426 lines
47 KiB
XML
426 lines
47 KiB
XML
<document id="0DB26E58F7DA4FC7DCBCF80F2EA384CB" ID-DOI="10.1649/0010-065X-71.4.781" ID-ISSN="1938-4394" ID-Zenodo-Dep="5378591" ID-ZooBank="urn:lsid:zoobank.org:pub:89F5244B-37FE-4A14-BF98-A7B1210DA4DA" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630286094741" checkinUser="diego" docAuthor="Ratcliffe, Brett C. & Tirant, Stephane Le" docDate="2017" docId="57093477FFC5FFFDFD74FEB863E8DE98" docLanguage="en" docName="0010-065X-71.4.781.pdf" docOrigin="The Coleopterists Bulletin 71 (4)" docSource="http://dx.doi.org/10.1649/0010-065x-71.4.781" docStyle="DocumentStyle:F824235CA89CB83023C7D5789122F870.3:TheColeopBull.2017-.journal_article.1cover" docStyleId="F824235CA89CB83023C7D5789122F870" docStyleName="TheColeopBull.2017-.journal_article.1cover" docStyleVersion="3" docTitle="Golofa limogesi Ratcliffe and Le Tirant 2017, new species" docType="treatment" docVersion="3" lastPageNumber="785" masterDocId="AB304C0FFFC7FFF8FFA7FFF1621DDB5C" masterDocTitle="A New Species of Golofa Hope, 1837 (Coleoptera: Scarabaeidae: Dynastinae) from Peru" masterLastPageNumber="785" masterPageNumber="781" pageNumber="782" updateTime="1699214410213" updateUser="plazi">
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<mods:title id="A94D4A827A22C3D2F0BC9642D8FE19CA">A New Species of Golofa Hope, 1837 (Coleoptera: Scarabaeidae: Dynastinae) from Peru</mods:title>
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<mods:namePart id="AF5D0E67886745F248D36F2AFBB8FB5C">Ratcliffe, Brett C.</mods:namePart>
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<mods:roleTerm id="08460F1C418578ABDF345BDF594B7BE8">Author</mods:roleTerm>
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<mods:namePart id="5347B78C1D11861FE37ED2656022438F">Tirant, Stephane Le</mods:namePart>
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<mods:typeOfResource id="655E7B0F41B0D7E5708BF8A4ABAFE202">text</mods:typeOfResource>
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<mods:title id="54CD726F903CBA92EA6130A8F35CE04D">The Coleopterists Bulletin</mods:title>
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</mods:titleInfo>
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<mods:part id="AF3A9019B5A3A59209F1EA770A44C5B7">
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<mods:date id="511E546150285AECD2F7A7699385F6B8">2017</mods:date>
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<mods:number id="FB6359D5256503624BCE01F39FE84040">2017-12-22</mods:number>
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<mods:number id="CCF64C4F73BA5EA5865A7C97E16EF66B">71</mods:number>
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<mods:url id="36781B6CCA26C61F5DBD4EA81D0B1D89">http://dx.doi.org/10.1649/0010-065x-71.4.781</mods:url>
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<mods:identifier id="04CED09F30829537BCBBFC59FFE00E71" type="DOI">10.1649/0010-065X-71.4.781</mods:identifier>
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<mods:identifier id="1BD20B6511E52DEDFF9F43CF9A045AEA" type="ISSN">1938-4394</mods:identifier>
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<subSubSection id="97BAD6EAFFC5FFFAFD74FEB861C6DA25" pageId="2" pageNumber="782" type="nomenclature">
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<paragraph id="DF1F8561FFC5FFFAFD74FEB861C6DA25" blockId="2.[723,1135,329,404]" pageId="2" pageNumber="782">
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<heading id="8457320DFFC5FFFAFD74FEB861C6DA25" centered="true" fontSize="9" level="2" pageId="2" pageNumber="782" reason="2">
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<emphasis id="EDD45973FFC5FFFAFD74FEB861C6DA25" bold="true" pageId="2" pageNumber="782">
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<taxonomicName id="18A0FEE2FFC5FFFAFD74FEB86674DA02" authority="Ratcliffe and Le Tirant" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[723,1129,329,351]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="782" phylum="Arthropoda" rank="species" species="limogesi" status="sp. nov.">
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<emphasis id="EDD45973FFC5FFFAFD74FEB8616CDA03" bold="true" box="[723,881,329,351]" italics="true" pageId="2" pageNumber="782">Golofa limogesi</emphasis>
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Ratcliffe and Le Tirant
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</taxonomicName>
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,
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<taxonomicNameLabel id="F6E7E408FFC5FFFAFCC1FE9561C6DA25" box="[870,987,356,377]" pageId="2" pageNumber="782" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="97BAD6EAFFC5FFFAFCF8FE8F61FFDACF" box="[863,994,382,404]" pageId="2" pageNumber="782" type="description">
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<paragraph id="DF1F8561FFC5FFFAFCF8FE8F61FFDACF" blockId="2.[723,1135,329,404]" box="[863,994,382,404]" pageId="2" pageNumber="782">
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(
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<figureCitation id="479B99E4FFC5FFFAFCC1FE8F61DEDACF" box="[870,963,382,404]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="2" pageNumber="782">Figs. 1–4</figureCitation>
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,
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<figureCitation id="479B99E4FFC5FFFAFC68FE8F61C6DACF" box="[975,987,382,403]" captionStart="Fig" captionStartId="4.[709,740,1711,1730]" captionTargetBox="[685,1172,1286,1694]" captionTargetId="figure-627@4.[685,1172,1286,1694]" captionTargetPageId="4" captionText="Fig. 6. Golofa limogesi, distribution." figureDoi="http://doi.org/10.5281/zenodo.5378597" httpUri="https://zenodo.org/record/5378597/files/figure.png" pageId="2" pageNumber="782">6</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="97BAD6EAFFC5FFFAFD62FE43666ED867" pageId="2" pageNumber="782" type="materials_examined">
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<paragraph id="DF1F8561FFC5FFFAFD62FE43666ED867" blockId="2.[685,1173,434,1730]" pageId="2" pageNumber="782">
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<emphasis id="EDD45973FFC5FFFAFD62FE436146DA9B" bold="true" box="[709,859,434,455]" pageId="2" pageNumber="782">Type Material.</emphasis>
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<materialsCitation id="6FC88F3CFFC5FFFAFCC5FE436173D910" collectingDate="2015-03-01" collectingDateMax="2015-03-05" collectingDateMin="2015-03-01" collectorName="Local" country="Peru" location="Ayabaca" pageId="2" pageNumber="782" specimenCount="16" specimenCount-female="10" specimenCount-male="1" stateProvince="Piura" typeStatus="holotype">
