239 lines
19 KiB
XML
239 lines
19 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/evolsyst.6.87158" ID-Pensoft-Pub="2535-0730-2-143" ID-Pensoft-UUID="DB80A4F877A753509B09A89648156DB4" ID-ZooBank="F5A5F50EC380414F838DA60AD251B2C6" ModsDocID="2535-0730-6-2-143" checkinTime="1658525555661" checkinUser="pensoft" docAuthor="Zamani, Alireza, Al-Yacoub, Ghassan A. Ali & Abdullah Najim, Shurooq" docDate="2022" docId="C389B070CAE15F098C5FBFA9DAC4D63A" docLanguage="en" docName="EvolutSyst 6(2): 143-150" docOrigin="Evolutionary Systematics 6 (2)" docPubDate="2022-07-22" docSource="http://dx.doi.org/10.3897/evolsyst.6.87158" docTitle="Pterotricha kovblyuki Zamani & Marusik 2018" docType="treatment" docVersion="1" id="DB80A4F877A753509B09A89648156DB4" lastPageNumber="143" masterDocId="DB80A4F877A753509B09A89648156DB4" masterDocTitle="New data on Gnaphosidae (Arachnida, Araneae) of Iraq" masterLastPageNumber="150" masterPageNumber="143" pageNumber="143" updateTime="1658525555661" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>New data on Gnaphosidae (Arachnida, Araneae) of Iraq</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Zamani, Alireza</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-8084-9666</mods:nameIdentifier>
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<mods:affiliation>Zoological Museum, Biodiversity Unit, University of Turku, Turku FI- 20014, Finland</mods:affiliation>
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<mods:nameIdentifier type="email">zamani.alireza5@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Al-Yacoub, Ghassan A. Ali</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-4931-855X</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, College of Education for Pure Sciences, University of Thi-Qar, Thi-Qar, Iraq</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Abdullah Najim, Shurooq</mods:namePart>
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<mods:affiliation>Department of Ecology, College of Science, University of Basrah, Basrah, Iraq</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Evolutionary Systematics</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>6</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>2</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>143</mods:start>
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<mods:end>150</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/evolsyst.6.87158</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/evolsyst.6.87158</mods:identifier>
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<mods:identifier type="Pensoft-Pub">2535-0730-2-143</mods:identifier>
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<mods:identifier type="ZooBank">F5A5F50EC380414F838DA60AD251B2C6</mods:identifier>
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<mods:identifier type="Pensoft-UUID">DB80A4F877A753509B09A89648156DB4</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:C389B070CAE15F098C5FBFA9DAC4D63A" httpUri="http://treatment.plazi.org/id/C389B070CAE15F098C5FBFA9DAC4D63A" lastPageNumber="143" pageId="0" pageNumber="143">
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<subSubSection pageId="0" pageNumber="143" type="nomenclature">
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<paragraph pageId="0" pageNumber="143">
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<taxonomicName LSID="C389B070-CAE1-5F09-8C5F-BFA9DAC4D63A" authority="Zamani & Marusik, 2018" authorityName="Zamani & Marusik" authorityYear="2018" class="Arachnida" family="Gnaphosidae" genus="Pterotricha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pterotricha kovblyuki" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="species" species="kovblyuki">Pterotricha kovblyuki Zamani & Marusik, 2018</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="description">
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<paragraph pageId="0" pageNumber="143">
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of male (A, D, E) and female (B, C, F) gnaphosids, dorsal view. A, B. Minosiella intermedia Denis, 1958; C. Marinarozelotes jaxartensis (Kroneberg, 1875); D. Odontodrassus aravaensis Levy, 1999; E. Pterotricha dalmasi Fage, 1929; F. Pterotricha kovblyuki Zamani & Marusik, 2018." httpUri="https://binary.pensoft.net/fig/718429" pageId="0" pageNumber="143">Figs 1F</figureCitation>
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigynes of Pterotricha dalmasi Fage, 1929 (A-C) and Pterotricha kovblyuki Zamani & Marusik, 2018 (D-F). A, D. intact epigyne, ventral view; B, E. macerated epigyne, ventral view; C, F. vulva, dorsal view. Abbreviations: Ag - accessorial gland; Ah - anterior hood; Ld - lateral duct; Lm - lateral margin; Re - receptacle." httpUri="https://binary.pensoft.net/fig/718433" pageId="0" pageNumber="143">, 5D-F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="identification">
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<paragraph pageId="0" pageNumber="143">Identification.</paragraph>
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<paragraph pageId="0" pageNumber="143">
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.777.26745" author="Zamani, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="143" pagination="17 - 41" refId="B31" refString="Zamani, A, Seiedy, M, Saboori, A, Marusik, YM, 2018. The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae). ZooKeys 777: 17 - 41, DOI: https://doi.org/10.3897/zookeys.777.26745" title="The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae)." url="https://doi.org/10.3897/zookeys.777.26745" volume="777" year="2018">Zamani et al. (2018)</bibRefCitation>
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,
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<bibRefCitation DOI="https://doi.org/10.3897/evolsyst.2.2998" author="Zamani, A" journalOrPublisher="Evolutionary Systematics" pageId="0" pageNumber="143" pagination="151 - 161" refId="B28" refString="Zamani, A, 2018. The spider genus Pterotricha Kulczynski, 1903 (Araneae, Gnaphosidae) in the United Arab Emirates. Evolutionary Systematics 2: 151 - 161, DOI: https://doi.org/10.3897/evolsyst.2.2998" title="The spider genus Pterotricha Kulczynski, 1903 (Araneae, Gnaphosidae) in the United Arab Emirates." url="https://doi.org/10.3897/evolsyst.2.2998" volume="2" year="2018">Zamani (2018)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="materials_examined">
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<paragraph pageId="0" pageNumber="143">Material.</paragraph>
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<paragraph pageId="0" pageNumber="143">
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<materialsCitation collectingDate="1979-01-18" collectorName="O. Jakes" country="IRAQ" elevation="260" latitude="30.8" location="Ash Shabakah" longLatPrecision="1220" longitude="43.65" specimenCount="15" specimenCount-female="1" specimenCount-male="14" stateProvince="Najaf Province">
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<collectingCountry name="Iraq">IRAQ</collectingCountry>
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:
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<emphasis bold="true" italics="true" pageId="0" pageNumber="143">
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<collectingRegion country="Iraq" name="An Najaf">Najaf Province</collectingRegion>
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</emphasis>
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:
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<specimenCount type="male">14♂</specimenCount>
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<specimenCount type="female">1♀</specimenCount>
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(MMBC),
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<location LSID="urn:lsid:plazi:treatment:C389B070CAE15F098C5FBFA9DAC4D63A:B9AF10EB3CBD0347E7BFDA5E467F2AFD" country="IRAQ" latitude="30.8" longLatPrecision="1220" longitude="43.65" name="Ash Shabakah" stateProvince="Najaf Province">Ash Shabakah</location>
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camp,
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<quantity metricMagnitude="5" metricUnit="m" metricValue="1.5" unit="km" value="150.0">150 km</quantity>
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SW of
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<collectingRegion country="Iraq" name="An Najaf">Najaf</collectingRegion>
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, [
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<geoCoordinate degrees="30" direction="north" minutes="48" orientation="latitude" precision="925" value="30.8">30°48'N</geoCoordinate>
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,
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<geoCoordinate degrees="43" direction="east" minutes="39" orientation="longitude" precision="925" value="43.65">43°39'E</geoCoordinate>
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],
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<quantity metricMagnitude="2" metricUnit="m" metricValue="2.6" unit="m" value="260.0">
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<elevation metricMagnitude="2" metricUnit="m" metricValue="2.6" unit="m" value="260.0">260 m</elevation>
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</quantity>
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, stone desert,
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<collectingDate value="1979-01-18">18.1.1979</collectingDate>
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(
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<collectorName>
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O.
