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<document id="FA14E23F8FD98DA9802342882661837E" ID-DOI="10.5281/zenodo.5392079" ID-ISSN="1638-9387" ID-Zenodo-Dep="5392079" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630316032044" checkinUser="marcus" docAuthor="Golovatch, Sergei I., Geoffroy, Jean-Jacques, Mauriès, Jean-Paul &amp; Spiegel, Didier Van Den" docDate="2007" docId="03F4C378A2383051FCB302D1FB9AFA99" docLanguage="en" docName="Zoosystema.2007.29.1.7-49.pdf" docOrigin="Zoosystema 29 (1)" docStyle="DocumentStyle:2A9C6F9EA5F90D98170D9E0D9CA29375.4:Zoosystema.2000-2008.journal_article.type1" docStyleId="2A9C6F9EA5F90D98170D9E0D9CA29375" docStyleName="Zoosystema.2000-2008.journal_article.type1" docStyleVersion="4" docTitle="Glyphiulus bedosae Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel 2007, n. sp." docType="treatment" docVersion="3" lastPageNumber="32" masterDocId="FFCDBB00A22F3048FF94075DFFE6FF91" masterDocTitle="Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group" masterLastPageNumber="49" masterPageNumber="7" pageNumber="30" updateTime="1699223690750" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0" zenodo-license-treatments="UNSPECIFIED">
<mods:mods id="CC51346C94768BF5949C69316AB9E3E0" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="6E6AC1CC742BEDAF5844B22A3137D8D6">
<mods:title id="D1AB137F98B1E9F6FA73719875B4CEE4">Review of the millipede genus Glyphiulus Gervais, 1847, with descriptions of new species from Southeast Asia (Diplopoda, Spirostreptida, Cambalopsidae). Part 1: the granulatus-group</mods:title>
</mods:titleInfo>
<mods:name id="8895E5A06B886E69084FD86C6D0C8710" type="personal">
<mods:role id="A55B47C4AC34AE40A70A731BD3BCC807">
<mods:roleTerm id="328FC580E9FC83F7D0BCE1EBF3BF6368">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="85836D58418B147809996CCCF4F7C2BF">Golovatch, Sergei I.</mods:namePart>
<mods:affiliation id="581E3DB2770155F4554A99F847F256EC">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky pr. 33, Moscow 119071 (Russia) sgol @ orc. ru</mods:affiliation>
<mods:nameIdentifier id="CE28189F453F77CA8E97DBD07EED141B" type="email">sgol@orc.ru</mods:nameIdentifier>
</mods:name>
<mods:name id="484CE0A1438EAC2FA5C9BFB8B54B9B76" type="personal">
<mods:role id="2D5AC9B612E37F7EB454C4E6B9F4D33B">
<mods:roleTerm id="A07F3B579C63BC5746C4275C8D4574E3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7180D9320AE8D700B7F6788F6797887D">Geoffroy, Jean-Jacques</mods:namePart>
<mods:affiliation id="723769C6D1A3FDA1FB11D46C6975DB76">Muséum national d’Histoire naturelle, Département Écologie et Gestion de la Biodiversité, USM 306 Réseaux trophiques du sol, 4 avenue du Petit Château, F- 91800 Brunoy (France) geoffroy @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="C2CA5B388232365BBBD8DEBB4B08405C" type="email">geoffroy@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="947297F06CA967554F3F7A1EDC61CC72" type="personal">
<mods:role id="58C135EB0476C02888555714EC945705">
<mods:roleTerm id="51B14CD2CD25BF6027F70ED239D635EB">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD02E8262255F9F147E97AD96117919E">Mauriès, Jean-Paul</mods:namePart>
<mods:affiliation id="E3A19C35867DDDCCD42C5F905FBDB2C8">Muséum national d’Histoire naturelle, Département Systématique et Évolution, USM 602, case postale 53, 57 rue Cuvier, F- 75231 Paris cedex 05 (France) collmill @ mnhn. fr</mods:affiliation>
<mods:nameIdentifier id="EC1CBD934440E407DFC54D4E1E7D5A46" type="email">collmill@mnhn.fr</mods:nameIdentifier>
</mods:name>
<mods:name id="6D6B9C4916730A316443D0A3A87E8BF5" type="personal">
<mods:role id="E58B6D0D3B1EF5D1C5BCA7BF263C33C1">
<mods:roleTerm id="65A6BC14CE0BD820470FB84A6FDCB3D3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7BDF5E6287B803887499EE190EDDB9DE">Spiegel, Didier Van Den</mods:namePart>
<mods:affiliation id="A0EAD7B505B927746DBE0079D361B0D3">Musée royal de l’Afrique centrale, Leuvensesteenweg 13, B- 3080 Tervuren (Belgium) didier. van. den. spiegel @ africamuseum. be</mods:affiliation>
</mods:name>
<mods:typeOfResource id="F605145891E4526CDFDB5916B57FC390">text</mods:typeOfResource>
<mods:relatedItem id="6E0D2783F5D12DA6779FFCAB9742AC49" type="host">
<mods:titleInfo id="52B0781B487E606C87CC62FBA0637BBB">
<mods:title id="750266F5CC3CAD0DA6D7A0D058CF1A73">Zoosystema</mods:title>
</mods:titleInfo>
<mods:part id="3EEE02EAE318243E0F0723EF5B8185B1">
<mods:date id="4957D062D871BF109EED86497763B0B2">2007</mods:date>
<mods:detail id="10D50327BE78CAF1725CAADD5814826A" type="volume">
<mods:number id="E15FCA9CE6B2D6532E476BFB5CDE9EF9">29</mods:number>
</mods:detail>
<mods:detail id="176179B78CD537F044199FF718378B69" type="issue">
<mods:number id="62FB65512388DDD2A3801E720E974171">1</mods:number>
</mods:detail>
<mods:extent id="D06F4CAF52DBCC7B8BB4BDC1AF7C785C" unit="page">
<mods:start id="F705014D208E99F46FADF5202FE2BF4C">7</mods:start>
<mods:end id="D5DACC60EA713B8D2AE2EDB7E192046A">49</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9468C18AD377B08F167A51A4DF0B4BD0">journal article</mods:classification>
<mods:identifier id="92C3C7B204446F40E907158367E12835" type="DOI">10.5281/zenodo.5392079</mods:identifier>
<mods:identifier id="F3D0A211A858FCE3AD36CE3215DEFE9A" type="ISSN">1638-9387</mods:identifier>
<mods:identifier id="11F5ACCDE114C4A08612E1CA27393D99" type="Zenodo-Dep">5392079</mods:identifier>
</mods:mods>
<treatment id="03F4C378A2383051FCB302D1FB9AFA99" LSID="urn:lsid:plazi:treatment:03F4C378A2383051FCB302D1FB9AFA99" httpUri="http://treatment.plazi.org/id/03F4C378A2383051FCB302D1FB9AFA99" lastPageId="25" lastPageNumber="32" pageId="23" pageNumber="30">
<subSubSection id="C34721E5A238305FFCB302D1FBD1FA37" box="[807,1079,1420,1446]" pageId="23" pageNumber="30" type="nomenclature">
<paragraph id="8BE2726EA238305FFCB302D1FBD1FA37" blockId="23.[807,1079,1420,1478]" box="[807,1079,1420,1446]" pageId="23" pageNumber="30">
<heading id="D0AAC502A238305FFCB302D1FBD1FA37" box="[807,1079,1420,1446]" centered="true" fontSize="11" level="2" pageId="23" pageNumber="30" reason="2">
<taxonomicName id="4C5D09EDA238305FFCB302D1FC10FA37" ID-CoL="3GGJH" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[807,1014,1420,1446]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="23" pageNumber="30" phylum="Arthropoda" rank="species" species="bedosae" status="sp. nov.">
<emphasis id="B929AE7CA238305FFCB302D1FC10FA37" bold="true" box="[807,1014,1420,1446]" italics="true" pageId="23" pageNumber="30">Glyphiulus bedosae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A238305FFC6902D1FBD1FA37" box="[1021,1079,1420,1446]" pageId="23" pageNumber="30" rank="species">n. sp.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A238305FFCFE02F1FC12FA57" box="[874,1012,1452,1478]" pageId="23" pageNumber="30" type="description">
<paragraph id="8BE2726EA238305FFCFE02F1FC12FA57" blockId="23.[807,1079,1420,1478]" box="[874,1012,1452,1478]" pageId="23" pageNumber="30">
(
<figureCitation id="13666EEBA238305FFCE502F1FC25FA57" box="[881,963,1452,1478]" captionStart="FIG" captionStartId="22.[113,124,1345,1361]" captionTargetBox="[121,1201,215,1301]" captionTargetId="figure-44@22.[391,664,758,940]" captionTargetPageId="22" captionText="FIG. 17. — Glyphiulus bedosae n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, lateral view; D, same, dorsal view; E, caudal part of body, lateral view; F, telson, ventral view. Scale bars: A, C, 0.5 mm; B, D, E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392115" httpUri="https://zenodo.org/record/5392115/files/figure.png" pageId="23" pageNumber="30">Figs 17</figureCitation>
;
<figureCitation id="13666EEBA238305FFC5B02F1FC0DFA57" box="[975,1003,1452,1478]" captionStart="FIG" captionStartId="23.[113,124,1189,1205]" captionTargetBox="[110,1225,182,1181]" captionTargetId="figure-14@23.[110,1225,182,1181]" captionTargetPageId="23" captionText="FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="23" pageNumber="30">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A238305FFD3102BDFD0DF932" pageId="23" pageNumber="30" type="materials_examined">
<paragraph id="8BE2726EA238305FFD3102BDFD0DF932" blockId="23.[674,1210,1504,1699]" pageId="23" pageNumber="30">
<materialsCitation id="3B357833A238305FFD3102BDFC86F9D5" collectingDate="2000-12-29" collectionCode="MNHN" collectorName="A. Bedos &amp; L. Deharveng" country="Laos" location="Tham Pha Kouang Cave" municipality="Nong Kiaw" pageId="23" pageNumber="30" specimenCode="GA 039" specimenCount="1" specimenCount-male="1" stateProvince="Louangphabang" typeStatus="holotype">
TYPE MATERIAL. —
<emphasis id="B929AE7CA238305FFCC102BCFC61FA69" bold="true" box="[853,903,1505,1528]" pageId="23" pageNumber="30">
<collectingCountry id="F34A32FEA238305FFCC102BCFC65FA69" box="[853,899,1505,1528]" name="Laos" pageId="23" pageNumber="30">Laos</collectingCountry>
.
</emphasis>
<collectingRegion id="4999BC8CA238305FFC1F02BCFBACFA69" box="[907,1098,1505,1528]" country="Laos" name="Louangphabang" pageId="23" pageNumber="30">Luang Prabang Prov.</collectingRegion>
,
<collectingMunicipality id="6B86E814A238305FFBC402BCFB5CFA69" box="[1104,1210,1505,1528]" pageId="23" pageNumber="30">Nong Kiaw</collectingMunicipality>
(= Muang Ngoy),
<location id="8E8224B5A238305FFCD802A7FBD2F980" LSID="urn:lsid:plazi:treatment:03F4C378A2383051FCB302D1FB9AFA99:8E8224B5A238305FFCD802A7FBD2F980" box="[844,1076,1530,1553]" country="Laos" municipality="Nong Kiaw" name="Tham Pha Kouang Cave" pageId="23" pageNumber="30" stateProvince="Louangphabang">Tham Pha Kouang Cave</location>
,
<date id="FFE354AEA238305FFBA802A7FB50F980" box="[1084,1206,1530,1553]" pageId="23" pageNumber="30" value="2000-12-29">
<collectingDate id="EFA7AD46A238305FFBA802A7FB50F980" box="[1084,1206,1530,1553]" pageId="23" pageNumber="30" value="2000-12-29">29.XII.2000</collectingDate>
</date>
, leg.
<collectorName id="26A817B8A238305FFD5E0149FCC7F9BA" box="[714,801,1556,1579]" pageId="23" pageNumber="30">A. Bedos</collectorName>
&amp;
<collectorName id="26A817B8A238305FFCD50149FC20F9BA" box="[833,966,1556,1579]" pageId="23" pageNumber="30">L. Deharveng</collectorName>
(LAO-099),
<typeStatus id="54E6CCCCA238305FFBDC0149FB78F9BA" box="[1096,1182,1556,1579]" pageId="23" pageNumber="30" type="holotype">holotype</typeStatus>
<specimenCount id="9D5BB9E7A238305FFB31014BFB5FF9BB" box="[1189,1209,1558,1578]" pageId="23" pageNumber="30" type="male">♂</specimenCount>
(
<collectionCode id="ED4CEAABA238305FFD380170FCE6F9D5" box="[684,768,1581,1604]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="30">MNHN</collectionCode>
<specimenCode id="DBFBDA15A238305FFC920170FCBEF9D5" box="[774,856,1581,1604]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="30">GA 039</specimenCode>
)
</materialsCitation>
;
<materialsCitation id="3B357833A238305FFCFE0170FD0EF932" collectionCode="MNHN, SEM" country="Laos" pageId="23" pageNumber="30" specimenCode="GA 039" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Louangphabang" typeStatus="paratype">
<typeStatus id="54E6CCCCA238305FFCFE0170FC20F9D5" box="[874,966,1581,1604]" pageId="23" pageNumber="30" type="paratype">paratypes</typeStatus>
<specimenCount id="9D5BB9E7A238305FFC590170FC12F9D5" box="[973,1012,1581,1604]" pageId="23" pageNumber="30" type="male">1 ♂</specimenCount>
(
<collectionCode id="ED4CEAABA238305FFB950170FBB3F9D5" box="[1025,1109,1581,1604]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="30">MNHN</collectionCode>
<specimenCode id="DBFBDA15A238305FFBC80170FB48F9D5" box="[1116,1198,1581,1604]" collectionCode="MNHN" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="23" pageNumber="30">GA 039</specimenCode>
),
<specimenCount id="9D5BB9E7A238305FFD31011AFD2FF9CC" box="[677,713,1607,1630]" pageId="23" pageNumber="30" type="female">1 ♀</specimenCount>
(
<collectionCode id="ED4CEAABA238305FFD42011AFCEBF9CF" box="[726,781,1607,1630]" pageId="23" pageNumber="30">SEM</collectionCode>
). ETYMOLOGY. — Honours Anne Bedos, one of the collectors
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF2222E6A2373050FFE5013EFE6FF930" ID-DOI="http://doi.org/10.5281/zenodo.5392119" ID-Zenodo-Dep="5392119" httpUri="https://zenodo.org/record/5392119/files/figure.png" pageId="24" pageNumber="31" startId="24.[113,124,1635,1651]" targetBox="[120,1199,216,1503]" targetPageId="24">
<paragraph id="8BE2726EA2373050FFE5013EFE6FF930" blockId="24.[113,1210,1635,1698]" pageId="24" pageNumber="31">
FIG. 19. —
<taxonomicName id="4C5D09EDA2373050FF5B013EFE6EF9E5" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[207,392,1635,1652]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="24" pageNumber="31" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B929AE7CA2373050FF5B013EFE6EF9E5" box="[207,392,1635,1652]" italics="true" pageId="24" pageNumber="31">Glyphiulus deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A2373050FE1A013EFE26F9E2" box="[398,448,1635,1651]" pageId="24" pageNumber="31" rank="species">n. sp.</taxonomicNameLabel>
, ♀ paratype:
<emphasis id="B929AE7CA2373050FDA6013EFDD9F9E5" bold="true" box="[562,575,1635,1652]" pageId="24" pageNumber="31">A</emphasis>
, anterior part of body, lateral view;
<emphasis id="B929AE7CA2373050FCF9013EFC9CF9E2" bold="true" box="[877,890,1635,1651]" pageId="24" pageNumber="31">B</emphasis>
, same, dorsal view;
<emphasis id="B929AE7CA2373050FBBE013EFBDEF9E5" bold="true" box="[1066,1080,1635,1652]" pageId="24" pageNumber="31">C</emphasis>
, middle part of body, dorsal view;
<emphasis id="B929AE7CA2373050FE9A0127FEFAF91B" bold="true" box="[270,284,1658,1674]" pageId="24" pageNumber="31">D</emphasis>
, telson, ventral view;
<emphasis id="B929AE7CA2373050FE460127FE38F91B" bold="true" box="[466,478,1658,1674]" pageId="24" pageNumber="31">E</emphasis>
, midbody segment section, caudal view;
<emphasis id="B929AE7CA2373050FCAF0127FCA0F91B" bold="true" box="[827,838,1658,1674]" pageId="24" pageNumber="31">F</emphasis>
, claw of a midbody leg. Scale bars: A, C, D, 0.5 mm; B, E, 1.0 mm; F, 0.1 mm.
</paragraph>
</caption>
<caption id="DF2222E6A2363051FFE50484FF00FB86" ID-DOI="http://doi.org/10.5281/zenodo.5392121" ID-Zenodo-Dep="5392121" httpUri="https://zenodo.org/record/5392121/files/figure.png" pageId="25" pageNumber="32" startId="25.[113,124,985,1001]" targetBox="[190,1126,202,956]" targetPageId="25">
<paragraph id="8BE2726EA2363051FFE50484FF00FB86" blockId="25.[112,1210,984,1047]" pageId="25" pageNumber="32">
FIG. 20. —
<taxonomicName id="4C5D09EDA2363051FF590485FE63FC78" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[205,389,984,1001]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="32" phylum="Arthropoda" rank="species" species="deharvengi" status="sp. nov.">
<emphasis id="B929AE7CA2363051FF590485FE63FC78" box="[205,389,984,1001]" italics="true" pageId="25" pageNumber="32">Glyphiulus deharvengi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A2363051FE1E0484FE5DFC78" box="[394,443,985,1001]" pageId="25" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
, ♂ paratype:
<emphasis id="B929AE7CA2363051FDB80485FDDFFC78" bold="true" box="[556,569,984,1001]" pageId="25" pageNumber="32">A</emphasis>
, antenna;
<emphasis id="B929AE7CA2363051FD050484FD78FC78" bold="true" box="[657,670,985,1001]" pageId="25" pageNumber="32">B</emphasis>
, gnathochilarium, caudal view;
<emphasis id="B929AE7CA2363051FC330485FC53FC78" bold="true" box="[935,949,984,1001]" pageId="25" pageNumber="32">C</emphasis>
, legs 1, caudal view;
<emphasis id="B929AE7CA2363051FBFE0484FB9EFC78" bold="true" box="[1130,1144,985,1001]" pageId="25" pageNumber="32">D</emphasis>
, legs 2, caudal view;
<emphasis id="B929AE7CA2363051FF4E04ADFF00FB91" bold="true" box="[218,230,1008,1024]" pageId="25" pageNumber="32">E</emphasis>
, leg 3;
<emphasis id="B929AE7CA2363051FE8B04ADFECCFB91" bold="true" box="[287,298,1008,1024]" pageId="25" pageNumber="32">F</emphasis>
, midbody leg;
<emphasis id="B929AE7CA2363051FE3504B2FE49FB91" bold="true" box="[417,431,1007,1024]" pageId="25" pageNumber="32">G</emphasis>
, anterior gonopods, caudal view;
<emphasis id="B929AE7CA2363051FD5404ADFD28FB91" bold="true" box="[704,718,1008,1024]" pageId="25" pageNumber="32">H</emphasis>
, posterior gonopod, subfrontal view.Scale bar: A-F, 0.6 mm; G, H, 0.3 mm.
</paragraph>
</caption>
<subSubSection id="C34721E5A2363051FFE50312FECCFB41" pageId="25" pageNumber="32" type="diagnosis">
<paragraph id="8BE2726EA2363051FFE50312FECCFB41" blockId="25.[111,648,1103,1232]" pageId="25" pageNumber="32">DIAGNOSIS. — Differs from other species by the paraprocts showing a clear median marginal ridge, combined with a plumose spatuliform distal process of the posterior gonopods and a subunciform distomedian corner of the anterior gonopods.</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2363051FFE503B3FB6AFB18" pageId="25" pageNumber="32" type="description">
<paragraph id="8BE2726EA2363051FFE503B3FEE4FA97" blockId="25.[113,648,1262,1700]" box="[113,258,1262,1288]" pageId="25" pageNumber="32">DESCRIPTION</paragraph>
<paragraph id="8BE2726EA2363051FFE50253FF1DFAD6" blockId="25.[113,648,1262,1700]" pageId="25" pageNumber="32">
Length of adults of both sexes
<quantity id="4CA5DF8BA2363051FE570253FDDDFAB9" box="[451,571,1294,1320]" metricMagnitude="-2" metricUnit="m" metricValue="1.8" metricValueMax="2.0" metricValueMin="1.6" pageId="25" pageNumber="32" unit="mm" value="18.0" valueMax="20.0" valueMin="16.0">16-20 mm</quantity>
, width 1.0-
<quantity id="4CA5DF8BA2363051FF0A0270FF11FAD6" box="[158,247,1325,1351]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="32" unit="mm" value="1.1">1.1 mm</quantity>
.
</paragraph>
<paragraph id="8BE2726EA2363051FF1C0210FD61FA37" blockId="25.[113,648,1262,1700]" pageId="25" pageNumber="32">
<typeStatus id="54E6CCCCA2363051FF1C0210FF12FAF6" box="[136,244,1357,1383]" pageId="25" pageNumber="32" type="holotype">Holotype</typeStatus>
<quantity id="4CA5DF8BA2363051FF6B0210FEB3FAF6" box="[255,341,1357,1383]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="25" pageNumber="32" unit="mm" value="16.0">16 mm</quantity>
long, 1.0 mm wide, with 40p+3a+T and 11 ocelli;
<typeStatus id="54E6CCCCA2363051FEEE0230FE33FA16" box="[378,469,1389,1415]" pageId="25" pageNumber="32" type="paratype">paratype</typeStatus>
<specimenCount id="9D5BB9E7A2363051FE4E0232FE16FA14" box="[474,496,1391,1413]" pageId="25" pageNumber="32" type="male">♂</specimenCount>
about
<quantity id="4CA5DF8BA2363051FDAC0230FD60FA16" box="[568,646,1389,1415]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="32" unit="mm" value="20.0">20 mm</quantity>
long,
<quantity id="4CA5DF8BA2363051FF3A02D1FEE5FA37" box="[174,259,1420,1446]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="25" pageNumber="32" unit="mm" value="1.1">1.1 mm</quantity>
wide, with 54p+2a+T and 14 ocelli.
</paragraph>
<paragraph id="8BE2726EA2363051FF1C02F1FE6DF9F5" blockId="25.[113,648,1262,1700]" pageId="25" pageNumber="32">
Coloration, antennae (
<figureCitation id="13666EEBA2363051FEE902F1FE04FA57" box="[381,482,1452,1478]" captionStart="FIG" captionStartId="22.[113,124,1345,1361]" captionTargetBox="[121,1201,215,1301]" captionTargetId="figure-44@22.[391,664,758,940]" captionTargetPageId="22" captionText="FIG. 17. — Glyphiulus bedosae n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, lateral view; D, same, dorsal view; E, caudal part of body, lateral view; F, telson, ventral view. Scale bars: A, C, 0.5 mm; B, D, E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392115" httpUri="https://zenodo.org/record/5392115/files/figure.png" pageId="25" pageNumber="32">Figs 17A</figureCitation>
;
<figureCitation id="13666EEBA2363051FE7902F1FDF8FA57" box="[493,542,1452,1478]" captionStart="FIG" captionStartId="23.[113,124,1189,1205]" captionTargetBox="[110,1225,182,1181]" captionTargetId="figure-14@23.[110,1225,182,1181]" captionTargetPageId="23" captionText="FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="25" pageNumber="32">18A</figureCitation>
), gnathochilarium (mentum divided, n = 1) (
<figureCitation id="13666EEBA2363051FD870291FD9EFA77" box="[531,632,1484,1510]" captionStart="FIG" captionStartId="23.[113,124,1189,1205]" captionTargetBox="[110,1225,182,1181]" captionTargetId="figure-14@23.[110,1225,182,1181]" captionTargetPageId="23" captionText="FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="25" pageNumber="32">Fig. 18B</figureCitation>
), carinotaxy (
<figureCitation id="13666EEBA2363051FF6B02B6FEB5F994" box="[255,339,1515,1541]" captionStart="FIG" captionStartId="22.[113,124,1345,1361]" captionTargetBox="[121,1201,215,1301]" captionTargetId="figure-44@22.[391,664,758,940]" captionTargetPageId="22" captionText="FIG. 17. — Glyphiulus bedosae n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, lateral view; D, same, dorsal view; E, caudal part of body, lateral view; F, telson, ventral view. Scale bars: A, C, 0.5 mm; B, D, E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392115" httpUri="https://zenodo.org/record/5392115/files/figure.png" pageId="25" pageNumber="32">Fig. 17</figureCitation>
A-E), segment shape, legs (
<figureCitation id="13666EEBA2363051FFED0156FF20F9B4" box="[121,198,1547,1573]" captionStart="FIG" captionStartId="23.[113,124,1189,1205]" captionTargetBox="[110,1225,182,1181]" captionTargetId="figure-14@23.[110,1225,182,1181]" captionTargetPageId="23" captionText="FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="25" pageNumber="32">Fig. 18</figureCitation>
C-F), etc., generally as in
<taxonomicName id="4C5D09EDA2363051FE790156FD99F9B4" baseAuthorityName="Gervais" baseAuthorityYear="1847" box="[493,639,1547,1573]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="32" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B929AE7CA2363051FE790156FD99F9B4" box="[493,639,1547,1573]" italics="true" pageId="25" pageNumber="32">G. granulatus</emphasis>
</taxonomicName>
, but paraprocts with a low, but evident, median marginal ridge (
<figureCitation id="13666EEBA2363051FE8B0117FE98F9F5" box="[287,382,1610,1636]" captionStart="FIG" captionStartId="22.[113,124,1345,1361]" captionTargetBox="[121,1201,215,1301]" captionTargetId="figure-44@22.[391,664,758,940]" captionTargetPageId="22" captionText="FIG. 17. — Glyphiulus bedosae n. sp., ♀ paratype: A, anterior part of body, lateral view; B, same, dorsal view; C, middle part of body, lateral view; D, same, dorsal view; E, caudal part of body, lateral view; F, telson, ventral view. Scale bars: A, C, 0.5 mm; B, D, E, 0.2 mm; F, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392115" httpUri="https://zenodo.org/record/5392115/files/figure.png" pageId="25" pageNumber="32">Fig. 17F</figureCitation>
).
</paragraph>
<paragraph id="8BE2726EA2363051FF1C0137FB6AFB18" blockId="25.[113,648,1262,1700]" lastBlockId="25.[677,1209,1103,1161]" pageId="25" pageNumber="32">
Anterior gonopods like those of
<taxonomicName id="4C5D09EDA2363051FE460137FD60F915" authority="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel, 2007" authorityName="Golovatch &amp; Geoffroy &amp; Mauriès &amp; Spiegel" authorityYear="2007" box="[466,646,1642,1668]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="32" phylum="Arthropoda" rank="species" species="semigranulatus" status="sp. nov.">
<emphasis id="B929AE7CA2363051FE460137FD60F915" box="[466,646,1642,1668]" italics="true" pageId="25" pageNumber="32">G. semigranulatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21A1307A2363051FFE501D7FF48F935" box="[113,174,1674,1700]" pageId="25" pageNumber="32" rank="species">n. sp.</taxonomicNameLabel>
(
<figureCitation id="13666EEBA2363051FF2A01D7FEC2F935" box="[190,292,1674,1700]" captionStart="FIG" captionStartId="23.[113,124,1189,1205]" captionTargetBox="[110,1225,182,1181]" captionTargetId="figure-14@23.[110,1225,182,1181]" captionTargetPageId="23" captionText="FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="25" pageNumber="32">Fig. 18G</figureCitation>
), but posterior gonopods with a plumose spatuliform distal process (
<figureCitation id="13666EEBA2363051FBB50312FB60FBF8" box="[1057,1158,1103,1129]" captionStart="FIG" captionStartId="23.[113,124,1189,1205]" captionTargetBox="[110,1225,182,1181]" captionTargetId="figure-14@23.[110,1225,182,1181]" captionTargetPageId="23" captionText="FIG. 18. — Glyphiulus bedosae n.sp., ♂ paratype:A, antenna; B, gnathochilarium, caudal view; C, legs 1, caudal view; D, legs 2, caudal view; E, leg 3; F, midbody leg; G, anterior gonopods, caudal view; H, posterior gonopod, subcaudal view. Scale bar: A-F, 0.2 mm; G, H, 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5392117" httpUri="https://zenodo.org/record/5392117/files/figure.png" pageId="25" pageNumber="32">Fig. 18H</figureCitation>
), i.e. much as in
<taxonomicName id="4C5D09EDA2363051FCB60332FC02FB18" authorityName="Chen &amp; Meng" authorityYear="1991" box="[802,996,1135,1161]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="32" phylum="Arthropoda" rank="species" species="quadrohamatus">
<emphasis id="B929AE7CA2363051FCB60332FC02FB18" box="[802,996,1135,1161]" italics="true" pageId="25" pageNumber="32">G. quadrohamatus</emphasis>
</taxonomicName>
or
<taxonomicName id="4C5D09EDA2363051FB9D0332FB63FB18" authorityName="Mauries &amp; Nguyen Duy-Jacquemin" authorityYear="1997" box="[1033,1157,1135,1161]" class="Diplopoda" family="Cambalopsidae" genus="Glyphiulus" kingdom="Animalia" order="Spirostreptida" pageId="25" pageNumber="32" phylum="Arthropoda" rank="species" species="lipsorum">
<emphasis id="B929AE7CA2363051FB9D0332FB63FB18" box="[1033,1157,1135,1161]" italics="true" pageId="25" pageNumber="32">G. lipsorum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C34721E5A2363051FD3103F3FB9AFA99" pageId="25" pageNumber="32" type="discussion">
<paragraph id="8BE2726EA2363051FD3103F3FCEFFB57" blockId="25.[674,1210,1198,1288]" box="[677,777,1198,1224]" pageId="25" pageNumber="32">REMARKS</paragraph>
<paragraph id="8BE2726EA2363051FD310393FB9AFA99" blockId="25.[674,1210,1198,1288]" pageId="25" pageNumber="32">In the absence of any troglomorphic traits, this species can only be considered troglophilic at most.</paragraph>
</subSubSection>
</treatment>
</document>
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