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<typeStatus id="001B3BC3FFC5FFFAFCC5FE4361A7DA9B" box="[866,954,434,455]" pageId="2" pageNumber="782" type="holotype">Holotype</typeStatus>
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male,
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||
<typeStatus id="001B3BC3FFC5FFFAFC5CFE43665ADA9B" box="[1019,1095,434,455]" pageId="2" pageNumber="782" type="allotype">allotype</typeStatus>
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female,
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<specimenCount id="C9A64EE8FFC5FFFAFD0AFE3D6105DABD" box="[685,792,460,481]" pageId="2" pageNumber="782" type="male">seven male</specimenCount>
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<typeStatus id="001B3BC3FFC5FFFAFCB9FE3D6166DABD" box="[798,891,460,481]" pageId="2" pageNumber="782" type="paratype">paratypes</typeStatus>
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,
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<specimenCount id="C9A64EE8FFC5FFFAFC22FE3D61F9DABD" box="[901,996,460,481]" pageId="2" pageNumber="782" type="female">10 female</specimenCount>
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<typeStatus id="001B3BC3FFC5FFFAFC4DFE3D6658DABD" box="[1002,1093,460,481]" pageId="2" pageNumber="782" type="paratype">paratypes</typeStatus>
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||
all with same data, labeled “
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||
<collectingCountry id="A7B7C5F1FFC5FFFAFCD5FE1661AFDAA0" box="[882,946,487,508]" name="Peru" pageId="2" pageNumber="782">PERU</collectingCountry>
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:
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<collectingRegion id="1D644B83FFC5FFFAFC1AFE1661F2DAA0" box="[957,1007,487,508]" country="Peru" name="Piura" pageId="2" pageNumber="782">Piura</collectingRegion>
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/
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<location id="DA7FD3BAFFC5FFFAFBA2FE166644DAA0" LSID="urn:lsid:plazi:treatment:57093477FFC5FFFDFD74FEB863E8DE98:DA7FD3BAFFC5FFFAFBA2FE166644DAA0" box="[1029,1113,487,508]" country="Peru" name="Ayabaca" pageId="2" pageNumber="782" stateProvince="Piura">Ayabaca</location>
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, 3000 mts, /
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<date id="AB1EA3A1FFC5FFFAFD4BFDF06143D94A" box="[748,862,513,534]" pageId="2" pageNumber="782" value="2015-03-01" valueMax="2015-03-05" valueMin="2015-03-01">
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<collectingDate id="BB5A5A49FFC5FFFAFD4BFDF06143D94A" box="[748,862,513,534]" pageId="2" pageNumber="782" value="2015-03-01" valueMax="2015-03-05" valueMin="2015-03-01">1-5.III.2015</collectingDate>
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</date>
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/
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<collectorName id="7255E0B7FFC5FFFAFCD2FDF061B6D94A" box="[885,939,513,534]" pageId="2" pageNumber="782">Local</collectorName>
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collector” and with our red
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||
<typeStatus id="001B3BC3FFC5FFFAFD7FFDED6133D96D" box="[728,814,540,561]" pageId="2" pageNumber="782" type="holotype">holotype</typeStatus>
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, red
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||
<typeStatus id="001B3BC3FFC5FFFAFCCFFDED61AAD96D" box="[872,951,540,561]" pageId="2" pageNumber="782" type="allotype">allotype</typeStatus>
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, and yellow
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<typeStatus id="001B3BC3FFC5FFFAFBE5FDED668ED96D" box="[1090,1171,540,561]" pageId="2" pageNumber="782" type="paratype">paratype</typeStatus>
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labels, respectively
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||
</materialsCitation>
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.
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<materialsCitation id="6FC88F3CFFC5FFFAFC27FDC66170D9DD" collectorName="Lincoln, NE" country="United States of America" location="Nebraska State" pageId="2" pageNumber="782" specimenCount="2" stateProvince="Nebraska" typeStatus="holotype">
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||
<typeStatus id="001B3BC3FFC5FFFAFC27FDC661C5D910" box="[896,984,567,588]" pageId="2" pageNumber="782" type="holotype">Holotype</typeStatus>
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and
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||
<typeStatus id="001B3BC3FFC5FFFAFBBDFDC6667BD910" box="[1050,1126,567,588]" pageId="2" pageNumber="782" type="allotype">allotype</typeStatus>
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||
deposited at the University of
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||
<collectingRegion id="1D644B83FFC5FFFAFC08FDA06621D93A" box="[943,1084,593,614]" country="United States of America" name="Nebraska" pageId="2" pageNumber="782">Nebraska State</collectingRegion>
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Museum (
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||
<collectorName id="7255E0B7FFC5FFFAFD13FD9D6130D9DD" box="[692,813,620,641]" pageId="2" pageNumber="782">Lincoln, NE</collectorName>
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,
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||
<collectingCountry id="A7B7C5F1FFC5FFFAFC90FD9D6175D9DD" box="[823,872,620,641]" name="United States of America" pageId="2" pageNumber="782">USA</collectingCountry>
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)
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||
</materialsCitation>
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<materialsCitation id="6FC88F3CFFC5FFFAFCD3FD9D6672D867" collectionCode="QC, DC" collectorName="de Montreal & US National Museum & Brett C. Ratcliffe Collection & Lincoln, NE" country="Canada" county="Terrebonne" location="Insectarium de Montreal" municipality="Stephane Le Tirant Collection" pageId="2" pageNumber="782" specimenCount="1" stateProvince="Washington" typeStatus="paratype">
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and
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||
<typeStatus id="001B3BC3FFC5FFFAFC3BFD9D61EAD9DD" box="[924,1015,620,641]" pageId="2" pageNumber="782" type="paratype">paratypes</typeStatus>
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||
deposited in the
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||
<location id="DA7FD3BAFFC5FFFAFD0AFD776183D9C7" LSID="urn:lsid:plazi:treatment:57093477FFC5FFFDFD74FEB863E8DE98:DA7FD3BAFFC5FFFAFD0AFD776183D9C7" box="[685,926,646,667]" country="Canada" county="Terrebonne" municipality="Stephane Le Tirant Collection" name="Insectarium de Montreal" pageId="2" pageNumber="782" stateProvince="Washington">
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Insectarium
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<collectorName id="7255E0B7FFC5FFFAFC81FD776183D9C7" box="[806,926,646,667]" pageId="2" pageNumber="782">de Montréal</collectorName>
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</location>
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(
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<location id="DA7FD3BAFFC5FFFAFC0AFD77661AD9C7" LSID="urn:lsid:plazi:treatment:57093477FFC5FFFDFD74FEB863E8DE98:DA7FD3BAFFC5FFFAFC0AFD77661AD9C7" box="[941,1031,646,667]" country="Canada" county="Terrebonne" municipality="Stephane Le Tirant Collection" name="Montreal" pageId="2" pageNumber="782" stateProvince="Washington">Montréal</location>
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,
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<collectionCode id="B9B11DA4FFC5FFFAFBB2FD776625D9C7" box="[1045,1080,646,667]" country="Zimbabwe" name="National Museum of Natural History, Bulawayo" pageId="2" pageNumber="782">QC</collectionCode>
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,
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<collectingCountry id="A7B7C5F1FFC5FFFAFBE1FD77668DD9C7" box="[1094,1168,646,667]" name="Canada" pageId="2" pageNumber="782">Canada</collectingCountry>
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; 2),
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<collectorName id="7255E0B7FFC5FFFAFD7FFD5061A3D9EA" box="[728,958,673,694]" pageId="2" pageNumber="782">
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US
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<location id="DA7FD3BAFFC5FFFAFCAFFD5061A3D9EA" LSID="urn:lsid:plazi:treatment:57093477FFC5FFFDFD74FEB863E8DE98:DA7FD3BAFFC5FFFAFCAFFD5061A3D9EA" box="[776,958,673,694]" country="Canada" county="Terrebonne" municipality="Stephane Le Tirant Collection" name="National Museum of Natural History" pageId="2" pageNumber="782" stateProvince="Washington">National Museum</location>
|
||
</collectorName>
|
||
of Natural History (
|
||
<collectingRegion id="1D644B83FFC5FFFAFD13FD4A6134D98C" box="[692,809,699,720]" country="United States of America" name="Washington" pageId="2" pageNumber="782">Washington</collectingRegion>
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||
,
|
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<collectionCode id="B9B11DA4FFC5FFFAFC96FD4A6149D98C" box="[817,852,699,720]" pageId="2" pageNumber="782">DC</collectionCode>
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||
, USA; 2),
|
||
<location id="DA7FD3BAFFC5FFFAFC1FFD4A6174D9B7" LSID="urn:lsid:plazi:treatment:57093477FFC5FFFDFD74FEB863E8DE98:DA7FD3BAFFC5FFFAFC1FFD4A6174D9B7" country="Canada" county="Terrebonne" municipality="Stephane Le Tirant Collection" name="Museo de Entomolog'ia Klaus Raven Buller" pageId="2" pageNumber="782" stateProvince="Washington">Museo de Entomolog´ıa Klaus Raven Büller</location>
|
||
(
|
||
<location id="DA7FD3BAFFC5FFFAFCD2FD276107D859" LSID="urn:lsid:plazi:treatment:57093477FFC5FFFDFD74FEB863E8DE98:DA7FD3BAFFC5FFFAFCD2FD276107D859" country="Canada" county="Terrebonne" municipality="Stephane Le Tirant Collection" name="Universidad Nacional Agraria La Molina" pageId="2" pageNumber="782" stateProvince="Washington">Universidad Nacional Agraria La Molina</location>
|
||
,
|
||
<collectingRegion id="1D644B83FFC5FFFAFC8DFD016143D859" box="[810,862,752,773]" country="Peru" name="Lima" pageId="2" pageNumber="782">Lima</collectingRegion>
|
||
,
|
||
<collectingCountry id="A7B7C5F1FFC5FFFAFCC9FD016181D859" box="[878,924,752,773]" name="Peru" pageId="2" pageNumber="782">Peru</collectingCountry>
|
||
; 2),
|
||
<collectingMunicipality id="3F7B1F1BFFC5FFFAFC75FD01610DD87C" pageId="2" pageNumber="782">Stephane Le Tirant Collection</collectingMunicipality>
|
||
(
|
||
<collectingCounty id="367EFDEDFFC5FFFAFCBAFCFA6190D87C" box="[797,909,779,800]" pageId="2" pageNumber="782">Terrebonne</collectingCounty>
|
||
,
|
||
<collectionCode id="B9B11DA4FFC5FFFAFC30FCFA61A7D87C" box="[919,954,779,800]" country="Zimbabwe" name="National Museum of Natural History, Bulawayo" pageId="2" pageNumber="782">QC</collectionCode>
|
||
,
|
||
<collectingCountry id="A7B7C5F1FFC5FFFAFC61FCFA660DD87C" box="[966,1040,779,800]" name="Canada" pageId="2" pageNumber="782">Canada</collectingCountry>
|
||
; 7), and
|
||
<collectorName id="7255E0B7FFC5FFFAFBC3FCFA6195D867" pageId="2" pageNumber="782">Brett C. Ratcliffe Collection</collectorName>
|
||
(
|
||
<collectorName id="7255E0B7FFC5FFFAFC32FCD7660DD867" box="[917,1040,806,827]" pageId="2" pageNumber="782">Lincoln, NE</collectorName>
|
||
, USA; 4)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="97BAD6EAFFC5FFFCFD62FCB1668CD80A" lastPageId="4" lastPageNumber="784" pageId="2" pageNumber="782" type="description">
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<paragraph id="DF1F8561FFC5FFFCFD62FCB162FED98C" blockId="2.[685,1173,434,1730]" lastBlockId="4.[151,639,194,1730]" lastPageId="4" lastPageNumber="784" pageId="2" pageNumber="782">
|
||
<emphasis id="EDD45973FFC5FFFAFD62FCB161FED809" bold="true" box="[709,995,832,853]" pageId="2" pageNumber="782">
|
||
Description.
|
||
<typeStatus id="001B3BC3FFC5FFFAFCEFFCB161B9D809" box="[840,932,832,853]" pageId="2" pageNumber="782" type="holotype">Holotype</typeStatus>
|
||
male.
|
||
</emphasis>
|
||
Length
|
||
<quantity id="18582884FFC5FFFAFB91FCB1668DD809" box="[1078,1168,832,853]" metricMagnitude="-2" metricUnit="m" metricValue="4.35" pageId="2" pageNumber="782" unit="mm" value="43.5">43.5 mm</quantity>
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||
; width
|
||
<quantity id="18582884FFC5FFFAFD4DFCAA615FD82C" box="[746,834,859,880]" metricMagnitude="-2" metricUnit="m" metricValue="1.88" pageId="2" pageNumber="782" unit="mm" value="18.8">18.8 mm</quantity>
|
||
. Color of pronotum and elytra dark reddish brown with fuscous and black margins and black longitudinally along center of pronotum (including horn) (
|
||
<figureCitation id="479B99E4FFC5FFFAFCFCFC5B6188D8E3" box="[859,917,938,959]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="2" pageNumber="782">Fig. 1</figureCitation>
|
||
). Head, horns, scutellum, elytral suture, pygidium, and legs black.
|
||
<emphasis id="EDD45973FFC5FFFAFBF1FC34668ED886" bold="true" box="[1110,1171,965,986]" pageId="2" pageNumber="782">Head:</emphasis>
|
||
Frons densely, coarsely rugopunctate and with moderately dense, long, pale yellow setae. Frontoclypeal region with moderately long (
|
||
<quantity id="18582884FFC5FFFAFB93FBE46694DF76" box="[1076,1161,1045,1066]" metricMagnitude="-3" metricUnit="m" metricValue="9.3" pageId="2" pageNumber="782" unit="mm" value="9.3">9.3 mm</quantity>
|
||
), slender, strongly recurved horn with acute apex (
|
||
<figureCitation id="479B99E4FFC5FFFAFD13FBBB60F2DF03" box="[692,751,1098,1119]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="2" pageNumber="782">Fig. 3</figureCitation>
|
||
); basal half of horn posteriorly with sparse, moderately long setae. Clypeus tapering to narrow, emarginate apex; surface with small, dense punctures. Interocular width equals 1.8 transverse eye diameters. Antennal club subequal in length to antennomeres 2–7. Mandible with apex distinctly notched.
|
||
<emphasis id="EDD45973FFC5FFFAFCAAFB186166DFA2" bold="true" box="[781,891,1257,1278]" pageId="2" pageNumber="782">Pronotum:</emphasis>
|
||
<collectingRegion id="1D644B83FFC5FFFAFC2EFB1861D5DFA2" box="[905,968,1257,1278]" country="France" name="Centre" pageId="2" pageNumber="782">Center</collectingRegion>
|
||
of disc with short (
|
||
<quantity id="18582884FFC5FFFAFD13FAF5611CDE45" box="[692,769,1284,1305]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="782" unit="mm" value="4.5">4.5 mm</quantity>
|
||
), parallel, slightly curved horn or elongate knob (
|
||
<figureCitation id="479B99E4FFC5FFFAFD4BFAEF613BDE6F" box="[748,806,1310,1331]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="2" pageNumber="782">Fig. 3</figureCitation>
|
||
) projecting forward and only slightly upward at about 22° from plane of pronotal disc, apex simply, bluntly rounded; anterior surface of horn with dense, moderately long, pale yellow setae, and a narrow sulcus extending from anterior base of horn to anterior margin of pronotum. Pronotal surface either side of horn finely shagreened, with moderately large, moderately dense, round and crescent-shaped punctures, punctures becoming denser along lateral margin. Base with complete marginal bead.
|
||
<emphasis id="EDD45973FFC5FFFAFCE3F9D96191DD61" bold="true" box="[836,908,1576,1597]" pageId="2" pageNumber="782">Elytra:</emphasis>
|
||
Surface finely shagreened, weakly shiny either side of suture and with large, moderately dense, shallow punctures; surface elsewhere dull, with small punctures in rows. Sutural stria a strongly impressed line.
|
||
<emphasis id="EDD45973FFC5FFFAFBA0F963668EDDFB" bold="true" box="[1031,1171,1682,1703]" pageId="2" pageNumber="782">Propygidium:</emphasis>
|
||
Stridulatory ridges converging towards apex.
|
||
<typeStatus id="001B3BC3FFC4FFFBFD10F93A611EDD82" box="[695,771,1739,1758]" pageId="3" pageNumber="783" type="holotype">holotype</typeStatus>
|
||
;
|
||
<emphasis id="EDD45973FFC4FFFBFCB5F93A613EDD82" bold="true" box="[786,803,1739,1758]" pageId="3" pageNumber="783">2)</emphasis>
|
||
Female
|
||
<typeStatus id="001B3BC3FFC4FFFBFCDFF93A61A3DD82" box="[888,958,1739,1758]" pageId="3" pageNumber="783" type="allotype">allotype</typeStatus>
|
||
. Lateral habitus:
|
||
<emphasis id="EDD45973FFC4FFFBFBC6F93A666FDD82" bold="true" box="[1121,1138,1739,1758]" pageId="3" pageNumber="783">3)</emphasis>
|
||
Male
|
||
<emphasis id="EDD45973FFC3FFFCFF30FF33631DDB8B" bold="true" box="[151,256,194,215]" pageId="4" pageNumber="784">Pygidium:</emphasis>
|
||
Surface on basal third with small, dense, setigerous punctures; setae long, dense, tawny. Surface on apical 2/3 glabrous, shagreened, with small, moderately dense punctures. In lateral view, basal third weakly convex, apical 2/3 nearly flat and almost “retracted” beneath apical third.
|
||
<emphasis id="EDD45973FFC3FFFCFDE1FEB76060DA07" bold="true" box="[582,637,326,347]" pageId="4" pageNumber="784">Legs:</emphasis>
|
||
Protibia tridentate, basal tooth distinctly removed from others. Basal tarsomere of protarsus shorter than next 2 tarsomeres combined. Mesotibia at apex with 2 small, sharp teeth; 2 transverse carinae at middle of tibia obsolete. Basal tarsomere of meso- and metatarsi with apex extended into small spine.
|
||
<emphasis id="EDD45973FFC3FFFCFF30FDF062FFD94A" bold="true" box="[151,226,513,534]" pageId="4" pageNumber="784">Venter:</emphasis>
|
||
Prosternal process laminate, tapering, partially obscured by long, tawny setae, apex narrowly subtruncate.
|
||
<emphasis id="EDD45973FFC3FFFCFEFAFDC763CCD917" bold="true" box="[349,465,566,587]" pageId="4" pageNumber="784">Parameres:</emphasis>
|
||
In caudal view, form asymmetrical with left paramere broadly emarginate at center on inner surface (
|
||
<figureCitation id="479B99E4FFC3FFFCFD97FD9A606CD9DC" box="[560,625,619,640]" captionStart="Fig" captionStartId="4.[709,740,609,628]" captionTargetBox="[685,1172,194,594]" captionTargetId="figure-616@4.[685,1172,194,594]" captionTargetPageId="4" captionText="Fig. 5. Golofa limogesi, parameres, lateral and caudal views." figureDoi="http://doi.org/10.5281/zenodo.5378595" httpUri="https://zenodo.org/record/5378595/files/figure.png" pageId="4" pageNumber="784">Fig. 5</figureCitation>
|
||
). Apices with long, dense, tawny setae. In lateral view, basal piece noticeably flattened on ventral surface.
|
||
</paragraph>
|
||
<caption id="8BDFD5E9FFC4FFFBFF62F93A63B7DDAA" ID-DOI="http://doi.org/10.5281/zenodo.5378593" ID-Zenodo-Dep="5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="3" pageNumber="783" startId="3.[197,236,1739,1758]" targetBox="[178,1187,194,1723]" targetPageId="3">
|
||
<paragraph id="DF1F8561FFC4FFFBFF62F93A63B7DDAA" blockId="3.[173,684,1738,1782]" pageId="3" pageNumber="783">
|
||
<emphasis id="EDD45973FFC4FFFBFF62F93A6302DD82" bold="true" box="[197,287,1739,1758]" pageId="3" pageNumber="783">Figs. 1–4.</emphasis>
|
||
<taxonomicName id="18A0FEE2FFC4FFFBFE92F93B63DFDD81" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[309,450,1738,1757]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="783" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC4FFFBFE92F93B63DFDD81" box="[309,450,1738,1757]" italics="true" pageId="3" pageNumber="783">Golofa limogesi</emphasis>
|
||
</taxonomicName>
|
||
. Dorsal habitus:
|
||
<emphasis id="EDD45973FFC4FFFBFDC3F93A6068DD82" bold="true" box="[612,629,1739,1758]" pageId="3" pageNumber="783">1)</emphasis>
|
||
Male holotype;
|
||
<emphasis id="EDD45973FFC4FFFBFEA4F9126309DDAA" bold="true" box="[259,276,1763,1782]" pageId="3" pageNumber="783">4)</emphasis>
|
||
Female allotype.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8BDFD5E9FFC3FFFCFD62FD90613ED9D0" ID-DOI="http://doi.org/10.5281/zenodo.5378595" ID-Zenodo-Dep="5378595" httpUri="https://zenodo.org/record/5378595/files/figure.png" pageId="4" pageNumber="784" startId="4.[709,740,609,628]" targetBox="[685,1172,194,594]" targetPageId="4">
|
||
<paragraph id="DF1F8561FFC3FFFCFD62FD90613ED9D0" blockId="4.[685,1172,609,652]" pageId="4" pageNumber="784">
|
||
<emphasis id="EDD45973FFC3FFFCFD62FD906115D928" bold="true" box="[709,776,609,628]" pageId="4" pageNumber="784">Fig. 5.</emphasis>
|
||
<taxonomicName id="18A0FEE2FFC3FFFCFCBAFD9061B2D928" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[797,943,609,628]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="784" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC3FFFCFCBAFD9061B2D928" box="[797,943,609,628]" italics="true" pageId="4" pageNumber="784">Golofa limogesi</emphasis>
|
||
</taxonomicName>
|
||
, parameres, lateral and caudal views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="DF1F8561FFC3FFFCFF08FD2462F9DFF2" blockId="4.[151,639,194,1730]" pageId="4" pageNumber="784">
|
||
<emphasis id="EDD45973FFC3FFFCFF08FD24637BD9B6" bold="true" box="[175,358,725,746]" pageId="4" pageNumber="784">
|
||
<typeStatus id="001B3BC3FFC3FFFCFF08FD246319D9B6" box="[175,260,725,746]" pageId="4" pageNumber="784" type="allotype">Allotype</typeStatus>
|
||
Female.
|
||
</emphasis>
|
||
Length
|
||
<quantity id="18582884FFC3FFFCFE69FD24602FD9B6" box="[462,562,725,746]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="4" pageNumber="784" unit="mm" value="38.5">38.5 mm</quantity>
|
||
; width
|
||
<quantity id="18582884FFC3FFFCFF30FD0162F2D859" box="[151,239,752,773]" metricMagnitude="-2" metricUnit="m" metricValue="1.68" pageId="4" pageNumber="784" unit="mm" value="16.8">16.8 mm</quantity>
|
||
. As
|
||
<typeStatus id="001B3BC3FFC3FFFCFEB0FD016377D859" box="[279,362,752,773]" pageId="4" pageNumber="784" type="holotype">holotype</typeStatus>
|
||
male except in the following respects. Color of head and pronotum black (
|
||
<figureCitation id="479B99E4FFC3FFFCFD9EFCFB606FD843" box="[569,626,778,799]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="4" pageNumber="784">Fig. 2</figureCitation>
|
||
).
|
||
<emphasis id="EDD45973FFC3FFFCFF30FCD462C8D866" bold="true" box="[151,213,805,826]" pageId="4" pageNumber="784">Head:</emphasis>
|
||
Frons and clypeus densely, coarsely rugopunctate; frons with a patch of long, reddish brown setae on both sides of middle. Horn absent (
|
||
<figureCitation id="479B99E4FFC3FFFCFD9FFCAB606FD833" box="[568,626,858,879]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="4" pageNumber="784">Fig. 4</figureCitation>
|
||
), but frons with low, transverse tubercle. Clypeus tapering to narrowly subtruncate apex. Interocular width equals 2.1 transverse eye diameters.
|
||
<emphasis id="EDD45973FFC3FFFCFDB4FC5B6063D8E3" bold="true" box="[531,638,938,959]" pageId="4" pageNumber="784">Pronotum:</emphasis>
|
||
Horn absent. Surface with large, dense (mostly confluent on apical half), deep, round punctures.
|
||
<emphasis id="EDD45973FFC3FFFCFF30FC0B631DDF53" bold="true" box="[151,256,1018,1039]" pageId="4" pageNumber="784">Pygidium:</emphasis>
|
||
Surface on apical 2/3 not as strongly “retracted” beneath apical third.
|
||
<emphasis id="EDD45973FFC3FFFCFE4AFBE56039DF75" bold="true" box="[493,548,1044,1065]" pageId="4" pageNumber="784">Legs:</emphasis>
|
||
Protibia quadridentate, all teeth subequally spaced from one another. Basal tarsomere of protarsus slightly longer than tarsomere 2. Mesotibia at apex with 2 large, sharp teeth; 2 transverse carinae at middle of tibia distinct.
|
||
</paragraph>
|
||
<paragraph id="DF1F8561FFC3FFFCFF08FB45668CD80A" blockId="4.[151,639,194,1730]" lastBlockId="4.[685,1173,700,1226]" pageId="4" pageNumber="784">
|
||
<emphasis id="EDD45973FFC3FFFCFF08FB456347DF95" bold="true" box="[175,346,1204,1225]" pageId="4" pageNumber="784">
|
||
<typeStatus id="001B3BC3FFC3FFFCFF08FB456309DF95" box="[175,276,1204,1225]" pageId="4" pageNumber="784" type="paratype">Paratypes</typeStatus>
|
||
. Males
|
||
</emphasis>
|
||
(
|
||
<emphasis id="EDD45973FFC3FFFCFEC1FB45636FDF95" box="[358,370,1204,1225]" italics="true" pageId="4" pageNumber="784">n</emphasis>
|
||
=7). Length 37.0–
|
||
<quantity id="18582884FFC3FFFCFD82FB456067DF95" box="[549,634,1204,1225]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="4" pageNumber="784" unit="mm" value="42.9">42.9 mm</quantity>
|
||
; width 17.3–20.0 mm. Very nearly identical with
|
||
<typeStatus id="001B3BC3FFC3FFFCFF30FB1862F1DFA2" box="[151,236,1257,1278]" pageId="4" pageNumber="784" type="holotype">holotype</typeStatus>
|
||
except in the following respects. Body length varies slightly. Color of pronotum dark reddish brown with black as type or only on horn and a spot on lateral margin at widest point.
|
||
<emphasis id="EDD45973FFC3FFFCFD98FAC86060DE12" bold="true" box="[575,637,1337,1358]" pageId="4" pageNumber="784">Head:</emphasis>
|
||
Frontoclypeal horn
|
||
<quantity id="18582884FFC3FFFCFEC7FAA263B3DE34" box="[352,430,1363,1384]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="4" pageNumber="784" unit="mm" value="4.9">4.9 mm</quantity>
|
||
in smallest male to
|
||
<quantity id="18582884FFC3FFFCFF30FA9F62F3DEDF" box="[151,238,1390,1411]" metricMagnitude="-2" metricUnit="m" metricValue="1.41" pageId="4" pageNumber="784" unit="mm" value="14.1">14.1 mm</quantity>
|
||
in largest male. Interocular width equals 1.5 transverse eye diameters.
|
||
<emphasis id="EDD45973FFC3FFFCFE68FA796022DEC1" bold="true" box="[463,575,1416,1437]" pageId="4" pageNumber="784">Pronotum:</emphasis>
|
||
Horn varies in length from
|
||
<quantity id="18582884FFC3FFFCFEC8FA5263A7DEE4" box="[367,442,1443,1464]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="784" unit="mm" value="2.5">2.5 mm</quantity>
|
||
in smallest male to
|
||
<quantity id="18582884FFC3FFFCFF30FA4F62F9DE8F" box="[151,228,1470,1491]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" pageId="4" pageNumber="784" unit="mm" value="8.3">8.3 mm</quantity>
|
||
in largest male and varies in angle from projecting forward and only slightly upward at about 20° from plane of pronotal disc in minor male to projecting at 45° from plane of pronotal disc in major male.
|
||
<emphasis id="EDD45973FFC3FFFCFEF8F9D963D7DD61" bold="true" box="[351,458,1576,1597]" pageId="4" pageNumber="784">Pygidium:</emphasis>
|
||
Surface on apical 2/3 not shagreened.
|
||
<emphasis id="EDD45973FFC3FFFCFEF0F9B26392DD04" bold="true" box="[343,399,1603,1624]" pageId="4" pageNumber="784">Legs:</emphasis>
|
||
Protibia tridentate, basal tooth only slightly removed from others. Mesotibia at apex with 1 large and 2 small, sharp teeth; 2 transverse carinae at middle of tibia distinct.
|
||
<emphasis id="EDD45973FFC3FFFCFF30F95C62F6DD9E" bold="true" box="[151,235,1709,1730]" pageId="4" pageNumber="784">Females</emphasis>
|
||
(
|
||
<emphasis id="EDD45973FFC3FFFCFF5BF95C6315DD9E" box="[252,264,1709,1730]" italics="true" pageId="4" pageNumber="784">n</emphasis>
|
||
= 10). Length 37.0–
|
||
<quantity id="18582884FFC3FFFCFE78F95C6027DD9E" box="[479,570,1709,1730]" metricMagnitude="-2" metricUnit="m" metricValue="4.29" pageId="4" pageNumber="784" unit="mm" value="42.9">42.9 mm</quantity>
|
||
; width 17.3–20.0 mm. The female
|
||
<typeStatus id="001B3BC3FFC3FFFCFC1EFD4D660BD98D" box="[953,1046,700,721]" pageId="4" pageNumber="784" type="paratype">paratypes</typeStatus>
|
||
do not differ significantly from the
|
||
<typeStatus id="001B3BC3FFC3FFFCFC3FFD2661F4D9B0" box="[920,1001,727,748]" pageId="4" pageNumber="784" type="allotype">allotype</typeStatus>
|
||
. Two specimens have reddish brown on the basal half of the pronotum, while the remainder is black. Tubercle on the frons varies from virtually obsolete to pronounced, conical to slightly, transversely curved.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="97BAD6EAFFC3FFFCFD62FCAD6690D89C" pageId="4" pageNumber="784" type="etymology">
|
||
<paragraph id="DF1F8561FFC3FFFCFD62FCAD6690D89C" blockId="4.[685,1173,700,1226]" pageId="4" pageNumber="784">
|
||
<emphasis id="EDD45973FFC3FFFCFD62FCAD6125D82D" bold="true" box="[709,824,860,881]" pageId="4" pageNumber="784">Etymology.</emphasis>
|
||
We are pleased to name this species for René Limoges in grateful recognition of his skill and generosity in providing superb images of beetles over the years to both of us for our work.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="97BAD6EAFFC3FFFDFD62FC3763BED81B" lastPageId="5" lastPageNumber="785" pageId="4" pageNumber="784" type="distribution">
|
||
<paragraph id="DF1F8561FFC3FFFDFD62FC3763BED81B" blockId="4.[685,1173,700,1226]" lastBlockId="5.[173,660,791,1477]" lastPageId="5" lastPageNumber="785" pageId="4" pageNumber="784">
|
||
<emphasis id="EDD45973FFC3FFFCFD62FC37615BD887" bold="true" box="[709,838,966,987]" pageId="4" pageNumber="784">Distribution.</emphasis>
|
||
<taxonomicName id="18A0FEE2FFC3FFFCFCF2FC3461EBD887" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[853,1014,965,987]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="784" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC3FFFCFCF2FC3461EBD887" box="[853,1014,965,987]" italics="true" pageId="4" pageNumber="784">Golofa limogesi</emphasis>
|
||
</taxonomicName>
|
||
is, at present, known only from above the town of Ayabaca in Ayabaca Province in the
|
||
<collectingRegion id="1D644B83FFC3FFFCFC09FC0A662DDF4C" box="[942,1072,1019,1040]" country="Peru" name="Piura" pageId="4" pageNumber="784">Piura Region</collectingRegion>
|
||
of northwestern
|
||
<collectingCountry id="A7B7C5F1FFC3FFFCFD5CFBE7613BDF77" box="[763,806,1046,1067]" name="Peru" pageId="4" pageNumber="784">Peru</collectingCountry>
|
||
(
|
||
<figureCitation id="479B99E4FFC3FFFCFC95FBE76177DF77" box="[818,874,1046,1067]" captionStart="Fig" captionStartId="4.[709,740,1711,1730]" captionTargetBox="[685,1172,1286,1694]" captionTargetId="figure-627@4.[685,1172,1286,1694]" captionTargetPageId="4" captionText="Fig. 6. Golofa limogesi, distribution." figureDoi="http://doi.org/10.5281/zenodo.5378597" httpUri="https://zenodo.org/record/5378597/files/figure.png" pageId="4" pageNumber="784">Fig. 6</figureCitation>
|
||
). Ayabaca is the highest town in the region at
|
||
<quantity id="18582884FFC3FFFCFC8EFBC16167DF19" box="[809,890,1072,1093]" metricMagnitude="3" metricUnit="m" metricValue="2.7" pageId="4" pageNumber="784" unit="m" value="2700.0">2,700 m</quantity>
|
||
. Most, though not all, species of
|
||
<taxonomicName id="18A0FEE2FFC3FFFCFD60FBBB6114DF3C" authorityName="Hope" authorityYear="1837" box="[711,777,1098,1120]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="784" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="EDD45973FFC3FFFCFD60FBBB6114DF3C" box="[711,777,1098,1120]" italics="true" pageId="4" pageNumber="784">Golofa</emphasis>
|
||
</taxonomicName>
|
||
live at elevations higher than
|
||
<quantity id="18582884FFC3FFFCFB85FBBA6672DF3C" box="[1058,1135,1099,1120]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="4" pageNumber="784" unit="m" value="1500.0">1,500 m</quantity>
|
||
but usually less than
|
||
<quantity id="18582884FFC3FFFCFCECFB946187DF26" box="[843,922,1125,1146]" metricMagnitude="3" metricUnit="m" metricValue="2.5" pageId="4" pageNumber="784" unit="m" value="2500.0">2,500 m</quantity>
|
||
. This locality is adjacent to
|
||
<collectingRegion id="1D644B83FFC3FFFCFD0AFB716127DFC9" box="[685,826,1152,1173]" country="Ecuador" name="Loja" pageId="4" pageNumber="784">Loja Province</collectingRegion>
|
||
in
|
||
<collectingCountry id="A7B7C5F1FFC3FFFCFCCFFB7161AADFC9" box="[872,951,1152,1173]" name="Ecuador" pageId="4" pageNumber="784">Ecuador</collectingCountry>
|
||
where BCR, Ronald Cave, and Aura Paucar are engaged in a biodiversity inventory of the Dynastinae of
|
||
<collectingCountry id="A7B7C5F1FFC3FFFCFB9BFB44668CDF96" box="[1084,1169,1205,1226]" name="Ecuador" pageId="4" pageNumber="784">Ecuador</collectingCountry>
|
||
. Accordingly,
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFE94FCE663B8D871" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[307,421,791,813]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC2FFFDFE94FCE663B8D871" box="[307,421,791,813]" italics="true" pageId="5" pageNumber="785">G. limogesi</emphasis>
|
||
</taxonomicName>
|
||
will be sought after in nearby southern
|
||
<collectingCountry id="A7B7C5F1FFC2FFFDFEE9FCC36382D81B" box="[334,415,818,839]" name="Ecuador" pageId="5" pageNumber="785">Ecuador</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="8BDFD5E9FFC3FFFCFD62F95E660DDD9E" ID-DOI="http://doi.org/10.5281/zenodo.5378597" ID-Zenodo-Dep="5378597" box="[709,1040,1710,1730]" httpUri="https://zenodo.org/record/5378597/files/figure.png" pageId="4" pageNumber="784" startId="4.[709,740,1711,1730]" targetBox="[685,1172,1286,1694]" targetPageId="4">
|
||
<paragraph id="DF1F8561FFC3FFFCFD62F95E660DDD9E" blockId="4.[709,1040,1710,1730]" box="[709,1040,1710,1730]" pageId="4" pageNumber="784">
|
||
<emphasis id="EDD45973FFC3FFFCFD62F95E60E3DD9E" bold="true" box="[709,766,1711,1730]" pageId="4" pageNumber="784">Fig. 6.</emphasis>
|
||
<taxonomicName id="18A0FEE2FFC3FFFCFCB3F95F6181DD9D" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[788,924,1710,1729]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="784" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC3FFFCFCB3F95F6181DD9D" box="[788,924,1710,1729]" italics="true" pageId="4" pageNumber="784">Golofa limogesi</emphasis>
|
||
</taxonomicName>
|
||
, distribution.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="8BDFD5E9FFC2FFFDFF62FD3E6013D9BE" ID-DOI="http://doi.org/10.5281/zenodo.5378599" ID-Zenodo-Dep="5378599" box="[197,526,718,738]" httpUri="https://zenodo.org/record/5378599/files/figure.png" pageId="5" pageNumber="785" startId="5.[197,228,719,738]" targetBox="[173,1195,194,703]" targetPageId="5">
|
||
<paragraph id="DF1F8561FFC2FFFDFF62FD3E6013D9BE" blockId="5.[197,526,718,738]" box="[197,526,718,738]" pageId="5" pageNumber="785">
|
||
<emphasis id="EDD45973FFC2FFFDFF62FD3E62E0D9BE" bold="true" box="[197,253,719,738]" pageId="5" pageNumber="785">Fig. 7.</emphasis>
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFEB4FD3F6399D9BD" box="[275,388,718,737]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="eacus">
|
||
<emphasis id="EDD45973FFC2FFFDFEB4FD3F6399D9BD" box="[275,388,718,737]" italics="true" pageId="5" pageNumber="785">Golofa eacus</emphasis>
|
||
</taxonomicName>
|
||
, lateral habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="97BAD6EAFFC2FFFDFF62FCBD63E8DE98" pageId="5" pageNumber="785" type="diagnosis">
|
||
<paragraph id="DF1F8561FFC2FFFDFF62FCBD63E8DE98" blockId="5.[173,660,791,1477]" pageId="5" pageNumber="785">
|
||
<emphasis id="EDD45973FFC2FFFDFF62FCBD6331D83D" bold="true" box="[197,300,844,865]" pageId="5" pageNumber="785">Diagnosis.</emphasis>
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFE95FCBD63D7D83E" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[306,458,844,866]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC2FFFDFE95FCBD63D7D83E" box="[306,458,844,866]" italics="true" pageId="5" pageNumber="785">Golofa limogesi</emphasis>
|
||
</taxonomicName>
|
||
males are distinctive because of the forward projecting pronotal horn or knob that is shorter than or equal to (in major males) the frontal horn, notched mandibular apex, elytra shiny next to the suture but dull elsewhere, black scutellum, and form of the parameres. The low, forward-projecting, curved pronotal horn resembles that of the South American
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFE61FBF762EFDF6A" authority="Burmeister" authorityName="Burmeister" authorityYear="1847" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="pelagon">
|
||
<emphasis id="EDD45973FFC2FFFDFE61FBF76043DF40" box="[454,606,1030,1052]" italics="true" pageId="5" pageNumber="785">Golofa pelagon</emphasis>
|
||
Burmeister
|
||
</taxonomicName>
|
||
and the Central American
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFE59FBD1637DDF0D" authority="Dechambre" authorityName="Dechambre" authorityYear="1975" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="obliquicornis">
|
||
<emphasis id="EDD45973FFC2FFFDFE59FBD162F5DF0D" italics="true" pageId="5" pageNumber="785">Golofa obliquicornis</emphasis>
|
||
Dechambre
|
||
</taxonomicName>
|
||
, but the parameres are different from those species. The parameres are nearly identical to those of the common and sympatric
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFF0AFB7B6360DFFC" authority="(Drury)" baseAuthorityName="Drury" box="[173,381,1162,1184]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="eacus">
|
||
<emphasis id="EDD45973FFC2FFFDFF0AFB7B6331DFFC" box="[173,300,1162,1184]" italics="true" pageId="5" pageNumber="785">Golofa eacus</emphasis>
|
||
(Drury)
|
||
</taxonomicName>
|
||
, but the form of the pronotal horn is different. In
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFEC1FB5463CFDFE7" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[358,466,1189,1211]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC2FFFDFEC1FB5463CFDFE7" box="[358,466,1189,1211]" italics="true" pageId="5" pageNumber="785">G. limogesi</emphasis>
|
||
</taxonomicName>
|
||
, the pronotal horn is stout, knob-like, and projects forwards in all but the most developed males where the horn projects forwards and slightly upwards (
|
||
<figureCitation id="479B99E4FFC2FFFDFE79FB076005DE57" box="[478,536,1270,1291]" captionStart="Figs" captionStartId="3.[197,236,1739,1758]" captionTargetBox="[178,1187,194,1723]" captionTargetId="figure-0@3.[178,1187,194,1723]" captionTargetPageId="3" captionText="Figs. 1–4. Golofa limogesi. Dorsal habitus: 1) Male holotype; 4) Female allotype." figureDoi="http://doi.org/10.5281/zenodo.5378593" httpUri="https://zenodo.org/record/5378593/files/figure.png" pageId="5" pageNumber="785">Fig. 3</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFF0AFAFE62E3DE79" box="[173,254,1295,1317]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="eacus">
|
||
<emphasis id="EDD45973FFC2FFFDFF0AFAFE62E3DE79" box="[173,254,1295,1317]" italics="true" pageId="5" pageNumber="785">G. eacus</emphasis>
|
||
</taxonomicName>
|
||
the pronotal horn is slender and erect (even in minor forms) (
|
||
<figureCitation id="479B99E4FFC2FFFDFEE9FADA639ADE1C" box="[334,391,1323,1344]" captionStart="Fig" captionStartId="5.[197,228,719,738]" captionTargetBox="[173,1195,194,703]" captionTargetId="figure-427@5.[173,1195,194,703]" captionTargetPageId="5" captionText="Fig. 7. Golofa eacus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.5378599" httpUri="https://zenodo.org/record/5378599/files/figure.png" pageId="5" pageNumber="785">Fig. 7</figureCitation>
|
||
). Both males and females of
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFF0AFAB56307DE06" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[173,282,1348,1370]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC2FFFDFF0AFAB56307DE06" box="[173,282,1348,1370]" italics="true" pageId="5" pageNumber="785">G. limogesi</emphasis>
|
||
</taxonomicName>
|
||
may be distinguished from
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFD85FAB56068DE06" box="[546,629,1348,1370]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="eacus">
|
||
<emphasis id="EDD45973FFC2FFFDFD85FAB56068DE06" box="[546,629,1348,1370]" italics="true" pageId="5" pageNumber="785">G. eacus</emphasis>
|
||
</taxonomicName>
|
||
by the color of the scutellum (black in
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFDBCFAAE6090DE29" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" box="[539,653,1375,1397]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC2FFFDFDBCFAAE6090DE29" box="[539,653,1375,1397]" italics="true" pageId="5" pageNumber="785">G. limogesi</emphasis>
|
||
</taxonomicName>
|
||
, reddish brown in
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFEC6FA8B63A4DECC" box="[353,441,1402,1424]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="eacus">
|
||
<emphasis id="EDD45973FFC2FFFDFEC6FA8B63A4DECC" box="[353,441,1402,1424]" italics="true" pageId="5" pageNumber="785">G. eacus</emphasis>
|
||
</taxonomicName>
|
||
), and the stridulatory bands on the propygidium (converging in
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFDDBFA6562E0DE99" authorityName="Ratcliffe and Le Tirant" authorityYear="2017" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="limogesi">
|
||
<emphasis id="EDD45973FFC2FFFDFDDBFA6562E0DE99" italics="true" pageId="5" pageNumber="785">G. limogesi</emphasis>
|
||
</taxonomicName>
|
||
but parallel in
|
||
<taxonomicName id="18A0FEE2FFC2FFFDFE34FA5E63F5DE99" box="[403,488,1455,1477]" class="Insecta" family="Dynastidae" genus="Golofa" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="785" phylum="Arthropoda" rank="species" species="eacus">
|
||
<emphasis id="EDD45973FFC2FFFDFE34FA5E63F5DE99" box="[403,488,1455,1477]" italics="true" pageId="5" pageNumber="785">G. eacus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |