9981FADA9621FFBAFC9996FFFC16F863" authorityName="Brown" authorityYear="2022" box="[802,1001,1935,1961]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="bisexta">
<emphasis id="6CF55D4B9621FFBAFC9996FFFC16F863" box="[802,1001,1935,1961]" italics="true" pageId="1" pageNumber="588">Thalleulia bisexta</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="9981FADA9621FFBAFAA996FFFA63F863" authorityName="Razowski and Wojtusiak" authorityYear="2009" box="[1298,1436,1935,1961]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="pondoana">
<emphasis id="6CF55D4B9621FFBAFAA996FFFA63F863" box="[1298,1436,1935,1961]" italics="true" pageId="1" pageNumber="588">T. pondoana</emphasis>
</taxonomicName>
and other congeners by the presence of dense, transverse, slender striae throughout the forewing (
<figureCitation id="C6BA9DDC9621FFBAFB6196C3FAE3F807" box="[1242,1308,1971,1997]" captionStart="FIGURES 13" captionStartId="2.[152,270,1418,1442]" captionTargetBox="[151,1435,270,1392]" captionTargetId="figure-70@2.[151,1435,270,1392]" captionTargetPageId="2" captionText="FIGURES 13. Thalleulia bisexta. 1. Adult male paratype. 2. Male genitalia, USNM slide 124,124, phallus removed. 3. Female genitalia, USNM slide 68,857, spermatophore in situ." figureDoi="http://doi.org/10.5281/zenodo.7381761" httpUri="https://zenodo.org/record/7381761/files/figure.png" pageId="1" pageNumber="588">Fig. 1</figureCitation>
). The male genitalia of
<taxonomicName id="9981FADA9621FFBAFEA596A7FE75F83B" authorityName="Brown" authorityYear="2022" box="[286,394,2007,2033]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="bisexta">
<emphasis id="6CF55D4B9621FFBAFEA596A7FE75F83B" box="[286,394,2007,2033]" italics="true" pageId="1" pageNumber="588">T. bisexta</emphasis>
</taxonomicName>
(
<figureCitation id="C6BA9DDC9621FFBAFE2196A7FE21F83B" box="[410,478,2007,2033]" captionStart="FIGURES 13" captionStartId="2.[152,270,1418,1442]" captionTargetBox="[151,1435,270,1392]" captionTargetId="figure-70@2.[151,1435,270,1392]" captionTargetPageId="2" captionText="FIGURES 13. Thalleulia bisexta. 1. Adult male paratype. 2. Male genitalia, USNM slide 124,124, phallus removed. 3. Female genitalia, USNM slide 68,857, spermatophore in situ." figureDoi="http://doi.org/10.5281/zenodo.7381761" httpUri="https://zenodo.org/record/7381761/files/figure.png" pageId="1" pageNumber="588">Fig. 2</figureCitation>
) are nearly identical to those of
<taxonomicName id="9981FADA9621FFBAFCF596A7FC09F83B" authorityName="Razowski" authorityYear="2004" box="[846,1014,2007,2033]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="588" phylum="Arthropoda" rank="species" species="gracilescens">
<emphasis id="6CF55D4B9621FFBAFCF596A7FC09F83B" box="[846,1014,2007,2033]" italics="true" pageId="1" pageNumber="588">T. gracilescens</emphasis>
</taxonomicName>
, but they can be distinguished by the slightly straighter sacculus (weakly undulate in
<taxonomicName id="9981FADA9622FFB9FD0D91E7FCA0FF7B" authorityName="Razowski" authorityYear="2004" box="[694,863,151,177]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="gracilescens">
<emphasis id="6CF55D4B9622FFB9FD0D91E7FCA0FF7B" box="[694,863,151,177]" italics="true" pageId="2" pageNumber="589">T. gracilescens</emphasis>
</taxonomicName>
), a hooked tip at the apex of the lateral process of the gnathos (slightly curved in the distal
<quantity id="99792CBC9622FFB9FDE591CBFD61FF1F" box="[606,670,187,213]" metricMagnitude="-3" metricUnit="m" metricValue="5.08" pageId="2" pageNumber="589" unit="in" value="0.2">0.2 in</quantity>
<taxonomicName id="9981FADA9622FFB9FD1E91CBFCB3FF1F" authorityName="Razowski" authorityYear="2004" box="[677,844,187,213]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="gracilescens">
<emphasis id="6CF55D4B9622FFB9FD1E91CBFCB3FF1F" box="[677,844,187,213]" italics="true" pageId="2" pageNumber="589">T. gracilescens</emphasis>
</taxonomicName>
), and a more uniform coverage of granulation on the vesica.
</paragraph>
</subSubSection>
<caption id="0AFED1D19622FFB9FF2394FAFD49FA0D" ID-DOI="http://doi.org/10.5281/zenodo.7381761" ID-Zenodo-Dep="7381761" httpUri="https://zenodo.org/record/7381761/files/figure.png" pageId="2" pageNumber="589" startId="2.[152,270,1418,1442]" targetBox="[151,1435,270,1392]" targetPageId="2">
<paragraph id="5E3E81599622FFB9FF2394FAFD49FA0D" blockId="2.[151,1437,1418,1479]" pageId="2" pageNumber="589">
<emphasis id="6CF55D4B9622FFB9FF2394FAFEC0FA69" bold="true" box="[152,319,1418,1443]" pageId="2" pageNumber="589">FIGURES 13.</emphasis>
<taxonomicName id="9981FADA9622FFB9FEFF94FBFE05FA69" authorityName="Brown" authorityYear="2022" box="[324,506,1419,1443]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="589" phylum="Arthropoda" rank="species" species="bisexta">
<emphasis id="6CF55D4B9622FFB9FEFF94FBFE05FA69" box="[324,506,1419,1443]" italics="true" pageId="2" pageNumber="589">Thalleulia bisexta</emphasis>
</taxonomicName>
. 1. Adult male paratype. 2. Male genitalia, USNM slide 124,124, phallus removed. 3. Female genitalia, USNM slide 68,857, spermatophore in situ.
</paragraph>
</caption>
<subSubSection id="169BD2D29622FFB8FF7C9485FEC1FE43" lastPageId="3" lastPageNumber="590" pageId="2" pageNumber="589" type="description">
<paragraph id="5E3E81599622FFB8FF7C9485FEC1FE43" blockId="2.[151,1437,1525,2019]" lastBlockId="3.[151,1437,151,645]" lastPageId="3" lastPageNumber="590" pageId="2" pageNumber="589">
<emphasis id="6CF55D4B9622FFB9FF7C9485FEAEF9C5" bold="true" box="[199,337,1525,1551]" pageId="2" pageNumber="589">Description</emphasis>
.
<emphasis id="6CF55D4B9622FFB9FEDB9485FE62F9C5" box="[352,413,1525,1551]" italics="true" pageId="2" pageNumber="589">Head</emphasis>
: Vertex dark rust, lower frons pale orange-ocherous; labial palpus pale rust laterally, cream on inner surface; antenna with rust scales basally, paler distally. Ocellus tiny. Chaetosemata present.
<emphasis id="6CF55D4B9622FFB9FABE9769FAA9F9F9" box="[1285,1366,1561,1587]" italics="true" pageId="2" pageNumber="589">Thorax</emphasis>
: Nota and tegula dark rust. Forewing length 7.010.0 mm; forewing (
<figureCitation id="C6BA9DDC9622FFB9FCE3974DFC67F99D" box="[856,920,1597,1623]" captionStart="FIGURES 13" captionStartId="2.[152,270,1418,1442]" captionTargetBox="[151,1435,270,1392]" captionTargetId="figure-70@2.[151,1435,270,1392]" captionTargetPageId="2" captionText="FIGURES 13. Thalleulia bisexta. 1. Adult male paratype. 2. Male genitalia, USNM slide 124,124, phallus removed. 3. Female genitalia, USNM slide 68,857, spermatophore in situ." figureDoi="http://doi.org/10.5281/zenodo.7381761" httpUri="https://zenodo.org/record/7381761/files/figure.png" pageId="2" pageNumber="589">Fig. 1</figureCitation>
) ground color pale cream, densely overscaled throughout with narrow, rust, mostly interrupted, transverse striae, usually with a single, slender, complete arched stria from postmedian costa to tornus and a slightly more concentrated region of scales just basal to end of discal cell; fringe rust-brown in termen, paler throughout tornus. Hindwing white with faint, pale tan overscaling, densest at outer 0.15; fringe pale cream-tan.
<emphasis id="6CF55D4B9622FFB9FD8F97BDFD61F92D" box="[564,670,1741,1767]" italics="true" pageId="2" pageNumber="589">Abdomen</emphasis>
: Scaling pale cream-tan. Male genitalia (
<figureCitation id="C6BA9DDC9622FFB9FBD497BDFB4CF92D" box="[1135,1203,1741,1767]" captionStart="FIGURES 13" captionStartId="2.[152,270,1418,1442]" captionTargetBox="[151,1435,270,1392]" captionTargetId="figure-70@2.[151,1435,270,1392]" captionTargetPageId="2" captionText="FIGURES 13. Thalleulia bisexta. 1. Adult male paratype. 2. Male genitalia, USNM slide 124,124, phallus removed. 3. Female genitalia, USNM slide 68,857, spermatophore in situ." figureDoi="http://doi.org/10.5281/zenodo.7381761" httpUri="https://zenodo.org/record/7381761/files/figure.png" pageId="2" pageNumber="589">Fig. 2</figureCitation>
) with uncus slender, rodlike, broadest basally; socius broad, pendant, membranous, with long fine setae, from a narrow, rounded stalk, creating appearance of a pair of sublateral humps; gnathos arms joined distally into long, slender, spine-like process, each arm with long, straight, free, latero-terminal process, broadest subbasally, attenuate distally, with hooked tip; transtilla a weakly sclerotized band with weak dorso-median lobe; valva broad, parallel sided, rounded apically, weakly upcurved, costa weakly sclerotized, a series of slightly oblique, parallel wrinkles in basal 0.5 (not associated with fine hairs from venter of valva), sacculus narrow, sclerotized, restricted to basal 0.5 of valva, ending in a free terminal tip; phallus relatively straight, broad, rounded at base, with a pair of slender, lateral, wing-like processes, vesica with a small patch of tiny, non-deciduous cornuti and large areas of fine granulation. Female genitalia (
<figureCitation id="C6BA9DDC9623FFB8FAD491E7FF5AFF1F" captionStart="FIGURES 13" captionStartId="2.[152,270,1418,1442]" captionTargetBox="[151,1435,270,1392]" captionTargetId="figure-70@2.[151,1435,270,1392]" captionTargetPageId="2" captionText="FIGURES 13. Thalleulia bisexta. 1. Adult male paratype. 2. Male genitalia, USNM slide 124,124, phallus removed. 3. Female genitalia, USNM slide 68,857, spermatophore in situ." figureDoi="http://doi.org/10.5281/zenodo.7381761" httpUri="https://zenodo.org/record/7381761/files/figure.png" pageId="3" pageNumber="590">Fig. 3</figureCitation>
) with papillae anales and apophyses unmodified; lamella postvaginalis a broad lateral plate with three or more, wrinkly, nearly parallel lines extending outward from median ovoid region; lamella antevaginalis a broadly Vshaped cup, truncate at anterior margin, with rounded excavation at posterior margin; ductus bursae short, stout, not differentiated from corpus bursae; origin of ductus seminalis undetermined; corpus bursae elongate-oblong, with a single long, subcylindrical signum bearing irregular rows of long slender spines. A single spermataphore present in one dissection.
</paragraph>
</subSubSection>
<subSubSection id="169BD2D29623FFB8FF7C90E2FE83FDF7" pageId="3" pageNumber="590" type="distribution">
<paragraph id="5E3E81599623FFB8FF7C90E2FE83FDF7" blockId="3.[151,1437,151,645]" pageId="3" pageNumber="590">
<emphasis id="6CF55D4B9623FFB8FF7C90E2FE0EFE66" bold="true" box="[199,497,402,429]" pageId="3" pageNumber="590">Distribution and Biology</emphasis>
.
<taxonomicName id="9981FADA9623FFB8FDBA90E3FD33FE67" authorityName="Brown" authorityYear="2022" box="[513,716,403,429]" class="Insecta" family="Tortricidae" genus="Thalleulia" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="590" phylum="Arthropoda" rank="species" species="bisexta">
<emphasis id="6CF55D4B9623FFB8FDBA90E3FD33FE67" box="[513,716,403,429]" italics="true" pageId="3" pageNumber="590">Thalleulia bisexta</emphasis>
</taxonomicName>
is known from the Costa Rican provinces of
<collectingRegion id="9C454FBB9623FFB8FB6E90E3FACFFE67" box="[1237,1328,403,429]" country="Costa Rica" name="Cartago" pageId="3" pageNumber="590">Cartago</collectingRegion>
,
<collectingRegion id="9C454FBB9623FFB8FA8690E3FA66FE67" box="[1341,1433,403,429]" country="Costa Rica" name="Heredia" pageId="3" pageNumber="590">Heredia</collectingRegion>
,
<collectingRegion id="9C454FBB9623FFB8FF2C90C7FEEAFE1B" box="[151,277,439,465]" country="Costa Rica" name="Puntarenas" pageId="3" pageNumber="590">Puntarenas</collectingRegion>
, and
<collectingRegion id="9C454FBB9623FFB8FEE290C7FE3CFE1B" box="[345,451,439,465]" country="Costa Rica" name="San Jose" pageId="3" pageNumber="590">San José</collectingRegion>
. Specimens have been collected mostly between 1000 and
<quantity id="99792CBC9623FFB8FB3F90C7FB21FE1B" box="[1156,1246,439,465]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="3" pageNumber="590" unit="m" value="1600.0">1600 m</quantity>
elevation;
<specimenCount id="48874AD09623FFB8FADF90C7FEF3FE3F" pageId="3" pageNumber="590" type="generic">three specimens</specimenCount>
from
<quantity id="99792CBC9623FFB8FEEE90ABFE0CFE3F" box="[341,499,475,501]" metricMagnitude="3" metricUnit="m" metricValue="2.7199999999999998" metricValueMax="2.84" metricValueMin="2.6" pageId="3" pageNumber="590" unit="m" value="2720.0" valueMax="2840.0" valueMin="2600.0">26002840 m</quantity>
are slightly larger than those collected at lower elevations. Specimens have been captured nearly throughout the year: March, April, May, June, July, August, September, and November. The early stages are unknown.
</paragraph>
</subSubSection>
<subSubSection id="169BD2D29623FFB8FF7C9337FDA9FD4F" pageId="3" pageNumber="590" type="etymology">
<paragraph id="5E3E81599623FFB8FF7C9337FDA9FD4F" blockId="3.[151,1437,151,645]" pageId="3" pageNumber="590">
<emphasis id="6CF55D4B9623FFB8FF7C9337FEB7FDAB" bold="true" box="[199,328,583,609]" pageId="3" pageNumber="590">Etymology</emphasis>
. The specific epithet refers to the fact that this species is represented by both males and females —the first species of the genus to be so.
</paragraph>
</subSubSection>
</treatment>
</document>