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<mods:title id="281F903E4077D591947D000D223A735F">Revision of the type species of Syllepte Hübner and other spilomeline genera recently synonymized (Lepidoptera: Crambidae)</mods:title>
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<subSubSection id="C341529A674BFFD3C0A043B354E49F8C" box="[151,495,978,1005]" pageId="3" pageNumber="346" type="nomenclature">
<paragraph id="8BE40111674BFFD3C0A043B354E49F8C" blockId="3.[151,495,978,1005]" box="[151,495,978,1005]" pageId="3" pageNumber="346">
<heading id="D0ACB67D674BFFD3C0A043B354E49F8C" bold="true" box="[151,495,978,1005]" fontSize="11" level="1" pageId="3" pageNumber="346" reason="1">
<emphasis id="B92FDD03674BFFD3C0A043B354E49F8C" bold="true" box="[151,495,978,1005]" pageId="3" pageNumber="346">
<taxonomicName id="4C5B7A92674BFFD3C0A043B354AD9F8C" ID-CoL="92NJG" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1823" box="[151,422,978,1005]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674BFFD3C0A043B355FD9F8D" bold="true" box="[151,246,978,1005]" italics="true" pageId="3" pageNumber="346">Syllepte</emphasis>
Hübner, 1819
</taxonomicName>
1821
</emphasis>
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</paragraph>
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<paragraph id="8BE40111674BFFD3C0A044765644982E" blockId="3.[151,1436,1047,1286]" pageId="3" pageNumber="346">
<taxonomicName id="4C5B7A92674BFFD3C0A044765474984F" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1823" box="[151,383,1047,1071]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674BFFD3C0A0447655EC984F" box="[151,231,1047,1071]" italics="true" pageId="3" pageNumber="346">Syllepte</emphasis>
Hübner, 1819
</taxonomicName>
1821.
<typeStatus id="54E0BFB3674BFFD3C1FA447954F49850" box="[461,511,1048,1072]" pageId="3" pageNumber="346">Type</typeStatus>
species:
<taxonomicName id="4C5B7A92674BFFD3C25544765686984F" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1819" box="[610,909,1047,1071]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="incomptalis">
<emphasis id="B92FDD03674BFFD3C255447657FF984F" box="[610,756,1047,1071]" italics="true" pageId="3" pageNumber="346">S. incomptalis</emphasis>
Hübner, 1819
</taxonomicName>
1821: pl. 50 figs. 285286; 1823: 18 (dated after
<bibRefCitation id="EFCA7CE0674BFFD3C0F044575461982E" author="Hemming, F." box="[199,362,1078,1102]" pageId="3" pageNumber="346" refId="ref11716" refString="Hemming, F. (1937) A bibliographical and systematic account of the entomological works of Jacob Hubner and of the supplements there to by Carl Geyer, Gottfried Franz von Frolich and Gottlieb August Wilhelm Herrich-Schaffer. Royal Entomological Society, London, xxxiv + 605 pp." type="book" year="1937">Hemming 1937</bibRefCitation>
). Synonym:
<taxonomicName id="4C5B7A92674BFFD3C1DE445757E3982E" authority="Dognin, 1905" authorityName="Dognin" authorityYear="1905" box="[489,744,1078,1102]" class="Insecta" family="Crambidae" genus="Neomabra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus" status="rev. stat.">
<emphasis id="B92FDD03674BFFD3C1DE4457575F982E" box="[489,596,1078,1102]" italics="true" pageId="3" pageNumber="346">Neomabra</emphasis>
<bibRefCitation id="EFCA7CE0674BFFD3C26D445757E3982E" author="Dognin, P." box="[602,744,1078,1102]" pageId="3" pageNumber="346" pagination="61 - 90" refId="ref11494" refString="Dognin, P. (1905) Heteroceres nouveaux de l'Amerique du Sud. Annales de la Societe Entomologique de Belgique, Bruxelles, 49, 61 - 90. Available from: https: // www. biodiversitylibrary. org / item / 44643 # page / 68 / mode / 2 up (accessed 29 November 2023)" type="journal article" year="1905">Dognin, 1905</bibRefCitation>
</taxonomicName>
,
<emphasis id="B92FDD03674BFFD3C2C544575644982E" bold="true" box="[754,847,1078,1102]" pageId="3" pageNumber="346">
<taxonomicNameLabel id="A21C6078674BFFD3C2C544575644982E" box="[754,847,1078,1102]" pageId="3" pageNumber="346">rev. stat.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BE40111674BFFD3C0A04435544C980C" blockId="3.[151,1436,1047,1286]" box="[151,327,1108,1132]" pageId="3" pageNumber="346">Included species:</paragraph>
<paragraph id="8BE40111674BFFD3C0A04412502A98EB" blockId="3.[151,1436,1047,1286]" box="[151,1313,1139,1164]" pageId="3" pageNumber="346">
<taxonomicName id="4C5B7A92674BFFD3C0A0441254FD98EB" authority="Hubner, 1819" authorityName="Hubner" authorityYear="1819" box="[151,502,1139,1163]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="incomptalis">
<emphasis id="B92FDD03674BFFD3C0A04412546F98EB" box="[151,356,1139,1163]" italics="true" pageId="3" pageNumber="346">Syllepte incomptalis</emphasis>
Hübner, 1819
</taxonomicName>
1821, type locality presumed to be
<collectingCountry id="F34C4181674BFFD3C351441256C198EB" box="[870,970,1139,1163]" name="Suriname" pageId="3" pageNumber="346">Suriname</collectingCountry>
,
<emphasis id="B92FDD03674BFFD3C3E3441251A798EA" bold="true" box="[980,1196,1139,1163]" pageId="3" pageNumber="346">
<typeStatus id="54E0BFB3674BFFD3C3E34412512698EB" box="[980,1069,1139,1163]" pageId="3" pageNumber="346" type="neotype">Neotype</typeStatus>
designated,
</emphasis>
<collectingCountry id="F34C4181674BFFD3C4844412502A98EB" box="[1203,1313,1139,1163]" name="Costa Rica" pageId="3" pageNumber="346">Costa Rica</collectingCountry>
</paragraph>
<paragraph id="8BE40111674BFFD3C0A044F3509998CA" blockId="3.[151,1436,1047,1286]" box="[151,1426,1170,1194]" pageId="3" pageNumber="346">
<taxonomicName id="4C5B7A92674BFFD3C0A044F354E898CA" authority="(Dognin, 1905)" baseAuthorityName="Dognin" baseAuthorityYear="1905" box="[151,483,1170,1194]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="nitidalis">
<emphasis id="B92FDD03674BFFD3C0A044F3544A98CA" box="[151,321,1170,1194]" italics="true" pageId="3" pageNumber="346">Syllepte nitidalis</emphasis>
(
<bibRefCitation id="EFCA7CE0674BFFD3C17944F354D798CA" author="Dognin, P." box="[334,476,1170,1194]" pageId="3" pageNumber="346" pagination="61 - 90" refId="ref11494" refString="Dognin, P. (1905) Heteroceres nouveaux de l'Amerique du Sud. Annales de la Societe Entomologique de Belgique, Bruxelles, 49, 61 - 90. Available from: https: // www. biodiversitylibrary. org / item / 44643 # page / 68 / mode / 2 up (accessed 29 November 2023)" type="journal article" year="1905">Dognin, 1905</bibRefCitation>
)
</taxonomicName>
(
<taxonomicName id="4C5B7A92674BFFD3C1C544F3575698CA" authorityName="Dognin" authorityYear="1905" box="[498,605,1170,1194]" class="Insecta" family="Crambidae" genus="Neomabra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674BFFD3C1C544F3575698CA" box="[498,605,1170,1194]" italics="true" pageId="3" pageNumber="346">Neomabra</emphasis>
</taxonomicName>
), type locality
<collectingCountry id="F34C4181674BFFD3C2C144F3564598CA" box="[758,846,1170,1194]" name="Ecuador" pageId="3" pageNumber="346">Ecuador</collectingCountry>
:
<collectingRegion id="499FCFF3674BFFD3C36F44F3568298CA" box="[856,905,1170,1194]" country="Ecuador" name="Loja" pageId="3" pageNumber="346">Loja</collectingRegion>
, Zamora valley,
<emphasis id="B92FDD03674BFFD3C40644F3509998CA" bold="true" box="[1073,1426,1170,1194]" pageId="3" pageNumber="346">
rev. comb.,
<typeStatus id="54E0BFB3674BFFD3C49A44F3501398CA" box="[1197,1304,1170,1194]" pageId="3" pageNumber="346" type="lectotype">Lectotype</typeStatus>
designated
</emphasis>
</paragraph>
<paragraph id="8BE40111674BFFD3C0A044D154099887" blockId="3.[151,1436,1047,1286]" pageId="3" pageNumber="346">
<taxonomicName id="4C5B7A92674BFFD3C0A044D1572C98A8" authority="(Lederer, 1863)" baseAuthorityName="Lederer" baseAuthorityYear="1863" box="[151,551,1200,1224]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="serratilinealis">
<emphasis id="B92FDD03674BFFD3C0A044D1547598A8" box="[151,382,1200,1224]" italics="true" pageId="3" pageNumber="346">Syllepte serratilinealis</emphasis>
(Lederer, 1863)
</taxonomicName>
(
<taxonomicName id="4C5B7A92674BFFD3C20E44D0577A98A8" authorityName="Latreille" authorityYear="1802" box="[569,625,1201,1224]" class="Insecta" family="Crambidae" genus="Botys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674BFFD3C20E44D0577A98A8" box="[569,625,1201,1224]" italics="true" pageId="3" pageNumber="346">Botys</emphasis>
</taxonomicName>
), type locality
<collectingCountry id="F34C4181674BFFD3C32644D1567798A8" box="[785,892,1200,1224]" name="Venezuela" pageId="3" pageNumber="346">Venezuela</collectingCountry>
,
<emphasis id="B92FDD03674BFFD3C3BF44D056EC98A9" bold="true" box="[904,999,1201,1225]" pageId="3" pageNumber="346">
<taxonomicNameLabel id="A21C6078674BFFD3C3BF44D056EC98A9" box="[904,999,1201,1225]" pageId="3" pageNumber="346">rev. stat.</taxonomicNameLabel>
</emphasis>
[type specimen not found, from figure of adult]
</paragraph>
<paragraph id="8BE40111674BFFD3C0A0448F503F9966" blockId="3.[151,1436,1047,1286]" box="[151,1332,1262,1286]" pageId="3" pageNumber="346">
<taxonomicName id="4C5B7A92674BFFD3C0A0448F54FC9966" authority="Hampson, 1912" authorityName="Hampson" authorityYear="1912" box="[151,503,1262,1286]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="species" species="leucinalis">
<emphasis id="B92FDD03674BFFD3C0A0448F545A9966" box="[151,337,1262,1286]" italics="true" pageId="3" pageNumber="346">Syllepte leucinalis</emphasis>
Hampson, 1912
</taxonomicName>
(
<taxonomicName id="4C5B7A92674BFFD3C231448F575A9966" authorityName="Hübner" authorityYear="1825" box="[518,593,1262,1286]" class="Insecta" family="Crambidae" genus="Sylepta" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="346" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674BFFD3C231448F575A9966" box="[518,593,1262,1286]" italics="true" pageId="3" pageNumber="346">Sylepta</emphasis>
</taxonomicName>
[sic]), type locality
<collectingCountry id="F34C4181674BFFD3C32A448F56449965" box="[797,847,1262,1285]" name="Peru" pageId="3" pageNumber="346">Peru</collectingCountry>
: Pozuzo,
<emphasis id="B92FDD03674BFFD3C387448F51079966" bold="true" box="[944,1036,1262,1286]" pageId="3" pageNumber="346">
<taxonomicNameLabel id="A21C6078674BFFD3C387448F51079966" box="[944,1036,1262,1286]" pageId="3" pageNumber="346">rev. stat.</taxonomicNameLabel>
</emphasis>
[see note in Remarks below]
</paragraph>
</subSubSection>
<subSubSection id="C341529A674BFFD3C0A04555571699BF" pageId="3" pageNumber="346" type="diagnosis">
<paragraph id="8BE40111674BFFD3C0A04555571699BF" blockId="3.[151,1437,1332,2043]" pageId="3" pageNumber="346">
<emphasis id="B92FDD03674BFFD3C0A04555541F992E" bold="true" box="[151,276,1332,1358]" pageId="3" pageNumber="346">Diagnosis:</emphasis>
In the forewing the reniform spot is constricted medially and often appears as a dark brown x-shaped mark in older specimens or a figure
<quantity id="4CA3ACF4674BFFD3C277453957659912" box="[576,622,1368,1394]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="3" pageNumber="346" unit="in" value="8.0">8 in</quantity>
recently collected specimens, and the postmedial line is a dark brown scalloped line (
<figureCitation id="13601D94674BFFD3C171451D54BE99F6" box="[326,437,1404,1430]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="3" pageNumber="346">Figs. 1, 2</figureCitation>
,
<figureCitation id="13601D94674BFFD3C1F3451D54E899F6" box="[452,483,1404,1430]" captionStart="FIGURES 1018" captionStartId="14.[151,260,1815,1839]" captionTargetBox="[230,1358,151,1796]" captionTargetId="figure-23@14.[230,1358,151,1803]" captionTargetPageId="14" captionText="FIGURES 1018. 10, Syllepte nitidalis (Dognin), lectotype adult male, Ecuador, Environs de Loja [USNMENT01433441]. 11, S. nitidalis,lectotypelabels.12,S.nitidalis,malegenitalia,paralectotype,Ecuador,Zamora,USNMslide#111085[USNMENT01769261]. 13, S. nitidalis, phallus, paralectotype, Ecuador, Zamora, USNM slide #111085 [USNMENT01769261]. 14, S. nitidalis, male modified structure and final segment, paralectotype, Ecuador, Environs de Loja, USNM slide #116219 [USNMENT01769438]. 15, Pantographa scripturalis (Guenée), lectotype adult female, Brazil [NHMUK01396720]. 16, P. scripturalis, lectotype labels. 17, P. scripturalis, male genitalia, Brazil, Parana, Castro, USNM slide #107232 [USNMENT01895419]. 18, P. scripturalis, female genitalia, Brazil, Rio de Janeiro, USNM slide #105696 [USNMENT01899000]." figureDoi="http://doi.org/10.5281/zenodo.10409851" httpUri="https://zenodo.org/record/10409851/files/figure.png" pageId="3" pageNumber="346">10</figureCitation>
). Males have a unique, lengthwise sclerotized structure with modified scales on 7
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tergite, which features a transverse comb-like central arch ending in two lateral thumb-like structures, medially with broad semi-circular scales (
<figureCitation id="13601D94674BFFD3C1FC45A5570499BF" box="[459,527,1476,1503]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="3" pageNumber="346">Fig. 5</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C341529A674BFFD4C0F0458956889951" lastPageId="4" lastPageNumber="347" pageId="3" pageNumber="346" type="description">
<paragraph id="8BE40111674BFFD4C0F0458954AA98A5" blockId="3.[151,1437,1332,2043]" lastBlockId="4.[151,1437,151,2013]" lastPageId="4" lastPageNumber="347" pageId="3" pageNumber="346">
<emphasis id="B92FDD03674BFFD3C0F0458954509A62" bold="true" box="[199,347,1512,1538]" pageId="3" pageNumber="346">Description:</emphasis>
<emphasis id="B92FDD03674BFFD3C156458954959A62" box="[353,414,1512,1538]" italics="true" pageId="3" pageNumber="346">Head</emphasis>
: Labial palpus upturned, 2
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segment 5 times as long as 3
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segment; maxillary palpus short, as long as 3
<superScript id="7C2EAC59674BFFD3C113466D54399A7A" attach="left" box="[292,306,1548,1562]" fontSize="6" pageId="3" pageNumber="346">rd</superScript>
labial palpus segment; ocellus with a dark, sclerotized band basally, clear apically; chaetosemata absent; male antenna with scales dorsally, without scales ventrally and with hairlike cilia, each hair slightly half as long as antennal segment, female similar but with very short cilia.
<emphasis id="B92FDD03674BFFD3C392463556FD9A0E" box="[933,1014,1620,1646]" italics="true" pageId="3" pageNumber="346">Thorax</emphasis>
: Average forewing length=
<quantity id="4CA3ACF4674BFFD3C51B463550969A0F" box="[1324,1437,1620,1647]" metricMagnitude="-2" metricUnit="m" metricValue="1.213" pageId="3" pageNumber="346" unit="mm" value="12.13">12.13 mm</quantity>
(n=15), range=
<quantity id="4CA3ACF4674BFFD3C175461954B19AF3" box="[322,442,1656,1683]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="346" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">1015 mm</quantity>
.
<emphasis id="B92FDD03674BFFD3C1F34618576F9AF2" box="[452,612,1657,1682]" italics="true" pageId="3" pageNumber="346">Wing venation</emphasis>
: Forewing Sc to 2/3 length of costa. R from distal third of discal cell; Rs
<subScript id="17DF0354674BFFD3C5BA46EB509E9AF8" attach="left" box="[1421,1429,1674,1688]" fontSize="6" pageId="3" pageNumber="346">1</subScript>
, Rs
<subScript id="17DF0354674BFFD3C08246CF55C59ADC" attach="left" box="[181,206,1710,1724]" fontSize="6" pageId="3" pageNumber="346">2+3</subScript>
, and Rs
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close together from anterior angle of cell, Rs
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running parallel for a short distance, then diverging toward outer margin. M
<subScript id="17DF0354674BFFD3C19546B354A19A80" attach="left" box="[418,426,1746,1760]" fontSize="6" pageId="3" pageNumber="346">1</subScript>
from anterior angle of discal cell to outer margin; base of M
<subScript id="17DF0354674BFFD3C46746B351539A80" attach="left" box="[1104,1112,1746,1760]" fontSize="6" pageId="3" pageNumber="346">2</subScript>
and M
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close to CuA
<subScript id="17DF0354674BFFD3C57E46B3505A9A80" attach="left" box="[1353,1361,1746,1760]" fontSize="6" pageId="3" pageNumber="346">1</subScript>
. CuA
<subScript id="17DF0354674BFFD3C5A346B350979A80" attach="left" box="[1428,1436,1746,1760]" fontSize="6" pageId="3" pageNumber="346">2</subScript>
from apical fifth of Cubitus. CuP absent. 1A+2A well marked from wing base to outer margin; 3A an extremely faint loop one half the length of 1A+2A. Hindwing Sc+R
<subScript id="17DF0354674BFFD3C2D2477B57E69B48" attach="left" box="[741,749,1818,1832]" fontSize="6" pageId="3" pageNumber="346">1</subScript>
and Rs stalked for about half length of Media beyond discal cell to outer margin. Origin of M
<subScript id="17DF0354674BFFD3C23B475F571F9B2C" attach="left" box="[524,532,1854,1868]" fontSize="6" pageId="3" pageNumber="346">1</subScript>
from Rs; origin of M
<subScript id="17DF0354674BFFD3C33D475F56199B2C" attach="left" box="[778,786,1854,1868]" fontSize="6" pageId="3" pageNumber="346">2</subScript>
and M
<subScript id="17DF0354674BFFD3C354475F56609B2C" attach="left" box="[867,875,1854,1868]" fontSize="6" pageId="3" pageNumber="346">3</subScript>
close to CuA
<subScript id="17DF0354674BFFD3C430475F51049B2C" attach="left" box="[1031,1039,1854,1868]" fontSize="6" pageId="3" pageNumber="346">1</subScript>
. CuA
<subScript id="17DF0354674BFFD3C464475F51509B2C" attach="left" box="[1107,1115,1854,1868]" fontSize="6" pageId="3" pageNumber="346">2</subScript>
from 2/3
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of Cubitus. Three Anal veins well marked. Forewing basal color white to pale yellow to light brown.
<emphasis id="B92FDD03674BFFD3C404473051FD9B0A" box="[1075,1270,1873,1898]" italics="true" pageId="3" pageNumber="346">Forewing pattern</emphasis>
(
<figureCitation id="13601D94674BFFD3C532473150429B0A" box="[1285,1353,1872,1898]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="3" pageNumber="346">Fig. 2</figureCitation>
): Basal line absent or as a spot near costa and posterior margin, antemedial line thin, light brown, complete or entire width of wing; orbicular spot round with a faint, brown outline, claviform spot in anal area faint, light brown if present; median line absent, sometimes present as a dark shade or spot after CuA
<subScript id="17DF0354674BFFD3C3FB47AF56DF9BBC" attach="left" box="[972,980,1998,2012]" fontSize="6" pageId="3" pageNumber="346">2</subScript>
, reniform spot from Sc to distal edge of discal area, constricted medially and often appearing as a dark, brown x-shaped mark; postmedial line dark brown appearing as a scalloped line, straight from costa and then inward at after M
<subScript id="17DF0354674CFFD4C3EC40C856E89CD7" attach="left" box="[987,995,169,183]" fontSize="6" pageId="4" pageNumber="347">3</subScript>
to posterior margin, less visible beyond CuA
<subScript id="17DF0354674CFFD4C0FB40AC55DF9CBB" attach="left" box="[204,212,205,219]" fontSize="6" pageId="4" pageNumber="347">1</subScript>
; subterminal line absent, dark brown shading entire width or most commonly from costa to M
<subScript id="17DF0354674CFFD4C4C740AC51F39CBB" attach="left" box="[1264,1272,205,219]" fontSize="6" pageId="4" pageNumber="347">3</subScript>
only; terminal line dark brown; fringe variable. Retinaculum scales anterior to CuP.
<emphasis id="B92FDD03674CFFD4C3B840BE515F9C98" box="[911,1108,223,249]" italics="true" pageId="4" pageNumber="347">Hindwing pattern</emphasis>
(
<figureCitation id="13601D94674CFFD4C45540BE51AD9C99" box="[1122,1190,223,249]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="4" pageNumber="347">Fig. 2</figureCitation>
): Postmedial line dark brown as double scalloped lines, curving to posterior margin, basally with variable dark brown shading, sometimes apically, from the costa to Rs
<subScript id="17DF0354674CFFD4C1EC415854E89D27" attach="left" box="[475,483,313,327]" fontSize="6" pageId="4" pageNumber="347">4</subScript>
with dark shading, subterminal line absent, terminal line dark brown, fringe variable.
<emphasis id="B92FDD03674CFFD4C0A0412A540F9D05" box="[151,260,331,357]" italics="true" pageId="4" pageNumber="347">Frenulum</emphasis>
: Female with 2 spines, male with 1 spine.
<emphasis id="B92FDD03674CFFD4C2C4412A56239D04" box="[755,808,331,356]" italics="true" pageId="4" pageNumber="347">Legs</emphasis>
: Foreleg with tibial epiphysis, midleg with one pair of subequal tibial spurs, hind leg with two pairs of tibial spurs, distal spurs equal to length midleg spurs, one species with hindleg modified setae (hair pencil) as long as the tibia.
<emphasis id="B92FDD03674CFFD4C30141F256E09DCC" box="[822,1003,403,429]" italics="true" pageId="4" pageNumber="347">Tympanal organ</emphasis>
(
<figureCitation id="13601D94674CFFD4C3CE41F251379DCD" box="[1017,1084,403,429]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="4" pageNumber="347">Fig. 8</figureCitation>
): Pons tympani prominent, post tympanal area with lightly sclerotized outer margin, tergo-sternal sclerite sclerotized, processus tympani small or not visible, 2
<superScript id="7C2EAC59674CFFD4C0C941BB54059D88" attach="left" box="[254,270,474,488]" fontSize="6" pageId="4" pageNumber="347">nd</superScript>
tergite segment with a prominent, lengthwise membranous fold, same width throughout with triangular, lateral pouches, praecinctorium bilobed, membranous. Males with a unique sclerotized structure of modified scales on 7
<superScript id="7C2EAC59674CFFD4C0FE424355DD9E50" attach="left" box="[201,214,546,560]" fontSize="6" pageId="4" pageNumber="347">th</superScript>
tergite (
<figureCitation id="13601D94674CFFD4C103424254889E5D" box="[308,387,546,573]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="4" pageNumber="347">Figs. 5</figureCitation>
,
<figureCitation id="13601D94674CFFD4C1A7424254A69E5D" box="[400,429,547,573]" captionStart="FIGURES 1018" captionStartId="14.[151,260,1815,1839]" captionTargetBox="[230,1358,151,1796]" captionTargetId="figure-23@14.[230,1358,151,1803]" captionTargetPageId="14" captionText="FIGURES 1018. 10, Syllepte nitidalis (Dognin), lectotype adult male, Ecuador, Environs de Loja [USNMENT01433441]. 11, S. nitidalis,lectotypelabels.12,S.nitidalis,malegenitalia,paralectotype,Ecuador,Zamora,USNMslide#111085[USNMENT01769261]. 13, S. nitidalis, phallus, paralectotype, Ecuador, Zamora, USNM slide #111085 [USNMENT01769261]. 14, S. nitidalis, male modified structure and final segment, paralectotype, Ecuador, Environs de Loja, USNM slide #116219 [USNMENT01769438]. 15, Pantographa scripturalis (Guenée), lectotype adult female, Brazil [NHMUK01396720]. 16, P. scripturalis, lectotype labels. 17, P. scripturalis, male genitalia, Brazil, Parana, Castro, USNM slide #107232 [USNMENT01895419]. 18, P. scripturalis, female genitalia, Brazil, Rio de Janeiro, USNM slide #105696 [USNMENT01899000]." figureDoi="http://doi.org/10.5281/zenodo.10409851" httpUri="https://zenodo.org/record/10409851/files/figure.png" pageId="4" pageNumber="347">14</figureCitation>
) lengthwise; comb-like with two lateral thumb-like structures, medially with broad semi-circular scales. Sixth segment often with light sclerotization and prominent scale sockets. Second sternite with lengthwise membranous fold, medially triangular, 7
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and 8
<superScript id="7C2EAC59674CFFD4C304420B564B9E18" attach="left" box="[819,832,618,632]" fontSize="6" pageId="4" pageNumber="347">th</superScript>
sternites (
<figureCitation id="13601D94674CFFD4C382420A510D9EE5" box="[949,1030,618,645]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="4" pageNumber="347">Figs. 5</figureCitation>
,
<figureCitation id="13601D94674CFFD4C424420A513B9EE5" box="[1043,1072,619,645]" captionStart="FIGURES 1018" captionStartId="14.[151,260,1815,1839]" captionTargetBox="[230,1358,151,1796]" captionTargetId="figure-23@14.[230,1358,151,1803]" captionTargetPageId="14" captionText="FIGURES 1018. 10, Syllepte nitidalis (Dognin), lectotype adult male, Ecuador, Environs de Loja [USNMENT01433441]. 11, S. nitidalis,lectotypelabels.12,S.nitidalis,malegenitalia,paralectotype,Ecuador,Zamora,USNMslide#111085[USNMENT01769261]. 13, S. nitidalis, phallus, paralectotype, Ecuador, Zamora, USNM slide #111085 [USNMENT01769261]. 14, S. nitidalis, male modified structure and final segment, paralectotype, Ecuador, Environs de Loja, USNM slide #116219 [USNMENT01769438]. 15, Pantographa scripturalis (Guenée), lectotype adult female, Brazil [NHMUK01396720]. 16, P. scripturalis, lectotype labels. 17, P. scripturalis, male genitalia, Brazil, Parana, Castro, USNM slide #107232 [USNMENT01895419]. 18, P. scripturalis, female genitalia, Brazil, Rio de Janeiro, USNM slide #105696 [USNMENT01899000]." figureDoi="http://doi.org/10.5281/zenodo.10409851" httpUri="https://zenodo.org/record/10409851/files/figure.png" pageId="4" pageNumber="347">14</figureCitation>
) with sclerotized patterns.
<emphasis id="B92FDD03674CFFD4C554420A55F09EC9" italics="true" pageId="4" pageNumber="347">Male genitalia</emphasis>
(
<figureCitation id="13601D94674CFFD4C13942EE548B9EC9" box="[270,384,655,681]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="4" pageNumber="347">Figs. 4, 6</figureCitation>
,
<figureCitation id="13601D94674CFFD4C1B842EE54D19EC9" box="[399,474,655,681]" captionStart="FIGURES 1018" captionStartId="14.[151,260,1815,1839]" captionTargetBox="[230,1358,151,1796]" captionTargetId="figure-23@14.[230,1358,151,1803]" captionTargetPageId="14" captionText="FIGURES 1018. 10, Syllepte nitidalis (Dognin), lectotype adult male, Ecuador, Environs de Loja [USNMENT01433441]. 11, S. nitidalis,lectotypelabels.12,S.nitidalis,malegenitalia,paralectotype,Ecuador,Zamora,USNMslide#111085[USNMENT01769261]. 13, S. nitidalis, phallus, paralectotype, Ecuador, Zamora, USNM slide #111085 [USNMENT01769261]. 14, S. nitidalis, male modified structure and final segment, paralectotype, Ecuador, Environs de Loja, USNM slide #116219 [USNMENT01769438]. 15, Pantographa scripturalis (Guenée), lectotype adult female, Brazil [NHMUK01396720]. 16, P. scripturalis, lectotype labels. 17, P. scripturalis, male genitalia, Brazil, Parana, Castro, USNM slide #107232 [USNMENT01895419]. 18, P. scripturalis, female genitalia, Brazil, Rio de Janeiro, USNM slide #105696 [USNMENT01899000]." figureDoi="http://doi.org/10.5281/zenodo.10409851" httpUri="https://zenodo.org/record/10409851/files/figure.png" pageId="4" pageNumber="347">1213</figureCitation>
): Uncus shape variable, v-shaped or pencil-like from tegumen, longer than valvae, two to three times as long as tegumen, uncus apex variable in shape, pointed or rounded with or without medial cleft distally, ventrally with long, non-deciduous setae; subscaphium membranous, or lightly or highly sclerotized, basally with or without microspines; valvae same width lengthwise or broader distally, approaching square shape in some species, fibula finger-like from dorsal valva base near costa base directed towards distal valva margin, slightly protruding, or flat with setae, saccular medial lobe variable in shape from small, round to elongate and flat, sacculus with dorsobasal process with setae, variable in shape, present or absent; transtilla medially flat with a suture, or medially broad, two sclerotized arms as wide as costa of valva, setae present on ventral edge of transtilla arms; juxta slightly sclerotized, plate-like, depth of medial gap/split of juxta &lt;10% of dorsoventral length of juxta; saccus v-shaped or rounded and shorter than wide, keel present dorsoanteriorly; coremata lateral to vinculum, round, about
<quantity id="4CA3ACF4674CFFD4C0A0439655E79871" box="[151,236,1015,1041]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" pageId="4" pageNumber="347" unit="mm" value="0.1">0.1 mm</quantity>
in diameter, bearing long, simple hair-like setae; phallus with one or several long cornuti, the longest a little less than half the length of the phallus, and without or without microspines posteriorly, ductus ejaculatorius close to anterior end.
<emphasis id="B92FDD03674CFFD4C17F445E570D9839" box="[328,518,1087,1113]" italics="true" pageId="4" pageNumber="347">Female genitalia</emphasis>
(
<figureCitation id="13601D94674CFFD4C222445E57519839" box="[533,602,1087,1113]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="4" pageNumber="347">Fig. 9</figureCitation>
): Anterior apophyses twice as long as posterior apophyses; antrum distally membranous or lightly sclerotized; ductus seminalis at anterior end of 7
<superScript id="7C2EAC59674CFFD4C3F3440356DA9810" attach="left" box="[964,977,1122,1136]" fontSize="6" pageId="4" pageNumber="347">th</superScript>
segment; ductus bursae twice as long or longer than corpus bursae; corpus bursae round, with very light scobination; signum single, round, not elevated, without transverse axis.
</paragraph>
<paragraph id="8BE40111674CFFD4C0F044AF56889951" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="347">
<emphasis id="B92FDD03674CFFD4C0F044AF54A39888" bold="true" box="[199,424,1230,1256]" pageId="4" pageNumber="347">Species examined:</emphasis>
<taxonomicName id="4C5B7A92674CFFD4C18144AE56339889" authority="(Dognin, 1905)" baseAuthorityName="Dognin" baseAuthorityYear="1905" box="[438,824,1230,1257]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="nitidalis">
<emphasis id="B92FDD03674CFFD4C18144AE57739889" box="[438,632,1231,1257]" italics="true" pageId="4" pageNumber="347">Syllepte nitidalis</emphasis>
(
<bibRefCitation id="EFCA7CE0674CFFD4C2BA44AE563B9889" author="Dognin, P." box="[653,816,1230,1257]" pageId="4" pageNumber="347" pagination="61 - 90" refId="ref11494" refString="Dognin, P. (1905) Heteroceres nouveaux de l'Amerique du Sud. Annales de la Societe Entomologique de Belgique, Bruxelles, 49, 61 - 90. Available from: https: // www. biodiversitylibrary. org / item / 44643 # page / 68 / mode / 2 up (accessed 29 November 2023)" type="journal article" year="1905">Dognin, 1905</bibRefCitation>
)
</taxonomicName>
[
<typeStatus id="54E0BFB3674CFFD4C37B44AE56749889" box="[844,895,1231,1257]" pageId="4" pageNumber="347">type</typeStatus>
specimens examined],
<taxonomicName id="4C5B7A92674CFFD4C4AE44AE5440996D" authority="(Lederer, 1863)" baseAuthorityName="Lederer" baseAuthorityYear="1863" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="serratilinealis">
<emphasis id="B92FDD03674CFFD4C4AE44AE50979889" box="[1177,1436,1231,1257]" italics="true" pageId="4" pageNumber="347">Syllepte serratilinealis</emphasis>
(Lederer, 1863)
</taxonomicName>
[
<typeStatus id="54E0BFB3674CFFD4C16A4492549B996D" box="[349,400,1267,1293]" pageId="4" pageNumber="347">type</typeStatus>
painting],
<taxonomicName id="4C5B7A92674CFFD4C222449256A2996D" authority="Hampson, 1912" authorityName="Hampson" authorityYear="1912" box="[533,937,1267,1293]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="leucinalis">
<emphasis id="B92FDD03674CFFD4C222449257ED996D" box="[533,742,1267,1293]" italics="true" pageId="4" pageNumber="347">Syllepte leucinalis</emphasis>
Hampson, 1912
</taxonomicName>
[
<typeStatus id="54E0BFB3674CFFD4C38C449256E5996D" box="[955,1006,1267,1293]" pageId="4" pageNumber="347">type</typeStatus>
examined externally]. Specimens of undescribed species were examined from
<collectingCountry id="F34C4181674CFFD4C25B457657C99951" box="[620,706,1303,1329]" name="Mexico" pageId="4" pageNumber="347">Mexico</collectingCountry>
south to
<collectingCountry id="F34C4181674CFFD4C31E457656749951" box="[809,895,1303,1329]" name="Bolivia" pageId="4" pageNumber="347">Bolivia</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C341529A674CFFD4C0F0455A54399ADD" pageId="4" pageNumber="347" type="discussion">
<paragraph id="8BE40111674CFFD4C0F0455A56FC99A1" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="347">
<emphasis id="B92FDD03674CFFD4C0F0455A543F9935" bold="true" box="[199,308,1339,1365]" pageId="4" pageNumber="347">Remarks</emphasis>
: This genus represents several undescribed species throughout the Western Hemisphere that are difficult to separate externally. Although there are long series of specimens for Costa Rican species, material from other areas in the Western Hemisphere, particularly
<collectingCountry id="F34C4181674CFFD4C2E445E2562299FD" box="[723,809,1411,1437]" name="Mexico" pageId="4" pageNumber="347">Mexico</collectingCountry>
and South America, is scarce. This genus needs revision, and more material needs to be located and subjected to comparative analysis.
</paragraph>
<paragraph id="8BE40111674CFFD4C0F045AA54399ADD" blockId="4.[151,1437,151,2013]" pageId="4" pageNumber="347">
<taxonomicName id="4C5B7A92674CFFD4C0F045AA54CF9985" baseAuthorityName="Lederer" baseAuthorityYear="1863" box="[199,452,1483,1509]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="serratilinealis">
<emphasis id="B92FDD03674CFFD4C0F045AA54CF9985" box="[199,452,1483,1509]" italics="true" pageId="4" pageNumber="347">Syllepte serratilinealis</emphasis>
</taxonomicName>
(described from
<collectingCountry id="F34C4181674CFFD4C2B345AA57F79985" box="[644,764,1483,1509]" name="Venezuela" pageId="4" pageNumber="347">Venezuela</collectingCountry>
),
<taxonomicName id="4C5B7A92674CFFD4C33845AA568C9985" baseAuthorityName="Dognin" baseAuthorityYear="1905" box="[783,903,1483,1509]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="nitidalis">
<emphasis id="B92FDD03674CFFD4C33845AA568C9985" box="[783,903,1483,1509]" italics="true" pageId="4" pageNumber="347">S. nitidalis</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C5B7A92674CFFD4C3F245AA51459985" authorityName="Hampson" authorityYear="1912" box="[965,1102,1483,1509]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="leucinalis">
<emphasis id="B92FDD03674CFFD4C3F245AA51459985" box="[965,1102,1483,1509]" italics="true" pageId="4" pageNumber="347">S. leucinalis</emphasis>
</taxonomicName>
were not included in
<bibRefCitation id="EFCA7CE0674CFFD4C57745AA55D29A69" author="Amsel, H. G." pageId="4" pageNumber="347" pagination="1 - 334" refId="ref11387" refString="Amsel, H. G. (1956 - 1957) Microlepidoptera Venezolana I and II. Boletin de Entomologia Venezolana, 10 (1 - 4), 1 - 334, pls. 1 - 110." type="journal article" year="1956">Amsels (1956</bibRefCitation>
7) work about Venezuelan microlepidoptera.
<bibRefCitation id="EFCA7CE0674CFFD4C2E9458E56839A69" author="Munroe, E." box="[734,904,1518,1545]" pageId="4" pageNumber="347" pagination="58 - 79" refId="ref12300" refString="Munroe, E. (1995) Spilomelinae. In: Heppner, J. B. (Ed.), Check List. Part 2. Atlas of Neotropical Lepidoptera. Brill / Flora and Fauna Books, Gainesville, Florida, pp. 58 - 79." type="book chapter" year="1995">Munroe (1995)</bibRefCitation>
synonymized
<taxonomicName id="4C5B7A92674CFFD4C41C458E51B89A69" authorityName="Hampson" authorityYear="1912" box="[1067,1203,1519,1545]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="leucinalis">
<emphasis id="B92FDD03674CFFD4C41C458E51B89A69" box="[1067,1203,1519,1545]" italics="true" pageId="4" pageNumber="347">S. leucinalis</emphasis>
</taxonomicName>
described from
<collectingCountry id="F34C4181674CFFD4C55E458E50979A69" box="[1385,1436,1519,1545]" name="Peru" pageId="4" pageNumber="347">Peru</collectingCountry>
with
<taxonomicName id="4C5B7A92674CFFD4C0E5467254869A4D" baseAuthorityName="Lederer" baseAuthorityYear="1863" box="[210,397,1555,1581]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="serratilinealis">
<emphasis id="B92FDD03674CFFD4C0E5467254869A4D" box="[210,397,1555,1581]" italics="true" pageId="4" pageNumber="347">S. serratilinealis</emphasis>
</taxonomicName>
and newly combined it under
<taxonomicName id="4C5B7A92674CFFD4C2DA4672568A9A4D" authorityName="Solis &amp; Mally &amp; Hayden &amp; Nuss" authorityYear="2023" box="[749,897,1555,1581]" class="Insecta" family="Crambidae" genus="Pantographa" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674CFFD4C2DA4672568A9A4D" box="[749,897,1555,1581]" italics="true" pageId="4" pageNumber="347">Pantographa</emphasis>
</taxonomicName>
.
<bibRefCitation id="EFCA7CE0674CFFD4C3A64672513F9A4D" author="Becker, V. O." box="[913,1076,1555,1581]" pageId="4" pageNumber="347" pagination="20220093" refId="ref11426" refString="Becker, V. O. (2023) The identity of Syllepte incomptalis Hubner (Lepidoptera: Crambidae: Spilomelinae) with synonymies, new combinations and new species. Revista Brasileira de Entomologia, Sao Paulo, 67 (1), e 20220093, 1 - 11. https: // doi. org / 10.1590 / 1806 - 9665 - rbent- 2022 - 0093" type="journal article" year="2023">Becker (2023)</bibRefCitation>
resurrected
<taxonomicName id="4C5B7A92674CFFD4C4F5467255C49A31" authority="Dognin, 1905" authorityName="Dognin" authorityYear="1905" class="Insecta" family="Crambidae" genus="Neomabra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674CFFD4C4F5467250339A4D" box="[1218,1336,1555,1581]" italics="true" pageId="4" pageNumber="347">Neomabra</emphasis>
<bibRefCitation id="EFCA7CE0674CFFD4C576467255C49A31" author="Dognin, P." pageId="4" pageNumber="347" pagination="61 - 90" refId="ref11494" refString="Dognin, P. (1905) Heteroceres nouveaux de l'Amerique du Sud. Annales de la Societe Entomologique de Belgique, Bruxelles, 49, 61 - 90. Available from: https: // www. biodiversitylibrary. org / item / 44643 # page / 68 / mode / 2 up (accessed 29 November 2023)" type="journal article" year="1905">Dognin, 1905</bibRefCitation>
</taxonomicName>
(
<typeStatus id="54E0BFB3674CFFD4C0E84656541A9A31" box="[223,273,1591,1617]" pageId="4" pageNumber="347">type</typeStatus>
species:
<taxonomicName id="4C5B7A92674CFFD4C14E465654F89A31" baseAuthorityName="Dognin" baseAuthorityYear="1905" box="[377,499,1591,1617]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="nitidalis">
<emphasis id="B92FDD03674CFFD4C14E465654F89A31" box="[377,499,1591,1617]" italics="true" pageId="4" pageNumber="347">S. nitidalis</emphasis>
</taxonomicName>
) as a valid genus, re-combined
<taxonomicName id="4C5B7A92674CFFD4C36A465656DD9A31" baseAuthorityName="Dognin" baseAuthorityYear="1905" box="[861,982,1591,1617]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="nitidalis">
<emphasis id="B92FDD03674CFFD4C36A465656DD9A31" box="[861,982,1591,1617]" italics="true" pageId="4" pageNumber="347">S. nitidalis</emphasis>
</taxonomicName>
, newly combined
<taxonomicName id="4C5B7A92674CFFD4C49F465650699A31" baseAuthorityName="Lederer" baseAuthorityYear="1863" box="[1192,1378,1591,1617]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="serratilinealis">
<emphasis id="B92FDD03674CFFD4C49F465650699A31" box="[1192,1378,1591,1617]" italics="true" pageId="4" pageNumber="347">S. serratilinealis</emphasis>
</taxonomicName>
with this genus, and newly synonymized
<taxonomicName id="4C5B7A92674CFFD4C219463A57BC9A15" authorityName="Hampson" authorityYear="1912" box="[558,695,1627,1653]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="leucinalis">
<emphasis id="B92FDD03674CFFD4C219463A57BC9A15" box="[558,695,1627,1653]" italics="true" pageId="4" pageNumber="347">S. leucinalis</emphasis>
</taxonomicName>
with the latter species. In this paper we remove
<taxonomicName id="4C5B7A92674CFFD4C4E1463A50549A15" authorityName="Hampson" authorityYear="1912" box="[1238,1375,1627,1653]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="leucinalis">
<emphasis id="B92FDD03674CFFD4C4E1463A50549A15" box="[1238,1375,1627,1653]" italics="true" pageId="4" pageNumber="347">S. leucinalis</emphasis>
</taxonomicName>
from synonymy based on reasons in the previous paragraph and the
<typeStatus id="54E0BFB3674CFFD4C36A461E56869AF9" box="[861,909,1663,1689]" pageId="4" pageNumber="347">type</typeStatus>
specimen needs to be dissected and studied in greater detail.
</paragraph>
</subSubSection>
<subSubSection id="C341529A674CFFD5C0F046A654B49D7D" lastPageId="5" lastPageNumber="348" pageId="4" pageNumber="347" type="materials_examined">
<paragraph id="8BE40111674CFFD5C0F046A654B49D7D" blockId="4.[151,1437,151,2013]" lastBlockId="5.[151,1437,151,285]" lastPageId="5" lastPageNumber="348" pageId="4" pageNumber="347">
The type material of
<taxonomicName id="4C5B7A92674CFFD4C18546A657909A81" authority="(Dognin)" baseAuthorityName="Dognin" baseAuthorityYear="1905" box="[434,667,1735,1761]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="species" species="nitidalis">
<emphasis id="B92FDD03674CFFD4C18546A657219A81" box="[434,554,1735,1761]" italics="true" pageId="4" pageNumber="347">S. nitidalis</emphasis>
(Dognin)
</taxonomicName>
,
<emphasis id="B92FDD03674CFFD4C29146A656119A81" bold="true" box="[678,794,1735,1761]" pageId="4" pageNumber="347">rev. comb</emphasis>
., is a series of males from
<collectingRegion id="499FCFF3674CFFD4C47F46A651709A81" box="[1096,1147,1735,1761]" country="Ecuador" name="Loja" pageId="4" pageNumber="347">Loja</collectingRegion>
and the valley of Zamora from Ecuador all with a small white label with a typewritten “Dognin Collection”, and with handwritten labels “
<taxonomicName id="4C5B7A92674CFFD4C096476E54139B49" authorityName="Dognin" authorityYear="1905" box="[161,280,1807,1833]" class="Insecta" family="Crambidae" genus="Neomabra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="genus">Neomabra</taxonomicName>
nitdalis, type
<specimenCount id="9D5DCA98674CFFD4C183476E54C29B49" box="[436,457,1807,1833]" count="1" pageId="4" pageNumber="347" type="male"></specimenCount>
Dgn”.
<location id="8E8457CA674CFFD4C220476E57379B49" LSID="urn:lsid:plazi:treatment:03F2B007674BFFD5C0A043B35069981D:8E8457CA674CFFD4C220476E57379B49" box="[535,572,1807,1833]" country="United States of America" county="Paul Dognin" municipality="William Schaus" name="We" pageId="4" pageNumber="347" stateProvince="Loja">We</location>
designate
<specimenCount id="9D5DCA98674CFFD4C285476E56419B49" box="[690,842,1807,1833]" count="1" pageId="4" pageNumber="347" type="generic">one specimen</specimenCount>
(
<figureCitation id="13601D94674CFFD4C36F476E56D59B49" box="[856,990,1807,1833]" captionStart="FIGURES 1018" captionStartId="14.[151,260,1815,1839]" captionTargetBox="[230,1358,151,1796]" captionTargetId="figure-23@14.[230,1358,151,1803]" captionTargetPageId="14" captionText="FIGURES 1018. 10, Syllepte nitidalis (Dognin), lectotype adult male, Ecuador, Environs de Loja [USNMENT01433441]. 11, S. nitidalis,lectotypelabels.12,S.nitidalis,malegenitalia,paralectotype,Ecuador,Zamora,USNMslide#111085[USNMENT01769261]. 13, S. nitidalis, phallus, paralectotype, Ecuador, Zamora, USNM slide #111085 [USNMENT01769261]. 14, S. nitidalis, male modified structure and final segment, paralectotype, Ecuador, Environs de Loja, USNM slide #116219 [USNMENT01769438]. 15, Pantographa scripturalis (Guenée), lectotype adult female, Brazil [NHMUK01396720]. 16, P. scripturalis, lectotype labels. 17, P. scripturalis, male genitalia, Brazil, Parana, Castro, USNM slide #107232 [USNMENT01895419]. 18, P. scripturalis, female genitalia, Brazil, Rio de Janeiro, USNM slide #105696 [USNMENT01899000]." figureDoi="http://doi.org/10.5281/zenodo.10409851" httpUri="https://zenodo.org/record/10409851/files/figure.png" pageId="4" pageNumber="347">Figs. 1014</figureCitation>
) [USNMENT01433441] to stabilize the name of this species with the following additional labels (
<figureCitation id="13601D94674CFFD4C338475256569B2D" box="[783,861,1843,1869]" captionStart="FIGURES 1018" captionStartId="14.[151,260,1815,1839]" captionTargetBox="[230,1358,151,1796]" captionTargetId="figure-23@14.[230,1358,151,1803]" captionTargetPageId="14" captionText="FIGURES 1018. 10, Syllepte nitidalis (Dognin), lectotype adult male, Ecuador, Environs de Loja [USNMENT01433441]. 11, S. nitidalis,lectotypelabels.12,S.nitidalis,malegenitalia,paralectotype,Ecuador,Zamora,USNMslide#111085[USNMENT01769261]. 13, S. nitidalis, phallus, paralectotype, Ecuador, Zamora, USNM slide #111085 [USNMENT01769261]. 14, S. nitidalis, male modified structure and final segment, paralectotype, Ecuador, Environs de Loja, USNM slide #116219 [USNMENT01769438]. 15, Pantographa scripturalis (Guenée), lectotype adult female, Brazil [NHMUK01396720]. 16, P. scripturalis, lectotype labels. 17, P. scripturalis, male genitalia, Brazil, Parana, Castro, USNM slide #107232 [USNMENT01895419]. 18, P. scripturalis, female genitalia, Brazil, Rio de Janeiro, USNM slide #105696 [USNMENT01899000]." figureDoi="http://doi.org/10.5281/zenodo.10409851" httpUri="https://zenodo.org/record/10409851/files/figure.png" pageId="4" pageNumber="347">Fig. 11</figureCitation>
) “Environs de
<collectingRegion id="499FCFF3674CFFD4C4344752513D9B2D" box="[1027,1078,1843,1869]" country="Ecuador" name="Loja" pageId="4" pageNumber="347">Loja</collectingRegion>
/ Equateur/1892”, “(
<taxonomicName id="4C5B7A92674CFFD4C525475250879B2D" authorityName="Dognin" authorityYear="1905" box="[1298,1420,1843,1869]" class="Insecta" family="Crambidae" genus="Neomabra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="4" pageNumber="347" phylum="Arthropoda" rank="genus">Neomabra</taxonomicName>
)/ (nitidalis)/nov. gen./nov. sp./
<specimenCount id="9D5DCA98674CFFD4C1E0473654E09B11" box="[471,491,1879,1905]" count="1" pageId="4" pageNumber="347" type="male"></specimenCount>
/ Warren -04”, “TypeNo./29669/
<collectingCountry id="F34C4181674CFFD4C355473656879B11" box="[866,908,1879,1905]" name="United States of America" pageId="4" pageNumber="347">U.S.</collectingCountry>
N.M (red label)”, undissected, as the
<typeStatus id="54E0BFB3674CFFD4C518473650929B11" box="[1327,1433,1879,1905]" pageId="4" pageNumber="347" type="lectotype">lectotype</typeStatus>
,
<typeStatus id="54E0BFB3674CFFD4C0A0471A55F59BF5" box="[151,254,1915,1941]" pageId="4" pageNumber="347" type="lectotype">lectotype</typeStatus>
designation, and the other
<specimenCount id="9D5DCA98674CFFD4C20E471A57FB9BF5" box="[569,752,1915,1941]" count="3" pageId="4" pageNumber="347" type="generic">three specimens</specimenCount>
are
<typeStatus id="54E0BFB3674CFFD4C313471A56CD9BF5" box="[804,966,1915,1941]" pageId="4" pageNumber="347" type="paralectotype">paralectotypes</typeStatus>
from
<collectingRegion id="499FCFF3674CFFD4C438471A514E9BF5" box="[1039,1093,1915,1941]" country="Ecuador" name="Loja" pageId="4" pageNumber="347">Loja</collectingRegion>
, 1889 [USNMENT01769438,
<collectionCode id="ED4A99D4674CFFD4C0A047FE55E39BD9" box="[151,232,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="347" type="Museum">USNM</collectionCode>
slide #116219],
<collectingRegion id="499FCFF3674CFFD4C19247FE54D09BD9" box="[421,475,1951,1977]" country="Ecuador" name="Loja" pageId="4" pageNumber="347">Loja</collectingRegion>
, 1893 [USNMENT01552814], and
<location id="8E8457CA674CFFD4C35847FE56CD9BD9" LSID="urn:lsid:plazi:treatment:03F2B007674BFFD5C0A043B35069981D:8E8457CA674CFFD4C35847FE56CD9BD9" box="[879,966,1951,1977]" country="United States of America" county="Paul Dognin" municipality="William Schaus" name="Zamora" pageId="4" pageNumber="347" stateProvince="Loja">Zamora</location>
(no year) [USNMENT01769251,
<collectionCode id="ED4A99D4674CFFD4C57C47FE50979BD9" box="[1355,1436,1951,1977]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="347" type="Museum">USNM</collectionCode>
slide #111085].
<location id="8E8457CA674CFFD4C17947A254D69BBD" LSID="urn:lsid:plazi:treatment:03F2B007674BFFD5C0A043B35069981D:8E8457CA674CFFD4C17947A254D69BBD" box="[334,477,1987,2013]" country="United States of America" county="Paul Dognin" municipality="William Schaus" name="Additionally" pageId="4" pageNumber="347" stateProvince="Loja">Additionally</location>
, there is another specimen [USNMENT01899001] from
<collectingRegion id="499FCFF3674CFFD4C45C47A251959BBD" box="[1131,1182,1987,2013]" country="Ecuador" name="Loja" pageId="4" pageNumber="347">Loja</collectingRegion>
that is not part of the
<location id="8E8457CA674DFFD5C0A040F655E09CD1" LSID="urn:lsid:plazi:treatment:03F2B007674BFFD5C0A043B35069981D:8E8457CA674DFFD5C0A040F655E09CD1" box="[151,235,151,177]" country="United States of America" county="Paul Dognin" municipality="William Schaus" name="Dognin" pageId="5" pageNumber="348" stateProvince="Loja">Dognin</location>
type series.
<location id="8E8457CA674DFFD5C14D40F654EF9CD1" LSID="urn:lsid:plazi:treatment:03F2B007674BFFD5C0A043B35069981D:8E8457CA674DFFD5C14D40F654EF9CD1" box="[378,484,151,177]" country="United States of America" county="Paul Dognin" municipality="William Schaus" name="Although" pageId="5" pageNumber="348" stateProvince="Loja">Although</location>
it has a similar locality label “Environs de
<collectingRegion id="499FCFF3674DFFD5C3E340F6510C9CD1" box="[980,1031,151,177]" country="Ecuador" name="Loja" pageId="5" pageNumber="348">Loja</collectingRegion>
/ Equateur/1889” and a handwritten label by
<location id="8E8457CA674DFFD5C13540DA545D9CB5" LSID="urn:lsid:plazi:treatment:03F2B007674BFFD5C0A043B35069981D:8E8457CA674DFFD5C13540DA545D9CB5" box="[258,342,187,213]" country="United States of America" county="Paul Dognin" municipality="William Schaus" name="Dognin" pageId="5" pageNumber="348" stateProvince="Loja">Dognin</location>
<taxonomicName id="4C5B7A92674DFFD5C15C40DA54EF9CB5" authorityName="Dognin" authorityYear="1905" box="[363,484,187,213]" class="Insecta" family="Crambidae" genus="Neomabra" higherTaxonomySource="GBIF,CoL" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">Neomabra</taxonomicName>
/nitidalis/Dgn”, it lacks the word “type” or male symbol, and there is a separate typewritten label that reveals this specimen was part of the “Schaus/Collection” and probably a gift to
<collectingMunicipality id="6B809B6B674DFFD5C57540BE55ED9D7D" pageId="5" pageNumber="348">William Schaus</collectingMunicipality>
from
<collectingCounty id="6285799D674DFFD5C11C416254B79D7D" box="[299,444,259,285]" pageId="5" pageNumber="348">Paul Dognin</collectingCounty>
.
</paragraph>
</subSubSection>
<subSubSection id="C341529A674DFFD5C0A0410E5069981D" pageId="5" pageNumber="348" type="discussion">
<paragraph id="8BE40111674DFFD5C0A0410E57439DE8" blockId="5.[151,584,366,393]" box="[151,584,366,393]" pageId="5" pageNumber="348">
<heading id="D0ACB67D674DFFD5C0A0410E57439DE8" bold="true" box="[151,584,366,393]" fontSize="11" level="1" pageId="5" pageNumber="348" reason="1">
<emphasis id="B92FDD03674DFFD5C0A0410E57439DE8" bold="true" box="[151,584,366,393]" pageId="5" pageNumber="348">
Placement of
<taxonomicName id="4C5B7A92674DFFD5C176410F54AB9DE9" ID-CoL="92NJG" authorityName="Hubner" authorityYear="1823" box="[321,416,366,393]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C176410F54AB9DE9" bold="true" box="[321,416,366,393]" italics="true" pageId="5" pageNumber="348">Syllepte</emphasis>
</taxonomicName>
in
<taxonomicName id="4C5B7A92674DFFD5C1F2410F57439DE8" box="[453,584,366,392]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BE40111674DFFD5C0A041D6500D9E79" blockId="5.[151,1437,439,1149]" pageId="5" pageNumber="348">
We place
<taxonomicName id="4C5B7A92674DFFD5C13E41D654699DB1" authorityName="Hubner" authorityYear="1823" box="[265,354,439,465]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C13E41D654699DB1" box="[265,354,439,465]" italics="true" pageId="5" pageNumber="348">Syllepte</emphasis>
</taxonomicName>
in
<taxonomicName id="4C5B7A92674DFFD5C1B041D657D49DB1" authority="(Mally et al. 2019)" baseAuthorityName="Mally" baseAuthorityYear="2019" box="[391,735,439,465]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">
Agroterini (
<bibRefCitation id="EFCA7CE0674DFFD5C23A41D657DC9DB1" author="Mally, R. &amp; Hayden, J. E. &amp; Neinhuis, C. &amp; Jordal, B. H. &amp; Nuss, M." box="[525,727,439,465]" pageId="5" pageNumber="348" pagination="141 - 204" refId="ref12102" refString="Mally, R., Hayden, J. E., Neinhuis, C., Jordal, B. H. &amp; Nuss, M. (2019) The phylogenetic systematics of Spilomelinae and Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) inferred from DNA and morphology. Arthropod Systematics and Phylogeny, 77 (1), 141 - 204. https: // doi. org / 10.26049 / ASP 77 - 1 - 2019 - 07" type="journal article" year="2019">
Mally
<emphasis id="B92FDD03674DFFD5C26D41D6579F9DB1" box="[602,660,439,465]" italics="true" pageId="5" pageNumber="348">et al.</emphasis>
2019
</bibRefCitation>
)
</taxonomicName>
based on morphological characteristics discussed below; this placement agrees with
<bibRefCitation id="EFCA7CE0674DFFD5C1AF41BA574D9D95" author="Munroe, E." box="[408,582,474,501]" pageId="5" pageNumber="348" pagination="58 - 79" refId="ref12300" refString="Munroe, E. (1995) Spilomelinae. In: Heppner, J. B. (Ed.), Check List. Part 2. Atlas of Neotropical Lepidoptera. Brill / Flora and Fauna Books, Gainesville, Florida, pp. 58 - 79." type="book chapter" year="1995">Munroe (1995)</bibRefCitation>
. Additionally,
<taxonomicName id="4C5B7A92674DFFD5C2DE41BA56499D95" authorityName="Hubner" authorityYear="1823" box="[745,834,475,501]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C2DE41BA56499D95" box="[745,834,475,501]" italics="true" pageId="5" pageNumber="348">Syllepte</emphasis>
</taxonomicName>
is the
<typeStatus id="54E0BFB3674DFFD5C3BC41BA56B09D95" box="[907,955,475,501]" pageId="5" pageNumber="348">type</typeStatus>
species of Syleptinae [sic] Swinhoe, 1900, and we therefore establish Syleptinae as a subjective synonym of
<taxonomicName id="4C5B7A92674DFFD5C342419E519C9E79" authority="Acloque, 1897" authorityName="Acloque" authorityYear="1897" box="[885,1175,511,537]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini Acloque, 1897</taxonomicName>
,
<emphasis id="B92FDD03674DFFD5C495419E51F49E79" bold="true" box="[1186,1279,511,537]" pageId="5" pageNumber="348">syn. rev</emphasis>
.
</paragraph>
<paragraph id="8BE40111674DFFD5C0F042425069981D" blockId="5.[151,1437,439,1149]" pageId="5" pageNumber="348">
Externally,the forewing maculation resembles that of some species of
<taxonomicName id="4C5B7A92674DFFD5C38F424251BE9E5D" authority="Snellen, 1875" authorityName="Snellen" authorityYear="1875" box="[952,1205,546,573]" class="Insecta" family="Crambidae" genus="Hileithia" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C38F424251179E5D" box="[952,1052,547,573]" italics="true" pageId="5" pageNumber="348">Hileithia</emphasis>
Snellen, 1875
</taxonomicName>
and
<taxonomicName id="4C5B7A92674DFFD5C4D4424254399E01" authority="Lederer, 1863" authorityName="Lederer" authorityYear="1863" class="Insecta" family="Crambidae" genus="Herpetogramma" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C4D4424250979E5C" box="[1251,1436,547,572]" italics="true" pageId="5" pageNumber="348">Herpetogramma</emphasis>
Lederer, 1863
</taxonomicName>
both placed in
<taxonomicName id="4C5B7A92674DFFD5C1EA422657B09E01" box="[477,699,583,609]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Herpetogrammatini">Herpetogrammatini</taxonomicName>
, but the four relatively straight hindwing lines in Hübners (1819 1823) illustration of
<taxonomicName id="4C5B7A92674DFFD5C149420A57169EE5" box="[382,541,619,645]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="incomptalis">
<emphasis id="B92FDD03674DFFD5C149420A57169EE5" box="[382,541,619,645]" italics="true" pageId="5" pageNumber="348">S. incomptalis</emphasis>
</taxonomicName>
are very unusual in
<taxonomicName id="4C5B7A92674DFFD5C336420A56939EE5" box="[769,920,619,645]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="345" phylum="Arthropoda" rank="subFamily" subFamily="Spilomelinae">Spilomelinae</taxonomicName>
; we therefore interpret them as artistic liberty. The morphology of the genitalia in particular shares several synapomorphies with
<taxonomicName id="4C5B7A92674DFFD5C47742EE51CD9EC9" authorityName="Warren" authorityYear="1890" box="[1088,1222,655,681]" class="Insecta" family="Crambidae" genus="Haritalodes" higherTaxonomySource="CoL,GBIF" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C47742EE51CD9EC9" box="[1088,1222,655,681]" italics="true" pageId="5" pageNumber="348">Haritalodes</emphasis>
</taxonomicName>
(also a member of the
<taxonomicName id="4C5B7A92674DFFD5C08942D2543E9EAD" authorityName="Acloque" authorityYear="1897" box="[190,309,691,717]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
) as reconstructed in
<bibRefCitation id="EFCA7CE0674DFFD5C22F42D257E89EAD" author="Mally, R. &amp; Hayden, J. E. &amp; Neinhuis, C. &amp; Jordal, B. H. &amp; Nuss, M." box="[536,739,691,717]" pageId="5" pageNumber="348" pagination="141 - 204" refId="ref12102" refString="Mally, R., Hayden, J. E., Neinhuis, C., Jordal, B. H. &amp; Nuss, M. (2019) The phylogenetic systematics of Spilomelinae and Pyraustinae (Lepidoptera: Pyraloidea: Crambidae) inferred from DNA and morphology. Arthropod Systematics and Phylogeny, 77 (1), 141 - 204. https: // doi. org / 10.26049 / ASP 77 - 1 - 2019 - 07" type="journal article" year="2019">
Mally
<emphasis id="B92FDD03674DFFD5C25642D257929EAD" box="[609,665,691,717]" italics="true" pageId="5" pageNumber="348">et al.</emphasis>
(2019
</bibRefCitation>
,
<figureCitation id="13601D94674DFFD5C2DA42D2563A9EAD" box="[749,817,691,717]" captionStart="FIGURES 19" captionStartId="13.[151,264,1837,1861]" captionTargetBox="[154,1433,170,1813]" captionTargetId="figure-23@13.[154,1433,170,1813]" captionTargetPageId="13" captionText="FIGURES 19. Syllepte incomptalis Hübner. 1, Original painting. 2, Neotype adult male, Costa Rica, Alajuela, Area de Conservación Guanacaste [USNMENT01899038]. 3, Neotype labels. 4, Male genitalia, Costa Rica,Alajuela,Area Conservación Guanacaste, USNM slide #116312 [USNMENT01894132]. 5, Sclerotized structure with modified scales on male 7th tergite, USNM slide #116223 [USNMENT01769416]. 6, Phallus, USNM slide #116312 [USNMENT01894132]. 7, Male 8th segment, USNM slide #116312 [USNMENT01894132]. 8, Tympanal organ, USNM slide #116312 [USNMENT01894132]. 9, Female genitalia, Costa Rica, Alajuela, Area Conservación Guanacaste, USNM slide #116314 [USNMENT01894003]." figureDoi="http://doi.org/10.5281/zenodo.10409849" httpUri="https://zenodo.org/record/10409849/files/figure.png" pageId="5" pageNumber="348">Fig. 2</figureCitation>
). The uncus is relatively membranous and non-capitate. The uncus setae are unsplit, thin, and distributed laterally on the uncus as well as dorsally. In
<taxonomicName id="4C5B7A92674DFFD5C4AA42B6501E9E91" authorityName="Acloque" authorityYear="1897" box="[1181,1301,727,753]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
, the valva is roughly rectangular and the costa is often straight. The base of the costa and the inner valva in general are setose. A small, setiferous protrusion is present on the face of the valva between the fibula and the dorsal edge of the sacculus. The saccus of
<taxonomicName id="4C5B7A92674DFFD5C10B4322549E9F3D" authorityName="Hubner" authorityYear="1823" box="[316,405,835,861]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="genus">
<emphasis id="B92FDD03674DFFD5C10B4322549E9F3D" box="[316,405,835,861]" italics="true" pageId="5" pageNumber="348">Syllepte</emphasis>
</taxonomicName>
is as wide as long, which fits in the range of variation of
<taxonomicName id="4C5B7A92674DFFD5C41F432251AB9F3D" authorityName="Acloque" authorityYear="1897" box="[1064,1184,835,861]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
; among
<taxonomicName id="4C5B7A92674DFFD5C536432250939F3D" box="[1281,1432,835,861]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="345" phylum="Arthropoda" rank="subFamily" subFamily="Spilomelinae">Spilomelinae</taxonomicName>
, such a broad saccus is found elsewhere. In the female genitalia,
<taxonomicName id="4C5B7A92674DFFD5C360430656FE9FE1" box="[855,1013,871,897]" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="incomptalis">
<emphasis id="B92FDD03674DFFD5C360430656FE9FE1" box="[855,1013,871,897]" italics="true" pageId="5" pageNumber="348">S. incomptalis</emphasis>
</taxonomicName>
has one round signum. The agroterine signum is variable, including many species with two round signa, but where it is single, it is round or slightly elongate, and never with a transverse ridge, which occurs in
<taxonomicName id="4C5B7A92674DFFD5C37943CE51229FA9" box="[846,1065,943,969]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Herpetogrammatini">Herpetogrammatini</taxonomicName>
and
<taxonomicName id="4C5B7A92674DFFD5C45443CE51C49FA9" box="[1123,1231,943,969]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Hydririni">Hydririni</taxonomicName>
. The antrum of
<taxonomicName id="4C5B7A92674DFFD5C5B043CE54119F8D" class="Insecta" family="Crambidae" genus="Syllepte" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="species" species="incomptalis">
<emphasis id="B92FDD03674DFFD5C5B043CE54119F8D" italics="true" pageId="5" pageNumber="348">S. incomptalis</emphasis>
</taxonomicName>
is broad, with a wide, poorly defined colliculum that does not have a longitudinal membranous strip, although such a colliculum occurs in many
<taxonomicName id="4C5B7A92674DFFD5C240439657E49871" authorityName="Acloque" authorityYear="1897" box="[631,751,1015,1041]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
. The ovoid corpus bursae is well-demarcated from the narrow ductus bursae. The papillae anales are directed ventrad at a 90-degree angle from the axis of the body, a conformation found in
<taxonomicName id="4C5B7A92674DFFD5C0CB445E547A9839" authorityName="Acloque" authorityYear="1897" box="[252,369,1087,1113]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
and some
<taxonomicName id="4C5B7A92674DFFD5C1DB445E57889839" box="[492,643,1087,1113]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Margaroniini">Margaroniini</taxonomicName>
. Finally, among
<taxonomicName id="4C5B7A92674DFFD5C309445E56DE9839" box="[830,981,1087,1113]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="345" phylum="Arthropoda" rank="subFamily" subFamily="Spilomelinae">Spilomelinae</taxonomicName>
, larval feeding on
<taxonomicName id="4C5B7A92674DFFD5C49F445E502A9839" box="[1192,1313,1087,1113]" class="Magnoliopsida" family="Malvaceae" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="348" phylum="Tracheophyta" rank="family">Malvaceae</taxonomicName>
(including
<taxonomicName id="4C5B7A92674DFFD5C0A044025427981D" box="[151,300,1123,1149]" class="Magnoliopsida" family="Sterculiaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="348" phylum="Tracheophyta" rank="family">Sterculiaceae</taxonomicName>
and
<taxonomicName id="4C5B7A92674DFFD5C155440254C1981D" box="[354,458,1123,1149]" class="Magnoliopsida" family="Tiliaceae" higherTaxonomySource="GBIF" kingdom="Plantae" order="Malvales" pageId="5" pageNumber="348" phylum="Tracheophyta" rank="family">Tiliaceae</taxonomicName>
) is common only in
<taxonomicName id="4C5B7A92674DFFD5C2984402562F981D" authorityName="Acloque" authorityYear="1897" box="[687,804,1123,1149]" class="Insecta" family="Crambidae" kingdom="Animalia" order="Lepidoptera" pageId="5" pageNumber="348" phylum="Arthropoda" rank="tribe" tribe="Agroterini">Agroterini</taxonomicName>
(
<bibRefCitation id="EFCA7CE0674DFFD5C30344025157981D" author="Janzen, D. H. &amp; Hallwachs, W." box="[820,1116,1123,1149]" pageId="5" pageNumber="348" refId="ref11966" refString="Janzen, D. H. &amp; Hallwachs, W. (2009) Dynamic database for an inventory of the macrocaterpillar fauna, and its food plants and parasitoids, of Area de Conservacion Guanacaste (ACG), northwestern Costa Rica (nn-SRNP-nnnnn voucher codes). Available from: http: // janzen. sas. upenn. edu (accessed 15 June 2023)" type="url" year="2009">Janzen &amp; Hallwachs 2009</bibRefCitation>
;
<bibRefCitation id="EFCA7CE0674DFFD5C45F4402505D981D" author="Robinson, G. S. &amp; Ackery, P. R. &amp; Kitching, I. &amp; Beccaloni, G. W. &amp; Hernandez, L. M." box="[1128,1366,1123,1149]" pageId="5" pageNumber="348" refId="ref12623" refString="Robinson, G. S., Ackery, P. R., Kitching, I., Beccaloni, G. W. &amp; Hernandez, L. M. (2023) HOSTS - a Database of the World's Lepidopteran Hostplants [Data set]. Natural History Museum. Available from: https: // data. nhm. ac. uk / dataset / hosts (accessed 18 October 2023) https: // doi. org / 10.5519 / havt 50 xw" type="url" year="2023">
Robinson
<emphasis id="B92FDD03674DFFD5C4ED44025019981D" box="[1242,1298,1123,1149]" italics="true" pageId="5" pageNumber="348">et al.</emphasis>
2023
</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>