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<normalizedToken originalValue="Jakeš">Jakes</normalizedToken>
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</collectorName>
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)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1978-02-11" collectorName="O. Jakes" country="Iraq" elevation="260" latitude="30.8" location="Ash Shabakah" longLatPrecision="1220" longitude="43.65" specimenCount="21" specimenCount-female="2" specimenCount-male="19" stateProvince="An Najaf">
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<specimenCount type="male">19♂</specimenCount>
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<specimenCount type="female">2♀</specimenCount>
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(MMBC), same locality,
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<collectingDate value="1978-02-11">11.2.1978</collectingDate>
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(
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<collectorName>
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O.
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<normalizedToken originalValue="Jakeš">Jakes</normalizedToken>
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</collectorName>
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)
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</materialsCitation>
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;
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<materialsCitation collectingDate="1972-10-16" collectorName="O. Jakes" country="Iraq" elevation="260" latitude="30.8" location="Ash Shabakah" longLatPrecision="1220" longitude="43.65" specimenCount="5" specimenCount-male="5" stateProvince="An Najaf">
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<specimenCount type="male">5♂</specimenCount>
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(MMBC), same locality,
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<collectingDate value="1972-10-16">16.10.1972</collectingDate>
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(
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<collectorName>
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O.
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<normalizedToken originalValue="Jakeš">Jakes</normalizedToken>
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</collectorName>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="emended diagnosis">
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<paragraph pageId="0" pageNumber="143">Emended diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="143">
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The female of
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<taxonomicName genus="P." lsidName="P. kovblyuki" pageId="0" pageNumber="143" rank="species" species="kovblyuki">
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<emphasis italics="true" pageId="0" pageNumber="143">P. kovblyuki</emphasis>
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</taxonomicName>
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differs from that of
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<taxonomicName genus="P." lsidName="P. dalmasi" pageId="0" pageNumber="143" rank="species" species="dalmasi">
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<emphasis italics="true" pageId="0" pageNumber="143">P. dalmasi</emphasis>
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</taxonomicName>
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by the round anterior hood (
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<emphasis italics="true" pageId="0" pageNumber="143">Ah</emphasis>
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) (
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<emphasis italics="true" pageId="0" pageNumber="143">vs.</emphasis>
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subdivided by a median pointed protrusion; Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigynes of Pterotricha dalmasi Fage, 1929 (A-C) and Pterotricha kovblyuki Zamani & Marusik, 2018 (D-F). A, D. intact epigyne, ventral view; B, E. macerated epigyne, ventral view; C, F. vulva, dorsal view. Abbreviations: Ag - accessorial gland; Ah - anterior hood; Ld - lateral duct; Lm - lateral margin; Re - receptacle." httpUri="https://binary.pensoft.net/fig/718433" pageId="0" pageNumber="143">5E</figureCitation>
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cf. Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigynes of Pterotricha dalmasi Fage, 1929 (A-C) and Pterotricha kovblyuki Zamani & Marusik, 2018 (D-F). A, D. intact epigyne, ventral view; B, E. macerated epigyne, ventral view; C, F. vulva, dorsal view. Abbreviations: Ag - accessorial gland; Ah - anterior hood; Ld - lateral duct; Lm - lateral margin; Re - receptacle." httpUri="https://binary.pensoft.net/fig/718433" pageId="0" pageNumber="143">5B</figureCitation>
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). Epigyne of
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<taxonomicName genus="P." lsidName="P. kovblyuki" pageId="0" pageNumber="143" rank="species" species="kovblyuki">
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<emphasis italics="true" pageId="0" pageNumber="143">P. kovblyuki</emphasis>
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</taxonomicName>
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is most similar to that of
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<taxonomicName genus="P." lsidName="P. algerica" pageId="0" pageNumber="143" rank="species" species="algerica">
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<emphasis italics="true" pageId="0" pageNumber="143">P. algerica</emphasis>
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</taxonomicName>
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Dalmas, 1921 and
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<taxonomicName genus="P." lsidName="P. egens" pageId="0" pageNumber="143" rank="species" species="egens">
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<emphasis italics="true" pageId="0" pageNumber="143">P. egens</emphasis>
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</taxonomicName>
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Denis, 1966 by having round anterior hood (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigynes of Pterotricha dalmasi Fage, 1929 (A-C) and Pterotricha kovblyuki Zamani & Marusik, 2018 (D-F). A, D. intact epigyne, ventral view; B, E. macerated epigyne, ventral view; C, F. vulva, dorsal view. Abbreviations: Ag - accessorial gland; Ah - anterior hood; Ld - lateral duct; Lm - lateral margin; Re - receptacle." httpUri="https://binary.pensoft.net/fig/718433" pageId="0" pageNumber="143">5E</figureCitation>
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cf.
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<bibRefCitation author="Denis, J" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle d'Afrique du Nord" pageId="0" pageNumber="143" pagination="103 - 144" refId="B12" refString="Denis, J, 1966. Les araignees du Fezzan. Bulletin de la Societe d'Histoire Naturelle d'Afrique du Nord 55: 103 - 144" title="Les araignees du Fezzan." volume="55" year="1966">Denis 1966</bibRefCitation>
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: fig. 8 and
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<bibRefCitation author="Dalmas, R de" journalOrPublisher="Annales de la Societe Entomologique de France" pageId="0" pageNumber="143" pagination="233 - 328" refId="B10" refString="Dalmas, R de, 1921. Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae). Annales de la Societe Entomologique de France 89: 233 - 328" title="Monographie des araignees de la section des Pterotricha (Aran. Gnaphosidae)." volume="89" year="1921">Dalmas 1921</bibRefCitation>
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: fig. 29), but can be distinguished from them by diverging lateral margins (
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<emphasis italics="true" pageId="0" pageNumber="143">Lm</emphasis>
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) of the fovea (
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<emphasis italics="true" pageId="0" pageNumber="143">vs.</emphasis>
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almost parallel), and anterior hood narrower than posterior part of fovea (
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<emphasis italics="true" pageId="0" pageNumber="143">vs.</emphasis>
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equal in width). For the male, see
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.777.26745" author="Zamani, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="143" pagination="17 - 41" refId="B31" refString="Zamani, A, Seiedy, M, Saboori, A, Marusik, YM, 2018. The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae). ZooKeys 777: 17 - 41, DOI: https://doi.org/10.3897/zookeys.777.26745" title="The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae)." url="https://doi.org/10.3897/zookeys.777.26745" volume="777" year="2018">Zamani et al. (2018)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="description">
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<paragraph pageId="0" pageNumber="143">Description.</paragraph>
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<paragraph pageId="0" pageNumber="143">
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Female. Habitus as in Fig.
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<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Habitus of male (A, D, E) and female (B, C, F) gnaphosids, dorsal view. A, B. Minosiella intermedia Denis, 1958; C. Marinarozelotes jaxartensis (Kroneberg, 1875); D. Odontodrassus aravaensis Levy, 1999; E. Pterotricha dalmasi Fage, 1929; F. Pterotricha kovblyuki Zamani & Marusik, 2018." httpUri="https://binary.pensoft.net/fig/718429" pageId="0" pageNumber="143">1F</figureCitation>
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. Total length 8.75. Carapace 2.90 long, 2.30 wide. Eye sizes: AME: 0.21, ALE: 0.21, PLE: 0.20, PME: 0.22. Carapace, chelicerae, sternum, maxillae and labium light brown. Chelicerae with a promarginal tooth and a retromarginal bifurcate serrated keel. Abdomen light beige, with 3 pairs of sigilla and, in some individuals, slightly darker cardiac mark. Spinnerets light yellowish brown, uniform in color. Legs colored as carapace, without annulations. Measurements of legs: I: 9.81 (2.75, 1.19, 2.16, 2.05, 1.66), II: 9.66 (2.56, 1.25, 2.00, 2.12, 1.73), III: 9.78 (2.35, 1.21, 1.88, 2.46, 1.88), IV: 12.53 (3.12, 1.35, 2.66, 3.23, 2.17).
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</paragraph>
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<paragraph pageId="0" pageNumber="143">
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Epigyne as in Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Epigynes of Pterotricha dalmasi Fage, 1929 (A-C) and Pterotricha kovblyuki Zamani & Marusik, 2018 (D-F). A, D. intact epigyne, ventral view; B, E. macerated epigyne, ventral view; C, F. vulva, dorsal view. Abbreviations: Ag - accessorial gland; Ah - anterior hood; Ld - lateral duct; Lm - lateral margin; Re - receptacle." httpUri="https://binary.pensoft.net/fig/718433" pageId="0" pageNumber="143">5D-F</figureCitation>
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; plate almost as long as wide; anterior hood with round margin and large lateral diverticula (
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<emphasis italics="true" pageId="0" pageNumber="143">Ld</emphasis>
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); lateral margin of fovea diverging; posterior part of fovea wider than anterior hood; receptacles (
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<emphasis italics="true" pageId="0" pageNumber="143">Re</emphasis>
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) and accessorial glands (
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<emphasis italics="true" pageId="0" pageNumber="143">Ag</emphasis>
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) diverging anteriorly.
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</paragraph>
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<paragraph pageId="0" pageNumber="143">
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Male. See
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<bibRefCitation DOI="https://doi.org/10.3897/zookeys.777.26745" author="Zamani, A" journalOrPublisher="ZooKeys" pageId="0" pageNumber="143" pagination="17 - 41" refId="B31" refString="Zamani, A, Seiedy, M, Saboori, A, Marusik, YM, 2018. The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae). ZooKeys 777: 17 - 41, DOI: https://doi.org/10.3897/zookeys.777.26745" title="The spider genus Pterotricha in Iran, with the description of a new genus (Araneae, Gnaphosidae)." url="https://doi.org/10.3897/zookeys.777.26745" volume="777" year="2018">Zamani et al. (2018)</bibRefCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="distribution">
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<paragraph pageId="0" pageNumber="143">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="143">Iraq, Iran and United Arab Emirates.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="143" type="comments">
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<paragraph pageId="0" pageNumber="143">Comment.</paragraph>
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<paragraph pageId="0" pageNumber="143">
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In Iraq, it was previously reported from Thi Qar Province (
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<bibRefCitation author="Al-Yacoub, GAA" journalOrPublisher="Serket" pageId="0" pageNumber="143" pagination="186 - 190" refId="B6" refString="Al-Yacoub, GAA, Al-Abbad, MYM, Kareem, DK, 2021b. First record of Pterotricha kovblyuki Zamani & Marusik, 2018 (Araneae: Gnaphosidae) from Iraq. Serket 18 (2): 186 - 190" title="First record of Pterotricha kovblyuki Zamani & Marusik, 2018 (Araneae: Gnaphosidae) from Iraq." volume="18" year="2021 b">Al-Yacoub et al. 2021b</bibRefCitation>
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). The current material represents the westernmost record of the species across its known range.
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</paragraph>
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<paragraph pageId="0" pageNumber="143">
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<emphasis bold="true" pageId="0" pageNumber="143">
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Genus
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<taxonomicName class="Arachnida" family="Gnaphosidae" genus="Zelotes" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zelotes" order="Araneae" pageId="0" pageNumber="143" phylum="Arthropoda" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="143">Zelotes</emphasis>
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</taxonomicName>
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Gistel, 1848
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</emphasis>
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |